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<document id="A29747F5CD2F57D54395C92D0B762306" ID-DOI="10.1080/00222930500415195" ID-ISSN="1464-5262" ID-Zenodo-Dep="5220912" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629366272137" checkinUser="felipe" docAuthor="Dick, Matthew H., Grischenko, Andrei V. &amp; Mawatari, Shunsuke F." docDate="2005" docId="03CE7B54FFF7FFF6DEB2186A8938BCD0" docLanguage="en" docName="JNatHist.39.43.3687-3784.pdf" docOrigin="Journal of Natural History 39 (43)" docSource="http://dx.doi.org/10.1080/00222930500415195" docStyle="DocumentStyle:5A9EB2ED5589A523F0236C66DE2A6BA8.4:JNatHist.2005-2007.journal_article.1cover" docStyleId="5A9EB2ED5589A523F0236C66DE2A6BA8" docStyleName="JNatHist.2005-2007.journal_article.1cover" docStyleVersion="4" docTitle="Celleporella nodasakae Dick &amp; Grischenko &amp; Mawatari 2005, new species" docType="treatment" docVersion="3" lastPageNumber="3732" masterDocId="FFF7032CFFDCFFD8DF1B1B7F8A29BE37" masterDocTitle="Intertidal Bryozoa (Cheilostomata) of Ketchikan, Alaska" masterLastPageNumber="3784" masterPageNumber="3687" pageNumber="3729" updateTime="1699069551647" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="913EA6AF0CDE8251EB4850A657E3CAC0">Intertidal Bryozoa (Cheilostomata) of Ketchikan, Alaska</mods:title>
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<mods:namePart id="604838F52654AF6C6227A2A3AC27BCB3">Dick, Matthew H.</mods:namePart>
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<mods:namePart id="CB53653FFE9809CDF2AF6298F1B63F80">Grischenko, Andrei V.</mods:namePart>
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<mods:namePart id="7A5FBB8DC84A65835FD3A76FD8E7E0D5">Mawatari, Shunsuke F.</mods:namePart>
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<mods:title id="30D5CC58833115EDC1FB5AD61E58D49F">Journal of Natural History</mods:title>
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<mods:date id="E1AE3ECA36721073CFB35620E834C1F8">2005</mods:date>
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<mods:number id="849013D7735CFFCA64AC56A419ABD644">2005-12-23</mods:number>
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<heading id="D0907D2EFFF7FFF3DEB2186A8946BD1A" box="[425,879,789,813]" centered="true" fontSize="10" level="2" pageId="43" pageNumber="3729" reason="2">
<taxonomicName id="4C67B1C1FFF7FFF3DEB2186A88FFBD1A" ID-CoL="69GWW" authority="Dick &amp; Grischenko &amp; Mawatari, 2005" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" box="[425,726,789,813]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="43" pageNumber="3729" phylum="Bryozoa" rank="species" species="nodasakae" status="sp. nov.">
<emphasis id="B9131650FFF7FFF3DEB2186A88FFBD1A" bold="true" box="[425,726,789,813]" italics="true" pageId="43" pageNumber="3729">Celleporella nodasakae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A220AB2BFFF7FFF3DDFC186A8946BD1A" box="[743,879,789,813]" pageId="43" pageNumber="3729" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8BD8CA42FFF7FFF3DD35184988C8BD79" blockId="43.[425,879,789,846]" box="[558,737,822,846]" pageId="43" pageNumber="3729">
(
<figureCitation id="135CD6C7FFF7FFF3DD2D184988F0BD79" box="[566,729,822,846]" captionStart="Figure 11" captionStartId="44.[125,185,1239,1258]" captionTargetBox="[125,1173,133,1203]" captionTargetId="figure-61@44.[125,1173,133,1203]" captionTargetPageId="44" captionText="Figure 11. Celleporella nodasakae n. sp.: (A) general colony view, showing uniserial branch giving rise to coherent sheet followed by oligoserial rami; (B) autozooids and an ovicellate zooid; (C) enlargement of autozooidal orifice; (D) enlargement of ovicell and female orifice; (E) photomicrograph of several autozooids from basal side, showing interzooidal connections; az, autozooid; o, orifice; tpc, tubular pore chamber; lc, interzooidal lacuna; (F) ancestrula (bottom, right of centre) and first daughter zooid (bottom, left of centre); from proximal margin of latter is budded a zooid directed perpendicularly. All specimens except (F) were bleached. Scale bars: 2 mm (A); 200 Mm (B); 50 Mm (C); 100 Mm (D); 250 Mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5220930" httpUri="https://zenodo.org/record/5220930/files/figure.png" pageId="43" pageNumber="3729">Figure 11AF</figureCitation>
)
</paragraph>
<paragraph id="8BD8CA42FFF7FFF3DF9118F88ADABDA8" blockId="43.[138,243,903,927]" box="[138,243,903,927]" pageId="43" pageNumber="3729">
<emphasis id="B9131650FFF7FFF3DF9118F88ADABDA8" box="[138,243,903,927]" italics="true" pageId="43" pageNumber="3729">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD8CA42FFF7FFF3DF9118C78B3BBA86" blockId="43.[138,1185,952,1201]" pageId="43" pageNumber="3729">Colony strictly unilaminar; growth form uniserial, biserial, oligoserial, aggregated, or coherent, often in same colony; only autozooids and ovicellate female zooids occur; male polymorphs and zooeciules lacking; zooids loosely connected by tubular pore chambers with conspicuous lacunae between them; ovicells with four to five pores, generally around perimeter; autozooidal orifice with a deep, U-shaped sinus and sharp condyles pointing distally, flanked by a notch; proximal margin of female orifice Mshaped, with a U- or rounded V-shaped sinus between triangular, distally directed projections.</paragraph>
<paragraph id="8BD8CA42FFF7FFF3DF911F958B0BBB35" blockId="43.[138,290,1258,1282]" box="[138,290,1258,1282]" pageId="43" pageNumber="3729">
<emphasis id="B9131650FFF7FFF3DF911F958B0BBB35" box="[138,290,1258,1282]" italics="true" pageId="43" pageNumber="3729">Type material</emphasis>
</paragraph>
<paragraph id="8BD8CA42FFF7FFF3DF911E658E55BBE4" blockId="43.[138,1185,1306,1491]" pageId="43" pageNumber="3729">
<materialsCitation id="3B0FC01FFFF7FFF3DF911E658E38BB05" accessionNumber="YPM 35839" box="[138,1041,1306,1330]" collectionCode="YPM" pageId="43" pageNumber="3729" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DC74E0FFF7FFF3DF911E658AD3BB05" box="[138,250,1306,1330]" pageId="43" pageNumber="3729" type="holotype">Holotype</typeStatus>
: SC, specimen KE-26 bleached and coated for SEM (
<collectionCode id="ED765287FFF7FFF3DC6B1E658998BB05" box="[880,945,1306,1330]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="43" pageNumber="3729" type="Museum">YPM</collectionCode>
35839)
</materialsCitation>
.
<materialsCitation id="3B0FC01FFFF7FFF3DB061E6589E2BB64" accessionNumber="YPM 35840" collectionCode="YPM" pageId="43" pageNumber="3729" specimenCount="1" typeStatus="paratype">
<typeStatus id="54DC74E0FFF7FFF3DB061E658EABBB05" box="[1053,1154,1306,1330]" pageId="43" pageNumber="3729" type="paratype">Paratype</typeStatus>
1: SC, specimen KE-24 bleached and coated for SEM (
<collectionCode id="ED765287FFF7FFF3DC3F1E44894CBB64" box="[804,869,1339,1363]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="43" pageNumber="3729" type="Museum">YPM</collectionCode>
35840)
</materialsCitation>
.
<typeStatus id="54DC74E0FFF7FFF3DCC61E448E6BBB64" box="[989,1090,1339,1363]" pageId="43" pageNumber="3729" type="paratype">Paratype</typeStatus>
2: SC, specimen KE-119 bleached and coated for SEM (NHM 2005.7.11.4).
<materialsCitation id="3B0FC01FFFF7FFF3DCF81E248EB4BBA4" accessionNumber="YPM 35841" collectionCode="YPM" pageId="43" pageNumber="3729" specimenCount="1" typeStatus="paratype">
<typeStatus id="54DC74E0FFF7FFF3DCF81E248E61BB44" box="[995,1096,1371,1395]" pageId="43" pageNumber="3729" type="paratype">Paratype</typeStatus>
3: SC, specimen KE-57 on bivalve fragment, lightly bleached and coated for SEM (
<collectionCode id="ED765287FFF7FFF3DCE61E048E17BBA4" box="[1021,1086,1403,1427]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="43" pageNumber="3729" type="Museum">YPM</collectionCode>
35841)
</materialsCitation>
.
<materialsCitation id="3B0FC01FFFF7FFF3DF911EE48EB4BB84" box="[138,1181,1435,1459]" collectionCode="ZIRAS" pageId="43" pageNumber="3729" specimenCode="ZIRAS 1" specimenCount="1" typeStatus="paratype">
<typeStatus id="54DC74E0FFF7FFF3DF911EE48AC6BB84" box="[138,239,1435,1459]" pageId="43" pageNumber="3729" type="paratype">Paratype</typeStatus>
4: SC, specimen KE-32 bleached and coated for SEM (
<specimenCode id="DBC16239FFF7FFF3DCD01EE48E11BB84" box="[971,1080,1435,1459]" collectionCode="ZIRAS" pageId="43" pageNumber="3729">ZIRAS 1</specimenCode>
/50530)
</materialsCitation>
.
<typeStatus id="54DC74E0FFF7FFF3DF911EC48AC6BBE4" box="[138,239,1467,1491]" pageId="43" pageNumber="3729" type="paratype">Paratype</typeStatus>
5: SC, specimen KE-13 bleached and coated for SEM (NHM 2005.7.11.5).
</paragraph>
<paragraph id="8BD8CA42FFF7FFF3DF911D738AD2B813" blockId="43.[138,251,1548,1572]" box="[138,251,1548,1572]" pageId="43" pageNumber="3729">
<emphasis id="B9131650FFF7FFF3DF911D738AD2B813" box="[138,251,1548,1572]" italics="true" pageId="43" pageNumber="3729">Etymology</emphasis>
</paragraph>
<paragraph id="8BD8CA42FFF7FFF3DF911D42884FB8A2" blockId="43.[138,1185,1597,1685]" pageId="43" pageNumber="3729">
The species is named for Dr Yoshinobu Nodasaka,
<collectingRegion id="49A304A0FFF7FFF3DC391D4289BDB862" box="[802,916,1597,1621]" country="Japan" name="Hokkaido" pageId="43" pageNumber="3729">Hokkaido</collectingRegion>
University School of Dentistry, to whom we are deeply indebted for cheerful assistance with scanning electron microscopy over a period of many years.
</paragraph>
<caption id="DF189ACAFFF0FFF4DF661FA88808BBBD" ID-DOI="http://doi.org/10.5281/zenodo.5220930" ID-Zenodo-Dep="5220930" httpUri="https://zenodo.org/record/5220930/files/figure.png" pageId="44" pageNumber="3730" startId="44.[125,185,1239,1258]" targetBox="[125,1173,133,1203]" targetPageId="44">
<paragraph id="8BD8CA42FFF0FFF4DF661FA88808BBBD" blockId="44.[125,1173,1239,1418]" pageId="44" pageNumber="3730">
Figure 11.
<taxonomicName id="4C67B1C1FFF0FFF4DFF31FA88B85BADD" authority="Dick &amp; Grischenko &amp; Mawatari, 2005" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" box="[232,428,1239,1258]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="44" pageNumber="3730" phylum="Bryozoa" rank="species" species="nodasakae" status="sp. nov.">
<emphasis id="B9131650FFF0FFF4DFF31FA88B85BADD" box="[232,428,1239,1258]" italics="true" pageId="44" pageNumber="3730">Celleporella nodasakae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220AB2BFFF0FFF4DEA91FA88BC1BADD" box="[434,488,1239,1258]" pageId="44" pageNumber="3730" rank="species">n. sp.</taxonomicNameLabel>
: (A) general colony view, showing uniserial branch giving rise to coherent sheet followed by oligoserial rami; (B) autozooids and an ovicellate zooid; (C) enlargement of autozooidal orifice; (D) enlargement of ovicell and female orifice; (E) photomicrograph of several autozooids from basal side, showing interzooidal connections; az, autozooid; o, orifice; tpc, tubular pore chamber; lc, interzooidal lacuna; (F) ancestrula (bottom, right of centre) and first daughter zooid (bottom, left of centre); from proximal margin of latter is budded a zooid directed perpendicularly. All specimens except (F) were bleached. Scale bars: 2 mm (A); 200 Mm (B); 50 Mm (C); 100 Mm (D); 250 Mm (E, F).
</paragraph>
</caption>
<paragraph id="8BD8CA42FFF0FFF4DF661ED18ADCBBF1" blockId="44.[125,245,1454,1478]" box="[125,245,1454,1478]" pageId="44" pageNumber="3730">
<emphasis id="B9131650FFF0FFF4DF661ED18ADCBBF1" box="[125,245,1454,1478]" italics="true" pageId="44" pageNumber="3730">Description</emphasis>
</paragraph>
<paragraph id="8BD8CA42FFF0FFF4DF661EA18AFEB8A2" blockId="44.[125,1172,1502,1685]" pageId="44" pageNumber="3730">
<emphasis id="B9131650FFF0FFF4DF661EA18AFBBBC1" box="[125,210,1502,1526]" italics="true" pageId="44" pageNumber="3730">Colony.</emphasis>
White, vitreous; strictly unilaminar; at Ketchikan found on rocks and dead bivalve shells. Zooids arranged in branching uniserial, biserial, or oligoserial series; an anastomosing, aggregated network; or a coherent crust; sometimes a single colony displays most or all of these forms in different parts (
<figureCitation id="135CD6C7FFF0FFF4DC651D428E2CB862" box="[894,1029,1597,1621]" captionStart="Figure 11" captionStartId="44.[125,185,1239,1258]" captionTargetBox="[125,1173,133,1203]" captionTargetId="figure-61@44.[125,1173,133,1203]" captionTargetPageId="44" captionText="Figure 11. Celleporella nodasakae n. sp.: (A) general colony view, showing uniserial branch giving rise to coherent sheet followed by oligoserial rami; (B) autozooids and an ovicellate zooid; (C) enlargement of autozooidal orifice; (D) enlargement of ovicell and female orifice; (E) photomicrograph of several autozooids from basal side, showing interzooidal connections; az, autozooid; o, orifice; tpc, tubular pore chamber; lc, interzooidal lacuna; (F) ancestrula (bottom, right of centre) and first daughter zooid (bottom, left of centre); from proximal margin of latter is budded a zooid directed perpendicularly. All specimens except (F) were bleached. Scale bars: 2 mm (A); 200 Mm (B); 50 Mm (C); 100 Mm (D); 250 Mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5220930" httpUri="https://zenodo.org/record/5220930/files/figure.png" pageId="44" pageNumber="3730">Figure 11A</figureCitation>
). Ovicellate female zooids interspersed with autozooids (
<figureCitation id="135CD6C7FFF0FFF4DD651D22892DB842" box="[638,772,1629,1653]" captionStart="Figure 11" captionStartId="44.[125,185,1239,1258]" captionTargetBox="[125,1173,133,1203]" captionTargetId="figure-61@44.[125,1173,133,1203]" captionTargetPageId="44" captionText="Figure 11. Celleporella nodasakae n. sp.: (A) general colony view, showing uniserial branch giving rise to coherent sheet followed by oligoserial rami; (B) autozooids and an ovicellate zooid; (C) enlargement of autozooidal orifice; (D) enlargement of ovicell and female orifice; (E) photomicrograph of several autozooids from basal side, showing interzooidal connections; az, autozooid; o, orifice; tpc, tubular pore chamber; lc, interzooidal lacuna; (F) ancestrula (bottom, right of centre) and first daughter zooid (bottom, left of centre); from proximal margin of latter is budded a zooid directed perpendicularly. All specimens except (F) were bleached. Scale bars: 2 mm (A); 200 Mm (B); 50 Mm (C); 100 Mm (D); 250 Mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5220930" httpUri="https://zenodo.org/record/5220930/files/figure.png" pageId="44" pageNumber="3730">Figure 11B</figureCitation>
); male polymorphs and zooeciules lacking.
</paragraph>
<paragraph id="8BD8CA42FFF1FFF5DF911BF6880DBFF3" blockId="45.[138,1185,137,452]" pageId="45" pageNumber="3731">
<emphasis id="B9131650FFF1FFF5DF911BF68AF0BE96" box="[138,217,137,161]" italics="true" pageId="45" pageNumber="3731">Zooids.</emphasis>
Autozooids oval or spindle-shaped, sometimes with a cauda one-third to two-fifths of zooid length, tapering to one-third to one-half of zooid width; separated by a groove, more-or-less loosely appressed to one another, with three to five conspicuous lacunae along each lateral margin between coarse tubular connections;
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long (average
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,
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530, 4) by
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wide (average
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,
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530, 4). Ovicellate zooids triangular, smaller than autozooids,
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long including ovicell (average
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,
<emphasis id="B9131650FFF1FFF5DE961A338BB2BF54" box="[397,411,332,355]" italics="true" pageId="45" pageNumber="3731">n</emphasis>
530, 4) by
<quantity id="4C9F67A7FFF1FFF5DD2C1A3488F7BF54" box="[567,734,331,355]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="45" pageNumber="3731" unit="mm" value="0.19" valueMax="0.25" valueMin="0.13">0.130.25 mm</quantity>
wide at orifice (average
<quantity id="4C9F67A7FFF1FFF5DB151A348EB5BF54" box="[1038,1180,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="5.0192" pageId="45" pageNumber="3731" unit="mm" value="50.192">50.192 mm</quantity>
,
<emphasis id="B9131650FFF1FFF5DF911A138AB1BFB4" box="[138,152,364,387]" italics="true" pageId="45" pageNumber="3731">n</emphasis>
530, 4). Zooids interconnect by two tubular pore chambers distally and four to six along each lateral margin (
<figureCitation id="135CD6C7FFF1FFF5DE941AF3883CBF93" box="[399,533,396,420]" captionStart="Figure 11" captionStartId="44.[125,185,1239,1258]" captionTargetBox="[125,1173,133,1203]" captionTargetId="figure-61@44.[125,1173,133,1203]" captionTargetPageId="44" captionText="Figure 11. Celleporella nodasakae n. sp.: (A) general colony view, showing uniserial branch giving rise to coherent sheet followed by oligoserial rami; (B) autozooids and an ovicellate zooid; (C) enlargement of autozooidal orifice; (D) enlargement of ovicell and female orifice; (E) photomicrograph of several autozooids from basal side, showing interzooidal connections; az, autozooid; o, orifice; tpc, tubular pore chamber; lc, interzooidal lacuna; (F) ancestrula (bottom, right of centre) and first daughter zooid (bottom, left of centre); from proximal margin of latter is budded a zooid directed perpendicularly. All specimens except (F) were bleached. Scale bars: 2 mm (A); 200 Mm (B); 50 Mm (C); 100 Mm (D); 250 Mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5220930" httpUri="https://zenodo.org/record/5220930/files/figure.png" pageId="45" pageNumber="3731">Figure 11E</figureCitation>
). Basal wall with a narrow, marginal calcified shelf surrounding the uncalcified centre.
</paragraph>
<paragraph id="8BD8CA42FFF1FFF5DF911A92893CBC12" blockId="45.[138,1184,493,549]" pageId="45" pageNumber="3731">
<emphasis id="B9131650FFF1FFF5DF911A928B34BC32" box="[138,285,493,517]" italics="true" pageId="45" pageNumber="3731">Frontal wall.</emphasis>
Thin, vitreous, translucent, imperforate, convex, with transverse striae or weak transverse ridges; umbo lacking in our specimens.
</paragraph>
<paragraph id="8BD8CA42FFF1FFF5DF9119318E88BD3F" blockId="45.[138,1185,590,776]" pageId="45" pageNumber="3731">
<emphasis id="B9131650FFF1FFF5DF9119318AF5BC51" box="[138,220,590,614]" italics="true" pageId="45" pageNumber="3731">Orifice.</emphasis>
Autozooidal orifice (
<figureCitation id="135CD6C7FFF1FFF5DEC019318848BC51" box="[475,609,590,614]" captionStart="Figure 11" captionStartId="44.[125,185,1239,1258]" captionTargetBox="[125,1173,133,1203]" captionTargetId="figure-61@44.[125,1173,133,1203]" captionTargetPageId="44" captionText="Figure 11. Celleporella nodasakae n. sp.: (A) general colony view, showing uniserial branch giving rise to coherent sheet followed by oligoserial rami; (B) autozooids and an ovicellate zooid; (C) enlargement of autozooidal orifice; (D) enlargement of ovicell and female orifice; (E) photomicrograph of several autozooids from basal side, showing interzooidal connections; az, autozooid; o, orifice; tpc, tubular pore chamber; lc, interzooidal lacuna; (F) ancestrula (bottom, right of centre) and first daughter zooid (bottom, left of centre); from proximal margin of latter is budded a zooid directed perpendicularly. All specimens except (F) were bleached. Scale bars: 2 mm (A); 200 Mm (B); 50 Mm (C); 100 Mm (D); 250 Mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5220930" httpUri="https://zenodo.org/record/5220930/files/figure.png" pageId="45" pageNumber="3731">Figure 11C</figureCitation>
) slightly longer than broad,
<quantity id="4C9F67A7FFF1FFF5DCA719318E4ABC51" box="[956,1123,590,614]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="45" pageNumber="3731" unit="mm" value="0.075" valueMax="0.09" valueMin="0.06">0.060.09 mm</quantity>
long (average
<quantity id="4C9F67A7FFF1FFF5DFF019118B50BCB0" box="[235,377,622,647]" metricMagnitude="-2" metricUnit="m" metricValue="5.0074" pageId="45" pageNumber="3731" unit="mm" value="50.074">50.074 mm</quantity>
,
<emphasis id="B9131650FFF1FFF5DE9019108BB0BCB1" box="[395,409,623,646]" italics="true" pageId="45" pageNumber="3731">n</emphasis>
530, 4) by
<quantity id="4C9F67A7FFF1FFF5DD2A191088F1BCB0" box="[561,728,623,647]" metricMagnitude="-5" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="45" pageNumber="3731" unit="mm" value="0.065" valueMax="0.08" valueMin="0.05">0.050.08 mm</quantity>
wide (average
<quantity id="4C9F67A7FFF1FFF5DC9219118E31BCB0" box="[905,1048,622,647]" metricMagnitude="-2" metricUnit="m" metricValue="5.0065" pageId="45" pageNumber="3731" unit="mm" value="50.065">50.065 mm</quantity>
,
<emphasis id="B9131650FFF1FFF5DB3119108E11BCB1" box="[1066,1080,623,646]" italics="true" pageId="45" pageNumber="3731">n</emphasis>
530, 4); proximal margin with a deep, broadly U-shaped sinus flanked by small, sharp, distally or distomedially directed condyles separated by a notch from lateral orificial margin. Female secondary orifice (
<figureCitation id="135CD6C7FFF1FFF5DE7319AF8BC6BCDF" box="[360,495,720,744]" captionStart="Figure 11" captionStartId="44.[125,185,1239,1258]" captionTargetBox="[125,1173,133,1203]" captionTargetId="figure-61@44.[125,1173,133,1203]" captionTargetPageId="44" captionText="Figure 11. Celleporella nodasakae n. sp.: (A) general colony view, showing uniserial branch giving rise to coherent sheet followed by oligoserial rami; (B) autozooids and an ovicellate zooid; (C) enlargement of autozooidal orifice; (D) enlargement of ovicell and female orifice; (E) photomicrograph of several autozooids from basal side, showing interzooidal connections; az, autozooid; o, orifice; tpc, tubular pore chamber; lc, interzooidal lacuna; (F) ancestrula (bottom, right of centre) and first daughter zooid (bottom, left of centre); from proximal margin of latter is budded a zooid directed perpendicularly. All specimens except (F) were bleached. Scale bars: 2 mm (A); 200 Mm (B); 50 Mm (C); 100 Mm (D); 250 Mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5220930" httpUri="https://zenodo.org/record/5220930/files/figure.png" pageId="45" pageNumber="3731">Figure 11D</figureCitation>
) semicircular; proximal margin M-shaped, with a U- or rounded V-shaped median sinus flanked by coarse, triangular, distally pointing projections.
</paragraph>
<paragraph id="8BD8CA42FFF1FFF5DF91184E8AFFBDFC" blockId="45.[138,1185,817,971]" pageId="45" pageNumber="3731">
<emphasis id="B9131650FFF1FFF5DF91184E8B8ABD7E" box="[138,419,817,841]" italics="true" pageId="45" pageNumber="3731">
Ovicell (
<figureCitation id="135CD6C7FFF1FFF5DFF1184E8BBCBD7E" box="[234,405,817,841]" captionStart="Figure 11" captionStartId="44.[125,185,1239,1258]" captionTargetBox="[125,1173,133,1203]" captionTargetId="figure-61@44.[125,1173,133,1203]" captionTargetPageId="44" captionText="Figure 11. Celleporella nodasakae n. sp.: (A) general colony view, showing uniserial branch giving rise to coherent sheet followed by oligoserial rami; (B) autozooids and an ovicellate zooid; (C) enlargement of autozooidal orifice; (D) enlargement of ovicell and female orifice; (E) photomicrograph of several autozooids from basal side, showing interzooidal connections; az, autozooid; o, orifice; tpc, tubular pore chamber; lc, interzooidal lacuna; (F) ancestrula (bottom, right of centre) and first daughter zooid (bottom, left of centre); from proximal margin of latter is budded a zooid directed perpendicularly. All specimens except (F) were bleached. Scale bars: 2 mm (A); 200 Mm (B); 50 Mm (C); 100 Mm (D); 250 Mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5220930" httpUri="https://zenodo.org/record/5220930/files/figure.png" pageId="45" pageNumber="3731">Figure 11B, D</figureCitation>
).
</emphasis>
Raised, globose, nearly circular in outline,
<quantity id="4C9F67A7FFF1FFF5DCA0184E8E4BBD7E" box="[955,1122,817,841]" metricMagnitude="-4" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="45" pageNumber="3731" unit="mm" value="0.195" valueMax="0.23" valueMin="0.16">0.160.23 mm</quantity>
long (average
<quantity id="4C9F67A7FFF1FFF5DFF0182E8B50BD5D" box="[235,377,849,874]" metricMagnitude="-2" metricUnit="m" metricValue="5.0185" pageId="45" pageNumber="3731" unit="mm" value="50.185">50.185 mm</quantity>
,
<emphasis id="B9131650FFF1FFF5DE90182D8BB0BD5E" box="[395,409,850,873]" italics="true" pageId="45" pageNumber="3731">n</emphasis>
530, 4) by
<quantity id="4C9F67A7FFF1FFF5DD2A182D88F1BD5D" box="[561,728,850,874]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="2.4" metricValueMin="1.6" pageId="45" pageNumber="3731" unit="mm" value="0.2" valueMax="0.24" valueMin="0.16">0.160.24 mm</quantity>
wide (average
<quantity id="4C9F67A7FFF1FFF5DC92182E8E31BD5D" box="[905,1048,849,874]" metricMagnitude="-2" metricUnit="m" metricValue="5.0204" pageId="45" pageNumber="3731" unit="mm" value="50.204">50.204 mm</quantity>
,
<emphasis id="B9131650FFF1FFF5DB31182D8E11BD5E" box="[1066,1080,850,873]" italics="true" pageId="45" pageNumber="3731">n</emphasis>
530, 4); with four to five pores scattered mostly around the sloping perimeter, generally away from the central area; sculptured with faint concentric growth lines and sometimes a slight umbo.
</paragraph>
<paragraph id="8BD8CA42FFF1FFF5DF91188B8BDABA3B" blockId="45.[138,499,1012,1036]" box="[138,499,1012,1036]" pageId="45" pageNumber="3731">
<emphasis id="B9131650FFF1FFF5DF91188B8BABBA3B" box="[138,386,1012,1036]" italics="true" pageId="45" pageNumber="3731">Spines and avicularia.</emphasis>
Lacking.
</paragraph>
<paragraph id="8BD8CA42FFF1FFF5DF911F4B8B9EBA99" blockId="45.[138,1185,1076,1198]" pageId="45" pageNumber="3731">
<emphasis id="B9131650FFF1FFF5DF911F4B8B2CBA7B" box="[138,261,1076,1100]" italics="true" pageId="45" pageNumber="3731">
<taxonomicName id="4C67B1C1FFF1FFF5DF911F4B8B28BA7B" authorityName="Delage &amp; Heroard" authorityYear="1897" box="[138,257,1076,1100]" genus="Ancestrula" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="3731" phylum="Bryozoa" rank="genus">Ancestrula</taxonomicName>
.
</emphasis>
One observed (
<figureCitation id="135CD6C7FFF1FFF5DEDF1F4B8861BA7B" box="[452,584,1076,1100]" captionStart="Figure 11" captionStartId="44.[125,185,1239,1258]" captionTargetBox="[125,1173,133,1203]" captionTargetId="figure-61@44.[125,1173,133,1203]" captionTargetPageId="44" captionText="Figure 11. Celleporella nodasakae n. sp.: (A) general colony view, showing uniserial branch giving rise to coherent sheet followed by oligoserial rami; (B) autozooids and an ovicellate zooid; (C) enlargement of autozooidal orifice; (D) enlargement of ovicell and female orifice; (E) photomicrograph of several autozooids from basal side, showing interzooidal connections; az, autozooid; o, orifice; tpc, tubular pore chamber; lc, interzooidal lacuna; (F) ancestrula (bottom, right of centre) and first daughter zooid (bottom, left of centre); from proximal margin of latter is budded a zooid directed perpendicularly. All specimens except (F) were bleached. Scale bars: 2 mm (A); 200 Mm (B); 50 Mm (C); 100 Mm (D); 250 Mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5220930" httpUri="https://zenodo.org/record/5220930/files/figure.png" pageId="45" pageNumber="3731">Figure 11F</figureCitation>
); similar in form to subsequent autozooids, though smaller,
<quantity id="4C9F67A7FFF1FFF5DFEB1F2A8B7FBA5A" box="[240,342,1109,1133]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="45" pageNumber="3731" unit="mm" value="0.3">0.30 mm</quantity>
long by
<quantity id="4C9F67A7FFF1FFF5DEDD1F2A8805BA5A" box="[454,556,1109,1133]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="45" pageNumber="3731" unit="mm" value="0.2">0.20 mm</quantity>
wide. This ancestrula produced one daughter zooid distally, which in turn produced two buds, one distally and the other at the proximolateral margin, directed laterally.
</paragraph>
<paragraph id="8BD8CA42FFF1FFF5DF911F998AC0BAC9" blockId="45.[138,233,1254,1278]" box="[138,233,1254,1278]" pageId="45" pageNumber="3731">
<emphasis id="B9131650FFF1FFF5DF911F998AC0BAC9" box="[138,233,1254,1278]" italics="true" pageId="45" pageNumber="3731">Remarks</emphasis>
</paragraph>
<paragraph id="8BD8CA42FFF1FFF6DF911E6888BBBF36" blockId="45.[138,1185,1303,1684]" lastBlockId="46.[125,1173,137,614]" lastPageId="46" lastPageNumber="3732" pageId="45" pageNumber="3731">
We have placed this species in
<taxonomicName id="4C67B1C1FFF1FFF5DEE21E68885DBB18" authorityName="Gray" authorityYear="1848" box="[505,628,1303,1327]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="45" pageNumber="3731" phylum="Bryozoa" rank="genus">
<emphasis id="B9131650FFF1FFF5DEE21E68885DBB18" box="[505,628,1303,1327]" italics="true" pageId="45" pageNumber="3731">Celleporella</emphasis>
</taxonomicName>
, rather than
<taxonomicName id="4C67B1C1FFF1FFF5DC0A1E6889AABB18" authorityName="Lamouroux" authorityYear="1821" box="[785,899,1303,1327]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" kingdom="Animalia" order="Cheilostomatida" pageId="45" pageNumber="3731" phylum="Bryozoa" rank="genus">
<emphasis id="B9131650FFF1FFF5DC0A1E6889AABB18" box="[785,899,1303,1327]" italics="true" pageId="45" pageNumber="3731">Hippothoa</emphasis>
</taxonomicName>
, because of its coherent growth form in at least parts of colonies, and its tubular pore chambers.
<taxonomicName id="4C67B1C1FFF1FFF5DB3D1E478AD5BB47" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="45" pageNumber="3731" phylum="Bryozoa" rank="species" species="nodasakae">
<emphasis id="B9131650FFF1FFF5DB3D1E478AD5BB47" italics="true" pageId="45" pageNumber="3731">Celleporella nodasakae</emphasis>
</taxonomicName>
is similar in growth form to
<taxonomicName id="4C67B1C1FFF1FFF5DD7C1E2789BEBB47" authority="Dick and Ross" authorityName="Dick and Ross" authorityYear="1988" box="[615,919,1368,1392]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="45" pageNumber="3731" phylum="Bryozoa" rank="species" species="reflexa">
<emphasis id="B9131650FFF1FFF5DD7C1E2788FCBB47" box="[615,725,1368,1392]" italics="true" pageId="45" pageNumber="3731">C. reflexa</emphasis>
Dick and Ross
</taxonomicName>
; both can range from uniserial to narrowly multiserial to coherent, and both can have female zooids in the primary layer. However, compared to
<taxonomicName id="4C67B1C1FFF1FFF5DD5D1EE68886BB86" authorityName="Dick and Ross" authorityYear="1988" box="[582,687,1433,1457]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="45" pageNumber="3731" phylum="Bryozoa" rank="species" species="reflexa">
<emphasis id="B9131650FFF1FFF5DD5D1EE68886BB86" box="[582,687,1433,1457]" italics="true" pageId="45" pageNumber="3731">C. reflexa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C67B1C1FFF1FFF5DDA41EE6897DBB86" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" box="[703,852,1433,1457]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="45" pageNumber="3731" phylum="Bryozoa" rank="species" species="nodasakae">
<emphasis id="B9131650FFF1FFF5DDA41EE6897DBB86" box="[703,852,1433,1457]" italics="true" pageId="45" pageNumber="3731">C. nodasakae</emphasis>
</taxonomicName>
has a greater tendency to be uniserial; lacks male zooids and zooeciules; was never observed to become multilaminar; has a distinctly smaller autozooidal orifice; has fewer pore chambers; has fewer, larger interzooidal lacunae; has fewer pores in the ovicell; and has the orifice of ovicellate zooids with a diagnostic, M-shaped proximal margin. The ancestrular budding pattern of
<taxonomicName id="4C67B1C1FFF1FFF5DB9D1D648AD5B864" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="45" pageNumber="3731" phylum="Bryozoa" rank="species" species="nodasakae">
<emphasis id="B9131650FFF1FFF5DB9D1D648AD5B864" italics="true" pageId="45" pageNumber="3731">C. nodasakae</emphasis>
</taxonomicName>
is somewhat similar to that of
<taxonomicName id="4C67B1C1FFF1FFF5DD461D4488ECB864" authorityName="Dick and Ross" authorityYear="1988" box="[605,709,1595,1619]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="45" pageNumber="3731" phylum="Bryozoa" rank="species" species="reflexa">
<emphasis id="B9131650FFF1FFF5DD461D4488ECB864" box="[605,709,1595,1619]" italics="true" pageId="45" pageNumber="3731">C. reflexa</emphasis>
</taxonomicName>
; in both species, the ancestrula first buds a distolateral daughter zooid, from which arises another zooid from the proximolateral margin. However, whereas this proximolaterally budded zooid grows back alongside the ancestrula in
<taxonomicName id="4C67B1C1FFF2FFF6DE0C1BF68B57BE96" authorityName="Dick and Ross" authorityYear="1988" box="[279,382,137,161]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="reflexa">
<emphasis id="B9131650FFF2FFF6DE0C1BF68B57BE96" box="[279,382,137,161]" italics="true" pageId="46" pageNumber="3732">C. reflexa</emphasis>
</taxonomicName>
, it grows laterally or proximolaterally in
<taxonomicName id="4C67B1C1FFF2FFF6DC541BF689CABE96" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" box="[847,995,137,161]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="nodasakae">
<emphasis id="B9131650FFF2FFF6DC541BF689CABE96" box="[847,995,137,161]" italics="true" pageId="46" pageNumber="3732">C. nodasakae</emphasis>
</taxonomicName>
, away from the ancestrula. We found both species in the Ketchikan vicinity; however, whereas
<taxonomicName id="4C67B1C1FFF2FFF6DB321BD68EBDBEF6" authorityName="Dick and Ross" authorityYear="1988" box="[1065,1172,169,193]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="reflexa">
<emphasis id="B9131650FFF2FFF6DB321BD68EBDBEF6" box="[1065,1172,169,193]" italics="true" pageId="46" pageNumber="3732">C. reflexa</emphasis>
</taxonomicName>
occurred at all three collecting sites,
<taxonomicName id="4C67B1C1FFF2FFF6DD2F1BB688E2BED6" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" box="[564,715,201,225]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="nodasakae">
<emphasis id="B9131650FFF2FFF6DD2F1BB688E2BED6" box="[564,715,201,225]" italics="true" pageId="46" pageNumber="3732">C. nodasakae</emphasis>
</taxonomicName>
occurred only at Settlers Cove, a silty environment with much freshwater influence.
</paragraph>
</subSubSection>
<subSubSection id="C37D99C9FFF2FFF6DF831A758E68BC51" pageId="46" pageNumber="3732" type="reference_group">
<paragraph id="8BD8CA42FFF2FFF6DF831A758B0ABFD3" blockId="46.[125,1173,137,614]" pageId="46" pageNumber="3732">
<taxonomicName id="4C67B1C1FFF2FFF6DF831A758BBABF15" ID-CoL="69GWW" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" box="[152,403,266,290]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="nodasakae">
<emphasis id="B9131650FFF2FFF6DF831A758BBABF15" box="[152,403,266,290]" italics="true" pageId="46" pageNumber="3732">Celleporella nodasakae</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C67B1C1FFF2FFF6DD5B1A758E2ABF15" ID-CoL="6LWLP" authority="Morris, 1980" authorityName="Morris" authorityYear="1980" box="[576,1027,266,290]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="minitumulosa">
<emphasis id="B9131650FFF2FFF6DD5B1A75897ABF15" box="[576,851,266,290]" italics="true" pageId="46" pageNumber="3732">Hippothoa minitumulosa</emphasis>
<bibRefCitation id="EFF6B7B3FFF2FFF6DC7B1A758E2ABF15" author="Morris PA" box="[864,1027,266,290]" pageId="46" pageNumber="3732" pagination="1 - 115" refId="ref48412" refString="Morris PA. 1980. The bryozoan family Hippothoidae (Cheilostomata-Ascophora) with emphasis on the genus Hippothoa. Allan Hancock Monographs in Marine Biology 10: 1 - 115." type="journal article" year="1980">Morris, 1980</bibRefCitation>
</taxonomicName>
, which also ranges from uniserial to coherent in form, displays only unilaminar growth, and lacks male polymorphs (the latter two characters were not specifically mentioned in the original description, but
<bibRefCitation id="EFF6B7B3FFF2FFF6DE201A148BC8BFB4" author="Morris PA" box="[315,481,363,387]" pageId="46" pageNumber="3732" pagination="1 - 115" refId="ref48412" refString="Morris PA. 1980. The bryozoan family Hippothoidae (Cheilostomata-Ascophora) with emphasis on the genus Hippothoa. Allan Hancock Monographs in Marine Biology 10: 1 - 115." type="journal article" year="1980">Morris (1980)</bibRefCitation>
gave no measurements for male polymorphs, and her figure shows only unilaminar growth). However, compared to
<taxonomicName id="4C67B1C1FFF2FFF6DC131AF489B4BF94" ID-CoL="69GWW" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" box="[776,925,395,419]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="nodasakae">
<emphasis id="B9131650FFF2FFF6DC131AF489B4BF94" box="[776,925,395,419]" italics="true" pageId="46" pageNumber="3732">C. nodasakae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C67B1C1FFF2FFF6DCB61AF38E4FBF94" ID-CoL="6LWLP" authorityName="Morris" authorityYear="1980" box="[941,1126,395,419]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="minitumulosa">
<emphasis id="B9131650FFF2FFF6DCB61AF38E4FBF94" box="[941,1126,395,419]" italics="true" pageId="46" pageNumber="3732">H. minitumulosa</emphasis>
</taxonomicName>
has smaller zooids with midline tubercles, ovicells with more pores, and a differently shaped female orifice.
</paragraph>
<paragraph id="8BD8CA42FFF2FFF6DF831A928E68BC51" blockId="46.[125,1173,137,614]" pageId="46" pageNumber="3732">
<taxonomicName id="4C67B1C1FFF2FFF6DF831A928BA5BC32" ID-CoL="69GWW" authorityName="Dick &amp; Grischenko &amp; Mawatari" authorityYear="2005" box="[152,396,493,517]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="nodasakae">
<emphasis id="B9131650FFF2FFF6DF831A928BA5BC32" box="[152,396,493,517]" italics="true" pageId="46" pageNumber="3732">Celleporella nodasakae</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C67B1C1FFF2FFF6DD4E1A918980BC32" ID-CoL="6LWM2" authority="Pinter, 1973" authorityName="Pinter" authorityYear="1973" box="[597,937,493,517]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="santacruzana">
<emphasis id="B9131650FFF2FFF6DD4E1A918927BC32" box="[597,782,494,517]" italics="true" pageId="46" pageNumber="3732">H. santacruzana</emphasis>
<bibRefCitation id="EFF6B7B3FFF2FFF6DC0E1A928980BC32" author="Pinter PA" box="[789,937,493,517]" pageId="46" pageNumber="3732" pagination="437 - 446" refId="ref49214" refString="Pinter PA. 1973. The Hippothoa hyalina (L.) complex with a new species from the Pacific coast of California. In: Larwood GP, editor. Living and fossil Bryozoa. London: Academic Press. p 437 - 446." type="book chapter" year="1973">Pinter, 1973</bibRefCitation>
</taxonomicName>
, in which the ovicell likewise can have a few peripheral pores, and in which the proximal margin of the female orifice has two rounded-triangular projections flanking a median sinus. However,
<taxonomicName id="4C67B1C1FFF2FFF6DB6D19518B39BC51" ID-CoL="6LWM2" authorityName="Pinter" authorityYear="1973" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" kingdom="Animalia" order="Cheilostomatida" pageId="46" pageNumber="3732" phylum="Bryozoa" rank="species" species="santacruzana">
<emphasis id="B9131650FFF2FFF6DB6D19518B39BC51" italics="true" pageId="46" pageNumber="3732">H. santacruzana</emphasis>
</taxonomicName>
forms only disc-shaped coherent crusts, and male polymorphs occur.
</paragraph>
</subSubSection>
<subSubSection id="C37D99C9FFF2FFF6DF6619E08938BCD0" box="[125,785,671,743]" pageId="46" pageNumber="3732" type="distribution">
<paragraph id="8BD8CA42FFF2FFF6DF6619E08AD4BC80" blockId="46.[125,253,671,695]" box="[125,253,671,695]" pageId="46" pageNumber="3732">
<emphasis id="B9131650FFF2FFF6DF6619E08AD4BC80" box="[125,253,671,695]" italics="true" pageId="46" pageNumber="3732">Distribution</emphasis>
</paragraph>
<paragraph id="8BD8CA42FFF2FFF6DF6619B08938BCD0" blockId="46.[125,785,719,743]" box="[125,785,719,743]" pageId="46" pageNumber="3732">Settlers Cove near Ketchikan is the only known locality.</paragraph>
</subSubSection>
</treatment>
</document>