273 lines
33 KiB
XML
273 lines
33 KiB
XML
<document id="191A07A36DC70DE5B88EF49ED0379735" ID-DOI="10.5281/zenodo.5188356" ID-ISSN="1942-1354" ID-Zenodo-Dep="5188356" ID-ZooBank="AC2597CC-301F-4E91-9711-5C17399C9AA2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628346141153" checkinUser="felipe" docAuthor="King, Jonas G. & Lago, Paul K." docDate="2012" docId="03E1561898270B22FF19FF3B4FFA1C19" docLanguage="en" docName="InsectaMundi.0275.1-53.pdf" docOrigin="Insecta Mundi 2012 (275)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Heterocerus schwarzi Horn" docType="treatment" docVersion="3" lastPageNumber="27" masterDocId="FFD82E60983B0B3EFFD9FFF94D431A43" masterDocTitle="The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Mississippi and Alabama, with discussion and keys to the species occurring in the southeastern United States" masterLastPageNumber="53" masterPageNumber="1" pageNumber="27" updateTime="1699045744038" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="06F2F7693CA763AEC002FF0A2196384E">
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<mods:title id="652C853E07224489D0A18ABBCDA19574">The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Mississippi and Alabama, with discussion and keys to the species occurring in the southeastern United States</mods:title>
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<mods:name id="91EE8CD1B196DFF81FA46DDEB51BA99E" type="personal">
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<mods:namePart id="CC4767036F955E80B1EA87808B117852">King, Jonas G.</mods:namePart>
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<mods:name id="59079DC762C4EE50C3A7C12B8D5C1069" type="personal">
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<mods:namePart id="9766079D5A7C4ADF84E73549058DC23E">Lago, Paul K.</mods:namePart>
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<mods:typeOfResource id="EF1DE6C3746ADA89D2D71045761D7793">text</mods:typeOfResource>
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<mods:title id="18DD02AB32599BAAF960CBFD7D61EF48">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="06C9DCE563C5B12FE10E9A5AB24CA6A9">
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<mods:date id="868A56DAB50AC0469E4FEF07164ED3FF">2012</mods:date>
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<mods:number id="4EA8392DE8B68684F76C55DEC9550B8F">2012-12-28</mods:number>
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<mods:detail id="7688FA91CE19EDCD6BD176EBB5B9C9E7" type="volume">
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<mods:number id="973E82F782485EA34F86E419EC4D188A">2012</mods:number>
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<mods:number id="5839748329FED48267D540B7AE2B8E2D">275</mods:number>
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<mods:classification id="993FB26F865B22FA798B27F17B13952D">journal article</mods:classification>
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<mods:identifier id="96E6010C3C79336DA0820C6607654169" type="DOI">10.5281/zenodo.5188356</mods:identifier>
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<mods:identifier id="20828861506675232310B6EB998DF34C" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="67C850186205352B8219238D90A061CF" type="Zenodo-Dep">5188356</mods:identifier>
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<treatment id="03E1561898270B22FF19FF3B4FFA1C19" LSID="urn:lsid:plazi:treatment:03E1561898270B22FF19FF3B4FFA1C19" httpUri="http://treatment.plazi.org/id/03E1561898270B22FF19FF3B4FFA1C19" lastPageNumber="27" pageId="28" pageNumber="27">
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<subSubSection id="C352B48598270B22FF19FF3B4F211A99" box="[192,610,194,218]" pageId="28" pageNumber="27" type="nomenclature">
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<paragraph id="8BF7E70E98270B22FF19FF3B4F211A99" blockId="28.[192,610,194,250]" box="[192,610,194,218]" pageId="28" pageNumber="27">
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<heading id="D0BF506298270B22FF19FF3B4F211A99" bold="true" box="[192,610,194,218]" fontSize="10" level="2" pageId="28" pageNumber="27" reason="3">
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<emphasis id="B93C3B1C98270B22FF19FF3B4F211A99" bold="true" box="[192,610,194,218]" pageId="28" pageNumber="27">
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15.
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<taxonomicName id="4C489C8D98270B22FF34FF3A4F211A99" authority="Horn" authorityName="Horn" box="[237,610,194,218]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
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<emphasis id="B93C3B1C98270B22FF34FF3A4F571A99" bold="true" box="[237,532,194,218]" italics="true" pageId="28" pageNumber="27">Heterocerus schwarzi</emphasis>
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Horn
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C352B48598270B22FF19FF1B4CC11AB9" box="[192,386,226,250]" pageId="28" pageNumber="27" type="description">
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<paragraph id="8BF7E70E98270B22FF19FF1B4CC11AB9" blockId="28.[192,610,194,250]" box="[192,386,226,250]" pageId="28" pageNumber="27">
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(
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<figureCitation id="1373FB8B98270B22FF10FF1B4C621AB9" box="[201,289,226,250]" captionStart="Figures 21-25" captionStartId="18.[192,286,1256,1278]" captionTargetBox="[192,1401,191,1233]" captionTargetPageId="18" captionText="Figures 21-25. Habiti of Heterocerus species of intermediate size with post-mesocoxal lines. 21) H. schwarzi. 22) H. tenuis. 23) H. undatus. 24) H. texanus. 25) H. collaris. Scale line = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5188374" httpUri="https://zenodo.org/record/5188374/files/figure.png" pageId="28" pageNumber="27">Fig. 21</figureCitation>
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,
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<figureCitation id="1373FB8B98270B22FEF7FF1B4C0D1AB9" box="[302,334,226,250]" captionStart="Figures 45-53" captionStartId="39.[192,286,843,865]" captionTargetBox="[203,1417,196,805]" captionTargetId="figure-157@39.[770,867,432,529]" captionTargetPageId="39" captionText="Figures 45-53. Male genitalia of southeastern species. The ventral view is shown unless otherwise stated. 45-50) Heterocerus species in the undulatus group. 45) H. undatus. 46) H. collaris. 47) H. texanus; left - ventral view, right - lateral view. 48) H. schwarzi; left - ventral view, right - lateral view. 49) H. tenuis. 50) H. selanderi. 51-53) Augyles and Tropicus species. Figures 52-53 are at roughly 2x relative scale of prior genitalia. 51) Augyles auromicans. 52) Tropicus nigrellus; left - dorsal view, center - ventral view, right - lateral view. 53) Tropicus pusillus; left - dorsal view, right - ventral view." figureDoi="http://doi.org/10.5281/zenodo.5188384" httpUri="https://zenodo.org/record/5188384/files/figure.png" pageId="28" pageNumber="27">48</figureCitation>
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,
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<figureCitation id="1373FB8B98270B22FE82FF1B4C3A1AB9" box="[347,377,226,250]" captionStart="Figure 66-72" captionStartId="41.[192,275,1784,1806]" captionTargetBox="[189,1438,191,1770]" captionTargetId="figure-840@41.[787,836,959,983]" captionTargetPageId="41" captionText="Figure 66-72. Distribution by county in the United States. 66) Heterocerus undatus. 67) Heterocerus texanus. 68) Heterocerus schwarzi. 69) Heterocerus tenuis. 70) Augyles auromicans. 71) Tropicus pusillus. 72) Tropicus nigrellus." figureDoi="http://doi.org/10.5281/zenodo.5188396" httpUri="https://zenodo.org/record/5188396/files/figure.png" pageId="28" pageNumber="27">68</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C352B48598270B22FF19FEDA4FB71B19" pageId="28" pageNumber="27" type="reference_group">
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<paragraph id="8BF7E70E98270B22FF19FEDA4FB71B19" blockId="28.[192,756,290,346]" pageId="28" pageNumber="27">
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<treatmentCitationGroup id="AB58C02098270B22FF19FEDA4FB71B19" pageId="28" pageNumber="27">
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<treatmentCitation id="0AE9C11F98270B22FF19FEDA4F3E1B79" author="Horn, G. H." box="[192,637,290,314]" page="11" pageId="28" pageNumber="27" year="1890">
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<taxonomicName id="4C489C8D98270B22FF19FEDA4F3E1B79" authority="Horn 1890: 11" authorityName="Horn" authorityPageNumber="11" authorityYear="1890" box="[192,637,290,314]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
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<emphasis id="B93C3B1C98270B22FF19FEDA4C801B79" box="[192,451,290,314]" italics="true" pageId="28" pageNumber="27">Heterocerus schwarzi</emphasis>
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<bibRefCitation id="EFD99AFF98270B22FE13FEDA4F3E1B79" author="Horn, G. H." box="[458,637,290,314]" pageId="28" pageNumber="27" pagination="1 - 16" refId="ref20843" refString="Horn, G. H. 1890. The species of Heterocerus of Boreal America. Transactions of the American Entomological Society 17: 1 - 16." type="journal article" year="1890">Horn 1890: 11</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<taxonomicName id="4C489C8D98270B22FF19FEBB4F471B19" authority="(Horn)" baseAuthorityName="Horn" baseAuthorityYear="1890" box="[192,516,322,346]" class="Insecta" family="Heteroceridae" genus="Peditatus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
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<emphasis id="B93C3B1C98270B22FF19FEBB4CE61B19" box="[192,421,322,346]" italics="true" pageId="28" pageNumber="27">Peditatus schwarzi</emphasis>
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(Horn)
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</taxonomicName>
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:
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<treatmentCitation id="0AE9C11F98270B22FDD7FEBB4FB31B19" author="Pacheco, F." box="[526,752,322,346]" page="123" pageId="28" pageNumber="27" year="1964">
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<bibRefCitation id="EFD99AFF98270B22FDD7FEBB4FB31B19" author="Pacheco, F." box="[526,752,322,346]" pageId="28" pageNumber="27" pagination="1 - 155" refId="ref22030" refString="Pacheco, F. 1964. Sistematica, filogenia y distribucion de los Heteroceridos de America (Coleoptera; Heteroceridae). Monografias del Colegio de Post-graduados, No. 1, Escuela Nacional de Agricultura, Chapingo, Mexico, Vol. 1: 1 - 155." type="journal article" year="1964">Pacheco 1964: 123</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C352B48598270B22FF19FE7B4F3218D9" pageId="28" pageNumber="27" type="description">
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<paragraph id="8BF7E70E98270B22FF19FE7B4F3218D9" blockId="28.[192,1441,386,666]" pageId="28" pageNumber="27">
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<emphasis id="B93C3B1C98270B22FF19FE7B4C2E1BD9" bold="true" box="[192,365,386,410]" pageId="28" pageNumber="27">Description.</emphasis>
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Length 3.0 -
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<quantity id="4CB04AEB98270B22FDC0FE7B4F391BD9" box="[537,634,386,410]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="28" pageNumber="27" unit="mm" value="3.7">3.7 mm</quantity>
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. Brown or reddish-brown. Elytra trifasciate, pale markings well defined, elytral surface with relatively dense pubescence; striae absent (
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<figureCitation id="1373FB8B98270B22FBCCFE5B49291BF9" box="[1045,1130,418,442]" captionStart="Figures 21-25" captionStartId="18.[192,286,1256,1278]" captionTargetBox="[192,1401,191,1233]" captionTargetPageId="18" captionText="Figures 21-25. Habiti of Heterocerus species of intermediate size with post-mesocoxal lines. 21) H. schwarzi. 22) H. tenuis. 23) H. undatus. 24) H. texanus. 25) H. collaris. Scale line = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5188374" httpUri="https://zenodo.org/record/5188374/files/figure.png" pageId="28" pageNumber="27">Fig. 21</figureCitation>
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). Post-metathoracic coxal lines absent, post-mesothoracic coxal lines present. Male genitalia (
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<figureCitation id="1373FB8B98270B22FBC9FE3B49291B99" box="[1040,1130,450,474]" captionStart="Figures 45-53" captionStartId="39.[192,286,843,865]" captionTargetBox="[203,1417,196,805]" captionTargetId="figure-157@39.[770,867,432,529]" captionTargetPageId="39" captionText="Figures 45-53. Male genitalia of southeastern species. The ventral view is shown unless otherwise stated. 45-50) Heterocerus species in the undulatus group. 45) H. undatus. 46) H. collaris. 47) H. texanus; left - ventral view, right - lateral view. 48) H. schwarzi; left - ventral view, right - lateral view. 49) H. tenuis. 50) H. selanderi. 51-53) Augyles and Tropicus species. Figures 52-53 are at roughly 2x relative scale of prior genitalia. 51) Augyles auromicans. 52) Tropicus nigrellus; left - dorsal view, center - ventral view, right - lateral view. 53) Tropicus pusillus; left - dorsal view, right - ventral view." figureDoi="http://doi.org/10.5281/zenodo.5188384" httpUri="https://zenodo.org/record/5188384/files/figure.png" pageId="28" pageNumber="27">Fig. 48</figureCitation>
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) of the
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<taxonomicName id="4C489C8D98270B22FB17FE3A48191B99" authorityName="Melsheimer" authorityYear="1844" box="[1230,1370,450,474]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="undatus">
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<emphasis id="B93C3B1C98270B22FB17FE3A48191B99" box="[1230,1370,450,474]" italics="true" pageId="28" pageNumber="27">H. undatus</emphasis>
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</taxonomicName>
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<typeStatus id="54F359AC98270B22FABDFE3A48DF1B98" box="[1380,1436,451,475]" pageId="28" pageNumber="27">type</typeStatus>
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, slightly more than three times as long as greatest width; body of phallobase dominated by median plate; each third of plate progressively wider anteriorly; anterior edge with a rounded central lobe and a dorsal spine-like projection on either side visible only in lateral view (
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<figureCitation id="1373FB8B98270B22FC7EFDDB4EBE1879" box="[935,1021,546,570]" captionStart="Figures 45-53" captionStartId="39.[192,286,843,865]" captionTargetBox="[203,1417,196,805]" captionTargetId="figure-157@39.[770,867,432,529]" captionTargetPageId="39" captionText="Figures 45-53. Male genitalia of southeastern species. The ventral view is shown unless otherwise stated. 45-50) Heterocerus species in the undulatus group. 45) H. undatus. 46) H. collaris. 47) H. texanus; left - ventral view, right - lateral view. 48) H. schwarzi; left - ventral view, right - lateral view. 49) H. tenuis. 50) H. selanderi. 51-53) Augyles and Tropicus species. Figures 52-53 are at roughly 2x relative scale of prior genitalia. 51) Augyles auromicans. 52) Tropicus nigrellus; left - dorsal view, center - ventral view, right - lateral view. 53) Tropicus pusillus; left - dorsal view, right - ventral view." figureDoi="http://doi.org/10.5281/zenodo.5188384" httpUri="https://zenodo.org/record/5188384/files/figure.png" pageId="28" pageNumber="27">Fig. 48</figureCitation>
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); lateral arms of phallobase greatly reduced; dorsal plate not distinct, instead dorsal surface with several distinct sclerites; parameres membranous and obscured by a sclerotized plate on the ventral side of the anterior tip of the genitalia. Hypermandibulate males unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C352B48598270B22FF19FD3B48491FF9" pageId="28" pageNumber="27" type="diagnosis">
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<paragraph id="8BF7E70E98270B22FF19FD3B4FCC19F9" blockId="28.[192,1441,706,954]" pageId="28" pageNumber="27">
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<emphasis id="B93C3B1C98270B22FF19FD3B4C131899" bold="true" box="[192,336,706,730]" pageId="28" pageNumber="27">Diagnosis.</emphasis>
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<taxonomicName id="4C489C8D98270B22FE80FD3A4F1C1899" box="[345,607,706,730]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
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<emphasis id="B93C3B1C98270B22FE80FD3A4F1C1899" box="[345,607,706,730]" italics="true" pageId="28" pageNumber="27">Heterocerus schwarzi</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="4C489C8D98270B22FCD4FD3A4ED21899" authorityName="King & Lago" authorityYear="2012" baseAuthorityName="King & Lago" baseAuthorityYear="2012" box="[781,913,707,730]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="texanus">
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<emphasis id="B93C3B1C98270B22FCD4FD3A4ED21899" box="[781,913,707,730]" italics="true" pageId="28" pageNumber="27">H. texanus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C489C8D98270B22FC0DFD3A49051899" authorityName="Miller" authorityYear="1988" box="[980,1094,707,730]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="tenuis">
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<emphasis id="B93C3B1C98270B22FC0DFD3A49051899" box="[980,1094,707,730]" italics="true" pageId="28" pageNumber="27">H. tenuis</emphasis>
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</taxonomicName>
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. It generally can be distinguished from
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<taxonomicName id="4C489C8D98270B22FEBDFD1A4CA018B9" authorityName="King & Lago" authorityYear="2012" baseAuthorityName="King & Lago" baseAuthorityYear="2012" box="[356,483,739,762]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="texanus">
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<emphasis id="B93C3B1C98270B22FEBDFD1A4CA018B9" box="[356,483,739,762]" italics="true" pageId="28" pageNumber="27">H. texanus</emphasis>
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</taxonomicName>
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using external characters if specimens of the two are compared simultaneously;
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<taxonomicName id="4C489C8D98270B22FF19FCFA4C101959" box="[192,339,770,794]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
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<emphasis id="B93C3B1C98270B22FF19FCFA4C101959" box="[192,339,770,794]" italics="true" pageId="28" pageNumber="27">H. schwarzi</emphasis>
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</taxonomicName>
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has elytra with narrower pale markings, a much more punctuate surface and has more pubescence than
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<taxonomicName id="4C489C8D98270B22FE4DFCDA4F5B1979" authorityName="King & Lago" authorityYear="2012" baseAuthorityName="King & Lago" baseAuthorityYear="2012" box="[404,536,803,826]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="texanus">
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<emphasis id="B93C3B1C98270B22FE4DFCDA4F5B1979" box="[404,536,803,826]" italics="true" pageId="28" pageNumber="27">H. texanus</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C489C8D98270B22FDFFFCDA4E681979" box="[550,811,802,826]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
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<emphasis id="B93C3B1C98270B22FDFFFCDA4E681979" box="[550,811,802,826]" italics="true" pageId="28" pageNumber="27">Heterocerus schwarzi</emphasis>
|
|
</taxonomicName>
|
|
is nearly identical to
|
|
<taxonomicName id="4C489C8D98270B22FBE2FCDA49E81979" authorityName="Miller" authorityYear="1988" box="[1083,1195,803,826]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="tenuis">
|
|
<emphasis id="B93C3B1C98270B22FBE2FCDA49E81979" box="[1083,1195,803,826]" italics="true" pageId="28" pageNumber="27">H. tenuis</emphasis>
|
|
</taxonomicName>
|
|
, and male genitalia must be used to distinguish them. The dorsal spine-like projections on either side of the anterior edge of the median plate point outward and are easily seen in the ventral view of the genitalia in
|
|
<taxonomicName id="4C489C8D98270B22FB2DFC9A48221939" authorityName="Miller" authorityYear="1988" box="[1268,1377,867,890]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="tenuis">
|
|
<emphasis id="B93C3B1C98270B22FB2DFC9A48221939" box="[1268,1377,867,890]" italics="true" pageId="28" pageNumber="27">H. tenuis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="1373FB8B98270B22FAB6FC9B4DA219D9" captionStart="Figures 45-53" captionStartId="39.[192,286,843,865]" captionTargetBox="[203,1417,196,805]" captionTargetId="figure-157@39.[770,867,432,529]" captionTargetPageId="39" captionText="Figures 45-53. Male genitalia of southeastern species. The ventral view is shown unless otherwise stated. 45-50) Heterocerus species in the undulatus group. 45) H. undatus. 46) H. collaris. 47) H. texanus; left - ventral view, right - lateral view. 48) H. schwarzi; left - ventral view, right - lateral view. 49) H. tenuis. 50) H. selanderi. 51-53) Augyles and Tropicus species. Figures 52-53 are at roughly 2x relative scale of prior genitalia. 51) Augyles auromicans. 52) Tropicus nigrellus; left - dorsal view, center - ventral view, right - lateral view. 53) Tropicus pusillus; left - dorsal view, right - ventral view." figureDoi="http://doi.org/10.5281/zenodo.5188384" httpUri="https://zenodo.org/record/5188384/files/figure.png" pageId="28" pageNumber="27">Fig. 49</figureCitation>
|
|
), but curve dorsally in
|
|
<taxonomicName id="4C489C8D98270B22FE25FC7A4FCE19D9" box="[508,653,898,922]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
|
|
<emphasis id="B93C3B1C98270B22FE25FC7A4FCE19D9" box="[508,653,898,922]" italics="true" pageId="28" pageNumber="27">H. schwarzi</emphasis>
|
|
</taxonomicName>
|
|
and thus are not visible in the ventral view. The ranges of these two species apparently do not overlap.
|
|
</paragraph>
|
|
<paragraph id="8BF7E70E98270B22FF19FC1B49A11E79" blockId="28.[192,1440,994,1082]" pageId="28" pageNumber="27">
|
|
<emphasis id="B93C3B1C98270B22FF19FC1B4C5419B9" bold="true" box="[192,279,994,1018]" pageId="28" pageNumber="27">Notes.</emphasis>
|
|
This species is rarely encountered in the Southeast. Only
|
|
<specimenCount id="9D4E2C8798270B22FC02FC1A49C319B9" box="[987,1152,994,1018]" pageId="28" pageNumber="27" type="generic">one specimen</specimenCount>
|
|
was examined from the study area and was taken along the banks of the
|
|
<collectingRegion id="498C29EC98270B22FCD5FBFB4ED71E59" box="[780,916,1026,1050]" country="United States of America" name="Mississippi" pageId="28" pageNumber="27">Mississippi</collectingRegion>
|
|
River in
|
|
<collectingRegion id="498C29EC98270B22FBDAFBFB491D1E59" box="[1027,1118,1026,1050]" country="Ecuador" name="Bolivar" pageId="28" pageNumber="27">Bolivar</collectingRegion>
|
|
County,
|
|
<collectingRegion id="498C29EC98270B22FB11FBFB48171E59" box="[1224,1364,1026,1050]" country="United States of America" name="Mississippi" pageId="28" pageNumber="27">Mississippi</collectingRegion>
|
|
. Additional specimens were seen from two localities along the
|
|
<collectingRegion id="498C29EC98270B22FCA9FBDB4EBB1E79" box="[880,1016,1058,1082]" country="United States of America" name="Mississippi" pageId="28" pageNumber="27">Mississippi</collectingRegion>
|
|
River in
|
|
<collectingRegion id="498C29EC98270B22FBBEFBDB499D1E79" box="[1127,1246,1058,1082]" country="United States of America" name="Arkansas" pageId="28" pageNumber="27">Arkansas</collectingRegion>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BF7E70E98270B22FF19FB9B4CFD1F19" blockId="28.[192,1441,1122,1466]" pageId="28" pageNumber="27">
|
|
<emphasis id="B93C3B1C98270B22FF19FB9B4F621E39" bold="true" box="[192,545,1122,1146]" pageId="28" pageNumber="27">
|
|
Designation of
|
|
<typeStatus id="54F359AC98270B22FE56FB9B4F581E39" box="[399,539,1122,1146]" pageId="28" pageNumber="27" type="lectotype">Lectotype</typeStatus>
|
|
.
|
|
</emphasis>
|
|
The Museum of Comparative Zoology (MCZ) at Harvard University inherited the Horn collection from the Academy of Natural Sciences in Philadelphia during the 1960s. The material held by the MCZ was examined and
|
|
<bibRefCitation id="EFD99AFF98270B22FD33FB5A4EC11EF9" author="Horn, G. H." box="[746,898,1186,1210]" pageId="28" pageNumber="27" pagination="1 - 16" refId="ref20843" refString="Horn, G. H. 1890. The species of Heterocerus of Boreal America. Transactions of the American Entomological Society 17: 1 - 16." type="journal article" year="1890">Horn (1890)</bibRefCitation>
|
|
did not establish a
|
|
<typeStatus id="54F359AC98270B22FBABFB5B499F1EF9" box="[1138,1244,1186,1210]" pageId="28" pageNumber="27" type="holotype">holotype</typeStatus>
|
|
for this species. One specimen was previously labeled as a
|
|
<typeStatus id="54F359AC98270B22FD0DFB3B4E001E99" box="[724,835,1218,1242]" pageId="28" pageNumber="27" type="lectotype">lectotype</typeStatus>
|
|
although this designation was apparently never published. The male specimen numbered MCZT_32152 (also labeled
|
|
<typeStatus id="54F359AC98270B22FC35FB1A49211EB8" box="[1004,1122,1251,1275]" pageId="28" pageNumber="27" type="lectotype">Lectotype</typeStatus>
|
|
3271) is
|
|
<emphasis id="B93C3B1C98270B22FB11FB1B48E31EB9" bold="true" box="[1224,1440,1250,1274]" pageId="28" pageNumber="27">here designated</emphasis>
|
|
as the
|
|
<emphasis id="B93C3B1C98270B22FED4FAFB4CCD1F59" bold="true" box="[269,398,1282,1306]" pageId="28" pageNumber="27">
|
|
<typeStatus id="54F359AC98270B22FED4FAFB4CCD1F59" box="[269,398,1282,1306]" pageId="28" pageNumber="27" type="lectotype">lectotype</typeStatus>
|
|
</emphasis>
|
|
of
|
|
<taxonomicName id="4C489C8D98270B22FE69FAFA4E7B1F59" authority="Horn. One" authorityName="Horn. One" box="[432,824,1282,1306]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
|
|
<emphasis id="B93C3B1C98270B22FE69FAFA4FF21F59" box="[432,689,1282,1306]" italics="true" pageId="28" pageNumber="27">Heterocerus schwarzi</emphasis>
|
|
Horn. One
|
|
</taxonomicName>
|
|
female specimen with the same collection data and labeled Para-Type 3271.2 is here designated
|
|
<typeStatus id="54F359AC98270B22FD3FFADB4ED21F79" box="[742,913,1314,1338]" pageId="28" pageNumber="27" type="paralectotype">paralectotype</typeStatus>
|
|
. Burnett County,
|
|
<collectingRegion id="498C29EC98270B22FBB5FADA49F01F79" box="[1132,1203,1315,1338]" country="United States of America" name="Texas" pageId="28" pageNumber="27">Texas</collectingRegion>
|
|
is fixed as the type locality (
|
|
<bibRefCitation id="EFD99AFF98270B22FEF0FABB4CF11F19" author="Horn, G. H." box="[297,434,1346,1370]" pageId="28" pageNumber="27" pagination="1 - 16" refId="ref20843" refString="Horn, G. H. 1890. The species of Heterocerus of Boreal America. Transactions of the American Entomological Society 17: 1 - 16." type="journal article" year="1890">Horn 1890</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BF7E70E98270B22FF29FA9B48491FF9" blockId="28.[192,1441,1122,1466]" pageId="28" pageNumber="27">
|
|
Labels attached to the
|
|
<typeStatus id="54F359AC98270B22FE26FA9B4F2D1F39" box="[511,622,1378,1402]" pageId="28" pageNumber="27" type="lectotype">lectotype</typeStatus>
|
|
are: Top - Burnett Co
|
|
<collectingRegion id="498C29EC98270B22FCA3FA9A4E871F39" box="[890,964,1379,1402]" country="United States of America" name="Texas" pageId="28" pageNumber="27">Texas</collectingRegion>
|
|
, 2
|
|
<superScript id="7C3D4A4698270B22FC04FA984EBB1F39" attach="none" box="[989,1016,1377,1402]" fontSize="6" pageId="28" pageNumber="27">nd-</superScript>
|
|
(male symbol), 3
|
|
<superScript id="7C3D4A4698270B22FB1DFA9849971F2C" attach="left" box="[1220,1236,1377,1391]" fontSize="6" pageId="28" pageNumber="27">rd</superScript>
|
|
-
|
|
<typeStatus id="54F359AC98270B22FB3EFA9A481C1F38" box="[1255,1375,1379,1403]" pageId="28" pageNumber="27" type="lectotype">Lectotype</typeStatus>
|
|
3271 (red), 4
|
|
<superScript id="7C3D4A4698270B22FEC0FA784C6B1FCC" attach="left" box="[281,296,1409,1423]" fontSize="6" pageId="28" pageNumber="27">th</superScript>
|
|
<emphasis id="B93C3B1C98270B22FEF7FA7A4C741FD9" box="[302,311,1411,1434]" italics="true" pageId="28" pageNumber="27">-</emphasis>
|
|
<taxonomicName id="4C489C8D98270B22FEE4FA7A4F5E1FD9" authority="Horn" authorityName="Horn" box="[317,541,1410,1434]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="10" phylum="Arthropoda" rank="species" species="schwarzi">
|
|
<emphasis id="B93C3B1C98270B22FEE4FA7A4C961FD9" box="[317,469,1410,1434]" italics="true" pageId="28" pageNumber="27">H. Schwarzi</emphasis>
|
|
Horn
|
|
</taxonomicName>
|
|
(hand written), 5
|
|
<superScript id="7C3D4A4698270B22FD2DFA784E401FCC" attach="left" box="[756,771,1409,1423]" fontSize="6" pageId="28" pageNumber="27">th</superScript>
|
|
<emphasis id="B93C3B1C98270B22FCD3FA7A4E501FD9" box="[778,787,1411,1434]" italics="true" pageId="28" pageNumber="27">-</emphasis>
|
|
M.C.Z. type 32152 (red),
|
|
<date id="FFF6C1CE98270B22FB9EFA7B48071FD9" box="[1095,1348,1409,1435]" pageId="28" pageNumber="27" value="2004-08-06" valueMax="2004-12-06" valueMin="2004-08-06">
|
|
6
|
|
<superScript id="7C3D4A4698270B22FB8FFA7849261FCC" attach="left" box="[1110,1125,1409,1423]" fontSize="6" pageId="28" pageNumber="27">th</superScript>
|
|
- Aug. - Dec. 2004
|
|
</date>
|
|
, M.C.Z. image Database, 7
|
|
<superScript id="7C3D4A4698270B22FE46FA584CED1FEC" attach="left" box="[415,430,1441,1455]" fontSize="6" pageId="28" pageNumber="27">th</superScript>
|
|
<emphasis id="B93C3B1C98270B22FE6AFA5A4CFF1FF9" box="[435,444,1443,1466]" italics="true" pageId="28" pageNumber="27">-</emphasis>
|
|
<typeStatus id="54F359AC98270B22FE1AFA5B4F2C1FF9" box="[451,623,1442,1466]" pageId="28" pageNumber="27" type="lectotype">LECTOTYPE</typeStatus>
|
|
<taxonomicName id="4C489C8D98270B22FDADFA5A49481FF9" authority="Horn 1890" authorityName="Horn" authorityYear="1890" box="[628,1035,1442,1466]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
|
|
Heterocerus schwarzi
|
|
<bibRefCitation id="EFD99AFF98270B22FC5DFA5A49481FF9" author="Horn, G. H." box="[900,1035,1442,1466]" pageId="28" pageNumber="27" pagination="1 - 16" refId="ref20843" refString="Horn, G. H. 1890. The species of Heterocerus of Boreal America. Transactions of the American Entomological Society 17: 1 - 16." type="journal article" year="1890">Horn 1890</bibRefCitation>
|
|
</taxonomicName>
|
|
, Jonas G. King (red).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C352B48598270B22FF19FA1B4FFA1C19" pageId="28" pageNumber="27" type="distribution">
|
|
<paragraph id="8BF7E70E98270B22FF19FA1B487F1C59" blockId="28.[192,1440,1506,1626]" pageId="28" pageNumber="27">
|
|
<emphasis id="B93C3B1C98270B22FF19FA1B4C361FB9" bold="true" box="[192,373,1506,1530]" pageId="28" pageNumber="27">Distribution.</emphasis>
|
|
<taxonomicName id="4C489C8D98270B22FEA6FA1A4FC71FB9" box="[383,644,1506,1530]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="schwarzi">
|
|
<emphasis id="B93C3B1C98270B22FEA6FA1A4FC71FB9" box="[383,644,1506,1530]" italics="true" pageId="28" pageNumber="27">Heterocerus schwarzi</emphasis>
|
|
</taxonomicName>
|
|
occurs from the northeastern
|
|
<collectingCountry id="F35FA79E98270B22FBDAFA1B49F31FB9" box="[1027,1200,1506,1530]" name="United States of America" pageId="28" pageNumber="27">United States</collectingCountry>
|
|
, southeast through
|
|
<collectingRegion id="498C29EC98270B22FF19F9FB4C631C59" box="[192,288,1538,1562]" country="United States of America" name="Indiana" pageId="28" pageNumber="27">Indiana</collectingRegion>
|
|
into northern
|
|
<collectingRegion id="498C29EC98270B22FE0FF9FA4F631C59" box="[470,544,1539,1562]" country="United States of America" name="Texas" pageId="28" pageNumber="27">Texas</collectingRegion>
|
|
,
|
|
<collectingRegion id="498C29EC98270B22FDF3F9FB4FE31C59" box="[554,672,1538,1562]" country="United States of America" name="Arkansas" pageId="28" pageNumber="27">Arkansas</collectingRegion>
|
|
, northern
|
|
<collectingRegion id="498C29EC98270B22FCF8F9FB4ED81C59" box="[801,923,1538,1562]" country="United States of America" name="Louisiana" pageId="28" pageNumber="27">Louisiana</collectingRegion>
|
|
and western
|
|
<collectingRegion id="498C29EC98270B22FBE6F9FB49841C59" box="[1087,1223,1538,1562]" country="United States of America" name="Mississippi" pageId="28" pageNumber="27">Mississippi</collectingRegion>
|
|
(
|
|
<figureCitation id="1373FB8B98270B22FB0FF9FB486D1C59" box="[1238,1326,1538,1562]" captionStart="Figure 66-72" captionStartId="41.[192,275,1784,1806]" captionTargetBox="[189,1438,191,1770]" captionTargetId="figure-840@41.[787,836,959,983]" captionTargetPageId="41" captionText="Figure 66-72. Distribution by county in the United States. 66) Heterocerus undatus. 67) Heterocerus texanus. 68) Heterocerus schwarzi. 69) Heterocerus tenuis. 70) Augyles auromicans. 71) Tropicus pusillus. 72) Tropicus nigrellus." figureDoi="http://doi.org/10.5281/zenodo.5188396" httpUri="https://zenodo.org/record/5188396/files/figure.png" pageId="28" pageNumber="27">Fig. 68</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BF7E70E98270B22FF19F9BB4FFA1C19" blockId="28.[192,1440,1506,1626]" box="[192,697,1602,1626]" pageId="28" pageNumber="27">
|
|
<emphasis id="B93C3B1C98270B22FF19F9BB4CA71C19" bold="true" box="[192,484,1602,1626]" pageId="28" pageNumber="27">Specimens examined.</emphasis>
|
|
7 (See Appendix).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |