330 lines
40 KiB
XML
330 lines
40 KiB
XML
<document id="A79DEEFC3A2C3E090B9882DDA2636BB1" ID-CLB-Dataset="25166" ID-DOI="10.11646/zootaxa.4688.3.1" ID-GBIF-Dataset="5644b1e6-649e-4bdb-9c2f-9547055eeb6f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3516745" ID-ZooBank="DC8D545C-9F75-4A4E-B6DD-22A547694B27" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571832611391" checkinUser="plazi" docAuthor="Schnabel, Kareen E. & Ahyong, Shane T." docDate="2019" docId="03E487857349FFDBFF25B766FED5FAB8" docLanguage="en" docName="zootaxa.4688.3.1.pdf" docOrigin="Zootaxa 4688 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phylladiorhynchus kermadecensis Schnabel & Ahyong 2019, n. sp." docType="treatment" docUuid="23B1772A-2398-405D-90E5-4D8C77D819ED" docUuidSource="ZooBank" docVersion="20" lastPageNumber="320" masterDocId="FFDDFFFD7358FFC8FFB2B617FFC7FFC3" masterDocTitle="The squat lobster genus Phylladiorhynchus Baba, 1969 in New Zealand and eastern Australia, with description of six new species" masterLastPageNumber="347" masterPageNumber="301" pageNumber="318" updateTime="1698760689878" updateUser="ExternalLinkService">
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<mods:titleInfo id="B8FDC6A9A6858CA55709AB3887045182">
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<mods:title id="FF2F01259C6CEA355E582F0BF834ABD6">The squat lobster genus Phylladiorhynchus Baba, 1969 in New Zealand and eastern Australia, with description of six new species</mods:title>
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<mods:namePart id="BCA5F26BD11296009CCE474057B473AC">Schnabel, Kareen E.</mods:namePart>
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<mods:namePart id="FBB022A3AC0B83803DEC9D69B498658F">Ahyong, Shane T.</mods:namePart>
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<mods:title id="CEC9AB3BFC1726955F0C8CED6B0B588E">Zootaxa</mods:title>
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<mods:part id="CA6E0D8060BC9FB8E212F2888218968D">
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<mods:date id="C60BD6317A1CB38EEF23D78451BA633B">2019</mods:date>
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<mods:number id="8BA1BCD9C3200A034510583305DA6FE4">2019-10-23</mods:number>
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<mods:number id="D11A445EE505DAFDCB2D634B2645DEBD">4688</mods:number>
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<treatment id="03E487857349FFDBFF25B766FED5FAB8" ID-DOI="http://doi.org/10.5281/zenodo.5030869" ID-GBIF-Taxon="159812084" ID-Zenodo-Dep="5030869" LSID="urn:lsid:zoobank.org:act:23B1772A-2398-405D-90E5-4D8C77D819ED" httpUri="http://treatment.plazi.org/id/03E487857349FFDBFF25B766FED5FAB8" lastPageId="19" lastPageNumber="320" pageId="17" pageNumber="318">
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<subSubSection id="C35765187349FFD9FF25B766FD47FE48" box="[151,640,369,396]" pageId="17" pageNumber="318" type="nomenclature">
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<paragraph id="8BF236937349FFD9FF25B766FD47FE48" blockId="17.[151,640,369,431]" box="[151,640,369,396]" pageId="17" pageNumber="318">
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<heading id="D0BA81FF7349FFD9FF25B766FD47FE48" bold="true" box="[151,640,369,396]" fontSize="11" level="1" pageId="17" pageNumber="318" reason="1">
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<emphasis id="B939EA817349FFD9FF25B766FD47FE48" bold="true" box="[151,640,369,396]" pageId="17" pageNumber="318">
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<taxonomicName id="4C4D4D107349FFD9FF25B766FDF1FE4F" ID-CoL="5bf68574-fb85-4c7e-9d87-6760c194c39c" authority="Schnabel & Ahyong, 2019" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[151,566,369,396]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="kermadecensis" status="sp. nov.">
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<emphasis id="B939EA817349FFD9FF25B766FDF1FE4F" bold="true" box="[151,566,369,396]" italics="true" pageId="17" pageNumber="318">Phylladiorhynchus kermadecensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20A57FA7349FFD9FD8FB766FD47FE48" box="[573,640,369,395]" pageId="17" pageNumber="318" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35765187349FFD9FF25B782FEC0FE6C" box="[151,263,404,431]" pageId="17" pageNumber="318" type="description">
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<paragraph id="8BF236937349FFD9FF25B782FEC0FE6C" blockId="17.[151,640,369,431]" box="[151,263,404,431]" pageId="17" pageNumber="318">
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(
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<figureCitation id="13762A167349FFD9FF12B782FF23FE6C" box="[160,228,404,431]" captionStart="FIGURE 5" captionStartId="13.[151,250,1975,2000]" captionTargetBox="[160,1424,1063,1943]" captionTargetId="figure@13.[151,1436,1057,1956]" captionTargetPageId="13" captionText="FIGURE 5. Distribution records for Phylladiorhynchus erebus n. sp. (white circles) and P. kermadecensis n. sp. (crossed circle). Lines demarcating the EEZ for both Australia and NZ shown." figureDoi="http://doi.org/10.5281/zenodo.3516755" httpUri="https://zenodo.org/record/3516755/files/figure.png" pageId="17" pageNumber="318">Fig. 5</figureCitation>
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,
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<figureCitation id="13762A167349FFD9FF42B782FF38FE6C" box="[240,255,405,431]" captionStart="FIGURE 7" captionStartId="18.[151,250,1939,1964]" captionTargetBox="[228,1354,181,1908]" captionTargetId="figure@18.[214,1373,166,1920]" captionTargetPageId="18" captionText="FIGURE 7. Phylladiorhynchus kermadecensis n. sp., holotype male, 2.9 mm (NIWA 135616): A, carapace and abdomen, dorsal view. B, sternal plastron. C, telson. D, left cephalic region, showing antennular and antennal peduncles, ventral view. E, left Mxp3, lateral view. F, left cheliped, dorsal view. G–I, left P2–4, lateral view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3516759" httpUri="https://zenodo.org/record/3516759/files/figure.png" pageId="17" pageNumber="318">7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35765187349FFD9FF25B7CAFE50FDA0" pageId="17" pageNumber="318" type="materials_examined">
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<paragraph id="8BF236937349FFD9FF25B7CAFE50FDA0" blockId="17.[151,1437,476,2015]" pageId="17" pageNumber="318">
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<emphasis id="B939EA817349FFD9FF25B7CAFE82FE34" bold="true" box="[151,325,476,503]" pageId="17" pageNumber="318">Type material.</emphasis>
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<materialsCitation id="3B253CCE7349FFD9FEFEB7CAFB37FDD8" ID-GBIF-Occurrence="2429072201" collectingDate="2016-10-30" collectionCode="NIWA, TAN" elevation="241" latitude="30.223" location="Macauley Island" longLatPrecision="73" longitude="-178.353" municipality="Kermadec Ridge" pageId="17" pageNumber="318" specimenCode="NIWA 135616" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54F688317349FFD9FEFEB7CAFE23FE34" box="[332,484,477,503]" pageId="17" pageNumber="318" type="holotype">HOLOTYPE</typeStatus>
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:
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<specimenCode id="DBEB9EE87349FFD9FE5DB7CAFD52FE34" box="[495,661,476,503]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="17" pageNumber="318" type="Other institutional type">NIWA 135616</specimenCode>
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, male (
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<quantity id="4CB59B767349FFD9FD54B7CAFC85FE34" box="[742,834,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="17" pageNumber="318" unit="mm" value="2.9">2.9 mm</quantity>
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),
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<location id="8E9260487349FFD9FCE7B7CAFBD5FE34" LSID="urn:lsid:plazi:treatment:03E487857349FFDBFF25B766FED5FAB8:8E9260487349FFD9FCE7B7CAFBD5FE34" box="[853,1042,477,503]" latitude="30.223" longLatPrecision="73" longitude="-178.353" municipality="Kermadec Ridge" name="Macauley Island" pageId="17" pageNumber="318">Macauley Island</location>
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,
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<collectingMunicipality id="6B96ACE97349FFD9FBAFB7CAFB19FE34" box="[1053,1246,477,503]" pageId="17" pageNumber="318">Kermadec Ridge</collectingMunicipality>
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,
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<geoCoordinate id="EE7950547349FFD9FB5BB7CAFF6FFDD8" pageId="17" pageNumber="318">30.223–30.219° S</geoCoordinate>
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,
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<geoCoordinate id="EE7950547349FFD9FF07B417FE62FDD8" box="[181,421,512,539]" pageId="17" pageNumber="318">178.354–178.353° W</geoCoordinate>
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,
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<quantity id="4CB59B767349FFD9FE02B417FDF6FDD8" box="[432,561,512,539]" metricMagnitude="2" metricUnit="m" metricValue="2.41" metricValueMax="2.87" metricValueMin="1.95" pageId="17" pageNumber="318" unit="m" value="241.0" valueMax="287.0" valueMin="195.0">
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<elevation id="0060D1A07349FFD9FE02B417FDF6FDD8" box="[432,561,512,539]" metricMagnitude="2" metricUnit="m" metricValue="2.41" metricValueMax="2.87" metricValueMin="1.95" pageId="17" pageNumber="318" unit="m" value="241.0" valueMax="287.0" valueMin="195.0">195–287 m</elevation>
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</quantity>
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,
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<date id="FFF310537349FFD9FD8FB416FD0DFDD8" box="[573,714,513,539]" pageId="17" pageNumber="318" value="2016-10-30">
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<collectingDate id="EFB7E9BB7349FFD9FD8FB416FD0DFDD8" box="[573,714,513,539]" pageId="17" pageNumber="318" value="2016-10-30">30 Oct 2016</collectingDate>
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</date>
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,
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||
<collectionCode id="ED5CAE567349FFD9FD67B416FCDAFDD8" box="[725,797,513,539]" collectionName="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="17" pageNumber="318" type="Other institutional type">NIWA</collectionCode>
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Kermadec-Rangitahua Stn.
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||
<collectionCode id="ED5CAE567349FFD9FBEBB416FB53FDD8" box="[1113,1172,513,539]" collectionName="TAN" pageId="17" pageNumber="318">TAN</collectionCode>
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1612/90
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</materialsCitation>
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.
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<materialsCitation id="3B253CCE7349FFD9FB48B416FCFCFDFC" ID-GBIF-Occurrence="2429072083" collectionCode="AWMM" pageId="17" pageNumber="318" specimenCode="MA73590" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<typeStatus id="54F688317349FFD9FB48B416FA5EFDD8" box="[1274,1433,513,539]" pageId="17" pageNumber="318" type="paratype">PARATYPES</typeStatus>
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:
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<collectionCode id="ED5CAE567349FFD9FF25B432FF32FDFC" box="[151,245,549,575]" collectionName="AWMM" pageId="17" pageNumber="318">AWMM</collectionCode>
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<specimenCode id="DBEB9EE87349FFD9FF4DB433FEB2FDFC" box="[255,373,548,575]" collectionCode="AWMM" pageId="17" pageNumber="318">MA73590</specimenCode>
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,
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<specimenCount id="9D4BFD1A7349FFD9FE30B432FE08FDFC" box="[386,463,549,575]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
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(
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<quantity id="4CB59B767349FFD9FE53B432FDF9FDFC" box="[481,574,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="318" unit="mm" value="2.6">2.6 mm</quantity>
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), station details as for
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</materialsCitation>
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<materialsCitation id="3B253CCE7349FFD9FCF6B432FE54FDA0" ID-GBIF-Occurrence="2429072082" collectionCode="NIWA" pageId="17" pageNumber="318" specimenCode="NIWA 118955" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="54F688317349FFD9FCF6B432FC6DFDFC" box="[836,938,549,575]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
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.
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<specimenCode id="DBEB9EE87349FFD9FC04B432FB9AFDFC" box="[950,1117,548,575]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="17" pageNumber="318" type="Other institutional type">NIWA 118955</specimenCode>
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,
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<specimenCount id="9D4BFD1A7349FFD9FBD8B432FB0AFDFC" box="[1130,1229,549,575]" pageId="17" pageNumber="318" type="female">1 female</specimenCount>
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(
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<quantity id="4CB59B767349FFD9FB6DB432FAFBFDFC" box="[1247,1340,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="318" unit="mm" value="2.4">2.4 mm</quantity>
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), station details as for
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<typeStatus id="54F688317349FFD9FE9FB45EFE54FDA0" box="[301,403,585,611]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35765187349FFDBFF75B47BFC90FC64" lastPageId="19" lastPageNumber="320" pageId="17" pageNumber="318" type="diagnosis">
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<paragraph id="8BF236937349FFD9FF75B47BFBECFC40" blockId="17.[151,1437,476,2015]" pageId="17" pageNumber="318">
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<emphasis id="B939EA817349FFD9FF75B47BFE85FD44" bold="true" box="[199,322,620,647]" pageId="17" pageNumber="318">Diagnosis.</emphasis>
|
||
Rostrum without subapical spines; lateral margins slightly concave. Carapace with transverse row of 5 epigastric spines (1 median and 2 on either side of midline); anterior branchial margin with 2 spines. Thoracic sternite 3 anterior margin sinuous, with shallow median concavity. Antennular article 1 with 5 well-developed spines: distomesial spine small, similar to lower distolateral spine, shorter than upper distolateral spine, well overreached by first lateral spine; second (proximal) lateral spine smallest but distinct. Antennal article 1 mesial process distally falling well short of second lateral antennular spine; article 2 distomesial spine longer than distolateral; article 3 with [small] to distinct distomesial spine. Maxilliped 3 merus with 2 prominent spines on flexor margin. P2–4 dactylus extensor margin without upright spines at bases of movable spines.
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||
</paragraph>
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||
<paragraph id="8BF236937349FFD9FF75B59AFCE0FAB8" blockId="17.[151,1437,476,2015]" pageId="17" pageNumber="318">
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<emphasis id="B939EA817349FFD9FF75B59AFF2FFC64" bold="true" box="[199,232,909,935]" pageId="17" pageNumber="318">De</emphasis>
|
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s
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||
<emphasis id="B939EA817349FFD9FF41B59BFE9EFC64" bold="true" box="[243,345,908,935]" pageId="17" pageNumber="318">cription.</emphasis>
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||
<emphasis id="B939EA817349FFD9FED0B59AFE17FC64" box="[354,464,909,935]" italics="true" pageId="17" pageNumber="318">Carapace</emphasis>
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: As long as wide; transverse ridges with dense short setae, and few scattered longer setae. Gastric region with epigastric ridge indistinct, with 5 spines: 1 median and 2 pairs of spines lateral, followed by 3, usually interrupted transverse ridges: anterior protogastric ridge [continuous] or with small medial gap, falling short laterally of carapace margin; anterior mesogastric ridge with small medial gap, interrupted laterally by cervical groove, laterally following cervical groove, then continuing uninterrupted to first anterior branchial spine; 1 medially interrupted metagastric ridge, laterally interrupted by cervical groove. Mid-transverse ridge uninterrupted, extending laterally to posterior branchial spine, followed by 2 uninterrupted ridges, intervened by 1 short lateral ridge, otherwise smooth. Lateral margins weakly convex, with 5 spines: 2 spines (anterolateral, hepatic) in front of, and 3 spines (2 anterior branchial, 1 posterior branchial) behind distinct anterior cervical groove. Anterolateral spine well-developed, nearly reaching lateral orbital spine, size about equal. Hepatic spine small, set slightly mesially from lateral margin. Anterior branchial spines largest, size diminishing posteriorly. Rostrum length nearly 0.5 × pcl, length-width ratio 1.4, distance between basal pair of rostral spines about 0.3 × carapace width; dorsal surface very slightly concave; lateral margins slightly concave, sub-apical spines absent. Pterygostomian flap anteriorly rounded, without spine; surface with series of short striae, unarmed.
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</paragraph>
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<paragraph id="8BF236937349FFD9FF75B392FB4EFA00" blockId="17.[151,1437,476,2015]" pageId="17" pageNumber="318">
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||
<emphasis id="B939EA817349FFD9FF75B392FE57FA5C" box="[199,400,1413,1439]" italics="true" pageId="17" pageNumber="318">Thoracic sternum</emphasis>
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: Sternal plastron as wide as long, lateral margins slightly divergent posteriorly. Sternite 3 width about 3 × length; anteriorly biconvex with shallow median notch; laterally rounded.
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</paragraph>
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<paragraph id="8BF236937349FFD9FF75B3DAFD5DF9C8" blockId="17.[151,1437,476,2015]" pageId="17" pageNumber="318">
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||
<emphasis id="B939EA817349FFD9FF75B3DAFEF6FA24" box="[199,305,1485,1511]" italics="true" pageId="17" pageNumber="318">Abdomen</emphasis>
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: Tergite 2 with anterior and posterior transverse ridge; tergites 3 and 4 with anterior ridge only; tergites 5 and 6 smooth, without elevated ridges.
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</paragraph>
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<paragraph id="8BF236937349FFD9FF75B001FD7AF990" blockId="17.[151,1437,476,2015]" pageId="17" pageNumber="318">
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||
<emphasis id="B939EA817349FFD9FF75B001FF36F9EC" box="[199,241,1558,1583]" italics="true" pageId="17" pageNumber="318">Eye</emphasis>
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: Eye length about 1.2 × width, peduncle distally setose, slightly expanded proximally, with few short transverse striae on lateral surface; cornea not dilated.
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</paragraph>
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<paragraph id="8BF236937349FFD9FF75B04AFAA2F958" blockId="17.[151,1437,476,2015]" pageId="17" pageNumber="318">
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||
<emphasis id="B939EA817349FFD9FF75B04AFEFEF9B4" box="[199,313,1629,1655]" italics="true" pageId="17" pageNumber="318">Antennule</emphasis>
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||
:Article 1 with 5 well-developed distal spines, distomesial spine small, shorter than distolateral spines; paired distolateral spines present; proximal lateral spine distinct. Short striae covering mesial parts of surface.
|
||
</paragraph>
|
||
<paragraph id="8BF236937349FFD9FF75B0B2FD55F8C4" blockId="17.[151,1437,476,2015]" pageId="17" pageNumber="318">
|
||
<emphasis id="B939EA817349FFD9FF75B0B2FEE3F97C" box="[199,292,1701,1727]" italics="true" pageId="17" pageNumber="318">Antenna</emphasis>
|
||
: Article 1 with prominent mesial process, distally falling well short of second (proximal) lateral antennular spine. Article 2 with distinct distal spines laterally and mesially, mesial spine larger. Article 3 with [small] to distinct distomesial spine. Article 4 unarmed.
|
||
</paragraph>
|
||
<paragraph id="8BF236937349FFD9FF75B106FACFF8B0" blockId="17.[151,1437,476,2015]" pageId="17" pageNumber="318">
|
||
<emphasis id="B939EA817349FFD9FF75B106FE69F8E8" box="[199,430,1809,1835]" italics="true" pageId="17" pageNumber="318">Maxilliped 3 (Mxp3)</emphasis>
|
||
: Ischium with small distal spine on extensor margin and larger distal spine on flexor margin; crista dentata with regular row of spines along entire margin. Merus 0.7 × length of ischium at midline, with small distal spines on extensor margin and 2 much larger spines at flexor margin, otherwise unarmed.
|
||
</paragraph>
|
||
<paragraph id="8BF236937349FFDBFF75B16AFE88FEA4" blockId="17.[151,1437,476,2015]" lastBlockId="19.[151,1437,152,1403]" lastPageId="19" lastPageNumber="320" pageId="17" pageNumber="318">
|
||
<emphasis id="B939EA817349FFD9FF75B16AFEEBF854" box="[199,300,1917,1943]" italics="true" pageId="17" pageNumber="318">Cheliped</emphasis>
|
||
: Total length 3.5–4.4 [4.2 and 4.4] × pcl (probably female); subcylindrical, spiny, with scattered long setae. Ischiomerus 1.4–1.8 [1.6 and 1.8] × pcl, just over twice length of carpus; with rows of spines, mesial spines strongest. Carpus with rows of spines around entire article, 1 or 2 mesial spines most prominent. Palm 1.0–[1.3] × carpus length, length-width ratio [1.5]–2.3, with 4 parallel rows of spines (2 mesial, 1 at about dorsal midline, 1 close to lateral margin); fingers gaping; outer base of dactylus with sharp spine. Pollex outer proximal margin with spine; occlusal margin with 1 proximal process and distinct triangular process distally when a gape is present (process absent when not gaping). Dactylus subequal in length to palm; outer proximal margin with spine; occlusal margin with 2 processes, distally with row of spines along curved margin; tip excavated; surface with short setiferous striae.
|
||
</paragraph>
|
||
<caption id="DF32661B734AFFDAFF25B184FB9CF837" ID-DOI="http://doi.org/10.5281/zenodo.3516759" ID-Zenodo-Dep="3516759" httpUri="https://zenodo.org/record/3516759/files/figure.png" pageId="18" pageNumber="319" startId="18.[151,250,1939,1964]" targetBox="[228,1354,181,1908]" targetPageId="18">
|
||
<paragraph id="8BF23693734AFFDAFF25B184FB9CF837" blockId="18.[151,1437,1939,2036]" pageId="18" pageNumber="319">
|
||
<emphasis id="B939EA81734AFFDAFF25B184FED4F86F" bold="true" box="[151,275,1939,1964]" pageId="18" pageNumber="319">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C4D4D10734AFFDAFEABB183FDB3F86F" authority="Schnabel & Ahyong, 2019" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[281,628,1940,1964]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="kermadecensis" status="sp. nov.">
|
||
<emphasis id="B939EA81734AFFDAFEABB183FDB3F86F" box="[281,628,1940,1964]" italics="true" pageId="18" pageNumber="319">Phylladiorhynchus kermadecensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA81734AFFDAFDC8B183FD74F86F" bold="true" box="[634,691,1940,1964]" pageId="18" pageNumber="319">
|
||
<taxonomicNameLabel id="A20A57FA734AFFDAFDC8B183FD74F86F" box="[634,691,1940,1964]" pageId="18" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male, 2.9 mm (NIWA 135616): A, carapace and abdomen, dorsal view. B, sternal plastron. C, telson. D, left cephalic region, showing antennular and antennal peduncles, ventral view. E, left Mxp3, lateral view. F, left cheliped, dorsal view. G–I, left P2–4, lateral view. Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23693734BFFDBFF75B766FB7AFD30" blockId="19.[151,1437,152,1403]" pageId="19" pageNumber="320">
|
||
<emphasis id="B939EA81734BFFDBFF75B766FE77FE48" box="[199,432,369,395]" italics="true" pageId="19" pageNumber="320">Walking legs (P2–4)</emphasis>
|
||
: Slender, subcylindrical, moderately setose and spinose. P2–4 meri subequal in length; length 5–7 × width; P4 merus slightly shorter (0.8–[0.9] ×P2 merus length). P2 merus [0.8]–1.0 × pcl, length [5.9] × width, [1.2] × longer than propodus. P3 merus 1.2 × P3 propodus length. P4 merus 0.9–[1.0] × P4 propodus length. Extensor margin of P2 and P3 with regular row of spines, proximally diminishing, with prominent distal spine; P4 extensor margin unarmed other than distal spine; ventral margins of all legs irregular and with strong distal spine; P4 lateral surface with median row of small spines, absent on P2 and P3. Carpus with 3 or 4 spines on extensor margin on P2–3 (very small in smaller specimens), irregular but unarmed on P4; distal spine prominent on all legs; 1–4 minute spines along row below extensor margin on lateral surface on all legs; flexor margin unarmed. P2–4 propodus length about 7–8 × height; extensor margin typically with 2 proximal spines on P2–3, unarmed on P4; flexor margin with 3 or 4 slender movable spines in addition to distal pair. Dactylus 0.5–0.6 × propodus length, ending in incurved, strong, sharp spine; flexor margin with 10 or 11 movable spines, otherwise unarmed.
|
||
</paragraph>
|
||
<paragraph id="8BF23693734BFFDBFF75B4EBFD1AFCD4" blockId="19.[151,1437,152,1403]" box="[199,733,764,791]" pageId="19" pageNumber="320">
|
||
<emphasis id="B939EA81734BFFDBFF75B4EBFF39FCD4" bold="true" box="[199,254,764,791]" pageId="19" pageNumber="320">Size.</emphasis>
|
||
Male pcl
|
||
<quantity id="4CB59B76734BFFDBFEDDB4EAFE3FFCD4" box="[367,504,765,791]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="19" pageNumber="320" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
|
||
, female pcl
|
||
<quantity id="4CB59B76734BFFDBFD32B4EAFD1EFCD4" box="[640,729,765,791]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="320" unit="mm" value="2.4">2.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF23693734BFFDBFF75B537FE2EFCF8" blockId="19.[151,1437,152,1403]" box="[199,489,800,827]" pageId="19" pageNumber="320">
|
||
<emphasis id="B939EA81734BFFDBFF75B537FEAAFCF8" bold="true" box="[199,365,800,827]" pageId="19" pageNumber="320">Colour in life.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BF23693734BFFDBFF75B553FC90FC64" blockId="19.[151,1437,152,1403]" pageId="19" pageNumber="320">
|
||
<emphasis id="B939EA81734BFFDBFF75B553FEA1FC9C" bold="true" box="[199,358,836,863]" pageId="19" pageNumber="320">Genetic data.</emphasis>
|
||
DNA extracted from two detached walking legs derived from the
|
||
<typeStatus id="54F68831734BFFDBFBFCB552FB7EFC9C" box="[1102,1209,837,863]" pageId="19" pageNumber="320" type="paratype">paratypes</typeStatus>
|
||
(NIWA 123247 and AWMM MA73590). COI sequence divergence between
|
||
<taxonomicName id="4C4D4D10734BFFDBFCA0B57DFC11FC40" authority="Schnabel & Ahyong, 2019" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[786,982,873,899]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="kermadecensis" status="sp. nov.">
|
||
<emphasis id="B939EA81734BFFDBFCA0B57DFCE4FC40" box="[786,803,874,899]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFC83B57EFC11FC40" box="[817,982,873,899]" italics="true" pageId="19" pageNumber="320">kermadecensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA81734BFFDBFC6FB57EFBDBFC40" bold="true" box="[989,1052,873,899]" pageId="19" pageNumber="320">
|
||
<taxonomicNameLabel id="A20A57FA734BFFDBFC6FB57EFBDBFC40" box="[989,1052,873,899]" pageId="19" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4D4D10734BFFDBFBE0B57DFB7BFC40" authority="Schnabel & Ahyong, 2019" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[1106,1212,873,899]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
|
||
<emphasis id="B939EA81734BFFDBFBE0B57DFBA4FC40" box="[1106,1123,874,899]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFBC0B57EFB7BFC40" box="[1138,1212,873,899]" italics="true" pageId="19" pageNumber="320">erebus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA81734BFFDBFB70B57EFAC6FC40" bold="true" box="[1218,1281,873,899]" pageId="19" pageNumber="320">
|
||
<taxonomicNameLabel id="A20A57FA734BFFDBFB70B57EFAC6FC40" box="[1218,1281,873,899]" pageId="19" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MA125788) is 17%, supporting interspecific levels of divergence (
|
||
<tableCitation id="C6CF0328734BFFDBFD40B59AFC82FC64" box="[754,837,909,935]" captionStart="TABLE 1" captionStartId="41.[151,244,511,538]" captionTargetBox="[165,2022,621,1679]" captionTargetId="graphics@41.[151,2038,1112,1687]" captionTargetPageId="41" captionText="TABLE 1. Pairwise percentage genetic distances (below diagonal) and genetic similarity (above diagonal) for the partial COI gene between specimens of Phylladi- orhynchus. Cells highlighted in grey indicate intraspecific levels of divergence. The sequence of P. integrirostris from Hawaii was kindly provided as a reference by P. Rodríguez-Flores (National Museum of Natural Sciences, Madrid)." httpUri="http://table.plazi.org/id/DF32661B7371FFE1FF25B7E8FC4FFDA1" pageId="19" pageNumber="320" tableUuid="DF32661B7371FFE1FF25B7E8FC4FFDA1">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576518734BFFDBFF75B5A6FBACFC08" box="[199,1131,945,971]" pageId="19" pageNumber="320" type="etymology">
|
||
<paragraph id="8BF23693734BFFDBFF75B5A6FBACFC08" blockId="19.[151,1437,152,1403]" box="[199,1131,945,971]" pageId="19" pageNumber="320">
|
||
<emphasis id="B939EA81734BFFDBFF75B5A6FE89FC08" bold="true" box="[199,334,945,971]" pageId="19" pageNumber="320">Etymology.</emphasis>
|
||
Named after its only known area of occurrence, the Kermadec Islands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576518734BFFDBFF75B5C3FBAFFC2C" box="[199,1128,980,1007]" pageId="19" pageNumber="320" type="distribution">
|
||
<paragraph id="8BF23693734BFFDBFF75B5C3FBAFFC2C" blockId="19.[151,1437,152,1403]" box="[199,1128,980,1007]" pageId="19" pageNumber="320">
|
||
<emphasis id="B939EA81734BFFDBFF75B5C3FEA6FC2C" bold="true" box="[199,353,980,1007]" pageId="19" pageNumber="320">Distribution.</emphasis>
|
||
Known from the Kermadec Ridge, off Macauley Island,
|
||
<quantity id="4CB59B76734BFFDBFC53B5C3FBA4FC2C" box="[993,1123,980,1007]" metricMagnitude="2" metricUnit="m" metricValue="2.41" metricValueMax="2.87" metricValueMin="1.95" pageId="19" pageNumber="320" unit="m" value="241.0" valueMax="287.0" valueMin="195.0">
|
||
195–
|
||
<specimenCount id="9D4BFD1A734BFFDBFBABB5C3FBA4FC2C" box="[1049,1123,980,1007]" pageId="19" pageNumber="320" type="generic">287 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576518734BFFDBFF75B5EEFED5FAB8" pageId="19" pageNumber="320" type="discussion">
|
||
<paragraph id="8BF23693734BFFDBFF75B5EEFED5FAB8" blockId="19.[151,1437,152,1403]" pageId="19" pageNumber="320">
|
||
<emphasis id="B939EA81734BFFDBFF75B5EEFEFCFBD0" bold="true" box="[199,315,1017,1043]" pageId="19" pageNumber="320">Remarks.</emphasis>
|
||
<taxonomicName id="4C4D4D10734BFFDBFEF6B5EEFD03FBD0" authority="Schnabel & Ahyong, 2019" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[324,708,1017,1043]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="kermadecensis" status="sp. nov.">
|
||
<emphasis id="B939EA81734BFFDBFEF6B5EEFD03FBD0" box="[324,708,1017,1043]" italics="true" pageId="19" pageNumber="320">Phylladiorhynchus kermadecensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA81734BFFDBFD7EB5EEFCCAFBD0" bold="true" box="[716,781,1017,1043]" pageId="19" pageNumber="320">
|
||
<taxonomicNameLabel id="A20A57FA734BFFDBFD7EB5EEFCCAFBD0" box="[716,781,1017,1043]" pageId="19" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
belongs to the ‘
|
||
<taxonomicName id="4C4D4D10734BFFDBFC75B5EEFBCCFBD0" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[967,1035,1017,1043]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="ikedai">
|
||
<emphasis id="B939EA81734BFFDBFC75B5EEFBCCFBD0" box="[967,1035,1017,1043]" italics="true" pageId="19" pageNumber="320">ikedai</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C4D4D10734BFFDBFBA6B5EEFB5FFBD0" authorityName="Tirmizi & Javed" authorityYear="1980" box="[1044,1176,1017,1043]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="bengalensis">
|
||
<emphasis id="B939EA81734BFFDBFBA6B5EEFB5FFBD0" box="[1044,1176,1017,1043]" italics="true" pageId="19" pageNumber="320">bengalensis</emphasis>
|
||
</taxonomicName>
|
||
group’ of species with five epigastric spines on the carapace and two spines on the flexor margin of the Mxp3 merus. Significant genetic divergence from congeners, however, supports
|
||
<taxonomicName id="4C4D4D10734BFFDBFD04B255FCBAFB98" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[694,893,1089,1115]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="kermadecensis">
|
||
<emphasis id="B939EA81734BFFDBFD04B255FD00FB98" box="[694,711,1090,1115]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFD6AB256FCBAFB98" box="[728,893,1089,1115]" italics="true" pageId="19" pageNumber="320">kermadecensis</emphasis>
|
||
</taxonomicName>
|
||
as a distinct species. Diagnostic characters are subtle but the presence of two anterior branchial carapace spines (rather than three as in both
|
||
<taxonomicName id="4C4D4D10734BFFDBFB2FB271FB38FBBC" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[1181,1279,1125,1151]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="ikedai">
|
||
<emphasis id="B939EA81734BFFDBFB2FB271FB69FBBC" box="[1181,1198,1126,1151]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFB09B272FB38FBBC" box="[1211,1279,1125,1151]" italics="true" pageId="19" pageNumber="320">ikedai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D10734BFFDBFA86B271FA5AFBBC" authority="Schnabel & Ahyong, 2019" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[1332,1437,1125,1151]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
|
||
<emphasis id="B939EA81734BFFDBFA86B271FA82FBBC" box="[1332,1349,1126,1151]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFAE1B272FA5AFBBC" box="[1363,1437,1125,1151]" italics="true" pageId="19" pageNumber="320">erebus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA81734BFFDBFF25B29EFF10FB60" bold="true" box="[151,215,1161,1187]" pageId="19" pageNumber="320">
|
||
<taxonomicNameLabel id="A20A57FA734BFFDBFF25B29EFF10FB60" box="[151,215,1161,1187]" pageId="19" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) and a single spine along the posterior branchial margin (instead of two or three spines in
|
||
<taxonomicName id="4C4D4D10734BFFDBFB57B29DFA8DFB60" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[1253,1354,1161,1187]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="ikedai">
|
||
<emphasis id="B939EA81734BFFDBFB57B29DFB31FB60" box="[1253,1270,1162,1187]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFAB4B29EFA8DFB60" box="[1286,1354,1161,1187]" italics="true" pageId="19" pageNumber="320">ikedai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D10734BFFDBFA36B29DFF26FB04" authority="Schnabel & Ahyong, 2019" authorityName="Schnabel & Ahyong" authorityYear="2019" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
|
||
<emphasis id="B939EA81734BFFDBFA36B29DFA52FB60" box="[1412,1429,1162,1187]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFF25B2BAFF26FB04" box="[151,225,1197,1223]" italics="true" pageId="19" pageNumber="320">erebus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA81734BFFDBFF55B2BAFEE2FB04" bold="true" box="[231,293,1197,1223]" pageId="19" pageNumber="320">
|
||
<taxonomicNameLabel id="A20A57FA734BFFDBFF55B2BAFEE2FB04" box="[231,293,1197,1223]" pageId="19" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) are constant in the material examined. Additionally, the lateral margin of the rostrum is concave in all specimens of
|
||
<taxonomicName id="4C4D4D10734BFFDBFE9DB2C5FE36FB28" authority="Schnabel & Ahyong, 2019" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[303,497,1233,1259]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="kermadecensis" status="sp. nov.">
|
||
<emphasis id="B939EA81734BFFDBFE9DB2C5FE87FB28" box="[303,320,1234,1259]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFEFEB2C6FE36FB28" box="[332,497,1233,1259]" italics="true" pageId="19" pageNumber="320">kermadecensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA81734BFFDBFE45B2C6FDF3FB28" bold="true" box="[503,564,1233,1259]" pageId="19" pageNumber="320">
|
||
<taxonomicNameLabel id="A20A57FA734BFFDBFE45B2C6FDF3FB28" box="[503,564,1233,1259]" pageId="19" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
examined, rather than straight as in
|
||
<taxonomicName id="4C4D4D10734BFFDBFC71B2C5FBE2FB28" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[963,1061,1233,1259]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="ikedai">
|
||
<emphasis id="B939EA81734BFFDBFC71B2C5FC13FB28" box="[963,980,1234,1259]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFC53B2C6FBE2FB28" box="[993,1061,1233,1259]" italics="true" pageId="19" pageNumber="320">ikedai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D10734BFFDBFBEBB2C5FB06FB28" authority="Schnabel & Ahyong, 2019" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[1113,1217,1233,1259]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
|
||
<emphasis id="B939EA81734BFFDBFBEBB2C5FBADFB28" box="[1113,1130,1234,1259]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFBC5B2C6FB06FB28" box="[1143,1217,1233,1259]" italics="true" pageId="19" pageNumber="320">erebus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA81734BFFDBFB74B2C6FAC3FB28" bold="true" box="[1222,1284,1233,1259]" pageId="19" pageNumber="320">
|
||
<taxonomicNameLabel id="A20A57FA734BFFDBFB74B2C6FAC3FB28" box="[1222,1284,1233,1259]" pageId="19" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The presence of a distinct distal process on the occlusal margin of the cheliped pollex in the male
|
||
<typeStatus id="54F68831734BFFDBFBF0B2E2FB63FACC" box="[1090,1188,1269,1295]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
|
||
is absent in the single loose cheliped that remains, which is presumably from the small
|
||
<typeStatus id="54F68831734BFFDBFC32B30EFC27FAF0" box="[896,992,1305,1331]" pageId="19" pageNumber="320" type="paratype">paratype</typeStatus>
|
||
female, indicating sexual dimorphism. This process, however, is absent from the chelipeds of
|
||
<taxonomicName id="4C4D4D10734BFFDBFD4FB329FCA7FA94" authorityName="Schnabel & Ahyong" authorityYear="2019" box="[765,864,1341,1367]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="ikedai">
|
||
<emphasis id="B939EA81734BFFDBFD4FB329FCC9FA94" box="[765,782,1342,1367]" italics="true" pageId="19" pageNumber="320">P</emphasis>
|
||
.
|
||
<emphasis id="B939EA81734BFFDBFCAEB32AFCA7FA94" box="[796,864,1341,1367]" italics="true" pageId="19" pageNumber="320">ikedai</emphasis>
|
||
</taxonomicName>
|
||
of similar size indicating that this feature might be diagnostic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |