185 lines
18 KiB
XML
185 lines
18 KiB
XML
<document id="BB6007E6CF521BBE1C06730985791376" ID-CLB-Dataset="37322" ID-DOI="10.11646/zootaxa.4211.1.1" ID-GBIF-Dataset="7af458af-d56d-44e3-a6f7-af97868175d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="272750" ID-ZooBank="6B86C6BA-6AFE-4AAD-870D-04794C138D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484386885843" checkinUser="plazi" docAuthor="Antić, Dragan Ž. & Makarov, Slobodan E." docDate="2016" docId="03B7878CFF85CD6CFF4CEA19B3A3ECF6" docLanguage="en" docName="zootaxa.4211.1.1.pdf.imf" docOrigin="Zootaxa 4211 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Paranotosoma Antić & Makarov, 2016, gen. nov." docType="treatment" docVersion="8" lastPageNumber="83" masterDocId="FF8EFFF4FFD7CD3EFFDBE875B173E949" masterDocTitle="The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" masterLastPageNumber="205" masterPageNumber="1" pageNumber="83" updateTime="1698435542289" updateUser="ExternalLinkService">
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<mods:mods id="0C1725D6F4FCA3FAE150188E832FE302" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C4875616A1139A3D10A7204294CC68E8">
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<mods:title id="A74EF0C6A58E4FA3334C977EF4072CEB">The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)</mods:title>
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<mods:name id="8C0CA71567F6B326369DCDFD0A759164" type="personal">
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<mods:roleTerm id="648E3A3D1E94B69CEF3F3D54060CBBAE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C52EA8CC2A47A5CFBC7A523F7FFBE6BE">Antić, Dragan Ž.</mods:namePart>
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</mods:name>
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<mods:name id="73193D070F21A915863DFBAC4FE41A9F" type="personal">
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<mods:role id="1C7966B01EA5B47D9D8CD9C543AE5484">
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<mods:roleTerm id="79D55FC87DC0761A952F68A6D4EA989B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="604E5A42CB5FDF128A24415B06067E21">Makarov, Slobodan E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DBAB491ADD8382CBAC6B51FF337F941E">text</mods:typeOfResource>
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<mods:titleInfo id="21F9FCFE81113D491A1E8A625A231256">
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<mods:title id="DABD9F4D3C87DC0F18F2D61EC166AF36">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8D448CAFF7E785E3E8A42037790F14D7">
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<mods:date id="3F6AD34CFBE741A729298A75E13E6A93">2016</mods:date>
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<mods:detail id="79EEA42E89440F236D30195BC7B2FF63" type="volume">
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<mods:number id="B8769255CB9940D424682DE9C88CE061">4211</mods:number>
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</mods:detail>
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<mods:detail id="A1F6CA797DB811ACFFAAD8D6DF0E3ABE" type="issue">
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<mods:number id="549833D40B24E9175B667230E80E871E">1</mods:number>
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<mods:classification id="4D9A36092750F3291078A08D7185468E">journal article</mods:classification>
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<mods:identifier id="52F8B5AC5D8042EAFFB97332A1DAD2BE" type="CLB-Dataset">37322</mods:identifier>
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<mods:identifier id="79FBB88C6E2DD687FB0D3B51492D8148" type="DOI">10.11646/zootaxa.4211.1.1</mods:identifier>
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<mods:identifier id="45E63CAC24774C3CCE7DAA18B682319E" type="GBIF-Dataset">7af458af-d56d-44e3-a6f7-af97868175d9</mods:identifier>
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<mods:identifier id="DA4AA07811D6D7AEDE887088648B3E91" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="EBE845E2FA488E7FC65721D1D7E0D5B6" type="Zenodo-Dep">272750</mods:identifier>
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</mods:mods>
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<treatment id="03B7878CFF85CD6CFF4CEA19B3A3ECF6" ID-DOI="http://doi.org/10.5281/zenodo.6074413" ID-GBIF-Taxon="127419159" ID-Zenodo-Dep="6074413" LSID="urn:lsid:plazi:treatment:03B7878CFF85CD6CFF4CEA19B3A3ECF6" httpUri="http://treatment.plazi.org/id/03B7878CFF85CD6CFF4CEA19B3A3ECF6" lastPageNumber="83" pageId="82" pageNumber="83">
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<subSubSection id="C3046511FF85CD6CFF4CEA19B360EBCF" box="[151,531,620,646]" pageId="82" pageNumber="83" type="nomenclature">
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<paragraph id="8BA1369AFF85CD6CFF4CEA19B360EBCF" blockId="82.[151,531,620,646]" box="[151,531,620,646]" pageId="82" pageNumber="83">
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<heading id="D0E981F6FF85CD6CFF4CEA19B360EBCF" bold="true" box="[151,531,620,646]" fontSize="11" level="1" pageId="82" pageNumber="83" reason="1">
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<emphasis id="B96AEA88FF85CD6CFF4CEA19B360EBCF" bold="true" box="[151,531,620,646]" pageId="82" pageNumber="83">
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Genus
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<taxonomicName id="4C1E4D19FF85CD6CFF35EA19B0EDEBCF" ID-CoL="845LX" box="[238,414,620,646]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B96AEA88FF85CD6CFF35EA19B0EDEBCF" bold="true" box="[238,414,620,646]" italics="true" pageId="82" pageNumber="83">Paranotosoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25957F3FF85CD6CFE7EEA19B360EBCF" box="[421,531,620,646]" pageId="82" pageNumber="83" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3046511FF85CD6CFF4CEAC7B4E6EBA6" pageId="82" pageNumber="83" type="diagnosis">
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<paragraph id="8BA1369AFF85CD6CFF4CEAC7B4E6EBA6" blockId="82.[151,1437,690,1471]" pageId="82" pageNumber="83">
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<emphasis id="B96AEA88FF85CD6CFF4CEAC7B062EB82" bold="true" box="[151,273,690,715]" pageId="82" pageNumber="83">Diagnosis.</emphasis>
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Differs from the superficially similar genus
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<taxonomicName id="4C1E4D19FF85CD6CFCD7EAC6B2EEEB83" box="[780,925,691,714]" class="Diplopoda" family="Anthroleucosomatidae" genus="Caucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="B96AEA88FF85CD6CFCD7EAC6B2EEEB83" box="[780,925,691,714]" italics="true" pageId="82" pageNumber="83">Caucaseuma</emphasis>
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</taxonomicName>
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in the structure of the coxal processes of the posterior gonopods, which are shorter and robust,
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<emphasis id="B96AEA88FF85CD6CFD12EAA2B393EBA7" box="[713,736,727,750]" italics="true" pageId="82" pageNumber="83">vs</emphasis>
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. flagelliform in
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<taxonomicName id="4C1E4D19FF85CD6CFC43EAA2B55AEBA7" box="[920,1065,727,750]" class="Diplopoda" family="Anthroleucosomatidae" genus="Caucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="B96AEA88FF85CD6CFC43EAA2B55AEBA7" box="[920,1065,727,750]" italics="true" pageId="82" pageNumber="83">Caucaseuma</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4C1E4D19FF85CD6CFB95EAA2B456EBA7" box="[1102,1317,727,750]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B96AEA88FF85CD6CFB95EAA2B456EBA7" box="[1102,1317,727,750]" italics="true" pageId="82" pageNumber="83">Heterocaucaseuma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25957F3FF85CD6CFAF7EAA3B4E6EBA6" box="[1324,1429,726,751]" pageId="82" pageNumber="83" rank="genus">
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<emphasis id="B96AEA88FF85CD6CFAF7EAA3B4E6EBA6" bold="true" box="[1324,1429,726,751]" pageId="82" pageNumber="83">gen. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3046511FF85CD6CFF1CEA8FB41DEA5B" box="[199,1390,762,787]" pageId="82" pageNumber="83" type="etymology">
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<paragraph id="8BA1369AFF85CD6CFF1CEA8FB41DEA5B" blockId="82.[151,1437,690,1471]" box="[199,1390,762,787]" pageId="82" pageNumber="83">
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<emphasis id="B96AEA88FF85CD6CFF1CEA8FB03DEA5A" bold="true" box="[199,334,762,787]" pageId="82" pageNumber="83">Etymology.</emphasis>
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To emphasize the well-developed lateral keels in the type-species. The name is a neuter noun.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3046511FF85CD6CFF1CEB6BB087EDAF" pageId="82" pageNumber="83" type="description">
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<paragraph id="8BA1369AFF85CD6CFF1CEB6BB46AEA8F" blockId="82.[151,1437,690,1471]" pageId="82" pageNumber="83">
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<emphasis id="B96AEA88FF85CD6CFF1CEB6BB02AEA7E" bold="true" box="[199,345,798,823]" pageId="82" pageNumber="83">Description.</emphasis>
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Body length
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<quantity id="4CE69B7FFF85CD6CFE22EB6BB3F9EA7F" box="[505,650,798,823]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.65" metricValueMin="1.2" pageId="82" pageNumber="83" unit="mm" value="14.25" valueMax="16.5" valueMin="12.0">12–16.5 mm</quantity>
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in males;
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<quantity id="4CE69B7FFF85CD6CFCDCEB6BB2EBEA7F" box="[775,920,798,823]" metricMagnitude="-2" metricUnit="m" metricValue="1.525" metricValueMax="1.85" metricValueMin="1.2" pageId="82" pageNumber="83" unit="mm" value="15.25" valueMax="18.5" valueMin="12.0">12–18.5 mm</quantity>
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in females; number of body segments
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<quantity id="4CE69B7FFF85CD6CFABBEB6AB4EFEA7E" box="[1376,1436,798,823]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="82" pageNumber="83" unit="in" value="30.0">30 in</quantity>
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adults (including telson). Body pigmented. Head slightly concave in males. Antennae and walking legs normal. Lateral keels well-developed, especially so in
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<taxonomicName id="4C1E4D19FF85CD6CFD7DEB12B21EEA36" box="[678,877,870,895]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="subrotundatum">
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<emphasis id="B96AEA88FF85CD6CFD7DEB12B21EEA36" box="[678,877,870,895]" italics="true" pageId="82" pageNumber="83">P. subrotundatum</emphasis>
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</taxonomicName>
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<emphasis id="B96AEA88FF85CD6CFCAEEB13B55AEA36" bold="true" box="[885,1065,870,895]" pageId="82" pageNumber="83">gen. et sp. nov.</emphasis>
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Macrochaetae short to mediumsized, trichoid. Leg-pairs
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<quantity id="4CE69B7FFF85CD6CFE65EBFEB37BEAEA" box="[446,520,906,931]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.778" metricValueMin="0.762" pageId="82" pageNumber="83" unit="in" value="5.0" valueMax="7.0" valueMin="3.0">3–7 in</quantity>
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males enlarged. Male leg-pair 7 with posterior coxal processes or at least their rudiments. Leg-pairs 10 and
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<quantity id="4CE69B7FFF85CD6CFE01EBDAB361EA8E" box="[474,530,942,967]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="82" pageNumber="83" unit="in" value="11.0">11 in</quantity>
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males with coxal glands, with neither any coxal horns nor processes.
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</paragraph>
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<paragraph id="8BA1369AFF85CD6CFF1CEBA7B05AED32" blockId="82.[151,1437,690,1471]" pageId="82" pageNumber="83">
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Anterior gonopods with a medial, subtriangular, hairy, sternal sac (
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<emphasis id="B96AEA88FF85CD6CFC76EBA1B2B0EAA2" box="[941,963,980,1003]" italics="true" pageId="82" pageNumber="83">ss</emphasis>
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) arising from sternal plate (
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<emphasis id="B96AEA88FF85CD6CFB21EBA1B460EAA2" box="[1274,1299,980,1003]" italics="true" pageId="82" pageNumber="83">sp</emphasis>
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) on anterior side. Coxal processes divided into an anterior and a posterior part. Anterior coxal processes (
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<emphasis id="B96AEA88FF85CD6CFB6EEB82B5BCED47" box="[1205,1231,1015,1038]" italics="true" pageId="82" pageNumber="83">cp</emphasis>
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) wide and shieldlike, fused, divided only distally. Posterior coxal processes (
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<emphasis id="B96AEA88FF85CD6CFCE0EC69B224ED7A" box="[827,855,1052,1075]" italics="true" pageId="82" pageNumber="83">pp</emphasis>
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) consisting of an anterior horn (
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<emphasis id="B96AEA88FF85CD6CFB1FEC6FB593ED7A" box="[1220,1248,1050,1075]" italics="true" pageId="82" pageNumber="83">ah</emphasis>
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) which abuts on posterior wall of anterior shields; and a posterior, more complex part. Medial part in posterior view with large coxal vesicles (
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<emphasis id="B96AEA88FF85CD6CFEDBEC11B06AED32" box="[256,281,1124,1147]" italics="true" pageId="82" pageNumber="83">cv</emphasis>
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).
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</paragraph>
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<paragraph id="8BA1369AFF85CD6CFF1CECF3B087EDAF" blockId="82.[151,1437,690,1471]" pageId="82" pageNumber="83">
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Posterior gonopods with completely divided coxites (
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<emphasis id="B96AEA88FF85CD6CFCFEECF2B242EDD7" box="[805,817,1159,1182]" italics="true" pageId="82" pageNumber="83">c</emphasis>
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). Posterolateral sides with small telopodites (
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<emphasis id="B96AEA88FF85CD6CFAE2ECF2B432EDD7" box="[1337,1345,1159,1182]" italics="true" pageId="82" pageNumber="83">t</emphasis>
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). Coxal vesicles (
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<emphasis id="B96AEA88FF85CD6CFEDBECD9B06AED8A" box="[256,281,1196,1219]" italics="true" pageId="82" pageNumber="83">cv</emphasis>
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) present on anterior side. Both coxites bearing a pair of coxal processes: a narrower anterior (
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<emphasis id="B96AEA88FF85CD6CFAEFECD9B423ED8A" box="[1332,1360,1196,1219]" italics="true" pageId="82" pageNumber="83">ap</emphasis>
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) and a more robust posterior (
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<emphasis id="B96AEA88FF85CD6CFE43ECBAB0C7EDAF" box="[408,436,1231,1254]" italics="true" pageId="82" pageNumber="83">pp</emphasis>
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) one.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3046511FF85CD6CFF1CEC87B21BEC42" box="[199,872,1266,1291]" pageId="82" pageNumber="83" type="distribution">
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<paragraph id="8BA1369AFF85CD6CFF1CEC87B21BEC42" blockId="82.[151,1437,690,1471]" box="[199,872,1266,1291]" pageId="82" pageNumber="83">
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<emphasis id="B96AEA88FF85CD6CFF1CEC87B013EC42" bold="true" box="[199,352,1266,1291]" pageId="82" pageNumber="83">Distribution.</emphasis>
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<collectingRegion id="49DAF878FF85CD6CFEBCEC87B0A6EC42" box="[359,469,1266,1291]" country="Georgia" name="Abkhazia" pageId="82" pageNumber="83">Abkhazia</collectingRegion>
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,
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<collectingRegion id="49DAF878FF85CD6CFE3BEC87B34EEC42" box="[480,573,1266,1291]" country="United States of America" name="Georgia" pageId="82" pageNumber="83">
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<collectingCountry id="F309760AFF85CD6CFE3BEC87B34EEC42" box="[480,573,1266,1291]" name="Georgia" pageId="82" pageNumber="83">Georgia</collectingCountry>
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</collectingRegion>
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,
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<collectingCountry id="F309760AFF85CD6CFD93EC87B3E0EC42" box="[584,659,1266,1291]" name="Russia" pageId="82" pageNumber="83">Russia</collectingCountry>
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(
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<figureCitation id="13252A1FFF85CD6CFD79EC87B271EC42" box="[674,770,1266,1291]" captionStart="FIGURE 167" captionStartId="196.[151,250,2007,2029]" captionTargetBox="[188,1398,1136,1986]" captionTargetId="figure@196.[188,1398,1136,1986]" captionTargetPageId="196" captionText="FIGURE 167. Distribution of the Caucasian Anthroleucosomatidae: the Caucaseuma complex." httpUri="https://zenodo.org/record/272916/files/figure.png" pageId="82" pageNumber="83">Fig. 167</figureCitation>
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, square).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3046511FF85CD6CFF1CED63B0F9EC1A" pageId="82" pageNumber="83" type="materials_examined">
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<paragraph id="8BA1369AFF85CD6CFF1CED63B5F7EC66" blockId="82.[151,1437,690,1471]" box="[199,1156,1302,1327]" pageId="82" pageNumber="83">
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<emphasis id="B96AEA88FF85CD6CFF1CED63B011EC66" bold="true" box="[199,354,1302,1327]" pageId="82" pageNumber="83">Type species.</emphasis>
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<taxonomicName id="4C1E4D19FF85CD6CFEB2ED62B3CEEC66" box="[361,701,1302,1327]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="subrotundatum">
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<emphasis id="B96AEA88FF85CD6CFEB2ED62B3CEEC66" box="[361,701,1302,1327]" italics="true" pageId="82" pageNumber="83">Paranotosoma subrotundatum</emphasis>
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</taxonomicName>
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<emphasis id="B96AEA88FF85CD6CFD1FED63B201EC66" bold="true" box="[708,882,1302,1327]" pageId="82" pageNumber="83">gen. et sp. nov.</emphasis>
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, by present designation.
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</paragraph>
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<paragraph id="8BA1369AFF85CD6CFF1CED4FB0F9EC1A" blockId="82.[151,1437,690,1471]" box="[199,394,1338,1363]" pageId="82" pageNumber="83">
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<emphasis id="B96AEA88FF85CD6CFF1CED4FB0F9EC1A" bold="true" box="[199,394,1338,1363]" pageId="82" pageNumber="83">Included species</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3046511FF85CD6CFF1CED2AB3A3ECF6" pageId="82" pageNumber="83" type="description">
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<paragraph id="8BA1369AFF85CD6CFF1CED2AB30AEC3E" blockId="82.[151,1437,690,1471]" box="[199,633,1374,1399]" pageId="82" pageNumber="83">
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<taxonomicName id="4C1E4D19FF85CD6CFF1CED2AB0B7EC3F" box="[199,452,1375,1398]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="attemsi">
|
||
<emphasis id="B96AEA88FF85CD6CFF1CED2AB0B7EC3F" box="[199,452,1375,1398]" italics="true" pageId="82" pageNumber="83">Paranotosoma attemsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA88FF85CD6CFE10ED2BB30AEC3E" bold="true" box="[459,633,1374,1399]" pageId="82" pageNumber="83">gen. et sp. nov.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA1369AFF85CD6CFF1CEDF1B3E1ECD2" blockId="82.[151,1437,690,1471]" box="[199,658,1410,1435]" pageId="82" pageNumber="83">
|
||
<taxonomicName id="4C1E4D19FF85CD6CFF1CEDF1B0AEECD2" box="[199,477,1410,1435]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="cordatum">
|
||
<emphasis id="B96AEA88FF85CD6CFF1CEDF1B0AEECD2" box="[199,477,1410,1435]" italics="true" pageId="82" pageNumber="83">Paranotosoma cordatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA88FF85CD6CFE3FEDF7B3E1ECD2" bold="true" box="[484,658,1410,1435]" pageId="82" pageNumber="83">gen. et sp. nov.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA1369AFF85CD6CFF1CEDD2B3A3ECF6" blockId="82.[151,1437,690,1471]" box="[199,720,1446,1471]" pageId="82" pageNumber="83">
|
||
<taxonomicName id="4C1E4D19FF85CD6CFF1CEDD2B36FECF6" box="[199,540,1446,1471]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="subrotundatum">
|
||
<emphasis id="B96AEA88FF85CD6CFF1CEDD2B36FECF6" box="[199,540,1446,1471]" italics="true" pageId="82" pageNumber="83">Paranotosoma subrotundatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA88FF85CD6CFDF9EDD3B3A3ECF6" bold="true" box="[546,720,1446,1471]" pageId="82" pageNumber="83">gen. et sp. nov.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |