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<mods:title id="6682DCF0A210AA5AB50CC8DC5BB34774">Redescriptions of North American Epidamaeus (Acari, Oribatida, Damaeidae) species proposed by N. Banks, H. E. Ewing, A. P. Jacot, and J. W. Wilson</mods:title>
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<taxonomicName id="4C6732C2FF99305AFF18A095FDC97E97" ID-CoL="8TBQR" authority="(Jacot, 1939)" baseAuthorityName="Jacot" baseAuthorityYear="1939" box="[151,604,151,178]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="craigheadi">
<emphasis id="B9139553FF99305AFF18A095FDC97E97" bold="true" box="[151,604,151,178]" pageId="49" pageNumber="50">
<emphasis id="B9139553FF99305AFF18A095FE207E94" bold="true" box="[151,437,151,178]" italics="true" pageId="49" pageNumber="50">Epidamaeus craigheadi</emphasis>
(
<bibRefCitation id="EFF634B0FF99305AFE4AA095FDC17E97" author="Jacot, A. P." box="[453,596,151,177]" pageId="49" pageNumber="50" pagination="321 - 332" refId="ref37962" refString="Jacot, A. P. (1939) New mites from the White Mountains. Occasional Papers of the Boston Society of Natural History, 8, 321 - 332." type="journal article" year="1939">Jacot, 1939</bibRefCitation>
)
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</paragraph>
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<subSubSection id="C37D1ACAFF99305AFF18A0B9FEB27EF0" box="[151,295,187,214]" pageId="49" pageNumber="50" type="description">
<paragraph id="8BD84941FF99305AFF18A0B9FEB27EF0" blockId="49.[151,604,151,214]" box="[151,295,187,214]" pageId="49" pageNumber="50">
(
<figureCitation id="135C55C4FF99305AFF10A0B9FE8A7EF0" box="[159,287,187,214]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStart-2="FIGURE 27" captionStart-3="FIGURE 28" captionStart-4="FIGURE 29" captionStart-5="FIGURE 30" captionStart-6="FIGURE 31" captionStart-7="FIGURE 32" captionStartId-0="51.[151,250,1943,1967]" captionStartId-1="52.[151,250,1822,1846]" captionStartId-2="53.[151,250,1858,1882]" captionStartId-3="55.[151,250,989,1013]" captionStartId-4="56.[151,250,1894,1918]" captionStartId-5="57.[151,250,1809,1833]" captionStartId-6="58.[151,250,1834,1858]" captionStartId-7="59.[151,250,1462,1486]" captionTargetBox-0="[220,1368,183,1907]" captionTargetBox-1="[171,1415,181,1798]" captionTargetBox-2="[157,1429,181,1834]" captionTargetBox-3="[159,1424,187,958]" captionTargetBox-4="[166,1419,194,1858]" captionTargetBox-5="[171,1415,181,1785]" captionTargetBox-6="[156,1430,380,1809]" captionTargetBox-7="[164,1423,844,1439]" captionTargetId-0="figure-17@51.[205,1382,181,1919]" captionTargetId-1="figure-17@52.[171,1415,181,1798]" captionTargetId-2="figure-17@53.[157,1429,181,1834]" captionTargetId-3="figure-17@55.[151,1436,181,966]" captionTargetId-4="figure-17@56.[151,1436,181,1871]" captionTargetId-5="figure-17@57.[171,1415,181,1785]" captionTargetId-6="figure-102@58.[156,1430,380,1809]" captionTargetId-7="figure-307@59.[164,1423,844,1439]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="53" captionTargetPageId-3="55" captionTargetPageId-4="56" captionTargetPageId-5="57" captionTargetPageId-6="58" captionTargetPageId-7="59" captionText-0="FIGURE 25. Epidamaeus craigheadi (Jacot 1939), adult: A—dorsal view (legs except right trochanters IIII not shown); B—anterior half of body, right lateral view (gnathosoma and legs not shown); C—posterior part of body, right lateral view; D—ventral view (legs represented only by trochanters). Scale bar 100 μm." captionText-1="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." captionText-2="FIGURE 27. Epidamaeus craigheadi (Jacot 1939), light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta lʹ labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E)." captionText-3="FIGURE 28. Epidamaeus craigheadi (Jacot 1939), adult: A—subcapitulum, ventral view; B—palp, right, abaxial view; C— chelicera, left, adaxial view (seta chb on opposite side, not labeled). Scale bars 20 μm." captionText-4="FIGURE 29. Epidamaeus craigheadi (Jacot 1939), adult:A—right leg I, except trochanter, abaxial view (femur slightly twisted, cf Fig. 27E); B—femur, genu and tibia of right leg II, abaxial view; C—femur, genu and tibia of left leg III, abaxial view; D— left leg IV, except trochanter, abaxial view. Scale bar 50 μm." captionText-5="FIGURE 30. Epidamaeus craigheadi (Jacot 1939), light photographs of nymphs: A—protonymph, lateral view; B—same, posterior region of gastronotum (left insert = enlargement of attachment cornicle k inserting in induration (ind) on underside of larval scalp; C—protonymph, dorsal view of hysterosoma with larval scalp intact (white labels on scalp setae; insert = larval scalp removed from different specimen, arrow to attachment induration); D—tritonymph, dorsal view (scalps removed; several setae broken; body artificially swollen by clearing); E—tritonymph, lateral view (scalps removed; lower left insert = enlargement of attachment cornicle k). Scale bars 20 μm (B, C), 50 μm (A, D, E)." captionText-6="FIGURE 31. Epidamaeus craigheadi (Jacot 1939), light photographs of protonymph (A), tritonymph (BG):A—lateral view of" captionText-7="FIGURE 32. Epidamaeus craigheadi (Jacot 1939), light photographs of deutonymph (A), tritonymph (BE):A—middle portion of leg II, abaxial view; B—middle portion of right leg I, dorsoadaxial view; C—portion of tibia I from different specimen, abaxial view (arrow to coupled seta d); D—bulb of left tarsus I, dorsoabaxial view, showing tarsal cluster (lower right insert = isolated famulus, arrow to basal root); E—isolated solenidion ω1 and cerotegument excrescences. Scale bars 5 μm (D, E), 10 μm (AC)." figureDoi-0="http://doi.org/10.5281/zenodo.5225245" figureDoi-1="http://doi.org/10.5281/zenodo.5225247" figureDoi-2="http://doi.org/10.5281/zenodo.5225249" figureDoi-3="http://doi.org/10.5281/zenodo.5225251" figureDoi-4="http://doi.org/10.5281/zenodo.5225253" figureDoi-5="http://doi.org/10.5281/zenodo.5225255" figureDoi-6="http://doi.org/10.5281/zenodo.5225257" figureDoi-7="http://doi.org/10.5281/zenodo.5225259" httpUri-0="https://zenodo.org/record/5225245/files/figure.png" httpUri-1="https://zenodo.org/record/5225247/files/figure.png" httpUri-2="https://zenodo.org/record/5225249/files/figure.png" httpUri-3="https://zenodo.org/record/5225251/files/figure.png" httpUri-4="https://zenodo.org/record/5225253/files/figure.png" httpUri-5="https://zenodo.org/record/5225255/files/figure.png" httpUri-6="https://zenodo.org/record/5225257/files/figure.png" httpUri-7="https://zenodo.org/record/5225259/files/figure.png" pageId="49" pageNumber="50">Figs 2532</figureCitation>
)
</paragraph>
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<paragraph id="8BD84941FF99305AFF18A0F8FDAB7F34" blockId="49.[151,868,250,336]" box="[151,574,250,274]" pageId="49" pageNumber="50">
<taxonomicName id="4C6732C2FF99305AFF18A0F8FE607F34" authority="Jacot, 1939" authorityName="Jacot" authorityYear="1939" box="[151,501,250,274]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="craigheadi">
<emphasis id="B9139553FF99305AFF18A0F8FEED7F34" box="[151,376,250,274]" italics="true" pageId="49" pageNumber="50">Metabelba craigheadi</emphasis>
<bibRefCitation id="EFF634B0FF99305AFEF1A0F9FE607F34" author="Jacot, A. P." box="[382,501,250,274]" pageId="49" pageNumber="50" pagination="321 - 332" refId="ref37962" refString="Jacot, A. P. (1939) New mites from the White Mountains. Occasional Papers of the Boston Society of Natural History, 8, 321 - 332." type="journal article" year="1939">Jacot, 1939</bibRefCitation>
</taxonomicName>
; p. 324
</paragraph>
<paragraph id="8BD84941FF99305AFF18A11BFCAC7F17" blockId="49.[151,868,250,336]" box="[151,825,281,305]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="AB776E6FFF99305AFF18A11BFCAC7F17" box="[151,825,281,305]" pageId="49" pageNumber="50">
<taxonomicName id="4C6732C2FF99305AFF18A11BFD837F17" ID-CoL="8TBQR" authority="(Jacot, 1939)" baseAuthorityName="Jacot" baseAuthorityYear="1939" box="[151,534,281,305]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="craigheadi">
<emphasis id="B9139553FF99305AFF18A11BFE1C7F17" box="[151,393,281,305]" italics="true" pageId="49" pageNumber="50">Epidamaeus craigheadi</emphasis>
(
<bibRefCitation id="EFF634B0FF99305AFE19A11BFD9B7F17" author="Jacot, A. P." box="[406,526,281,305]" pageId="49" pageNumber="50" pagination="321 - 332" refId="ref37962" refString="Jacot, A. P. (1939) New mites from the White Mountains. Occasional Papers of the Boston Society of Natural History, 8, 321 - 332." type="journal article" year="1939">Jacot, 1939</bibRefCitation>
)
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;
<treatmentCitation id="0AC66F50FF99305AFDAFA11BFCAC7F17" author="Marshall, V. G. &amp; Reeves, R. M. &amp; Norton, R. A." box="[544,825,281,305]" page="139" pageId="49" pageNumber="50" year="1987">
<bibRefCitation id="EFF634B0FF99305AFDAFA11BFD7A7F17" author="Marshall, V. G. &amp; Reeves, R. M. &amp; Norton, R. A." box="[544,751,281,305]" pageId="49" pageNumber="50" pagination="1 - 418" refId="ref38119" refString="Marshall, V. G., Reeves, R. M. &amp; Norton, R. A. (1987) Catalogue of the Oribatida of continental United States and Canada. Memoirs of the Entomological Society of Canada, 139, 1 - 418. https: // doi. org / 10.4039 / entm 119139 fv" type="journal article" year="1987">
Marshall
<emphasis id="B9139553FF99305AFD0EA118FD387F17" box="[641,685,281,305]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. 1987
</bibRefCitation>
, p. 139
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</paragraph>
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<treatmentCitationGroup id="AB776E6FFF99305AFF18A13AFCF17F76" box="[151,868,312,336]" pageId="49" pageNumber="50">
<taxonomicName id="4C6732C2FF99305AFF18A13AFD187F76" ID-CoL="7SZP8" authority="(Jacot, 1939)" baseAuthorityName="Jacot" baseAuthorityYear="1939" box="[151,653,312,336]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="craigheadi" subGenus="Epidamaeus">
<emphasis id="B9139553FF99305AFF18A13AFF627F69" box="[151,247,312,335]" italics="true" pageId="49" pageNumber="50">Damaeus</emphasis>
(
<emphasis id="B9139553FF99305AFE89A13AFE167F76" box="[262,387,312,336]" italics="true" pageId="49" pageNumber="50">Epidamaeus</emphasis>
)
<emphasis id="B9139553FF99305AFE1DA13AFD957F76" box="[402,512,312,336]" italics="true" pageId="49" pageNumber="50">craigheadi</emphasis>
(
<bibRefCitation id="EFF634B0FF99305AFD82A13AFD107F76" author="Jacot, A. P." box="[525,645,312,336]" pageId="49" pageNumber="50" pagination="321 - 332" refId="ref37962" refString="Jacot, A. P. (1939) New mites from the White Mountains. Occasional Papers of the Boston Society of Natural History, 8, 321 - 332." type="journal article" year="1939">Jacot, 1939</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="0AC66F50FF99305AFD18A13AFCC87F76" author="Subias, L. S." box="[663,861,312,336]" page="79" pageId="49" pageNumber="50" year="2004">
<bibRefCitation id="EFF634B0FF99305AFD18A13AFC8B7F76" author="Subias, L. S." box="[663,798,312,336]" pageId="49" pageNumber="50" pagination="3 - 305" refId="ref40172" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60 (Numero Extraordinario), 3 - 305. https: // doi. org / 10.3989 / graellsia. 2004. v 60. iExtra. 218" type="journal article" year="2004">Subías (2004</bibRefCitation>
, p. 79
</treatmentCitation>
)
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</paragraph>
</subSubSection>
<subSubSection id="C37D1ACAFF99305DFF18A17AFC837C1B" lastPageId="54" lastPageNumber="55" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="8BD84941FF99305AFF18A17AFE9F7FB4" blockId="49.[151,1437,376,725]" box="[151,266,376,402]" pageId="49" pageNumber="50">
<emphasis id="B9139553FF99305AFF18A17AFE9F7FB4" bold="true" box="[151,266,376,402]" pageId="49" pageNumber="50">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD84941FF99305AFF18A19EFE437CF3" blockId="49.[151,1437,376,725]" pageId="49" pageNumber="50">
<taxonomicName id="4C6732C2FF99305AFF18A19EFEEC7F90" box="[151,377,412,438]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9139553FF99305AFF18A19EFEB47F90" box="[151,289,412,438]" italics="true" pageId="49" pageNumber="50">Epidamaeus</emphasis>
species
</taxonomicName>
with total length of adult 365426. Cerotegument excrescences ranging from granular to short, vermiform filaments; procuticle distinctly microtuberculate on much of body; without exuvial scalps or adherent debris. Propodolateral apophysis absent; postbothridial enantiophysis unbalanced:
<emphasis id="B9139553FF99305AFBD1A1E6FBE87FDB" box="[1118,1149,484,509]" italics="true" pageId="49" pageNumber="50">Ba</emphasis>
prominent,
<emphasis id="B9139553FF99305AFA9EA1E6FAA57FDB" box="[1297,1328,484,509]" italics="true" pageId="49" pageNumber="50">Bp</emphasis>
virtually absent, represented only by vague, ridge-like contour. Bothridial seta subflagellate, smooth, inclined posterolaterad. Notogaster with spina adnata straight, narrow to triangular, not quite reaching level of tubercle
<emphasis id="B9139553FF99305AFB3DA22EFB447C63" box="[1202,1233,556,581]" italics="true" pageId="49" pageNumber="50">Ba</emphasis>
; setae of
<emphasis id="B9139553FF99305AFAB5A22EFAD37C63" box="[1338,1350,556,581]" italics="true" pageId="49" pageNumber="50">c</emphasis>
-,
<emphasis id="B9139553FF99305AFAD2A229FAF07C63" box="[1373,1381,555,581]" italics="true" pageId="49" pageNumber="50">l</emphasis>
- and
<emphasis id="B9139553FF99305AFF18A24DFF307C4F" box="[151,165,591,617]" italics="true" pageId="49" pageNumber="50">h</emphasis>
-series oriented longitudinally, thin, acuminate, strongly curved near base. Parastigmatic enantiophysis dimorphic,
<emphasis id="B9139553FF99305AFF18A271FF267CAB" box="[151,179,627,653]" italics="true" pageId="49" pageNumber="50">Sa</emphasis>
horn-like,
<emphasis id="B9139553FF99305AFEBEA271FED87CAB" box="[305,333,627,653]" italics="true" pageId="49" pageNumber="50">Sp</emphasis>
short, blunt. Ventral enantiophyses
<emphasis id="B9139553FF99305AFD6BA271FC967CAB" box="[740,771,627,653]" italics="true" pageId="49" pageNumber="50">E2</emphasis>
and
<emphasis id="B9139553FF99305AFCB3A271FCC67CAA" box="[828,851,627,652]" italics="true" pageId="49" pageNumber="50">M</emphasis>
absent;
<emphasis id="B9139553FF99305AFC3DA271FC567CAA" box="[946,963,627,652]" italics="true" pageId="49" pageNumber="50">V</emphasis>
strongly developed; discidium in form of thick, slightly arched claw. Seta
<emphasis id="B9139553FF99305AFE73A294FD8D7C96" box="[508,536,662,688]" italics="true" pageId="49" pageNumber="50">ad</emphasis>
<subScript id="17E34B04FF99305AFD97A2ABFDB57C91" attach="left" box="[536,544,681,695]" fontSize="6" pageId="49" pageNumber="50">3</subScript>
close to anal plate. Leg IV 1.21.3 times body length; femur IV distinctly longer (1.51.8) than trochanter IV.
</paragraph>
<paragraph id="8BD84941FF99305AFF18A2FFFF4E7D31" blockId="49.[151,1437,765,2043]" box="[151,219,765,791]" pageId="49" pageNumber="50">
<emphasis id="B9139553FF99305AFF18A2FFFF4E7D31" bold="true" box="[151,219,765,791]" pageId="49" pageNumber="50">Adult</emphasis>
</paragraph>
<paragraph id="8BD84941FF99305AFF18A323FDEB7D79" blockId="49.[151,1437,765,2043]" pageId="49" pageNumber="50">
<emphasis id="B9139553FF99305AFF18A323FE8E7D1C" box="[151,283,801,826]" italics="true" pageId="49" pageNumber="50">Dimensions</emphasis>
. Total length and maximum width (n =
<specimenCount id="9D6182C8FF99305AFD5AA323FCD67D1D" box="[725,835,801,827]" pageId="49" pageNumber="50" type="female">7 females</specimenCount>
,
<specimenCount id="9D6182C8FF99305AFCC1A323FC3C7D1A" box="[846,937,801,828]" pageId="49" pageNumber="50" type="male">7 males</specimenCount>
): 365426 × 221260; males usually slightly smaller (365392) than females (387426).
</paragraph>
<paragraph id="8BD84941FF99305AFF48A36BFF2F7AE3" blockId="49.[151,1437,765,2043]" pageId="49" pageNumber="50">
<emphasis id="B9139553FF99305AFF48A36BFED07DA4" box="[199,325,873,898]" italics="true" pageId="49" pageNumber="50">Integument</emphasis>
. Body color of mature adults medium brown in life; preserved specimens medium to yellowish brown. Cerotegument excrescences forming conspicuous layer on legs and body; all excrescences pulverulent, coated with dusting of fine particles (
<figureCitation id="135C55C4FF99305AFE62A3B2FDC07DEC" box="[493,597,943,970]" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">Fig. 26C</figureCitation>
). Excrescences of prodorsum and notogaster mostly granular, columnar to short vermiform filaments (
<figureCitation id="135C55C4FF99305AFE42A3D1FDC17DCB" box="[461,596,979,1005]" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">Fig. 26B, C</figureCitation>
); those of lateral regions and basal leg segments largely capitate-columnar, appearing globular viewed
<emphasis id="B9139553FF99305AFE5FA3FAFDB77A37" box="[464,546,1015,1041]" italics="true" pageId="49" pageNumber="50">en face</emphasis>
(
<figureCitation id="135C55C4FF99305AFDBBA3F5FD297A34" box="[564,700,1015,1042]" captionStart="FIGURE 27" captionStartId="53.[151,250,1858,1882]" captionTargetBox="[157,1429,181,1834]" captionTargetId="figure-17@53.[157,1429,181,1834]" captionTargetPageId="53" captionText="FIGURE 27. Epidamaeus craigheadi (Jacot 1939), light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta lʹ labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E)." figureDoi="http://doi.org/10.5281/zenodo.5225249" httpUri="https://zenodo.org/record/5225249/files/figure.png" pageId="49" pageNumber="50">Fig. 27B, F</figureCitation>
); setae bearing granular to capitate-columnar excrescences (
<figureCitation id="135C55C4FF99305AFAE0A3F5FF5C7A10" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">Fig. 26G</figureCitation>
) or not (
<figureCitation id="135C55C4FF99305AFEB7A419FE347A10" box="[312,417,1051,1078]" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">Fig. 26F</figureCitation>
). Procuticle conspicuously microtuberculate on prodorsum, subcapitulum, coxisternum, projecting tubercles and basal leg segments (
<figureCitation id="135C55C4FF99305AFD05A43CFD607A7E" box="[650,757,1086,1112]" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">Figs 26D</figureCitation>
,
<figureCitation id="135C55C4FF99305AFC8FA43DFCC37A7F" box="[768,854,1087,1113]" captionStart="FIGURE 27" captionStartId="53.[151,250,1858,1882]" captionTargetBox="[157,1429,181,1834]" captionTargetId="figure-17@53.[157,1429,181,1834]" captionTargetPageId="53" captionText="FIGURE 27. Epidamaeus craigheadi (Jacot 1939), light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta lʹ labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E)." figureDoi="http://doi.org/10.5281/zenodo.5225249" httpUri="https://zenodo.org/record/5225249/files/figure.png" pageId="49" pageNumber="50">27AD</figureCitation>
), more weakly developed on genital and anal plates, latter as well as subcapitulum with regions of elongated microtubercles; microtubercles inconspicuous, but usually present on notogaster (
<figureCitation id="135C55C4FF99305AFE16A484FE6A7A87" box="[409,511,1158,1185]" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">Fig. 26C</figureCitation>
) and ventral plate. Without exuvial stack or noticeable adherent debris (
<figureCitation id="135C55C4FF99305AFABFA484FF397AE3" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">Fig. 26A, B</figureCitation>
).
</paragraph>
<paragraph id="8BD84941FF99305AFF48A4CCFB447868" blockId="49.[151,1437,765,2043]" pageId="49" pageNumber="50">
<emphasis id="B9139553FF99305AFF48A4CCFED17ACE" box="[199,324,1230,1256]" italics="true" pageId="49" pageNumber="50">Prodorsum</emphasis>
(
<figureCitation id="135C55C4FF99305AFED8A4CCFE7F7ACE" box="[343,490,1230,1257]" captionStart="FIGURE 25" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[220,1368,183,1907]" captionTargetId="figure-17@51.[205,1382,181,1919]" captionTargetPageId="51" captionText="FIGURE 25. Epidamaeus craigheadi (Jacot 1939), adult: A—dorsal view (legs except right trochanters IIII not shown); B—anterior half of body, right lateral view (gnathosoma and legs not shown); C—posterior part of body, right lateral view; D—ventral view (legs represented only by trochanters). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5225245" httpUri="https://zenodo.org/record/5225245/files/figure.png" pageId="49" pageNumber="50">Figs 25A, B</figureCitation>
;
<figureCitation id="135C55C4FF99305AFE75A4CCFDC77ACE" box="[506,594,1230,1256]" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">26B, D</figureCitation>
). Contour of body wall immediately anterior to acetabulum I oblique in dorsal view, nearly straight; contour anterior to acetabulum II broadly rounded (propodolateral apophysis absent). Without noticeable transverse contour anterior to bothridium. Postbothridial enantiophysis represented only by distally rounded tubercle
<emphasis id="B9139553FF99305AFE32A538FE497B75" box="[445,476,1338,1363]" italics="true" pageId="49" pageNumber="50">Ba</emphasis>
;
<emphasis id="B9139553FF99305AFE61A538FD987B75" box="[494,525,1338,1363]" italics="true" pageId="49" pageNumber="50">Bp</emphasis>
absent, but usually replaced by strong contour, appearing like short transverse ridge in dorsal view. Tubercle
<emphasis id="B9139553FF99305AFE7EA55FFD867B50" box="[497,531,1373,1398]" italics="true" pageId="49" pageNumber="50">Da</emphasis>
and laterosejugal enantiophysis absent. Bothridium with projecting part funnelshaped but relatively short, without noticeable vertical banding; mutual distance of pair slightly greater than that of tubercles
<emphasis id="B9139553FF99305AFE8CA5A7FEB77B98" box="[259,290,1445,1470]" italics="true" pageId="49" pageNumber="50">Ba</emphasis>
. Bothridial seta angled posterolaterad; long (182196), smooth, subflagellate but usually only slightly curved or nearly straight in proximal two-thirds; without cerotegument. Setae
<emphasis id="B9139553FF99305AFC77A5CBFB857BC4" box="[1016,1040,1481,1506]" italics="true" pageId="49" pageNumber="50">ro</emphasis>
(5765) and
<emphasis id="B9139553FF99305AFB2CA5CAFB227BC4" box="[1187,1207,1480,1506]" italics="true" pageId="49" pageNumber="50">le</emphasis>
(6573), both finely attenuate, curved anteromediad;
<emphasis id="B9139553FF99305AFD86A5EEFDB47823" box="[521,545,1516,1541]" italics="true" pageId="49" pageNumber="50">ro</emphasis>
usually smooth,
<emphasis id="B9139553FF99305AFD6AA5EEFD6C7820" box="[741,761,1516,1542]" italics="true" pageId="49" pageNumber="50">le</emphasis>
with few barbs at base, slightly thicker than
<emphasis id="B9139553FF99305AFB72A5EEFA807823" box="[1277,1301,1516,1541]" italics="true" pageId="49" pageNumber="50">ro</emphasis>
;
<emphasis id="B9139553FF99305AFAAAA5EEFAAE7823" box="[1317,1339,1516,1541]" italics="true" pageId="49" pageNumber="50">in</emphasis>
(3234) thin, acuminate, smooth, nearly straight, angled distinctly posterolaterad. Seta
<emphasis id="B9139553FF99305AFBABA612FBA8780F" box="[1060,1085,1552,1577]" italics="true" pageId="49" pageNumber="50">ex</emphasis>
(~20) attenuate, thin, smooth; vestige
<emphasis id="B9139553FF99305AFF62A636FE87786B" box="[237,274,1588,1613]" italics="true" pageId="49" pageNumber="50">exv</emphasis>
present immediately posterior to and appearing to merge with
<emphasis id="B9139553FF99305AFC5CA636FC79786B" box="[979,1004,1588,1613]" italics="true" pageId="49" pageNumber="50">ex</emphasis>
alveolus (
<figureCitation id="135C55C4FF99305AFBD0A631FB507868" box="[1119,1221,1587,1614]" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">Fig. 26E</figureCitation>
).
</paragraph>
<paragraph id="8BD84941FF99305AFF42A65AFA9679B6" blockId="49.[151,1437,765,2043]" pageId="49" pageNumber="50">
<emphasis id="B9139553FF99305AFF42A65AFEDC7857" box="[205,329,1624,1649]" italics="true" pageId="49" pageNumber="50">Notogaster</emphasis>
(
<figureCitation id="135C55C4FF99305AFED7A655FE717854" box="[344,484,1623,1650]" captionStart="FIGURE 25" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[220,1368,183,1907]" captionTargetId="figure-17@51.[205,1382,181,1919]" captionTargetPageId="51" captionText="FIGURE 25. Epidamaeus craigheadi (Jacot 1939), adult: A—dorsal view (legs except right trochanters IIII not shown); B—anterior half of body, right lateral view (gnathosoma and legs not shown); C—posterior part of body, right lateral view; D—ventral view (legs represented only by trochanters). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5225245" httpUri="https://zenodo.org/record/5225245/files/figure.png" pageId="49" pageNumber="50">Figs 25AC</figureCitation>
;
<figureCitation id="135C55C4FF99305AFE61A655FDAA7857" box="[494,575,1623,1649]" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">26A, F</figureCitation>
). Nearly circular in outline, about 1.1 times longer than broad; hemispherical, uniformly curved in lateral view, with maximum height at or slightly posterior to middle (
<figureCitation id="135C55C4FF99305AFB15A678FB6078B3" box="[1178,1269,1658,1685]" captionStart="FIGURE 7" captionStartId="22.[151,250,1474,1498]" captionTargetBox="[151,1435,181,1449]" captionTargetId="figure-17@22.[151,1435,181,1449]" captionTargetPageId="22" captionText="FIGURE 7. Epidamaeus michaeli (Ewing, 1909), light and SEM photographs of adult:A—habitus, lateral view; B—bothridial" figureDoi="http://doi.org/10.5281/zenodo.5225209" httpUri="https://zenodo.org/record/5225209/files/figure.png" pageId="49" pageNumber="50">Fig. 7A</figureCitation>
). Spina adnata directed toward but not reaching tubercle
<emphasis id="B9139553FF99305AFD0EA69DFD35789E" box="[641,672,1695,1720]" italics="true" pageId="49" pageNumber="50">Ba</emphasis>
in dorsal view; of medium length (1820), elongate-triangular to almost parallel-sided, straight to slightly curved laterad; relatively closely spaced, mutual distance of pair ~ 0.2 times maximum notogastral width. Setae of
<emphasis id="B9139553FF99305AFDC0A6E5FDCE7926" box="[591,603,1767,1792]" italics="true" pageId="49" pageNumber="50">c</emphasis>
-,
<emphasis id="B9139553FF99305AFDF9A6E4FDEB7926" box="[630,638,1766,1792]" italics="true" pageId="49" pageNumber="50">l</emphasis>
- and
<emphasis id="B9139553FF99305AFD4DA6E4FD457926" box="[706,720,1766,1792]" italics="true" pageId="49" pageNumber="50">h</emphasis>
-series oriented longitudinally, with
<emphasis id="B9139553FF99305AFBE0A6E5FBEE7926" box="[1135,1147,1767,1792]" italics="true" pageId="49" pageNumber="50">c</emphasis>
<subScript id="17E34B04FF99305AFBF3A6FAFB117920" attach="left" box="[1148,1156,1784,1798]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
directed anteriad, others posteriad; pairs inserted in nearly straight, closely parallel rows such that mutual distance of lyrifissures
<emphasis id="B9139553FF99305AFAA1A708FADF7905" box="[1326,1354,1802,1827]" italics="true" pageId="49" pageNumber="50">im</emphasis>
almost twice that of pair
<emphasis id="B9139553FF99305AFED6A72FFEE07961" box="[345,373,1837,1863]" italics="true" pageId="49" pageNumber="50">lm</emphasis>
in dorsal view; setae (5361) with light greyish brown pigmentation, smooth, thinly acuminate, with strong bend in basal third (
<figureCitation id="135C55C4FF99305AFE72A753FDF4794A" box="[509,609,1873,1900]" captionStart="FIGURE 26" captionStartId="52.[151,250,1822,1846]" captionTargetBox="[171,1415,181,1798]" captionTargetId="figure-17@52.[171,1415,181,1798]" captionTargetPageId="52" captionText="FIGURE 26. Epidamaeus craigheadi (Jacot 1939), light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae le, ro (cotype); E—alveolus of seta ex, with arrow to vestige exv (anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta lʹ of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="49" pageNumber="50">Fig. 26F</figureCitation>
). Setae of
<emphasis id="B9139553FF99305AFD59A750FD71794D" box="[726,740,1874,1899]" italics="true" pageId="49" pageNumber="50">p</emphasis>
-row finely attenuate, strongly bent in proximal third, smooth; relatively small, shorter than distance between alveoli;
<emphasis id="B9139553FF99305AFC8EA777FC9A79A8" box="[769,783,1909,1934]" italics="true" pageId="49" pageNumber="50">p</emphasis>
<subScript id="17E34B04FF99305AFC80A785FC8279B3" attach="left" box="[783,791,1927,1941]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
(3641) slightly thicker than
<emphasis id="B9139553FF99305AFBEBA777FBE779A8" box="[1124,1138,1909,1934]" italics="true" pageId="49" pageNumber="50">p</emphasis>
<subScript id="17E34B04FF99305AFBFDA785FBEF79B3" attach="left" box="[1138,1146,1927,1941]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
,
<emphasis id="B9139553FF99305AFB09A777FB0179A8" box="[1158,1172,1909,1934]" italics="true" pageId="49" pageNumber="50">p</emphasis>
<subScript id="17E34B04FF99305AFB1BA785FB0979B3" attach="left" box="[1172,1180,1927,1941]" fontSize="6" pageId="49" pageNumber="50">3</subScript>
(3236).
</paragraph>
<paragraph id="8BD84941FF993059FF48A79BFB417F43" blockId="49.[151,1437,765,2043]" lastBlockId="50.[151,1437,151,1005]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<emphasis id="B9139553FF99305AFF48A79BFDC77994" box="[199,594,1944,1971]" italics="true" pageId="49" pageNumber="50">Coxisternum and lateral podosoma</emphasis>
(
<figureCitation id="135C55C4FF99305AFDEEA79AFD787995" box="[609,749,1944,1971]" captionStart="FIGURE 25" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[220,1368,183,1907]" captionTargetId="figure-17@51.[205,1382,181,1919]" captionTargetPageId="51" captionText="FIGURE 25. Epidamaeus craigheadi (Jacot 1939), adult: A—dorsal view (legs except right trochanters IIII not shown); B—anterior half of body, right lateral view (gnathosoma and legs not shown); C—posterior part of body, right lateral view; D—ventral view (legs represented only by trochanters). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5225245" httpUri="https://zenodo.org/record/5225245/files/figure.png" pageId="49" pageNumber="50">Figs 25B, D</figureCitation>
,
<figureCitation id="135C55C4FF99305AFD77A79BFCDA7995" box="[760,847,1945,1971]" captionStart="FIGURE 27" captionStartId="53.[151,250,1858,1882]" captionTargetBox="[157,1429,181,1834]" captionTargetId="figure-17@53.[157,1429,181,1834]" captionTargetPageId="53" captionText="FIGURE 27. Epidamaeus craigheadi (Jacot 1939), light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta lʹ labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E)." figureDoi="http://doi.org/10.5281/zenodo.5225249" httpUri="https://zenodo.org/record/5225249/files/figure.png" pageId="49" pageNumber="50">27AD</figureCitation>
). Tectum of podocephalic fossa normally developed, without lateral tooth-like projection. Distinct coxisternal pit absent from epimere I but inverted-Y-shaped depression in its place (Fig, 27A, arrow). Enantiophyses
<emphasis id="B9139553FF99305AFD07A7E3FD3D79DD" box="[648,680,2017,2043]" italics="true" pageId="49" pageNumber="50">E2</emphasis>
and
<emphasis id="B9139553FF99305AFD55A7E3FD6479DC" box="[730,753,2017,2042]" italics="true" pageId="49" pageNumber="50">M</emphasis>
absent,
<emphasis id="B9139553FF99305AFCC5A7E3FCCE79DC" box="[842,859,2017,2042]" italics="true" pageId="49" pageNumber="50">V</emphasis>
strongly developed; tubercle
<emphasis id="B9139553FF99305AFB2FA7E3FB2979DC" box="[1184,1212,2017,2042]" italics="true" pageId="49" pageNumber="50">Va</emphasis>
triangular in ventral view, distally rounded;
<emphasis id="B9139553FF9A3059FE14A095FE2F7E96" box="[411,442,151,176]" italics="true" pageId="50" pageNumber="51">Vp</emphasis>
of similar size but broadly rounded to subquadrate (
<figureCitation id="135C55C4FF9A3059FC73A095FBF77E97" box="[1020,1122,151,177]" captionStart="FIGURE 27" captionStartId="53.[151,250,1858,1882]" captionTargetBox="[157,1429,181,1834]" captionTargetId="figure-17@53.[157,1429,181,1834]" captionTargetPageId="53" captionText="FIGURE 27. Epidamaeus craigheadi (Jacot 1939), light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta lʹ labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E)." figureDoi="http://doi.org/10.5281/zenodo.5225249" httpUri="https://zenodo.org/record/5225249/files/figure.png" pageId="50" pageNumber="51">Fig. 27C</figureCitation>
). Tubercles of parastigmatic enantiophysis dimorphic:
<emphasis id="B9139553FF9A3059FE30A0B9FE4E7EF3" box="[447,475,187,213]" italics="true" pageId="50" pageNumber="51">Sa</emphasis>
large, conspicuous, horn-like,
<emphasis id="B9139553FF9A3059FCB0A0B9FCCE7EF3" box="[831,859,187,213]" italics="true" pageId="50" pageNumber="51">Sp</emphasis>
short, subconical, perpendicular to
<emphasis id="B9139553FF9A3059FB77A0B9FA817EF3" box="[1272,1300,187,213]" italics="true" pageId="50" pageNumber="51">Sa</emphasis>
(Fig, 27B). Epimeral setae finely attenuate, almost smooth, those of epimeres 1, 2 on small but distinct tubercles (
<figureCitation id="135C55C4FF9A3059FAA9A0DDFA047EDF" box="[1318,1425,223,249]" captionStart="FIGURE 27" captionStartId="53.[151,250,1858,1882]" captionTargetBox="[157,1429,181,1834]" captionTargetId="figure-17@53.[157,1429,181,1834]" captionTargetPageId="53" captionText="FIGURE 27. Epidamaeus craigheadi (Jacot 1939), light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta lʹ labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E)." figureDoi="http://doi.org/10.5281/zenodo.5225249" httpUri="https://zenodo.org/record/5225249/files/figure.png" pageId="50" pageNumber="51">Fig. 27A</figureCitation>
). Setation: 3134; approximate lengths
<emphasis id="B9139553FF9A3059FDE7A101FD107F3B" box="[616,645,259,285]" italics="true" pageId="50" pageNumber="51">1a</emphasis>
,
<emphasis id="B9139553FF9A3059FD1AA101FD257F3B" box="[661,688,259,285]" italics="true" pageId="50" pageNumber="51">1c</emphasis>
,
<emphasis id="B9139553FF9A3059FD4FA101FD497F3B" box="[704,732,259,285]" italics="true" pageId="50" pageNumber="51">2a</emphasis>
,
<emphasis id="B9139553FF9A3059FD62A101FC9C7F3B" box="[749,777,259,285]" italics="true" pageId="50" pageNumber="51">3a</emphasis>
(20),
<emphasis id="B9139553FF9A3059FCDDA101FCFA7F3B" box="[850,879,259,285]" italics="true" pageId="50" pageNumber="51">1b</emphasis>
,
<emphasis id="B9139553FF9A3059FCF0A101FC0E7F3B" box="[895,923,259,285]" italics="true" pageId="50" pageNumber="51">3b</emphasis>
,
<emphasis id="B9139553FF9A3059FC23A101FC5D7F3B" box="[940,968,259,285]" italics="true" pageId="50" pageNumber="51">4a</emphasis>
(2832)
<emphasis id="B9139553FF9A3059FBBBA101FBDB7F3B" box="[1076,1102,259,285]" italics="true" pageId="50" pageNumber="51">3c</emphasis>
,
<emphasis id="B9139553FF9A3059FBD0A101FBE97F3B" box="[1119,1148,259,285]" italics="true" pageId="50" pageNumber="51">4b</emphasis>
,
<emphasis id="B9139553FF9A3059FB03A101FB337F3B" box="[1164,1190,259,285]" italics="true" pageId="50" pageNumber="51">4c</emphasis>
,
<emphasis id="B9139553FF9A3059FB38A101FB467F3B" box="[1207,1235,259,285]" italics="true" pageId="50" pageNumber="51">4d</emphasis>
(4145). Seta
<emphasis id="B9139553FF9A3059FA0FA101FA097F3B" box="[1408,1436,259,285]" italics="true" pageId="50" pageNumber="51">3b</emphasis>
inserted near end of
<emphasis id="B9139553FF9A3059FEF1A125FE087F66" box="[382,413,295,320]" italics="true" pageId="50" pageNumber="51">Vp</emphasis>
,
<emphasis id="B9139553FF9A3059FE23A125FE5D7F67" box="[428,456,295,321]" italics="true" pageId="50" pageNumber="51">4a</emphasis>
inserted at base of
<emphasis id="B9139553FF9A3059FD2BA125FD567F66" box="[676,707,295,320]" italics="true" pageId="50" pageNumber="51">Vp</emphasis>
or slightly removed,
<emphasis id="B9139553FF9A3059FC39A125FC457F67" box="[950,976,295,321]" italics="true" pageId="50" pageNumber="51">3c</emphasis>
inserted between
<emphasis id="B9139553FF9A3059FB12A125FB2C7F67" box="[1181,1209,295,321]" italics="true" pageId="50" pageNumber="51">Sp</emphasis>
and acetabulum III. Discidium usually sharply pointed, curved slightly posteriad like broad claw or thorn (
<figureCitation id="135C55C4FF9A3059FBD1A149FB527F43" box="[1118,1223,331,357]" captionStart="FIGURE 27" captionStartId="53.[151,250,1858,1882]" captionTargetBox="[157,1429,181,1834]" captionTargetId="figure-17@53.[157,1429,181,1834]" captionTargetPageId="53" captionText="FIGURE 27. Epidamaeus craigheadi (Jacot 1939), light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta lʹ labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E)." figureDoi="http://doi.org/10.5281/zenodo.5225249" httpUri="https://zenodo.org/record/5225249/files/figure.png" pageId="50" pageNumber="51">Fig. 27D</figureCitation>
).
</paragraph>
<paragraph id="8BD84941FF9A3059FF48A16DFF767C3F" blockId="50.[151,1437,151,1005]" pageId="50" pageNumber="51">
<emphasis id="B9139553FF9A3059FF48A16DFE047FAE" box="[199,401,367,393]" italics="true" pageId="50" pageNumber="51">Anogenital region</emphasis>
(
<figureCitation id="135C55C4FF9A3059FE2FA16DFDBC7FAF" box="[416,553,366,393]" captionStart="FIGURE 25" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[220,1368,183,1907]" captionTargetId="figure-17@51.[205,1382,181,1919]" captionTargetPageId="51" captionText="FIGURE 25. Epidamaeus craigheadi (Jacot 1939), adult: A—dorsal view (legs except right trochanters IIII not shown); B—anterior half of body, right lateral view (gnathosoma and legs not shown); C—posterior part of body, right lateral view; D—ventral view (legs represented only by trochanters). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5225245" httpUri="https://zenodo.org/record/5225245/files/figure.png" pageId="50" pageNumber="51">Fig. 25C, D</figureCitation>
). Genital setae thin, attenuate, smooth or weakly barbed; setae
<emphasis id="B9139553FF9A3059FB63A16DFA9D7FAE" box="[1260,1288,367,392]" italics="true" pageId="50" pageNumber="51">ag</emphasis>
similar to
<emphasis id="B9139553FF9A3059FA0FA16DFA097FAF" box="[1408,1436,367,393]" italics="true" pageId="50" pageNumber="51">ad</emphasis>
(~ 20), thin, attenuate, smooth;
<emphasis id="B9139553FF9A3059FE71A191FD8F7F8B" box="[510,538,403,429]" italics="true" pageId="50" pageNumber="51">ad</emphasis>
<subScript id="17E34B04FF9A3059FD95A1A7FDB77F95" attach="left" box="[538,546,421,435]" fontSize="6" pageId="50" pageNumber="51">3</subScript>
inserted close to anal aperture, nearly aligned between
<emphasis id="B9139553FF9A3059FB14A191FB227F8B" box="[1179,1207,403,429]" italics="true" pageId="50" pageNumber="51">ad</emphasis>
<subScript id="17E34B04FF9A3059FB38A1A7FB2A7F95" attach="left" box="[1207,1215,421,435]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
and
<emphasis id="B9139553FF9A3059FB77A191FA817F8A" box="[1272,1300,403,428]" italics="true" pageId="50" pageNumber="51">ag</emphasis>
. Lyrifissure
<emphasis id="B9139553FF9A3059FF18A1B5FF2E7FF6" box="[151,187,439,464]" italics="true" pageId="50" pageNumber="51">ian</emphasis>
obliquely oriented; located near anal plate at level of seta
<emphasis id="B9139553FF9A3059FCB3A1B5FCCD7FF6" box="[828,856,439,464]" italics="true" pageId="50" pageNumber="51">an</emphasis>
<subScript id="17E34B04FF9A3059FCD7A1CBFCF57FF1" attach="left" box="[856,864,457,471]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
;
<emphasis id="B9139553FF9A3059FCE2A1B5FC047FF6" box="[877,913,439,464]" italics="true" pageId="50" pageNumber="51">ian</emphasis>
anterolateral to
<emphasis id="B9139553FF9A3059FBCBA1B5FBF57FF6" box="[1092,1120,439,464]" italics="true" pageId="50" pageNumber="51">an</emphasis>
<subScript id="17E34B04FF9A3059FBEFA1CBFBFD7FF1" attach="left" box="[1120,1128,457,471]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
, minute, length about equal to setal alveolus diameter. Medial edge of anal plate with usual narrow longitudinal band, sharply demarcated by carina.
</paragraph>
<paragraph id="8BD84941FF9A3059FF48A221FAED7C8F" blockId="50.[151,1437,151,1005]" pageId="50" pageNumber="51">
<taxonomicName id="4C6732C2FF9A3059FF48A221FEC37C1B" authorityName="Krefft" authorityYear="1871" box="[199,342,547,573]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="50" pageNumber="51" phylum="Nematoda" rank="genus">
<emphasis id="B9139553FF9A3059FF48A221FEC37C1B" box="[199,342,547,573]" italics="true" pageId="50" pageNumber="51">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="135C55C4FF9A3059FEE7A221FE557C1B" box="[360,448,547,573]" captionStart="FIGURE 27" captionStartId="53.[151,250,1858,1882]" captionTargetBox="[157,1429,181,1834]" captionTargetId="figure-17@53.[157,1429,181,1834]" captionTargetPageId="53" captionText="FIGURE 27. Epidamaeus craigheadi (Jacot 1939), light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta lʹ labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E)." figureDoi="http://doi.org/10.5281/zenodo.5225249" httpUri="https://zenodo.org/record/5225249/files/figure.png" pageId="50" pageNumber="51">Fig. 27</figureCitation>
). Subcapitulum longer than wide (98102 × 7382). Subcapitular setae (
<emphasis id="B9139553FF9A3059FA84A221FA8C7C1A" box="[1291,1305,547,572]" italics="true" pageId="50" pageNumber="51">a</emphasis>
, 1820;
<emphasis id="B9139553FF9A3059FA0EA221FA007C1A" box="[1409,1429,547,572]" italics="true" pageId="50" pageNumber="51">m</emphasis>
,
<emphasis id="B9139553FF9A3059FF18A245FF307C47" box="[151,165,583,609]" italics="true" pageId="50" pageNumber="51">h</emphasis>
, 3436) attenuate, barbed. Adoral setae (810) thin, slightly barbed. Palp (8286) with typical setation: 021 39(+ω). Solenidion baculiform, appressed to surface. Postpalpal seta (6) spiniform, smooth. Chelicera (9094) typical of family; attenuate seta
<emphasis id="B9139553FF9A3059FE70A28DFDB27C8F" box="[511,551,655,681]" italics="true" pageId="50" pageNumber="51">cha</emphasis>
(2830) barbed throughout,
<emphasis id="B9139553FF9A3059FCE5A28DFC077C8F" box="[874,914,655,681]" italics="true" pageId="50" pageNumber="51">chb</emphasis>
(1820) barbed unilaterally in middle part.
</paragraph>
<paragraph id="8BD84941FF9A3059FF48A2B1FDE67DCB" blockId="50.[151,1437,151,1005]" pageId="50" pageNumber="51">
<emphasis id="B9139553FF9A3059FF48A2B1FF697CEA" box="[199,252,691,716]" italics="true" pageId="50" pageNumber="51">Legs</emphasis>
(
<figureCitation id="135C55C4FF9A3059FE81A2B1FE0C7CEB" box="[270,409,691,717]" captionStart="FIGURE 27" captionStartId="53.[151,250,1858,1882]" captionTargetBox="[157,1429,181,1834]" captionTargetId="figure-17@53.[157,1429,181,1834]" captionTargetPageId="53" captionText="FIGURE 27. Epidamaeus craigheadi (Jacot 1939), light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta lʹ labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E)." figureDoi="http://doi.org/10.5281/zenodo.5225249" httpUri="https://zenodo.org/record/5225249/files/figure.png" pageId="50" pageNumber="51">Figs 27E, F</figureCitation>
,
<figureCitation id="135C55C4FF9A3059FE28A2B1FE527CEB" box="[423,455,691,717]" captionStart="FIGURE 28" captionStartId="55.[151,250,989,1013]" captionTargetBox="[159,1424,187,958]" captionTargetId="figure-17@55.[151,1436,181,966]" captionTargetPageId="55" captionText="FIGURE 28. Epidamaeus craigheadi (Jacot 1939), adult: A—subcapitulum, ventral view; B—palp, right, abaxial view; C— chelicera, left, adaxial view (seta chb on opposite side, not labeled). Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.5225251" httpUri="https://zenodo.org/record/5225251/files/figure.png" pageId="50" pageNumber="51">28</figureCitation>
). Relative lengths (I to IV) 1.0: 0.8: 1.0: 1.31.4; leg IV 1.21.3 times body length. Femur IV distinctly longer (1.51.8) than trochanter IV; approximate length of leg segments shown in
<tableCitation id="C6E57CFAFF9A3059FAB0A2D5FA037CD7" box="[1343,1430,727,753]" captionStart="TABLE 1" captionStartId="7.[151,239,404,428]" captionText="TABLE 1. Leg segment length in Epidamaeus species1" pageId="50" pageNumber="51">Table 1</tableCitation>
. Proximal stalk of femur IV distinctly longer (1.41.5) than distal bulb. Claw of each leg smooth, without basal tooth. Formulas of leg setation and solenidia: I (174420) [122], II (164417) [112], III (253317) [110], IV (143314) [010]. Notably: all tarsi with proximal accessory pair (
<emphasis id="B9139553FF9A3059FBB6A341FBD07D7A" box="[1081,1093,835,860]" italics="true" pageId="50" pageNumber="51">v</emphasis>
) present,
<emphasis id="B9139553FF9A3059FB3DA341FB2B7D7A" box="[1202,1214,835,860]" italics="true" pageId="50" pageNumber="51">v</emphasis>
<subScript id="17E34B04FF9A3059FB31A357FB537D45" attach="both" box="[1214,1222,853,867]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
<emphasis id="B9139553FF9A3059FB48A346FB587D79" box="[1223,1229,836,863]" italics="true" pageId="50" pageNumber="51">ʹ</emphasis>
and
<emphasis id="B9139553FF9A3059FA8EA341FA987D7A" box="[1281,1293,835,860]" italics="true" pageId="50" pageNumber="51">v</emphasis>
<subScript id="17E34B04FF9A3059FA82A357FA807D45" attach="both" box="[1293,1301,853,867]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
<emphasis id="B9139553FF9A3059FA9AA346FAB47D79" box="[1301,1313,836,863]" italics="true" pageId="50" pageNumber="51">ʺ</emphasis>
;
<emphasis id="B9139553FF9A3059FAA0A341FAAE7D7A" box="[1327,1339,835,860]" italics="true" pageId="50" pageNumber="51">v</emphasis>
<subScript id="17E34B04FF9A3059FAB4A357FAD67D45" attach="both" box="[1339,1347,853,867]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
<emphasis id="B9139553FF9A3059FACCA346FADA7D79" box="[1347,1359,836,863]" italics="true" pageId="50" pageNumber="51">ʺ</emphasis>
absent from femora I, II; seta
<emphasis id="B9139553FF9A3059FE10A36AFE227DA5" box="[415,439,872,899]" italics="true" pageId="50" pageNumber="51"></emphasis>
present on femur III. Setae
<emphasis id="B9139553FF9A3059FD73A365FC9F7DA7" box="[764,778,871,897]" italics="true" pageId="50" pageNumber="51">d</emphasis>
(if uncoupled) and
<emphasis id="B9139553FF9A3059FC61A365FC637DA7" box="[1006,1014,871,897]" italics="true" pageId="50" pageNumber="51">l</emphasis>
barbed, acuminate to attenuate,
<emphasis id="B9139553FF9A3059FAE6A365FAE27DA7" box="[1385,1399,871,897]" italics="true" pageId="50" pageNumber="51">d</emphasis>
on femora larger, thicker than
<emphasis id="B9139553FF9A3059FE41A389FE437D83" box="[462,470,907,933]" italics="true" pageId="50" pageNumber="51">l</emphasis>
; coupled setae
<emphasis id="B9139553FF9A3059FD07A389FD037D83" box="[648,662,907,933]" italics="true" pageId="50" pageNumber="51">d</emphasis>
of genua IIII as long or slightly longer than respective solenidion. Solenidia φ
<subScript id="17E34B04FF9A3059FE97A3C3FEB57DE9" attach="left" box="[280,288,961,975]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
on tibia I flagellate, φ on tibia IV subflagellate; others tapered but blunt (ceratiform), erect or slightly curved. Famulus of tarsus I acicular, erect.
</paragraph>
<paragraph id="8BD84941FF9A3059FF18A418FE087A13" blockId="50.[151,1437,1050,2013]" box="[151,413,1050,1077]" pageId="50" pageNumber="51">
<emphasis id="B9139553FF9A3059FF18A418FE907A12" bold="true" box="[151,261,1050,1076]" pageId="50" pageNumber="51">Juveniles</emphasis>
(
<figureCitation id="135C55C4FF9A3059FE9BA419FE007A13" box="[276,405,1051,1077]" captionStart-0="FIGURE 30" captionStart-1="FIGURE 31" captionStart-2="FIGURE 32" captionStartId-0="57.[151,250,1809,1833]" captionStartId-1="58.[151,250,1834,1858]" captionStartId-2="59.[151,250,1462,1486]" captionTargetBox-0="[171,1415,181,1785]" captionTargetBox-1="[156,1430,380,1809]" captionTargetBox-2="[164,1423,844,1439]" captionTargetId-0="figure-17@57.[171,1415,181,1785]" captionTargetId-1="figure-102@58.[156,1430,380,1809]" captionTargetId-2="figure-307@59.[164,1423,844,1439]" captionTargetPageId-0="57" captionTargetPageId-1="58" captionTargetPageId-2="59" captionText-0="FIGURE 30. Epidamaeus craigheadi (Jacot 1939), light photographs of nymphs: A—protonymph, lateral view; B—same, posterior region of gastronotum (left insert = enlargement of attachment cornicle k inserting in induration (ind) on underside of larval scalp; C—protonymph, dorsal view of hysterosoma with larval scalp intact (white labels on scalp setae; insert = larval scalp removed from different specimen, arrow to attachment induration); D—tritonymph, dorsal view (scalps removed; several setae broken; body artificially swollen by clearing); E—tritonymph, lateral view (scalps removed; lower left insert = enlargement of attachment cornicle k). Scale bars 20 μm (B, C), 50 μm (A, D, E)." captionText-1="FIGURE 31. Epidamaeus craigheadi (Jacot 1939), light photographs of protonymph (A), tritonymph (BG):A—lateral view of" captionText-2="FIGURE 32. Epidamaeus craigheadi (Jacot 1939), light photographs of deutonymph (A), tritonymph (BE):A—middle portion of leg II, abaxial view; B—middle portion of right leg I, dorsoadaxial view; C—portion of tibia I from different specimen, abaxial view (arrow to coupled seta d); D—bulb of left tarsus I, dorsoabaxial view, showing tarsal cluster (lower right insert = isolated famulus, arrow to basal root); E—isolated solenidion ω1 and cerotegument excrescences. Scale bars 5 μm (D, E), 10 μm (AC)." figureDoi-0="http://doi.org/10.5281/zenodo.5225255" figureDoi-1="http://doi.org/10.5281/zenodo.5225257" figureDoi-2="http://doi.org/10.5281/zenodo.5225259" httpUri-0="https://zenodo.org/record/5225255/files/figure.png" httpUri-1="https://zenodo.org/record/5225257/files/figure.png" httpUri-2="https://zenodo.org/record/5225259/files/figure.png" pageId="50" pageNumber="51">Figs 3032</figureCitation>
)
</paragraph>
<paragraph id="8BD84941FF9A3059FF18A43DFC9F7A87" blockId="50.[151,1437,1050,2013]" pageId="50" pageNumber="51">
<emphasis id="B9139553FF9A3059FF18A43DFE8E7A7E" box="[151,283,1087,1112]" italics="true" pageId="50" pageNumber="51">Dimensions</emphasis>
. Larva unknown except as exuvial scalp. Total length of PN (n = 3) 245255, DN (n = 3) 269308, TN (n = 4) 343377. Maximum width of hysterosoma about half body length. Setal lengths given below only for one average TN: proportional in other nymphs unless noted.
</paragraph>
<paragraph id="8BD84941FF9A3059FF48A4A9FD107B17" blockId="50.[151,1437,1050,2013]" pageId="50" pageNumber="51">
<emphasis id="B9139553FF9A3059FF48A4A9FED07AE2" box="[199,325,1195,1220]" italics="true" pageId="50" pageNumber="51">Integument</emphasis>
. Excrescences of cerotegument, short, strongly pulverulent; dense, adjacent or slightly separated on most of body and legs; mostly bottle-shaped to capitate-columnar, but appearing granular (diameter 23) seen faceon (
<figureCitation id="135C55C4FF9A3059FF4DA4F1FED87B2B" box="[194,333,1267,1293]" captionStart="FIGURE 31" captionStartId="58.[151,250,1834,1858]" captionTargetBox="[156,1430,380,1809]" captionTargetId="figure-102@58.[156,1430,380,1809]" captionTargetPageId="58" captionText="FIGURE 31. Epidamaeus craigheadi (Jacot 1939), light photographs of protonymph (A), tritonymph (BG):A—lateral view of" figureDoi="http://doi.org/10.5281/zenodo.5225257" httpUri="https://zenodo.org/record/5225257/files/figure.png" pageId="50" pageNumber="51">Figs 31B, C</figureCitation>
,
<figureCitation id="135C55C4FF9A3059FED7A4F1FE187B2B" box="[344,397,1267,1293]" captionStart="FIGURE 32" captionStartId="59.[151,250,1462,1486]" captionTargetBox="[164,1423,844,1439]" captionTargetId="figure-307@59.[164,1423,844,1439]" captionTargetPageId="59" captionText="FIGURE 32. Epidamaeus craigheadi (Jacot 1939), light photographs of deutonymph (A), tritonymph (BE):A—middle portion of leg II, abaxial view; B—middle portion of right leg I, dorsoadaxial view; C—portion of tibia I from different specimen, abaxial view (arrow to coupled seta d); D—bulb of left tarsus I, dorsoabaxial view, showing tarsal cluster (lower right insert = isolated famulus, arrow to basal root); E—isolated solenidion ω1 and cerotegument excrescences. Scale bars 5 μm (D, E), 10 μm (AC)." figureDoi="http://doi.org/10.5281/zenodo.5225259" httpUri="https://zenodo.org/record/5225259/files/figure.png" pageId="50" pageNumber="51">32D</figureCitation>
). Setae of untreated specimens usually with cerotegument; excrescences smaller, shaped like those of body surface or minute (1) spheres.
</paragraph>
<paragraph id="8BD84941FF9A3059FF48A539FB007BE7" blockId="50.[151,1437,1050,2013]" pageId="50" pageNumber="51">
<emphasis id="B9139553FF9A3059FF48A539FED17B73" box="[199,324,1339,1365]" italics="true" pageId="50" pageNumber="51">Prodorsum</emphasis>
. About half length of gastronotum; rostrum broadly rounded. Setae
<emphasis id="B9139553FF9A3059FBB1A539FBC37B72" box="[1086,1110,1339,1364]" italics="true" pageId="50" pageNumber="51">ro</emphasis>
(
<quantity id="4C9FE4A4FF9A3059FBEBA538FB0A7B73" box="[1124,1183,1338,1365]" metricMagnitude="0" metricUnit="m" metricValue="1.3716" pageId="50" pageNumber="51" unit="in" value="54.0">54 in</quantity>
TN) and
<emphasis id="B9139553FF9A3059FA85A539FA8B7B73" box="[1290,1310,1339,1365]" italics="true" pageId="50" pageNumber="51">le</emphasis>
(
<quantity id="4C9FE4A4FF9A3059FAA2A539FAF27B73" box="[1325,1383,1339,1365]" metricMagnitude="0" metricUnit="m" metricValue="1.5748" pageId="50" pageNumber="51" unit="in" value="62.0">62 in</quantity>
TN) attenuate, barbed, curved anteromediad (
<figureCitation id="135C55C4FF9A3059FDECA55DFD5F7B5F" box="[611,714,1375,1401]" captionStart="FIGURE 31" captionStartId="58.[151,250,1834,1858]" captionTargetBox="[156,1430,380,1809]" captionTargetId="figure-102@58.[156,1430,380,1809]" captionTargetPageId="58" captionText="FIGURE 31. Epidamaeus craigheadi (Jacot 1939), light photographs of protonymph (A), tritonymph (BG):A—lateral view of" figureDoi="http://doi.org/10.5281/zenodo.5225257" httpUri="https://zenodo.org/record/5225257/files/figure.png" pageId="50" pageNumber="51">Fig. 31B</figureCitation>
);
<emphasis id="B9139553FF9A3059FD6FA55DFD6C7B5E" box="[736,761,1375,1400]" italics="true" pageId="50" pageNumber="51">ex</emphasis>
(
<quantity id="4C9FE4A4FF9A3059FC86A55DFCD17B5F" box="[777,836,1375,1401]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" pageId="50" pageNumber="51" unit="in" value="37.0">37 in</quantity>
TN) attenuate, weakly barbed (
<figureCitation id="135C55C4FF9A3059FB25A55DFA847B5F" box="[1194,1297,1375,1401]" captionStart="FIGURE 31" captionStartId="58.[151,250,1834,1858]" captionTargetBox="[156,1430,380,1809]" captionTargetId="figure-102@58.[156,1430,380,1809]" captionTargetPageId="58" captionText="FIGURE 31. Epidamaeus craigheadi (Jacot 1939), light photographs of protonymph (A), tritonymph (BG):A—lateral view of" figureDoi="http://doi.org/10.5281/zenodo.5225257" httpUri="https://zenodo.org/record/5225257/files/figure.png" pageId="50" pageNumber="51">Fig. 31E</figureCitation>
); seta
<emphasis id="B9139553FF9A3059FAD6A55DFAFA7B5E" box="[1369,1391,1375,1400]" italics="true" pageId="50" pageNumber="51">in</emphasis>
(
<quantity id="4C9FE4A4FF9A3059FAF0A55DFF387BBB" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="50" pageNumber="51" unit="in" value="20.0">20 in</quantity>
TN) spiniform—smooth, straight, with several minute terminal teeth (
<figureCitation id="135C55C4FF9A3059FC51A581FBDC7BBB" box="[990,1097,1411,1437]" captionStart="FIGURE 31" captionStartId="58.[151,250,1834,1858]" captionTargetBox="[156,1430,380,1809]" captionTargetId="figure-102@58.[156,1430,380,1809]" captionTargetPageId="58" captionText="FIGURE 31. Epidamaeus craigheadi (Jacot 1939), light photographs of protonymph (A), tritonymph (BG):A—lateral view of" figureDoi="http://doi.org/10.5281/zenodo.5225257" httpUri="https://zenodo.org/record/5225257/files/figure.png" pageId="50" pageNumber="51">Fig. 31D</figureCitation>
). Bothridial seta (
<quantity id="4C9FE4A4FF9A3059FA96A581FAF17BBB" box="[1305,1380,1411,1437]" metricMagnitude="0" metricUnit="m" metricValue="4.8006" pageId="50" pageNumber="51" unit="in" value="189.0">189 in</quantity>
TN) subflagellate, smooth, usually covered in cerotegument over whole length (
<figureCitation id="135C55C4FF9A3059FC6DA5A5FBDC7BE7" box="[994,1097,1447,1473]" captionStart="FIGURE 30" captionStartId="57.[151,250,1809,1833]" captionTargetBox="[171,1415,181,1785]" captionTargetId="figure-17@57.[171,1415,181,1785]" captionTargetPageId="57" captionText="FIGURE 30. Epidamaeus craigheadi (Jacot 1939), light photographs of nymphs: A—protonymph, lateral view; B—same, posterior region of gastronotum (left insert = enlargement of attachment cornicle k inserting in induration (ind) on underside of larval scalp; C—protonymph, dorsal view of hysterosoma with larval scalp intact (white labels on scalp setae; insert = larval scalp removed from different specimen, arrow to attachment induration); D—tritonymph, dorsal view (scalps removed; several setae broken; body artificially swollen by clearing); E—tritonymph, lateral view (scalps removed; lower left insert = enlargement of attachment cornicle k). Scale bars 20 μm (B, C), 50 μm (A, D, E)." figureDoi="http://doi.org/10.5281/zenodo.5225255" httpUri="https://zenodo.org/record/5225255/files/figure.png" pageId="50" pageNumber="51">Fig. 30D</figureCitation>
,
<figureCitation id="135C55C4FF9A3059FBDAA5A5FB1D7BE7" box="[1109,1160,1447,1473]" captionStart="FIGURE 31" captionStartId="58.[151,250,1834,1858]" captionTargetBox="[156,1430,380,1809]" captionTargetId="figure-102@58.[156,1430,380,1809]" captionTargetPageId="58" captionText="FIGURE 31. Epidamaeus craigheadi (Jacot 1939), light photographs of protonymph (A), tritonymph (BG):A—lateral view of" figureDoi="http://doi.org/10.5281/zenodo.5225257" httpUri="https://zenodo.org/record/5225257/files/figure.png" pageId="50" pageNumber="51">31A</figureCitation>
).
</paragraph>
<paragraph id="8BD84941FF9A3059FF48A5C9FAE4790F" blockId="50.[151,1437,1050,2013]" pageId="50" pageNumber="51">
<emphasis id="B9139553FF9A3059FF48A5C9FE0F7BC2" box="[199,410,1483,1509]" italics="true" pageId="50" pageNumber="51">Gastronotic region</emphasis>
. Larval scalp with strong topography of broad transverse grooves (
<figureCitation id="135C55C4FF9A3059FB09A5C9FA997BC3" box="[1158,1292,1483,1509]" captionStart="FIGURE 30" captionStartId="57.[151,250,1809,1833]" captionTargetBox="[171,1415,181,1785]" captionTargetId="figure-17@57.[171,1415,181,1785]" captionTargetPageId="57" captionText="FIGURE 30. Epidamaeus craigheadi (Jacot 1939), light photographs of nymphs: A—protonymph, lateral view; B—same, posterior region of gastronotum (left insert = enlargement of attachment cornicle k inserting in induration (ind) on underside of larval scalp; C—protonymph, dorsal view of hysterosoma with larval scalp intact (white labels on scalp setae; insert = larval scalp removed from different specimen, arrow to attachment induration); D—tritonymph, dorsal view (scalps removed; several setae broken; body artificially swollen by clearing); E—tritonymph, lateral view (scalps removed; lower left insert = enlargement of attachment cornicle k). Scale bars 20 μm (B, C), 50 μm (A, D, E)." figureDoi="http://doi.org/10.5281/zenodo.5225255" httpUri="https://zenodo.org/record/5225255/files/figure.png" pageId="50" pageNumber="51">Fig. 30A, B</figureCitation>
); attachment induration (
<emphasis id="B9139553FF9A3059FE91A5EDFED7782F" box="[286,322,1519,1545]" italics="true" pageId="50" pageNumber="51">ind</emphasis>
;
<figureCitation id="135C55C4FF9A3059FEDBA5EDFE20782F" box="[340,437,1519,1545]" captionStart="FIGURE 30" captionStartId="57.[151,250,1809,1833]" captionTargetBox="[171,1415,181,1785]" captionTargetId="figure-17@57.[171,1415,181,1785]" captionTargetPageId="57" captionText="FIGURE 30. Epidamaeus craigheadi (Jacot 1939), light photographs of nymphs: A—protonymph, lateral view; B—same, posterior region of gastronotum (left insert = enlargement of attachment cornicle k inserting in induration (ind) on underside of larval scalp; C—protonymph, dorsal view of hysterosoma with larval scalp intact (white labels on scalp setae; insert = larval scalp removed from different specimen, arrow to attachment induration); D—tritonymph, dorsal view (scalps removed; several setae broken; body artificially swollen by clearing); E—tritonymph, lateral view (scalps removed; lower left insert = enlargement of attachment cornicle k). Scale bars 20 μm (B, C), 50 μm (A, D, E)." figureDoi="http://doi.org/10.5281/zenodo.5225255" httpUri="https://zenodo.org/record/5225255/files/figure.png" pageId="50" pageNumber="51">Fig 30B</figureCitation>
, insert) conspicuously darkened in transmitted light (
<figureCitation id="135C55C4FF9A3059FBAEA5EDFB1C782F" box="[1057,1161,1519,1545]" captionStart="FIGURE 30" captionStartId="57.[151,250,1809,1833]" captionTargetBox="[171,1415,181,1785]" captionTargetId="figure-17@57.[171,1415,181,1785]" captionTargetPageId="57" captionText="FIGURE 30. Epidamaeus craigheadi (Jacot 1939), light photographs of nymphs: A—protonymph, lateral view; B—same, posterior region of gastronotum (left insert = enlargement of attachment cornicle k inserting in induration (ind) on underside of larval scalp; C—protonymph, dorsal view of hysterosoma with larval scalp intact (white labels on scalp setae; insert = larval scalp removed from different specimen, arrow to attachment induration); D—tritonymph, dorsal view (scalps removed; several setae broken; body artificially swollen by clearing); E—tritonymph, lateral view (scalps removed; lower left insert = enlargement of attachment cornicle k). Scale bars 20 μm (B, C), 50 μm (A, D, E)." figureDoi="http://doi.org/10.5281/zenodo.5225255" httpUri="https://zenodo.org/record/5225255/files/figure.png" pageId="50" pageNumber="51">Fig. 30C</figureCitation>
, insert). Setal ontogeny normal. Except as noted, setae pigmented, directed dorsad and curved slightly posteriad, barbed on outer curvature; finely attenuated to threadlike hyaline tip, inserted on distinct tubercles (measurements for TN only): seta
<emphasis id="B9139553FF9A3059FAC4A635FACC7877" box="[1355,1369,1591,1617]" italics="true" pageId="50" pageNumber="51">h</emphasis>
<subScript id="17E34B04FF9A3059FAD6A64BFAF47871" attach="left" box="[1369,1377,1609,1623]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
(220 245) subflagellate, pair nearly straight, diverging from nearly adjacent insertions on pygidial process; pair
<emphasis id="B9139553FF9A3059FACFA659FAD97852" box="[1344,1356,1627,1652]" italics="true" pageId="50" pageNumber="51">c</emphasis>
<subScript id="17E34B04FF9A3059FAC2A66FFAC0785D" attach="left" box="[1357,1365,1645,1659]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
(170 183) nearly adjacent on single weak sclerite, posterior bend strongest near base;
<emphasis id="B9139553FF9A3059FBBAA67DFBD478BE" box="[1077,1089,1663,1688]" italics="true" pageId="50" pageNumber="51">c</emphasis>
<subScript id="17E34B04FF9A3059FBCEA693FBDC78B9" attach="left" box="[1089,1097,1681,1695]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
,
<emphasis id="B9139553FF9A3059FBD6A67DFBFA78BF" box="[1113,1135,1663,1689]" italics="true" pageId="50" pageNumber="51">la</emphasis>
,
<emphasis id="B9139553FF9A3059FBF0A67DFB0E78BF" box="[1151,1179,1663,1689]" italics="true" pageId="50" pageNumber="51">lm</emphasis>
,
<emphasis id="B9139553FF9A3059FB24A67DFB5478BF" box="[1195,1217,1663,1689]" italics="true" pageId="50" pageNumber="51">lp</emphasis>
similar (147170),
<emphasis id="B9139553FF9A3059FF18A6A1FF30789B" box="[151,165,1699,1725]" italics="true" pageId="50" pageNumber="51">h</emphasis>
<subScript id="17E34B04FF9A3059FF2AA6B7FF3878E5" attach="left" box="[165,173,1717,1731]" fontSize="6" pageId="50" pageNumber="51">3</subScript>
(110 135) and
<emphasis id="B9139553FF9A3059FEE1A6A1FEE9789B" box="[366,380,1699,1725]" italics="true" pageId="50" pageNumber="51">h</emphasis>
<subScript id="17E34B04FF9A3059FEF3A6B7FE1178E5" attach="left" box="[380,388,1717,1731]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
(98110) slightly smaller; seta
<emphasis id="B9139553FF9A3059FD7EA6A1FD68789A" box="[753,765,1699,1724]" italics="true" pageId="50" pageNumber="51">c</emphasis>
<subScript id="17E34B04FF9A3059FD71A6B7FC9378E5" attach="left" box="[766,774,1717,1731]" fontSize="6" pageId="50" pageNumber="51">3</subScript>
(50) smallest;
<emphasis id="B9139553FF9A3059FC39A6A1FC51789A" box="[950,964,1699,1724]" italics="true" pageId="50" pageNumber="51">p</emphasis>
<subScript id="17E34B04FF9A3059FC4BA6B7FC5978E5" attach="left" box="[964,972,1717,1731]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
(6174) and
<emphasis id="B9139553FF9A3059FBE3A6A1FBEF789A" box="[1132,1146,1699,1724]" italics="true" pageId="50" pageNumber="51">p</emphasis>
<subScript id="17E34B04FF9A3059FBF5A6B7FB1778E5" attach="left" box="[1146,1154,1717,1731]" fontSize="6" pageId="50" pageNumber="51">3</subScript>
(5060) relatively close together,
<emphasis id="B9139553FF9A3059FE8FA6C5FE9B78C6" box="[256,270,1735,1760]" italics="true" pageId="50" pageNumber="51">p</emphasis>
<subScript id="17E34B04FF9A3059FE81A6DBFE8378C1" attach="left" box="[270,278,1753,1767]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
(6178) more distant, inserted lateral to
<emphasis id="B9139553FF9A3059FD6BA6C5FD6778C7" box="[740,754,1735,1761]" italics="true" pageId="50" pageNumber="51">h</emphasis>
<subScript id="17E34B04FF9A3059FD7DA6DBFD6F78C1" attach="left" box="[754,762,1753,1767]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
on weak sclerite of pygidial process. Attachment cornicle
<emphasis id="B9139553FF9A3059FA1FA6C5FA0978C7" box="[1424,1436,1735,1761]" italics="true" pageId="50" pageNumber="51">k</emphasis>
of nymphs positioned between setal pair
<emphasis id="B9139553FF9A3059FDD3A6E9FDE77923" box="[604,626,1771,1797]" italics="true" pageId="50" pageNumber="51">lp</emphasis>
(
<figureCitation id="135C55C4FF9A3059FD0FA6E9FD707923" box="[640,741,1771,1797]" captionStart="FIGURE 30" captionStartId="57.[151,250,1809,1833]" captionTargetBox="[171,1415,181,1785]" captionTargetId="figure-17@57.[171,1415,181,1785]" captionTargetPageId="57" captionText="FIGURE 30. Epidamaeus craigheadi (Jacot 1939), light photographs of nymphs: A—protonymph, lateral view; B—same, posterior region of gastronotum (left insert = enlargement of attachment cornicle k inserting in induration (ind) on underside of larval scalp; C—protonymph, dorsal view of hysterosoma with larval scalp intact (white labels on scalp setae; insert = larval scalp removed from different specimen, arrow to attachment induration); D—tritonymph, dorsal view (scalps removed; several setae broken; body artificially swollen by clearing); E—tritonymph, lateral view (scalps removed; lower left insert = enlargement of attachment cornicle k). Scale bars 20 μm (B, C), 50 μm (A, D, E)." figureDoi="http://doi.org/10.5281/zenodo.5225255" httpUri="https://zenodo.org/record/5225255/files/figure.png" pageId="50" pageNumber="51">Fig. 30E</figureCitation>
); relatively short (~
<quantity id="4C9FE4A4FF9A3059FC32A6E8FC637923" box="[957,1014,1770,1797]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="50" pageNumber="51" unit="in" value="35.0">35 in</quantity>
TN), mostly straight, with slight bend in distal third; in PN, distal third inserted in sleeve-like induration on ventral surface of larval scalp (
<figureCitation id="135C55C4FF9A3059FB71A70DFAF0790F" box="[1278,1381,1807,1833]" captionStart="FIGURE 30" captionStartId="57.[151,250,1809,1833]" captionTargetBox="[171,1415,181,1785]" captionTargetId="figure-17@57.[171,1415,181,1785]" captionTargetPageId="57" captionText="FIGURE 30. Epidamaeus craigheadi (Jacot 1939), light photographs of nymphs: A—protonymph, lateral view; B—same, posterior region of gastronotum (left insert = enlargement of attachment cornicle k inserting in induration (ind) on underside of larval scalp; C—protonymph, dorsal view of hysterosoma with larval scalp intact (white labels on scalp setae; insert = larval scalp removed from different specimen, arrow to attachment induration); D—tritonymph, dorsal view (scalps removed; several setae broken; body artificially swollen by clearing); E—tritonymph, lateral view (scalps removed; lower left insert = enlargement of attachment cornicle k). Scale bars 20 μm (B, C), 50 μm (A, D, E)." figureDoi="http://doi.org/10.5281/zenodo.5225255" httpUri="https://zenodo.org/record/5225255/files/figure.png" pageId="50" pageNumber="51">Fig. 30B</figureCitation>
).
</paragraph>
<paragraph id="8BD84941FF9A3059FF48A731FE2979B3" blockId="50.[151,1437,1050,2013]" pageId="50" pageNumber="51">
<emphasis id="B9139553FF9A3059FF48A731FE9A796A" box="[199,271,1843,1868]" italics="true" pageId="50" pageNumber="51">Venter</emphasis>
. Epimeral setae acuminate to attenuate, smooth or weakly barbed,
<emphasis id="B9139553FF9A3059FB95A731FBA2796B" box="[1050,1079,1843,1869]" italics="true" pageId="50" pageNumber="51">1b</emphasis>
longest (TN 28; others mostly 1720); genitals and aggenital (15 20) similar in form; anals (2025) and adanals (3050,
<emphasis id="B9139553FF9A3059FB00A755FB3E7957" box="[1167,1195,1879,1905]" italics="true" pageId="50" pageNumber="51">ad</emphasis>
<subScript id="17E34B04FF9A3059FB24A76BFB267951" attach="left" box="[1195,1203,1897,1911]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
longest) extended in threadlike tip when intact.
</paragraph>
<paragraph id="8BD84941FF9A3059FF48A79DFE2479FB" blockId="50.[151,1437,1050,2013]" pageId="50" pageNumber="51">
<taxonomicName id="4C6732C2FF9A3059FF48A79DFEC3799F" authorityName="Krefft" authorityYear="1871" box="[199,342,1951,1977]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="50" pageNumber="51" phylum="Nematoda" rank="genus">
<emphasis id="B9139553FF9A3059FF48A79DFEC3799F" box="[199,342,1951,1977]" italics="true" pageId="50" pageNumber="51">Gnathosoma</emphasis>
</taxonomicName>
. Subcapitulum length and width about equal. Setae of mentum (
<emphasis id="B9139553FF9A3059FBBDA79DFBD5799F" box="[1074,1088,1951,1977]" italics="true" pageId="50" pageNumber="51">h</emphasis>
~20) and gena (
<emphasis id="B9139553FF9A3059FB74A79DFA9C799E" box="[1275,1289,1951,1976]" italics="true" pageId="50" pageNumber="51">a</emphasis>
~20,
<emphasis id="B9139553FF9A3059FAC3A79DFAF5799E" box="[1356,1376,1951,1976]" italics="true" pageId="50" pageNumber="51">m</emphasis>
~25) attenuate, nearly smooth.
</paragraph>
<caption id="DF1819C9FF9B3058FF18A795FC1E79DE" ID-DOI="http://doi.org/10.5281/zenodo.5225245" ID-Zenodo-Dep="5225245" httpUri="https://zenodo.org/record/5225245/files/figure.png" pageId="51" pageNumber="52" startId="51.[151,250,1943,1967]" targetBox="[220,1368,183,1907]" targetPageId="51">
<paragraph id="8BD84941FF9B3058FF18A795FC1E79DE" blockId="51.[151,1436,1943,2040]" pageId="51" pageNumber="52">
<emphasis id="B9139553FF9B3058FF18A795FEB67989" bold="true" box="[151,291,1943,1967]" pageId="51" pageNumber="52">FIGURE 25.</emphasis>
<taxonomicName id="4C6732C2FF9B3058FEA2A79AFD3A7996" authority="(Jacot 1939)" baseAuthorityName="Jacot" baseAuthorityYear="1939" box="[301,687,1944,1968]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="craigheadi">
<emphasis id="B9139553FF9B3058FEA2A79AFDB77996" box="[301,546,1944,1968]" italics="true" pageId="51" pageNumber="52">Epidamaeus craigheadi</emphasis>
(
<bibRefCitation id="EFF634B0FF9B3058FDBDA79AFD3D7996" author="Jacot, A. P." box="[562,680,1944,1968]" pageId="51" pageNumber="52" pagination="321 - 332" refId="ref37962" refString="Jacot, A. P. (1939) New mites from the White Mountains. Occasional Papers of the Boston Society of Natural History, 8, 321 - 332." type="journal article" year="1939">Jacot 1939</bibRefCitation>
)
</taxonomicName>
, adult: A—dorsal view (legs except right trochanters IIII not shown); B—anterior half of body, right lateral view (gnathosoma and legs not shown); C—posterior part of body, right lateral view; D—ventral view (legs represented only by trochanters). Scale bar 100 μm.
</paragraph>
</caption>
<caption id="DF1819C9FF9C305FFF18A71CFAED79CC" ID-DOI="http://doi.org/10.5281/zenodo.5225247" ID-Zenodo-Dep="5225247" httpUri="https://zenodo.org/record/5225247/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,250,1822,1846]" targetBox="[171,1415,181,1798]" targetPageId="52">
<paragraph id="8BD84941FF9C305FFF18A71CFAED79CC" blockId="52.[151,1437,1822,2028]" pageId="52" pageNumber="53">
<emphasis id="B9139553FF9C305FFF18A71CFEB57910" bold="true" box="[151,288,1822,1846]" pageId="52" pageNumber="53">FIGURE 26.</emphasis>
<taxonomicName id="4C6732C2FF9C305FFEA9A71CFD0B7910" authority="(Jacot 1939)" baseAuthorityName="Jacot" baseAuthorityYear="1939" box="[294,670,1822,1846]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="craigheadi">
<emphasis id="B9139553FF9C305FFEA9A71CFD8D7910" box="[294,536,1822,1846]" italics="true" pageId="52" pageNumber="53">Epidamaeus craigheadi</emphasis>
(
<bibRefCitation id="EFF634B0FF9C305FFDAAA71CFD027910" author="Jacot, A. P." box="[549,663,1822,1846]" pageId="52" pageNumber="53" pagination="321 - 332" refId="ref37962" refString="Jacot, A. P. (1939) New mites from the White Mountains. Occasional Papers of the Boston Society of Natural History, 8, 321 - 332." type="journal article" year="1939">Jacot 1939</bibRefCitation>
)
</taxonomicName>
, light and SEM photographs of adult: A—habitus, lateral view; B (SEM)— anterior half, dorsal view; C (SEM)—cuticle of notogaster, near sejugal furrow, showing weak procuticular microtubercles (short arrows) and two forms of overlying cerotegument excrescences, vermiform and granular (long arrows), all pulverulent (covered with minute, dust-like particles); D—microtuberculate procuticle of rostrum in vicinity of right setae
<emphasis id="B9139553FF9C305FFA8BA788FA837984" box="[1284,1302,1930,1954]" italics="true" pageId="52" pageNumber="53">le</emphasis>
,
<emphasis id="B9139553FF9C305FFAABA789FAAF7984" box="[1316,1338,1931,1954]" italics="true" pageId="52" pageNumber="53">ro</emphasis>
(cotype); E—alveolus of seta
<emphasis id="B9139553FF9C305FFEE3A7ADFE1779E0" box="[364,386,1967,1990]" italics="true" pageId="52" pageNumber="53">ex</emphasis>
, with arrow to vestige
<emphasis id="B9139553FF9C305FFDF5A7ADFD0979E0" box="[634,668,1967,1990]" italics="true" pageId="52" pageNumber="53">exv</emphasis>
(anterior to right); F—partial notogaster, lateral view (upper left insert = enlarged spina adnata, dorsal view); G—seta
<emphasis id="B9139553FF9C305FFDECA7D1FDFA79CA" box="[611,623,2003,2028]" italics="true" pageId="52" pageNumber="53">lʹ</emphasis>
of genu IV. Scale bars 2 μm (C, E), 10 μm (D, G), 20 μm (F), 50 μm (A, B).
</paragraph>
</caption>
<caption id="DF1819C9FF9D305EFF18A740FA9879CC" ID-DOI="http://doi.org/10.5281/zenodo.5225249" ID-Zenodo-Dep="5225249" httpUri="https://zenodo.org/record/5225249/files/figure.png" pageId="53" pageNumber="54" startId="53.[151,250,1858,1882]" targetBox="[157,1429,181,1834]" targetPageId="53">
<paragraph id="8BD84941FF9D305EFF18A740FA9879CC" blockId="53.[151,1436,1858,2027]" pageId="53" pageNumber="54">
<emphasis id="B9139553FF9D305EFF18A740FEB4797C" bold="true" box="[151,289,1858,1882]" pageId="53" pageNumber="54">FIGURE 27.</emphasis>
<taxonomicName id="4C6732C2FF9D305EFEA7A740FD37797C" authority="(Jacot 1939)" baseAuthorityName="Jacot" baseAuthorityYear="1939" box="[296,674,1858,1882]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="craigheadi">
<emphasis id="B9139553FF9D305EFEA7A740FD8F797C" box="[296,538,1858,1882]" italics="true" pageId="53" pageNumber="54">Epidamaeus craigheadi</emphasis>
(
<bibRefCitation id="EFF634B0FF9D305EFDA7A740FD0E797C" author="Jacot, A. P." box="[552,667,1858,1882]" pageId="53" pageNumber="54" pagination="321 - 332" refId="ref37962" refString="Jacot, A. P. (1939) New mites from the White Mountains. Occasional Papers of the Boston Society of Natural History, 8, 321 - 332." type="journal article" year="1939">Jacot 1939</bibRefCitation>
)
</taxonomicName>
, light photographs of adult: A—central epimere I and base of subcapitular mentum; B—right ventrosejugal region with insert (upper right) at higher focus on trochanter III, showing cerotegument (seta
<emphasis id="B9139553FF9D305EFF18A78EFF367983" box="[151,163,1932,1957]" italics="true" pageId="53" pageNumber="54">lʹ</emphasis>
labeled for orientation); C—ventrosejugal enantiophysis, ventrolateral view (anterior to right); D—region of left discidium, ventral view (cotype); E—legs IIV; F—partial femur IV, with inserts showing enlargements of cerotegument excrescences on dorsal (lower left) and ventral (lower right) section of stalk. Scale bars 5 μm (C, D), 10 μm (A, B, F), 50 μm (E).
</paragraph>
</caption>
<paragraph id="8BD84941FF9E305DFF48A095FB347F67" blockId="54.[151,1437,150,573]" pageId="54" pageNumber="55">
<emphasis id="B9139553FF9E305DFF48A095FE917E96" box="[199,260,151,176]" italics="true" pageId="54" pageNumber="55">Legs.</emphasis>
Formulas of leg setation and solenidia counts: PN, I (023316) [112], II (023313) [111], III (122313) [110], IV (00007) [000]; DN, I (144516) [122], II (144413) [112], III (233413) [110], IV (122312) [010]; TN, I (154518) [122], II (144515) [112], III (2 33415) [110], IV (133412) [010]. Ontogeny of setae and solenidia in nymphs as in
<taxonomicName id="4C6732C2FF9E305DFB40A101FADB7F3B" baseAuthorityName="Ewing" baseAuthorityYear="1909" box="[1231,1358,259,285]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis id="B9139553FF9E305DFB40A101FADB7F3B" box="[1231,1358,259,285]" italics="true" pageId="54" pageNumber="55">E. michaeli</emphasis>
</taxonomicName>
(Table 4) except as follows: seta
<emphasis id="B9139553FF9E305DFE36A12AFE5E7F65" box="[441,459,296,323]" italics="true" pageId="54" pageNumber="55">vʹ</emphasis>
of trochanter IV deutonymphal; seta
<emphasis id="B9139553FF9E305DFCE0A125FCEE7F66" box="[879,891,295,320]" italics="true" pageId="54" pageNumber="55">v</emphasis>
ʹ of genu IV tritonymphal.
</paragraph>
<paragraph id="8BD84941FF9E305DFF48A149FC837C1B" blockId="54.[151,1437,150,573]" pageId="54" pageNumber="55">
Setal pair (
<emphasis id="B9139553FF9E305DFEC5A149FECD7F42" box="[330,344,331,356]" italics="true" pageId="54" pageNumber="55">p</emphasis>
) appears eupathidial on tarsus I of all observed instars, seta
<emphasis id="B9139553FF9E305DFBABA149FBBA7F42" box="[1060,1071,331,356]" italics="true" pageId="54" pageNumber="55">s</emphasis>
normal in juveniles (becoming eupathidial in adult). Famulus (
<figureCitation id="135C55C4FF9E305DFE77A16DFDCB7FAF" box="[504,606,367,393]" captionStart="FIGURE 32" captionStartId="59.[151,250,1462,1486]" captionTargetBox="[164,1423,844,1439]" captionTargetId="figure-307@59.[164,1423,844,1439]" captionTargetPageId="59" captionText="FIGURE 32. Epidamaeus craigheadi (Jacot 1939), light photographs of deutonymph (A), tritonymph (BE):A—middle portion of leg II, abaxial view; B—middle portion of right leg I, dorsoadaxial view; C—portion of tibia I from different specimen, abaxial view (arrow to coupled seta d); D—bulb of left tarsus I, dorsoabaxial view, showing tarsal cluster (lower right insert = isolated famulus, arrow to basal root); E—isolated solenidion ω1 and cerotegument excrescences. Scale bars 5 μm (D, E), 10 μm (AC)." figureDoi="http://doi.org/10.5281/zenodo.5225259" httpUri="https://zenodo.org/record/5225259/files/figure.png" pageId="54" pageNumber="55">Fig. 32D</figureCitation>
) acicular, emergent in all nymphs (
<quantity id="4C9FE4A4FF9E305DFC63A16DFB8D7FAF" box="[1004,1048,367,393]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="54" pageNumber="55" unit="in" value="8.0">8 in</quantity>
TN); without sunken cup but base internalized as short root (2) at least in TN (no opportunity to study in PN or DN; see R23). Solenidion ω
<subScript id="17E34B04FF9E305DFAFCA1A7FAEE7F95" attach="left" box="[1395,1403,421,435]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
of tarsus I ceratiform, distinctly curved laterad, slightly S-shaped or with third weak bend distally (
<figureCitation id="135C55C4FF9E305DFB6EA1B5FAFE7FF7" box="[1249,1387,439,465]" captionStart="FIGURE 32" captionStartId="59.[151,250,1462,1486]" captionTargetBox="[164,1423,844,1439]" captionTargetId="figure-307@59.[164,1423,844,1439]" captionTargetPageId="59" captionText="FIGURE 32. Epidamaeus craigheadi (Jacot 1939), light photographs of deutonymph (A), tritonymph (BE):A—middle portion of leg II, abaxial view; B—middle portion of right leg I, dorsoadaxial view; C—portion of tibia I from different specimen, abaxial view (arrow to coupled seta d); D—bulb of left tarsus I, dorsoabaxial view, showing tarsal cluster (lower right insert = isolated famulus, arrow to basal root); E—isolated solenidion ω1 and cerotegument excrescences. Scale bars 5 μm (D, E), 10 μm (AC)." figureDoi="http://doi.org/10.5281/zenodo.5225259" httpUri="https://zenodo.org/record/5225259/files/figure.png" pageId="54" pageNumber="55">Fig. 32D, E</figureCitation>
); ω
<subScript id="17E34B04FF9E305DFA1BA1CBFA097FF1" attach="left" box="[1428,1436,457,471]" fontSize="6" pageId="54" pageNumber="55">2</subScript>
smaller, piliform. Coupled solenidia of genua (
<figureCitation id="135C55C4FF9E305DFD14A1D9FC8B7FD3" box="[667,798,475,501]" captionStart="FIGURE 32" captionStartId="59.[151,250,1462,1486]" captionTargetBox="[164,1423,844,1439]" captionTargetId="figure-307@59.[164,1423,844,1439]" captionTargetPageId="59" captionText="FIGURE 32. Epidamaeus craigheadi (Jacot 1939), light photographs of deutonymph (A), tritonymph (BE):A—middle portion of leg II, abaxial view; B—middle portion of right leg I, dorsoadaxial view; C—portion of tibia I from different specimen, abaxial view (arrow to coupled seta d); D—bulb of left tarsus I, dorsoabaxial view, showing tarsal cluster (lower right insert = isolated famulus, arrow to basal root); E—isolated solenidion ω1 and cerotegument excrescences. Scale bars 5 μm (D, E), 10 μm (AC)." figureDoi="http://doi.org/10.5281/zenodo.5225259" httpUri="https://zenodo.org/record/5225259/files/figure.png" pageId="54" pageNumber="55">Fig. 32A, B</figureCitation>
) almost isodiametric, about 1/2 length of seta
<emphasis id="B9139553FF9E305DFA93A1D9FABF7FD3" box="[1308,1322,475,501]" italics="true" pageId="54" pageNumber="55">d</emphasis>
on genua; those of tibiae IIIII similar or ceratiform but longer, about 3/4 length of
<emphasis id="B9139553FF9E305DFC42A1FDFC4E7C3F" box="[973,987,511,537]" italics="true" pageId="54" pageNumber="55">d</emphasis>
, that of tibiae I (φ
<subScript id="17E34B04FF9E305DFB26A213FB247C39" attach="left" box="[1193,1201,529,543]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
) and IV flagellate or subflagellate (tactile;
<figureCitation id="135C55C4FF9E305DFE13A221FDB47C1B" box="[412,545,547,573]" captionStart="FIGURE 32" captionStartId="59.[151,250,1462,1486]" captionTargetBox="[164,1423,844,1439]" captionTargetId="figure-307@59.[164,1423,844,1439]" captionTargetPageId="59" captionText="FIGURE 32. Epidamaeus craigheadi (Jacot 1939), light photographs of deutonymph (A), tritonymph (BE):A—middle portion of leg II, abaxial view; B—middle portion of right leg I, dorsoadaxial view; C—portion of tibia I from different specimen, abaxial view (arrow to coupled seta d); D—bulb of left tarsus I, dorsoabaxial view, showing tarsal cluster (lower right insert = isolated famulus, arrow to basal root); E—isolated solenidion ω1 and cerotegument excrescences. Scale bars 5 μm (D, E), 10 μm (AC)." figureDoi="http://doi.org/10.5281/zenodo.5225259" httpUri="https://zenodo.org/record/5225259/files/figure.png" pageId="54" pageNumber="55">Fig. 32B, C</figureCitation>
), much longer than
<emphasis id="B9139553FF9E305DFC8EA221FC9A7C1B" box="[769,783,547,573]" italics="true" pageId="54" pageNumber="55">d</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>