treatments-xml/data/03/B7/87/03B787C7FF5676A3FCEAD52C4F72F0F8.xml

<document id="D3EDE2EE48520B238AA0456FB2C0F41B" ID-CLB-Dataset="115951" ID-DOI="10.1093/zoolinnean/zlab033" ID-GBIF-Dataset="75b10efe-a17d-4d10-b46e-094c6a7ebf01" ID-ISSN="0024-4082" ID-Zenodo-Dep="6967774" ID-ZooBank="BE2B3859-8F6A-4543-8A69-91840F82377B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1653991067759" checkinUser="plazi" docAuthor="Lin, Shou-Wang, Lopardo, Lara &amp; Uhl, Gabriele" docDate="2022" docId="03B787C7FF5676A3FCEAD52C4F72F0F8" docLanguage="en" docName="zlab033.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Oedothorax uncus Tanasevitch 2015, INCERTAE" docType="treatment" docVersion="8" lastPageNumber="559" masterDocId="FF8EFFBFFFDD762DFFABD4584A34F51A" masterDocTitle="Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)" masterLastPageNumber="584" masterPageNumber="417" pageNumber="556" updateTime="1699338434129" updateUser="ExternalLinkService">
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<mods:title id="C0D58B4918FD4D6A31002FCDA77B9654">Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)</mods:title>
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<mods:namePart id="14B23A9E9C94EFB40ADDD0658A86C213">Lin, Shou-Wang</mods:namePart>
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<mods:namePart id="941078C8839AC94DC1D5824CB1B7FB02">Lopardo, Lara</mods:namePart>
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<mods:namePart id="27B0860EF40C08F0A3F12F17CC4D588E">Uhl, Gabriele</mods:namePart>
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<subSubSection id="C304655AFF5676A6FCEAD52C4EA4F4B0" pageId="139" pageNumber="556" type="nomenclature">
<paragraph id="8BA136D1FF5676A6FCEAD52C4EA4F4B0" blockId="139.[833,1437,371,426]" pageId="139" pageNumber="556">
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<taxonomicName id="4C1E4D52FF5676A6FCEAD52C4F11F496" ID-CoL="48WVR" authority="TANASEVITCH, 2015 INCERTAE" authorityName="Tanasevitch" authorityYear="2015" box="[833,1317,371,396]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="139" pageNumber="556" phylum="Arthropoda" rank="species" species="uncus" status="INCERTAE">
<emphasis id="B96AEAC3FF5676A6FCEAD52C4E08F491" box="[833,1084,372,395]" italics="true" pageId="139" pageNumber="556">‘OEDOTHORAX’ UNCUS</emphasis>
<bibRefCitation id="EF8F4B20FF5676A6FBE9D52B4F11F496" author="Tanasevitch AV" box="[1090,1317,371,396]" pageId="139" pageNumber="556" pagination="381 - 398" refId="ref95840" refString="Tanasevitch AV. 2015. Notes on the spider genus Oedothorax Bertkau, 1883 with description of eleven new species from India (Linyphiidae: Erigoninae). Revue suisse de Zoologie 122: 381 - 398." type="journal article" year="2015">TANASEVITCH, 2015</bibRefCitation>
</taxonomicName>
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SEDIS
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5676A6FC4DD5E14EC3F4C8" box="[998,1271,441,466]" pageId="139" pageNumber="556" type="description">
<paragraph id="8BA136D1FF5676A6FC4DD5E14EC3F4C8" blockId="139.[998,1271,441,466]" box="[998,1271,441,466]" pageId="139" pageNumber="556">
(
<figureCitation id="13252A54FF5676A6FC45D5E14E62F4CA" box="[1006,1110,441,466]" captionStart="Figure 19" captionStartId="37.[164,241,1114,1136]" captionTargetBox="[166,1446,198,1071]" captionTargetId="figure-288@37.[163,1449,195,1074]" captionTargetPageId="37" captionText="Figure 19. Male prosomal morphology, lateral. A, ‘Oedothorax’ meghalaya Tanasevitch, 2015 (traced from photograph). B, ‘Oe.’ uncus Tanasevitch, 2015 (traced from photograph). C, Holmelgonia basalis Jocqué &amp; Scharff, 1986 (Jocqué &amp; Scharff, 1986, fig. 18). D, Callitrichia holmi (Wunderlich, 1978) (traced from photograph). E, Ca. picta (Caporiacco, 1949) (Holm 1962, fig. 58). F, Ca. gloriosa (Jocqué, 1984) (traced from photograph). G, Ca. convector Tanasevitch, 2014 (traced from photograph). H, Ca. sellafrontis Scharff, 1990 (traced from photograph). I, Ca. juguma (Scharff, 1990) (Scharff 1990a, fig. 164). J, Ca. uncata (Jocqué &amp; Scharff, 1986) (Jocqué &amp; Scharff, 1986, fig. 130). K, Ca. pilosa (Wunderlich, 1978) (traced from photograph). L, Ca. usitata (Jocqué &amp; Scharff, 1986) (traced from photograph). M, Ca. legrandi (Jocqué, 1985) (Jocqué, 1985, fig. 21). N, Ca. macrophthalma (Locket &amp; Russell-Smith, 1980) (traced from photograph). O, Ca. muscicola (Bosmans, 1988) (traced from photograph). P, Ca. latitibialis (Bosmans, 1988) (traced from photograph). Q, Ca. longiducta (Bosmans, 1988) (traced from photograph). R, Nasoona setifera (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 3). S, N. crucifera (Thorell, 1895) (Tu &amp; Li, 2004, fig. 6A)." figureDoi="http://doi.org/10.5281/zenodo.6967840" httpUri="https://zenodo.org/record/6967840/files/figure.png" pageId="139" pageNumber="556">FIGS 19B</figureCitation>
, 
<figureCitation id="13252A54FF5676A6FBC8D5E14EBAF4CA" box="[1123,1166,441,466]" captionStart="Figure 22" captionStartId="40.[145,224,1771,1793]" captionTargetBox="[209,1362,197,1730]" captionTargetId="figure-25@40.[207,1362,195,1731]" captionTargetPageId="40" captionText="Figure 22. Male chelicerae, stridulatory striae on lateral side. A, ‘Oedothorax’ meghalaya. B, ‘Oe.’ uncus. C, Holmelgonia basalis. D, Callitrichia holmi. E, Ca. picta. F, Ca. gloriosa. G, Ca. convector. H, Ca. sellafrontis. I, Ca. juguma. J, Ca. uncata. K, Ca. pilosa. L, Ca. usitata. M, Ca. legrandi. N, Ca. macrophthalma. O, Ca. muscicola. P, Ca. latitibialis. Q, Ca. longiducta. R. Nasoona setifera. S, N. crucifera. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967850" httpUri="https://zenodo.org/record/6967850/files/figure.png" pageId="139" pageNumber="556">22B</figureCitation>
, 
<figureCitation id="13252A54FF5676A6FB30D5E14EF2F4CA" box="[1179,1222,441,466]" captionStart="Figure 24" captionStartId="44.[145,223,1135,1157]" captionTargetBox="[147,1428,197,1094]" captionTargetId="figure-293@44.[145,1428,195,1096]" captionTargetPageId="44" captionText="Figure 24. Male opisthosoma, dorsal view. A, ‘Oedothorax’ meghalaya Tanasevitch, 2015. B, ‘Oe.’ uncus Tanasevitch, 2015. C, Holmelgonia basalis Jocqué &amp; Scharff, 1986. D, Callitrichia holmi (Wunderlich, 1978). E, Ca. picta (Caporiacco, 1949). F, Ca. gloriosa (Jocqué, 1984). G, Ca. convector Tanasevitch, 2014. H, Ca. sellafrontis Scharff, 1990. I, Ca. juguma (Scharff, 1990). J, Ca. uncata (Jocqué &amp; Scharff, 1986). K, Ca. pilosa (Wunderlich, 1978). L, Ca. usitata (Jocqué &amp; Scharff, 1986). M, Ca. legrandi (Jocqué, 1985). N, Ca. macrophthalma (Locket &amp; Russell-Smith, 1980). O, Ca. muscicola (Bosmans, 1988). P, Ca. latitibialis (Bosmans, 1988). Q, Ca. longiducta (Bosmans, 1988). R, Nasoona setifera (Tanasevitch, 1998). S, N. crucifera (Thorell, 1895). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967856" httpUri="https://zenodo.org/record/6967856/files/figure.png" pageId="139" pageNumber="556">24B</figureCitation>
, 
<figureCitation id="13252A54FF5676A6FB78D5E14EDBF4C8" box="[1235,1263,441,466]" captionStart="Figure 81" captionStartId="140.[145,223,1610,1632]" captionTargetBox="[148,1428,198,1567]" captionTargetId="figure-91@140.[145,1431,195,1570]" captionTargetPageId="140" captionText="Figure 81. ‘Oedothorax’ uncus Tanasevitch, 2015. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="139" pageNumber="556">81</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5676A6FC90D5B94990F70C" pageId="139" pageNumber="556" type="reference_group">
<paragraph id="8BA136D1FF5676A6FC90D5B94990F70C" blockId="139.[827,1442,481,534]" pageId="139" pageNumber="556">
<treatmentCitationGroup id="AB0E11FFFF5676A6FC90D5B94990F70C" pageId="139" pageNumber="556">
<treatmentCitation id="0ABF10C0FF5676A6FC90D5B94F14F4EC" author="Tanasevitch AV" box="[827,1312,481,503]" page="393" pageId="139" pageNumber="556" year="2015">
<taxonomicName id="4C1E4D52FF5676A6FC90D5B94F14F4EC" ID-CoL="48WVR" authority="Tanasevitch, 2015: 393" authorityName="Tanasevitch" authorityPageNumber="393" authorityYear="2015" box="[827,1312,481,503]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="139" pageNumber="556" phylum="Arthropoda" rank="species" species="uncus">
<emphasis id="B96AEAC3FF5676A6FC90D5B94E38F4ED" box="[827,1036,481,503]" italics="true" pageId="139" pageNumber="556">Oedothorax uncus</emphasis>
<bibRefCitation id="EF8F4B20FF5676A6FBB8D5B94F14F4EC" author="Tanasevitch AV" box="[1043,1312,481,503]" pageId="139" pageNumber="556" pagination="381 - 398" refId="ref95840" refString="Tanasevitch AV. 2015. Notes on the spider genus Oedothorax Bertkau, 1883 with description of eleven new species from India (Linyphiidae: Erigoninae). Revue suisse de Zoologie 122: 381 - 398." type="journal article" year="2015">Tanasevitch, 2015: 393</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 86–88 (Dmf).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5676A6FC90D6674EBCF7D5" pageId="139" pageNumber="556" type="materials_examined">
<paragraph id="8BA136D1FF5676A6FC90D6674EBCF7D5" blockId="139.[827,1442,574,719]" pageId="139" pageNumber="556">
<emphasis id="B96AEAC3FF5676A6FC90D66749DDF749" box="[827,1001,574,596]" italics="true" pageId="139" pageNumber="556">Type material:</emphasis>
<materialsCitation id="3B763C8CFF5676A6FC5DD666498CF7AA" ID-GBIF-Occurrence="3880945327" collectingDate="1978-10-28" collectionCode="MHNG" collectorName="C. Besuchet &amp; I. Lobl" country="India" elevation="1800" location="India" pageId="139" pageNumber="556" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A58873FF5676A6FC5DD6664E50F74E" box="[1014,1124,574,596]" pageId="139" pageNumber="556" type="holotype">Holotype</typeStatus>
: 
<emphasis id="B96AEAC3FF5676A6FBDAD6664EF5F74E" bold="true" box="[1137,1217,574,596]" pageId="139" pageNumber="556">
<collectingCountry id="F3097641FF5676A6FBDAD6664E8EF74E" box="[1137,1210,574,596]" name="India" pageId="139" pageNumber="556">India</collectingCountry>
:
</emphasis>
<taxonomicName id="4C1E4D52FF5676A6FB61D6664E58F768" authority=", Khasi Hills, Mawphlang, 1800" authorityName=", Khasi Hills, Mawphlang" authorityYear="1800" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="139" pageNumber="556" phylum="Arthropoda" rank="species" species="meghalaya">
Meghalaya, Khasi Hills, Mawphlang, 
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<elevation id="0033D1E2FF5676A6FB9AD6054E58F768" box="[1073,1132,605,626]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="139" pageNumber="556" unit="m" value="1800.0">1800</elevation>
</quantity>
</taxonomicName>
m, forest, sifting litter, 
<specimenCount id="9D18FD58FF5676A6FC90D627497FF78B" box="[827,843,639,657]" pageId="139" pageNumber="556" type="male">♂</specimenCount>
<date id="FFA01011FF5676A6FCFCD62449E6F78B" box="[855,978,636,657]" pageId="139" pageNumber="556" value="1978-10-28">
<collectingDate id="EFE4E9F9FF5676A6FCFCD62449E6F78B" box="[855,978,636,657]" pageId="139" pageNumber="556" value="1978-10-28">28.x.1978</collectingDate>
</date>
, leg. 
<collectorName id="26EB5307FF5676A6FBB2D6244E9BF788" box="[1049,1199,636,658]" pageId="139" pageNumber="556">C. Besuchet</collectorName>
&amp; 
<collectorName id="26EB5307FF5676A6FB70D6244F1BF788" box="[1243,1327,636,658]" pageId="139" pageNumber="556">I. Löbl</collectorName>
(
<collectionCode id="ED0FAE14FF5676A6FAE8D6244FA9F78B" box="[1347,1437,636,657]" country="Switzerland" name="Museum d'Histoire Naturelle" pageId="139" pageNumber="556" type="Museum">MHNG</collectionCode>
, examined)
</materialsCitation>
. 
<materialsCitation id="3B763C8CFF5676A6FC68D6C34EB0F7D5" ID-GBIF-Occurrence="3880945372" collectionCode="MHNG" country="India" pageId="139" pageNumber="556" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54A58873FF5676A6FC68D6C34E04F7AA" box="[963,1072,667,688]" pageId="139" pageNumber="556" type="paratype">Paratype</typeStatus>
: 
<specimenCount id="9D18FD58FF5676A6FB97D6C34E6DF7B5" box="[1084,1113,667,688]" pageId="139" pageNumber="556" type="female">1♀</specimenCount>
, collected together with the 
<typeStatus id="54A58873FF5676A6FC90D6E149A8F7D5" box="[827,924,697,719]" pageId="139" pageNumber="556" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED0FAE14FF5676A6FC01D6E149CBF7D4" box="[938,1023,697,718]" country="Switzerland" name="Museum d'Histoire Naturelle" pageId="139" pageNumber="556" type="Museum">MHNG</collectionCode>
, examined)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5676A6FC90D6A04E98F6FD" pageId="139" pageNumber="556" type="diagnosis">
<paragraph id="8BA136D1FF5676A6FC90D6A04986F617" blockId="139.[827,946,760,781]" box="[827,946,760,781]" pageId="139" pageNumber="556">
<emphasis id="B96AEAC3FF5676A6FC90D6A04986F617" box="[827,946,760,781]" italics="true" pageId="139" pageNumber="556">Diagnosis:</emphasis>
</paragraph>
<paragraph id="8BA136D1FF5676A6FC90D76E4F90F690" blockId="139.[827,1444,822,906]" pageId="139" pageNumber="556">
<emphasis id="B96AEAC3FF5676A6FC90D76E49B2F651" box="[827,902,822,843]" italics="true" pageId="139" pageNumber="556">Males:</emphasis>
This species can be distinguished by the long, hook-shaped VRP, the long, palpal tibial apophysis with distal dentation and the lack of prosomal modification.
</paragraph>
<paragraph id="8BA136D1FF5676A6FC90D7EA4E98F6FD" blockId="139.[827,1443,946,999]" pageId="139" pageNumber="556">
<emphasis id="B96AEAC3FF5676A6FC90D7EA49AAF6DD" box="[827,926,946,967]" italics="true" pageId="139" pageNumber="556">Females:</emphasis>
This species can be identified by the enlarged median part of copulatory ducts.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5676A1FC90D04B4FA4F279" lastPageId="140" lastPageNumber="557" pageId="139" pageNumber="556" type="description">
<paragraph id="8BA136D1FF5676A6FC90D04B49F7F132" blockId="139.[827,963,1043,1064]" box="[827,963,1043,1064]" pageId="139" pageNumber="556">
<emphasis id="B96AEAC3FF5676A6FC90D04B49F7F132" box="[827,963,1043,1064]" italics="true" pageId="139" pageNumber="556">Description:</emphasis>
</paragraph>
<paragraph id="8BA136D1FF5676A1FC90D00C4B67F279" blockId="139.[827,1444,1108,1896]" lastBlockId="140.[145,762,1778,1892]" lastPageId="140" lastPageNumber="557" pageId="139" pageNumber="556">
<emphasis id="B96AEAC3FF5676A6FC90D00C49CAF173" box="[827,1022,1108,1129]" italics="true" pageId="139" pageNumber="556">
Male (
<typeStatus id="54A58873FF5676A6FC22D00C49C4F173" box="[905,1008,1108,1129]" pageId="139" pageNumber="556" type="holotype">holotype</typeStatus>
):
</emphasis>
Total length: 2.20. Prosoma: 0.99 long, 0.8 wide, unmodified; interocular region with short stout setae directed upwards (
<figureCitation id="13252A54FF5676A6FB42D0C94F78F1BC" box="[1257,1356,1169,1191]" captionStart="Figure 19" captionStartId="37.[164,241,1114,1136]" captionTargetBox="[166,1446,198,1071]" captionTargetId="figure-288@37.[163,1449,195,1074]" captionTargetPageId="37" captionText="Figure 19. Male prosomal morphology, lateral. A, ‘Oedothorax’ meghalaya Tanasevitch, 2015 (traced from photograph). B, ‘Oe.’ uncus Tanasevitch, 2015 (traced from photograph). C, Holmelgonia basalis Jocqué &amp; Scharff, 1986 (Jocqué &amp; Scharff, 1986, fig. 18). D, Callitrichia holmi (Wunderlich, 1978) (traced from photograph). E, Ca. picta (Caporiacco, 1949) (Holm 1962, fig. 58). F, Ca. gloriosa (Jocqué, 1984) (traced from photograph). G, Ca. convector Tanasevitch, 2014 (traced from photograph). H, Ca. sellafrontis Scharff, 1990 (traced from photograph). I, Ca. juguma (Scharff, 1990) (Scharff 1990a, fig. 164). J, Ca. uncata (Jocqué &amp; Scharff, 1986) (Jocqué &amp; Scharff, 1986, fig. 130). K, Ca. pilosa (Wunderlich, 1978) (traced from photograph). L, Ca. usitata (Jocqué &amp; Scharff, 1986) (traced from photograph). M, Ca. legrandi (Jocqué, 1985) (Jocqué, 1985, fig. 21). N, Ca. macrophthalma (Locket &amp; Russell-Smith, 1980) (traced from photograph). O, Ca. muscicola (Bosmans, 1988) (traced from photograph). P, Ca. latitibialis (Bosmans, 1988) (traced from photograph). Q, Ca. longiducta (Bosmans, 1988) (traced from photograph). R, Nasoona setifera (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 3). S, N. crucifera (Thorell, 1895) (Tu &amp; Li, 2004, fig. 6A)." figureDoi="http://doi.org/10.5281/zenodo.6967840" httpUri="https://zenodo.org/record/6967840/files/figure.png" pageId="139" pageNumber="556">Fig. 19B</figureCitation>
). Eyes: AME-AME: 0.04, AME width: 0.05, AME-ALE: 0.03, ALE width: 0.08, ALE-PLE: 0.01, PLE width: 0.08, PLE-PME: 0.06, PME width: 0.06, PME-PME: 0.08. Clypeus: not hirsute, one sub-AME seta. Sternum: 0.61 long, 0.59 wide. Chelicerae: mastidia absent; stridulatory striae ridged, rows compressed and evenly spaced (
<figureCitation id="13252A54FF5676A6FC3ED13049C6F067" box="[917,1010,1384,1406]" captionStart="Figure 22" captionStartId="40.[145,224,1771,1793]" captionTargetBox="[209,1362,197,1730]" captionTargetId="figure-25@40.[207,1362,195,1731]" captionTargetPageId="40" captionText="Figure 22. Male chelicerae, stridulatory striae on lateral side. A, ‘Oedothorax’ meghalaya. B, ‘Oe.’ uncus. C, Holmelgonia basalis. D, Callitrichia holmi. E, Ca. picta. F, Ca. gloriosa. G, Ca. convector. H, Ca. sellafrontis. I, Ca. juguma. J, Ca. uncata. K, Ca. pilosa. L, Ca. usitata. M, Ca. legrandi. N, Ca. macrophthalma. O, Ca. muscicola. P, Ca. latitibialis. Q, Ca. longiducta. R. Nasoona setifera. S, N. crucifera. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967850" httpUri="https://zenodo.org/record/6967850/files/figure.png" pageId="139" pageNumber="556">Fig. 22B</figureCitation>
). Legs: tibia chaetotaxy 2-2-1-1, dorsal proximal macroseta on tibia I, III and IV 1.35, 2.55 and 2.94 times diameter of tibia, respectively; Tm I: 0.65. All metatarsi with trichobothrium. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, two retrolateral trichobothria; TPS absent; TPA prominent, dorso-retrolaterally directed, with several enlarged setal bases at tip; TRA absent (
<figureCitation id="13252A54FF5676A6FC31D20549C8F368" box="[922,1020,1629,1651]" captionStart="Figure 81" captionStartId="140.[145,223,1610,1632]" captionTargetBox="[148,1428,198,1567]" captionTargetId="figure-91@140.[145,1431,195,1570]" captionTargetPageId="140" captionText="Figure 81. ‘Oedothorax’ uncus Tanasevitch, 2015. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="139" pageNumber="556">Fig. 81C</figureCitation>
); PC median-sized, base not visible from dorsal view, distal setae close to distal clasp, distal clasp without striae, clasp directed apically (
<figureCitation id="13252A54FF5676A6FAD3D2C24953F3D4" captionStart="Figure 81" captionStartId="140.[145,223,1610,1632]" captionTargetBox="[148,1428,198,1567]" captionTargetId="figure-91@140.[145,1431,195,1570]" captionTargetPageId="140" captionText="Figure 81. ‘Oedothorax’ uncus Tanasevitch, 2015. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="139" pageNumber="556">Fig. 81A</figureCitation>
); T without papillae; PT median-sized papillae; TS absent (
<figureCitation id="13252A54FF5676A6FC30D28049CBF3F7" box="[923,1023,1752,1774]" captionStart="Figure 81" captionStartId="140.[145,223,1610,1632]" captionTargetBox="[148,1428,198,1567]" captionTargetId="figure-91@140.[145,1431,195,1570]" captionTargetPageId="140" captionText="Figure 81. ‘Oedothorax’ uncus Tanasevitch, 2015. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="139" pageNumber="556">Fig. 81D</figureCitation>
); MSA present; DSA tip wavy (
<figureCitation id="13252A54FF5676A6FADCD280495CF216" captionStart="Figure 81" captionStartId="140.[145,223,1610,1632]" captionTargetBox="[148,1428,198,1567]" captionTargetId="figure-91@140.[145,1431,195,1570]" captionTargetPageId="140" captionText="Figure 81. ‘Oedothorax’ uncus Tanasevitch, 2015. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="139" pageNumber="556">Fig. 81A</figureCitation>
); EM flat, length approximately equals ARP, with long papillae (
<figureCitation id="13252A54FF5676A6FC4BD34D4E75F230" box="[992,1089,1813,1835]" captionStart="Figure 81" captionStartId="140.[145,223,1610,1632]" captionTargetBox="[148,1428,198,1567]" captionTargetId="figure-91@140.[145,1431,195,1570]" captionTargetPageId="140" captionText="Figure 81. ‘Oedothorax’ uncus Tanasevitch, 2015. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="139" pageNumber="556">Fig. 81D</figureCitation>
); ARP flat, weakly sclerotized; LER absent; VRP long, dorsally curved, tip pointed, proximally directed; TP tip round, dorsally directed; E retrolaterally spiral, anterior margin at base slightly wavy (
<figureCitation id="13252A54FF5176A1FF77D3484B0FF23C" box="[220,315,1808,1830]" captionStart="Figure 79" captionStartId="135.[164,242,1819,1841]" captionTargetBox="[166,1428,663,1777]" captionTargetId="figure-165@135.[163,1449,655,1780]" captionTargetPageId="135" captionText="Figure 79. ‘Oedothorax’ paracymbialis Tanasevitch, 2015. A–E, male right palp, images flipped horizontally.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, embolic division, prolateral view. F, male spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968024" httpUri="https://zenodo.org/record/6968024/files/figure.png" pageId="140" pageNumber="557">Fig. 79B</figureCitation>
). Opisthosoma: dorsal pattern see 
<figureCitation id="13252A54FF5176A1FD65D3484A8AF25E" captionStart="Figure 24" captionStartId="44.[145,223,1135,1157]" captionTargetBox="[147,1428,197,1094]" captionTargetId="figure-293@44.[145,1428,195,1096]" captionTargetPageId="44" captionText="Figure 24. Male opisthosoma, dorsal view. A, ‘Oedothorax’ meghalaya Tanasevitch, 2015. B, ‘Oe.’ uncus Tanasevitch, 2015. C, Holmelgonia basalis Jocqué &amp; Scharff, 1986. D, Callitrichia holmi (Wunderlich, 1978). E, Ca. picta (Caporiacco, 1949). F, Ca. gloriosa (Jocqué, 1984). G, Ca. convector Tanasevitch, 2014. H, Ca. sellafrontis Scharff, 1990. I, Ca. juguma (Scharff, 1990). J, Ca. uncata (Jocqué &amp; Scharff, 1986). K, Ca. pilosa (Wunderlich, 1978). L, Ca. usitata (Jocqué &amp; Scharff, 1986). M, Ca. legrandi (Jocqué, 1985). N, Ca. macrophthalma (Locket &amp; Russell-Smith, 1980). O, Ca. muscicola (Bosmans, 1988). P, Ca. latitibialis (Bosmans, 1988). Q, Ca. longiducta (Bosmans, 1988). R, Nasoona setifera (Tanasevitch, 1998). S, N. crucifera (Thorell, 1895). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967856" httpUri="https://zenodo.org/record/6967856/files/figure.png" pageId="140" pageNumber="557">Fig. 24B</figureCitation>
; PMS with mAP, AC absent; PLS with triad, one AC (
<figureCitation id="13252A54FF5176A1FF68D3164B70F279" box="[195,324,1870,1892]" captionStart="Figure 81" captionStartId="140.[145,223,1610,1632]" captionTargetBox="[148,1428,198,1567]" captionTargetId="figure-91@140.[145,1431,195,1570]" captionTargetPageId="140" captionText="Figure 81. ‘Oedothorax’ uncus Tanasevitch, 2015. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="140" pageNumber="557">Fig. 81G, H</figureCitation>
).
</paragraph>
<caption id="DF616659FF5176A1FF3AD212494CF3A2" ID-DOI="http://doi.org/10.5281/zenodo.6968032" ID-Zenodo-Dep="6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="140" pageNumber="557" startId="140.[145,223,1610,1632]" targetBox="[148,1428,198,1567]" targetPageId="140">
<paragraph id="8BA136D1FF5176A1FF3AD212494CF3A2" blockId="140.[145,1425,1610,1720]" pageId="140" pageNumber="557">
<emphasis id="B96AEAC3FF5176A1FF3AD2124B31F345" bold="true" box="[145,261,1610,1632]" pageId="140" pageNumber="557">Figure 81.</emphasis>
‘ 
<taxonomicName id="4C1E4D52FF5176A1FEBED2124BB9F37A" box="[277,397,1610,1632]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="140" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAC3FF5176A1FEBED2124BB9F37A" box="[277,397,1610,1632]" italics="true" pageId="140" pageNumber="557">Oedothorax</emphasis>
</taxonomicName>
’ 
<taxonomicName id="4C1E4D52FF5176A1FE33D21348ADF37A" authority="Tanasevitch, 2015" authorityName="Tanasevitch" authorityYear="2015" box="[408,665,1610,1632]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="140" pageNumber="557" phylum="Arthropoda" rank="species" species="uncus">
<emphasis id="B96AEAC3FF5176A1FE33D2134BE2F37A" box="[408,470,1611,1632]" italics="true" pageId="140" pageNumber="557">uncus</emphasis>
<bibRefCitation id="EF8F4B20FF5176A1FE71D21248ADF37A" author="Tanasevitch AV" box="[474,665,1610,1632]" pageId="140" pageNumber="557" pagination="381 - 398" refId="ref95840" refString="Tanasevitch AV. 2015. Notes on the spider genus Oedothorax Bertkau, 1883 with description of eleven new species from India (Linyphiidae: Erigoninae). Revue suisse de Zoologie 122: 381 - 398." type="journal article" year="2015">Tanasevitch, 2015</bibRefCitation>
</taxonomicName>
. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA136D1FF5176A1FC82D2AA4FA4F279" blockId="140.[809,1425,1778,1892]" pageId="140" pageNumber="557">
<emphasis id="B96AEAC3FF5176A1FC82D2AA4E27F21D" box="[809,1043,1778,1799]" italics="true" pageId="140" pageNumber="557">
Female (
<typeStatus id="54A58873FF5176A1FC3ED2AA4E30F21D" box="[917,1028,1778,1799]" pageId="140" pageNumber="557" type="paratype">paratype</typeStatus>
):
</emphasis>
Total length: 2.28. Prosoma: 1.01 long, 0.78 wide. Eyes: AME-AME: 0.04, AME width: 0.04, AME-ALE: 0.02, ALE width: 0.09, ALE- PLE: 0.01, PLE width: 0.08, PLE-PME: 0.05, PME
</paragraph>
</subSubSection>
<caption id="DF616659FF5076A0FF08D22F4BD7F3DD" ID-DOI="http://doi.org/10.5281/zenodo.6968036" ID-Zenodo-Dep="6968036" httpUri="https://zenodo.org/record/6968036/files/figure.png" pageId="141" pageNumber="558" startId="141.[163,243,1655,1677]" targetBox="[180,1430,198,1597]" targetPageId="141">
<paragraph id="8BA136D1FF5076A0FF08D22F4BD7F3DD" blockId="141.[163,1443,1655,1735]" pageId="141" pageNumber="558">
<emphasis id="B96AEAC3FF5076A0FF08D22F4B2EF396" bold="true" box="[163,282,1655,1677]" pageId="141" pageNumber="558">Figure 82.</emphasis>
<taxonomicName id="4C1E4D52FF5076A0FE8DD22F48BAF397" authority="(Thorell, 1895)" baseAuthorityName="Thorell" baseAuthorityYear="1895" box="[294,654,1655,1677]" class="Arachnida" family="Linyphiidae" genus="Nasoona" kingdom="Animalia" order="Araneae" pageId="141" pageNumber="558" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="B96AEAC3FF5076A0FE8DD22F4BD3F397" box="[294,487,1655,1677]" italics="true" pageId="141" pageNumber="558">Nasoona crucifera</emphasis>
(Thorell, 1895)
</taxonomicName>
. A–D, male left palp. A, retrolateral view. B, tibia and patella, retrolateral view. C, retrolateral view, with copulatory bulb half-expanded. D, ventral view. E, male left spinnerets. F, female right spinnerets. Scale bars 0.1 mm.
</paragraph>
</caption>
<caption id="DF616659FF5376A3FF3AD0D14E64F1A7" ID-DOI="http://doi.org/10.5281/zenodo.6968038" ID-Zenodo-Dep="6968038" httpUri="https://zenodo.org/record/6968038/files/figure.png" pageId="142" pageNumber="559" startId="142.[145,223,1161,1183]" targetBox="[145,1428,195,1122]" targetPageId="142">
<paragraph id="8BA136D1FF5376A3FF3AD0D14E64F1A7" blockId="142.[145,1425,1161,1213]" pageId="142" pageNumber="559">
<emphasis id="B96AEAC3FF5376A3FF3AD0D14B32F184" bold="true" box="[145,262,1161,1183]" pageId="142" pageNumber="559">Figure 83.</emphasis>
<taxonomicName id="4C1E4D52FF5376A3FEBAD0D2486DF1BA" authority="Kulczynski, 1908" authorityName="Kulczynski" authorityYear="1908" box="[273,601,1162,1184]" class="Arachnida" family="Linyphiidae" genus="Tmeticus" kingdom="Animalia" order="Araneae" pageId="142" pageNumber="559" phylum="Arthropoda" rank="species" species="tolli">
<emphasis id="B96AEAC3FF5376A3FEBAD0D24B94F1BA" box="[273,416,1162,1184]" italics="true" pageId="142" pageNumber="559">Tmeticus tolli</emphasis>
Kulczyński, 1908
</taxonomicName>
. A–C, male left palp. A, retrolateral view. B, embolic division, prolateral view. C, distal part of tibia. D, E, epigyne. D, dorsal view. E, external morphology. Scale bars 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C304655AFF5376A3FF3AD16B487EF361" pageId="142" pageNumber="559" type="discussion">
<paragraph id="8BA136D1FF5376A3FF3AD16B487EF361" blockId="142.[145,762,1331,1659]" pageId="142" pageNumber="559">
width: 0.08, PME-PME: 0.06. Clypeus: not hirsute, one sub-AME seta. Sternum: 0.58 long; 0.61 wide. Legs: tibia chaetotaxy 2-2-1-1, dorsal proximal macroseta on tibia I and IV 2.28 and 3.44 times diameter of tibia, respectively; Tm I: 0.60. Epigyne: Clade 13 characteristic morphology, CO posteriorly oriented, middle part of copulatory ducts enlarged, entrances of copulatory ducts into spermathecae ectal to exits of fertilization ducts (
<figureCitation id="13252A54FF5376A3FED8D2704BC7F324" box="[371,499,1576,1598]" captionStart="Figure 81" captionStartId="140.[145,223,1610,1632]" captionTargetBox="[148,1428,198,1567]" captionTargetId="figure-91@140.[145,1431,195,1570]" captionTargetPageId="140" captionText="Figure 81. ‘Oedothorax’ uncus Tanasevitch, 2015. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="142" pageNumber="559">Fig. 81E, F</figureCitation>
). Opisthosoma: dorsal pattern same as male; PMS with mAP, one CY; PLS with triad, two CY, one AC (
<figureCitation id="13252A54FF5376A3FE60D23D480FF361" box="[459,571,1637,1659]" captionStart="Figure 81" captionStartId="140.[145,223,1610,1632]" captionTargetBox="[148,1428,198,1567]" captionTargetId="figure-91@140.[145,1431,195,1570]" captionTargetPageId="140" captionText="Figure 81. ‘Oedothorax’ uncus Tanasevitch, 2015. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, H, male posterior median spinnerets and posterior lateral spinnerets. G, lateral view. H, dorsal view. I, female spinnerets. J, female posterior median spinnerets and posterior lateral spinnerets. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968032" httpUri="https://zenodo.org/record/6968032/files/figure.png" pageId="142" pageNumber="559">Fig. 81I, J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5376A3FF3AD2FC48CEF3A0" box="[145,762,1700,1722]" pageId="142" pageNumber="559" type="distribution">
<paragraph id="8BA136D1FF5376A3FF3AD2FC48CEF3A0" blockId="142.[145,762,1700,1722]" box="[145,762,1700,1722]" pageId="142" pageNumber="559">
<emphasis id="B96AEAC3FF5376A3FF3AD2FC4B17F3A3" box="[145,291,1700,1721]" italics="true" pageId="142" pageNumber="559">Distribution:</emphasis>
<collectingCountry id="F3097641FF5376A3FE84D2FC4B59F3A0" box="[303,365,1700,1722]" name="India" pageId="142" pageNumber="559">India</collectingCountry>
, only known from the 
<typeStatus id="54A58873FF5376A3FDC0D2FD48A8F3A0" box="[619,668,1701,1722]" pageId="142" pageNumber="559">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5376A3FF3AD2BB4BA4F3E3" box="[145,400,1763,1785]" pageId="142" pageNumber="559" type="biology_ecology">
<paragraph id="8BA136D1FF5376A3FF3AD2BB4BA4F3E3" blockId="142.[145,400,1763,1785]" box="[145,400,1763,1785]" pageId="142" pageNumber="559">
<emphasis id="B96AEAC3FF5376A3FF3AD2BB4AC4F3E2" box="[145,240,1763,1784]" italics="true" pageId="142" pageNumber="559">Habitat:</emphasis>
Forest litter.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5376A3FF3AD3794F72F0F8" pageId="142" pageNumber="559" type="discussion">
<paragraph id="8BA136D1FF5376A3FF3AD3794F72F0F8" blockId="142.[145,760,1825,1878]" lastBlockId="142.[809,1425,1331,1506]" pageId="142" pageNumber="559">
<emphasis id="B96AEAC3FF5376A3FF3AD3794AC8F22C" box="[145,252,1825,1846]" italics="true" pageId="142" pageNumber="559">Remarks.</emphasis>
According to the results of our phylogenetic analysis and the lack of the distinctive characteristics of 
<emphasis id="B96AEAC3FF5376A3FCEED16B49C7F052" box="[837,1011,1331,1352]" italics="true" pageId="142" pageNumber="559">
<taxonomicName id="4C1E4D52FF5376A3FCEED16B49FEF052" box="[837,970,1331,1352]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="142" pageNumber="559" phylum="Arthropoda" rank="genus" sensu="stricto">Oedothorax</taxonomicName>
<taxonomicNameLabel id="A25957B8FF5376A3FC7AD16B49C7F052" box="[977,1011,1331,1352]" pageId="142" pageNumber="559" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
, this species is more closely related to 
<taxonomicName id="4C1E4D52FF5376A3FCE3D10A499CF07D" baseAuthorityName="Jocque" baseAuthorityYear="1983" box="[840,936,1362,1383]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="142" pageNumber="559" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAC3FF5376A3FCE3D10A499CF07D" box="[840,936,1362,1383]" italics="true" pageId="142" pageNumber="559">Atypena</emphasis>
</taxonomicName>
, some ‘ 
<taxonomicName id="4C1E4D52FF5376A3FC54D1094EB3F07C" authority="Lin &amp; Lopardo &amp; Uhl, 2022" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[1023,1159,1361,1382]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="142" pageNumber="559" phylum="Arthropoda" rank="genus" status="incertae sedis">
<emphasis id="B96AEAC3FF5376A3FC54D1094EB3F07C" box="[1023,1159,1361,1382]" italics="true" pageId="142" pageNumber="559">Oedothorax</emphasis>
</taxonomicName>
’ 
<emphasis id="B96AEAC3FF5376A3FB3ED10A4F01F07C" box="[1173,1333,1361,1383]" italics="true" pageId="142" pageNumber="559">
<taxonomicNameLabel id="A25957B8FF5376A3FB3ED10A4F01F07C" box="[1173,1333,1361,1383]" pageId="142" pageNumber="559">incertae sedis</taxonomicNameLabel>
</emphasis>
species and 
<taxonomicName id="4C1E4D52FF5376A3FCF1D129498AF09C" authorityName="van Helsdingen" authorityYear="1985" box="[858,958,1393,1414]" class="Arachnida" family="Linyphiidae" genus="Mitrager" kingdom="Animalia" order="Araneae" pageId="142" pageNumber="559" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAC3FF5376A3FCF1D129498AF09C" box="[858,958,1393,1414]" italics="true" pageId="142" pageNumber="559">Mitrager</emphasis>
</taxonomicName>
, which, as in this species, are distributed in the Himalayan region and the 
<collectingRegion id="49DAF833FF5376A3FB1AD1D74F20F0BF" box="[1201,1300,1423,1445]" country="Morocco" name="Oriental" pageId="142" pageNumber="559">Oriental</collectingRegion>
realm. We provisionally leave the taxonomic status of this species unchanged until further data become available.
</paragraph>
</subSubSection>
</treatment>
</document>