treatments-xml/data/03/B1/87/03B187F8FFD4FFF62F3FBB89B682FA12.xml

<document id="643C993054ACBF3158F312C03A5CB6FF" ID-DOI="10.1653/024.100.0419" ID-ISSN="1938-5102" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721857016275" checkinUser="felipe" docAuthor="Smolis, Adrian &amp; Bernard, Ernest C." docDate="2017" docId="03B187F8FFD4FFF62F3FBB89B682FA12" docLanguage="en" docName="FlaEntomol.100.4.725-730.pdf" docOrigin="Florida Entomologist 100 (4)" docSource="http://www.bioone.org/doi/10.1653/024.100.0419" docStyle="DocumentStyle:C0CD22107794FB65089F701CBA118AA5.3:FlaEntomol.2015-.journal_article" docStyleId="C0CD22107794FB65089F701CBA118AA5" docStyleName="FlaEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Intricatonura Smolis &amp; Bernard, 2017, gen. nov." docType="treatment" docVersion="1" lastPageNumber="728" masterDocId="FF88FF80FFD7FFF22C67BF34B353FF86" masterDocTitle="Intricatonura fjellbergi, a new peculiar genus and species of Neanurini (Collembola: Neanuridae: Neanurinae) from Great Smoky Mountains National Park" masterLastPageNumber="730" masterPageNumber="725" pageNumber="727" updateTime="1721864226650" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CFDFFBEBDCB9BFAF1610E6CAE63100CB">Intricatonura fjellbergi, a new peculiar genus and species of Neanurini (Collembola: Neanuridae: Neanurinae) from Great Smoky Mountains National Park</mods:title>
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<mods:namePart id="28407481927886F0BE38B807991043CA">Smolis, Adrian</mods:namePart>
<mods:affiliation id="B715D1C13D6CA23F17EEE56B3E1B3389">University of Wrocław, Department of Invertebrate Biology, Evolution and Conservation, Institute of Environmental Biology, Przybyszewskiego 65, 51 - 148 Wrocław, Poland, E-mail: adrian. smolis @ uwr. edu. pl (A. S.)</mods:affiliation>
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<mods:namePart id="F59C7EB5D53202A326C419A9487FE3F7">Bernard, Ernest C.</mods:namePart>
<mods:affiliation id="CAD6C54400575F6B042A4BD8EF4A4F74">University of Tennessee, Department of Entomology &amp; Plant Pathology, 2505 E. J. Chapman Drive, 370 Plant Biotechnology, Knoxville, Tennessee 377996 - 4560, USA, E-mail: ebernard @ utk. edu (E. C. B.)</mods:affiliation>
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<paragraph id="8BA736EEFFD4FFF12F3FBB89B764FB57" blockId="3.[856,1079,1213,1233]" box="[856,1079,1213,1233]" pageId="3" pageNumber="727">
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<taxonomicName id="4C184D6DFFD4FFF12F3FBB89B084FB57" ID-CoL="8ZNRP" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[856,983,1213,1233]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="727" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAFCFFD4FFF12F3FBB89B084FB57" box="[856,983,1213,1233]" italics="true" pageId="3" pageNumber="727">Intricatonura</emphasis>
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<taxonomicNameLabel id="A25F5787FFD4FFF12FBABB89B764FB57" box="[989,1079,1213,1233]" pageId="3" pageNumber="727" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="8BA736EEFFD4FFF12F3FBBD2B659FB7C" blockId="3.[856,1290,1254,1274]" box="[856,1290,1254,1274]" pageId="3" pageNumber="727">
<typeStatus id="54A3884CFFD4FFF12F3FBBD2B0D5FB7C" box="[856,902,1254,1274]" pageId="3" pageNumber="727">Type</typeStatus>
species: 
<taxonomicName id="4C184D6DFFD4FFF12FB9BBD2B7EBFB7C" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[990,1208,1254,1274]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="727" phylum="Arthropoda" rank="species" species="fjellbergi" status="sp. nov.">
<emphasis id="B96CEAFCFFD4FFF12FB9BBD2B7EBFB7C" box="[990,1208,1254,1274]" italics="true" pageId="3" pageNumber="727">Intricatonura fjellbergi</emphasis>
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<emphasis id="B96CEAFCFFD4FFF128D9BBD2B659FB7C" bold="true" box="[1214,1290,1254,1274]" pageId="3" pageNumber="727">
<taxonomicNameLabel id="A25F5787FFD4FFF128D9BBD2B659FB7C" box="[1214,1290,1254,1274]" pageId="3" pageNumber="727" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BA736EEFFD4FFF12F1BBA24B74CF95A" blockId="3.[856,1538,1296,2020]" pageId="3" pageNumber="727">
<emphasis id="B96CEAFCFFD4FFF12F1BBA24B089FAA2" box="[892,986,1296,1316]" italics="true" pageId="3" pageNumber="727">Diagnosis</emphasis>
. Blue hypodermic pigment present on the body. Tubercles on body well developed. Three pigmented eyes on each side of head. Mouth parts reduced, maxilla styliform, mandible thin, triden- tate. Labral chaetotaxy 2/2, 2. Chaetotaxy of labium reduced with 2 basal, 2 distal, and 2 lateral setae, and without papillae x. Eight tubercles on head, with Di fused to De, and DL fused to (L + So). Chaetotaxy of tubercles Oc and (Di + De) reduced on head, with only 6 setae (Ocm, Di1 and De2) and setae Oca, Ocp, Di2 and De2 absent. Tubercle DL of Th. 
<collectionCode id="ED09AE2BFFD4FFF12F1CBACEB0D5F988" box="[891,902,1530,1550]" pageId="3" pageNumber="727">II</collectionCode>
– 
<collectionCode id="ED09AE2BFFD4FFF12FF5BACEB0F6F988" box="[914,933,1530,1550]" pageId="3" pageNumber="727">III</collectionCode>
divided into 2 components. Chaetotaxy of tubercles Di strongly reduced on Th. 
<collectionCode id="ED09AE2BFFD4FFF12861B92CB742F9AA" box="[1030,1041,1560,1580]" pageId="3" pageNumber="727">II</collectionCode>
– 
<collectionCode id="ED09AE2BFFD4FFF1287AB92CB763F9AA" box="[1053,1072,1560,1580]" pageId="3" pageNumber="727">III</collectionCode>
and Abd. I– 
<collectionCode id="ED09AE2BFFD4FFF128C6B92CB7EAF9AA" box="[1185,1209,1560,1580]" pageId="3" pageNumber="727">IV</collectionCode>
, with seta Di1 only. Tubercle Di of Abd. I– 
<collectionCode id="ED09AE2BFFD4FFF12FFDB901B0E2F9CF" box="[922,945,1589,1609]" pageId="3" pageNumber="727">IV</collectionCode>
rectangle-shaped. Tubercles De on Abd. I– 
<collectionCode id="ED09AE2BFFD4FFF12928B901B635F9CF" box="[1359,1382,1589,1609]" pageId="3" pageNumber="727">IV</collectionCode>
not developed. Chaetotaxy of groups De strongly reduced on Abd. I– 
<collectionCode id="ED09AE2BFFD4FFF12939B966B625F9E0" box="[1374,1398,1618,1638]" pageId="3" pageNumber="727">IV</collectionCode>
, without ordinary setae. Abd. 
<collectionCode id="ED09AE2BFFD4FFF12F90B944B758F902" box="[1015,1035,1648,1668]" pageId="3" pageNumber="727">IV</collectionCode>
and 
<collectionCode id="ED09AE2BFFD4FFF1285FB944B715F902" box="[1080,1094,1648,1668]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="727" type="Museum">V</collectionCode>
with 6 and 2 tubercles, respectively. Tubercles (Di + De + 
<collectionCode id="ED09AE2BFFD4FFF12FDDB9B9B086F927" box="[954,981,1677,1697]" pageId="3" pageNumber="727">DL</collectionCode>
) on abd. 
<collectionCode id="ED09AE2BFFD4FFF12849B9B9B76FF927" box="[1070,1084,1677,1697]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="727" type="Museum">V</collectionCode>
separate along midline. Cryptopygy present as Abd. 
<collectionCode id="ED09AE2BFFD4FFF12FECB99EB0CCF938" box="[907,927,1706,1726]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="727">VI</collectionCode>
nearly invisible from dorsal side. Tibiotarsal chaetotaxy 18, 18, 17. Claw untoothed.
</paragraph>
<paragraph id="8BA736EEFFD4FFF12F1BB9D1B635F80A" blockId="3.[856,1538,1296,2020]" pageId="3" pageNumber="727">
The new genus constitutes a monophyletic group within 
<taxonomicName id="4C184D6DFFD4FFF129CCB9D1B03EF890" authorityName="Salmon" authorityYear="1951" class="Collembola" family="Neanuridae" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="727" phylum="Arthropoda" rank="tribe" tribe="Neanurini">Neanurini</taxonomicName>
, with the following autapomorphies: reduction of labial chaetotaxy with 6 setae only, division of DL-tubercles on 2nd and 3rd thoracic tergites, presence of only 1 Di-seta on each side of Th. 
<collectionCode id="ED09AE2BFFD4FFF1292CB809B605F8D7" box="[1355,1366,1853,1873]" pageId="3" pageNumber="727">II</collectionCode>
– 
<collectionCode id="ED09AE2BFFD4FFF12905B809B626F8D7" box="[1378,1397,1853,1873]" pageId="3" pageNumber="727">III</collectionCode>
and Abd. I– 
<collectionCode id="ED09AE2BFFD4FFF12982B809B6AEF8D7" box="[1509,1533,1853,1873]" pageId="3" pageNumber="727">IV</collectionCode>
, rectangular and transverse shape of Di-tubercles on Abd. I– 
<collectionCode id="ED09AE2BFFD4FFF129F1B86EB6FDF8E8" box="[1430,1454,1882,1902]" pageId="3" pageNumber="727">IV</collectionCode>
, and absence of tubercles and ordinary De setae on Abd. I– 
<collectionCode id="ED09AE2BFFD4FFF1292DB84CB632F80A" box="[1354,1377,1912,1932]" pageId="3" pageNumber="727">IV</collectionCode>
.
</paragraph>
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Some of above mentioned features, i.e., division of DL-tubercles on Th. 
<collectionCode id="ED09AE2BFFD4FFF12FFCB886B0F5F840" box="[923,934,1970,1990]" pageId="3" pageNumber="727">II</collectionCode>
– 
<collectionCode id="ED09AE2BFFD4FFF12FD5B886B096F840" box="[946,965,1970,1990]" pageId="3" pageNumber="727">III</collectionCode>
and rectangular shape of Di-tubercles on Abd. I– 
<collectionCode id="ED09AE2BFFD4FFF129F9B886B6E6F840" box="[1438,1461,1970,1990]" pageId="3" pageNumber="727">IV</collectionCode>
are un- known in other members of the subfamily. 
<taxonomicName id="4C184D6DFFD4FFF12977B8E4B6DCF862" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[1296,1423,2000,2020]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="727" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAFCFFD4FFF12977B8E4B6DCF862" box="[1296,1423,2000,2020]" italics="true" pageId="3" pageNumber="727">Intricatonura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F5787FFD4FFF129F0B8E4B68FF862" box="[1431,1500,2000,2020]" pageId="3" pageNumber="727" rank="genus">gen. n.</taxonomicNameLabel>
resembles most closely 
<taxonomicName id="4C184D6DFFD3FFF62D5DBC3DB1CFFC9B" authority="Deharveng &amp; Bedos, 2000" authorityName="Deharveng &amp; Bedos" authorityYear="2000" box="[314,668,777,797]" class="Collembola" family="Neanuridae" genus="Vietnura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAFCFFD3FFF62D5DBC3DB2DDFC9B" box="[314,398,777,797]" italics="true" pageId="4" pageNumber="728">Vietnura</emphasis>
<bibRefCitation id="EF894B1FFFD3FFF62DF2BC3DB1CFFC9B" author="Deharveng L &amp; Bedos A." box="[405,668,777,797]" pageId="4" pageNumber="728" pagination="209 - 214" refId="ref4435" refString="Deharveng L, Bedos A. 2000. Vietnura caerulea new genus, new species, from Vietnam: first record of the Palearctic tribe Neanurini in tropical Asia (Collembola: Neanuridae). The Raffles Bulletin of Zoology 48: 209 - 214." type="journal article" year="2000">Deharveng &amp; Bedos, 2000</bibRefCitation>
</taxonomicName>
, the genus created for 
<taxonomicName id="4C184D6DFFD3FFF62CA9BC12B26AFCBC" authorityName="Deharveng &amp; Bedos" authorityYear="2000" box="[206,313,806,826]" class="Collembola" family="Neanuridae" genus="Vietnura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis id="B96CEAFCFFD3FFF62CA9BC12B38FFCBC" box="[206,220,806,826]" italics="true" pageId="4" pageNumber="728">V</emphasis>
. 
<emphasis id="B96CEAFCFFD3FFF62C81BC12B26AFCBC" box="[230,313,806,826]" italics="true" pageId="4" pageNumber="728">caerulea</emphasis>
</taxonomicName>
from 
<collectingCountry id="F30F767EFFD3FFF62D16BC12B290FCBC" box="[369,451,806,826]" name="Vietnam" pageId="4" pageNumber="728">Vietnam</collectingCountry>
(
<bibRefCitation id="EF894B1FFFD3FFF62DAABC12B19EFCBC" author="Deharveng L &amp; Bedos A." box="[461,717,806,826]" pageId="4" pageNumber="728" pagination="209 - 214" refId="ref4435" refString="Deharveng L, Bedos A. 2000. Vietnura caerulea new genus, new species, from Vietnam: first record of the Palearctic tribe Neanurini in tropical Asia (Collembola: Neanuridae). The Raffles Bulletin of Zoology 48: 209 - 214." type="journal article" year="2000">Deharveng &amp; Bedos,2000</bibRefCitation>
). Both genera share the fusion of all lateral tubercles on the head into a single mass, fusion of tubercles Di and De on head, reduction of the chaetotaxy of tubercles Di, De and Oc on the head, and presence of cryptopygy. Besides the apomorphies outlined above, they differ additionally in the number of eyes (in 
<taxonomicName id="4C184D6DFFD3FFF62D3CBC8DB2FCFC4B" authorityName="Deharveng &amp; Bedos" authorityYear="2000" box="[347,431,953,973]" class="Collembola" family="Neanuridae" genus="Vietnura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAFCFFD3FFF62D3CBC8DB2FCFC4B" box="[347,431,953,973]" italics="true" pageId="4" pageNumber="728">Vietnura</emphasis>
</taxonomicName>
2, in 
<taxonomicName id="4C184D6DFFD3FFF62D83BC8DB130FC4B" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[484,611,953,973]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAFCFFD3FFF62D83BC8DB130FC4B" box="[484,611,953,973]" italics="true" pageId="4" pageNumber="728">Intricatonura</emphasis>
</taxonomicName>
<emphasis id="B96CEAFCFFD3FFF62E0EBC8DB190FC4B" bold="true" box="[617,707,953,973]" pageId="4" pageNumber="728">
<taxonomicNameLabel id="A25F5787FFD3FFF62E0EBC8DB190FC4B" box="[617,707,953,973]" pageId="4" pageNumber="728" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
3), the length of dorsal macrosetae (in 
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<emphasis id="B96CEAFCFFD3FFF62DFDBCE2B2BDFC6C" box="[410,494,982,1002]" italics="true" pageId="4" pageNumber="728">Vietnura</emphasis>
</taxonomicName>
long, at least 3 times longer than length of tubercles; in 
<taxonomicName id="4C184D6DFFD3FFF62D09BCC0B2BEFB8E" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[366,493,1012,1032]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAFCFFD3FFF62D09BCC0B2BEFB8E" box="[366,493,1012,1032]" italics="true" pageId="4" pageNumber="728">Intricatonura</emphasis>
</taxonomicName>
<emphasis id="B96CEAFCFFD3FFF62D94BCC0B11DFB8E" bold="true" box="[499,590,1012,1032]" pageId="4" pageNumber="728">
<taxonomicNameLabel id="A25F5787FFD3FFF62D94BCC0B11DFB8E" box="[499,590,1012,1032]" pageId="4" pageNumber="728" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
notably short, 2 or 3 times shorter than length of tubercles), the arrangement of tubercles in the central area of the head (in 
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<emphasis id="B96CEAFCFFD3FFF62DC9BB1AB151FBC4" box="[430,514,1070,1090]" italics="true" pageId="4" pageNumber="728">Vietnura</emphasis>
</taxonomicName>
tubercles Af and Oc fused, in 
<taxonomicName id="4C184D6DFFD3FFF62F25BC3DB092FC9B" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[834,961,777,797]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAFCFFD3FFF62F25BC3DB092FC9B" box="[834,961,777,797]" italics="true" pageId="4" pageNumber="728">Intricatonura</emphasis>
</taxonomicName>
<emphasis id="B96CEAFCFFD3FFF62FAFBC3DB776FC9B" bold="true" box="[968,1061,777,797]" pageId="4" pageNumber="728">
<taxonomicNameLabel id="A25F5787FFD3FFF62FAFBC3DB776FC9B" box="[968,1061,777,797]" pageId="4" pageNumber="728" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
separate), the number of tubercles on the penultimate abdominal tergum (in 
<taxonomicName id="4C184D6DFFD3FFF628E1BC12B789FCBC" authorityName="Deharveng &amp; Bedos" authorityYear="2000" box="[1158,1242,806,826]" class="Collembola" family="Neanuridae" genus="Vietnura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAFCFFD3FFF628E1BC12B789FCBC" box="[1158,1242,806,826]" italics="true" pageId="4" pageNumber="728">Vietnura</emphasis>
</taxonomicName>
3 tubercles: 2 (De + 
<collectionCode id="ED09AE2BFFD3FFF629D7BC12B698FCBC" box="[1456,1483,806,826]" pageId="4" pageNumber="728">DL</collectionCode>
) and (Di + Di), in 
<taxonomicName id="4C184D6DFFD3FFF62FADBC70B71AFCDE" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[970,1097,836,856]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAFCFFD3FFF62FADBC70B71AFCDE" box="[970,1097,836,856]" italics="true" pageId="4" pageNumber="728">Intricatonura</emphasis>
</taxonomicName>
<emphasis id="B96CEAFCFFD3FFF62837BC70B7FFFCDE" bold="true" box="[1104,1196,836,856]" pageId="4" pageNumber="728">
<taxonomicNameLabel id="A25F5787FFD3FFF62837BC70B7FFFCDE" box="[1104,1196,836,856]" pageId="4" pageNumber="728" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
2 tubercles: 2 (Di + De + 
<collectionCode id="ED09AE2BFFD3FFF629CEBC70B697FCDE" box="[1449,1476,836,856]" pageId="4" pageNumber="728">DL</collectionCode>
)) and the presence of seta M on the tibiotarsi (in 
<taxonomicName id="4C184D6DFFD3FFF6296CBC55B60CFCF3" authorityName="Deharveng &amp; Bedos" authorityYear="2000" box="[1291,1375,865,885]" class="Collembola" family="Neanuridae" genus="Vietnura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAFCFFD3FFF6296CBC55B60CFCF3" box="[1291,1375,865,885]" italics="true" pageId="4" pageNumber="728">Vietnura</emphasis>
</taxonomicName>
present, in 
<taxonomicName id="4C184D6DFFD3FFF62F4FBC4AB0F4FC14" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[808,935,894,914]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAFCFFD3FFF62F4FBC4AB0F4FC14" box="[808,935,894,914]" italics="true" pageId="4" pageNumber="728">Intricatonura</emphasis>
</taxonomicName>
<emphasis id="B96CEAFCFFD3FFF62FCABC4AB754FC14" bold="true" box="[941,1031,894,914]" pageId="4" pageNumber="728">
<taxonomicNameLabel id="A25F5787FFD3FFF62FCABC4AB754FC14" box="[941,1031,894,914]" pageId="4" pageNumber="728" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
absent).
</paragraph>
<caption id="DF676666FFD3FFF62C07BFE1B185FF61" box="[96,726,213,231]" pageId="4" pageNumber="728" startId="4.[96,143,213,231]" targetType="table">
<paragraph id="8BA736EEFFD3FFF62C07BFE1B185FF61" blockId="4.[96,726,213,231]" box="[96,726,213,231]" pageId="4" pageNumber="728">
<emphasis id="B96CEAFCFFD3FFF62C07BFE1B3F7FF61" bold="true" box="[96,164,213,231]" pageId="4" pageNumber="728">Table 1.</emphasis>
Cephalic chaetotaxy of 
<taxonomicName id="4C184D6DFFD3FFF62D15BFE1B167FF61" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[370,564,213,231]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="species" species="fjellbergi" status="gen. nov., sp. nov.">
<emphasis id="B96CEAFCFFD3FFF62D15BFE1B167FF61" box="[370,564,213,231]" italics="true" pageId="4" pageNumber="728">Intricatonura fjellbergi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F5787FFD3FFF62E5FBFE1B185FF61" box="[568,726,213,231]" pageId="4" pageNumber="728" rank="species">
<emphasis id="B96CEAFCFFD3FFF62E5FBFE1B1DAFF61" bold="true" box="[568,649,213,231]" pageId="4" pageNumber="728">gen. nov.</emphasis>
, 
<emphasis id="B96CEAFCFFD3FFF62EF5BFE1B181FF61" bold="true" box="[658,722,213,231]" pageId="4" pageNumber="728">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BA736EEFFD3FFF62C07BE35B67AFD4B" pageId="4" pageNumber="728">
<table id="F918C44EFFD3000D2C07BE35B6DCFD4B" box="[96,1423,257,717]" gridcols="4" gridrows="13" pageId="4" pageNumber="728">
<tr id="352834ACFFD3000D2C07BE35B6DCFE95" box="[96,1423,257,275]" gridrow="0" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BE35B226FE95" box="[96,373,257,275]" gridcol="0" gridrow="0" pageId="4" pageNumber="728">Tubercle</th>
<th id="76F95DD0FFD3000D2E70BE35B1FAFE95" box="[535,681,257,275]" gridcol="1" gridrow="0" pageId="4" pageNumber="728">Number of setae</th>
<th id="76F95DD0FFD3000D2FE5BE35B750FE95" box="[898,1027,257,275]" gridcol="2" gridrow="0" pageId="4" pageNumber="728">Types of setae</th>
<th id="76F95DD0FFD3000D28BABE35B6DCFE95" box="[1245,1423,257,275]" gridcol="3" gridrow="0" pageId="4" pageNumber="728">Names of setae</th>
</tr>
<tr id="352834ACFFD3000D2C07BE1FB6DCFEBB" box="[96,1423,299,317]" gridrow="1" pageId="4" pageNumber="728" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76F95DD0FFD3000D2C07BE1FB226FEBB" box="[96,373,299,317]" gridcol="0" gridrow="1" pageId="4" pageNumber="728">Cephalic chaetotaxy–dorsal side</th>
</tr>
<tr id="352834ACFFD3000D2C07BE72B6DCFEF6" box="[96,1423,326,368]" gridrow="2" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BE72B226FEF6" box="[96,373,326,368]" gridcol="0" gridrow="2" pageId="4" pageNumber="728">Cl</th>
<td id="76F95DD0FFD3000D2E70BE72B1FAFEF6" box="[535,681,326,368]" gridcol="1" gridrow="2" pageId="4" pageNumber="728">4</td>
<td id="76F95DD0FFD3000D2FE5BE72B750FEF6" box="[898,1027,326,368]" gridcol="2" gridrow="2" pageId="4" pageNumber="728">Ml Mc</td>
<td id="76F95DD0FFD3000D28BABE72B6DCFEF6" box="[1245,1423,326,368]" gridcol="3" gridrow="2" pageId="4" pageNumber="728">F G</td>
</tr>
<tr id="352834ACFFD3000D2C07BE4EB6DCFE3A" box="[96,1423,378,444]" gridrow="3" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BE4EB226FE3A" box="[96,373,378,444]" gridcol="0" gridrow="3" pageId="4" pageNumber="728">Af</th>
<td id="76F95DD0FFD3000D2E70BE4EB1FAFE3A" box="[535,681,378,444]" gridcol="1" gridrow="3" pageId="4" pageNumber="728">6-7</td>
<td id="76F95DD0FFD3000D2FE5BE4EB750FE3A" box="[898,1027,378,444]" gridcol="2" gridrow="3" pageId="4" pageNumber="728">Ml Mc me</td>
<td id="76F95DD0FFD3000D28BABE4EB6DCFE3A" box="[1245,1423,378,444]" gridcol="3" gridrow="3" pageId="4" pageNumber="728">B A, O D</td>
</tr>
<tr id="352834ACFFD3000D2C07BEF1B6DCFE51" box="[96,1423,453,471]" gridrow="4" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BEF1B226FE51" box="[96,373,453,471]" gridcol="0" gridrow="4" pageId="4" pageNumber="728">Oc</th>
<td id="76F95DD0FFD3000D2E70BEF1B1FAFE51" box="[535,681,453,471]" gridcol="1" gridrow="4" pageId="4" pageNumber="728">1</td>
<td id="76F95DD0FFD3000D2FE5BEF1B750FE51" box="[898,1027,453,471]" gridcol="2" gridrow="4" pageId="4" pageNumber="728">Ml</td>
<td id="76F95DD0FFD3000D28BABEF1B6DCFE51" box="[1245,1423,453,471]" gridcol="3" gridrow="4" pageId="4" pageNumber="728">Ocm</td>
</tr>
<tr id="352834ACFFD3000D2C07BED6B6DCFE72" box="[96,1423,482,500]" gridrow="5" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BED6B226FE72" box="[96,373,482,500]" gridcol="0" gridrow="5" pageId="4" pageNumber="728">(Di + De)</th>
<td id="76F95DD0FFD3000D2E70BED6B1FAFE72" box="[535,681,482,500]" gridcol="1" gridrow="5" pageId="4" pageNumber="728">2</td>
<td id="76F95DD0FFD3000D2FE5BED6B750FE72" box="[898,1027,482,500]" gridcol="2" gridrow="5" pageId="4" pageNumber="728">Ml or Mc</td>
<td id="76F95DD0FFD3000D28BABED6B6DCFE72" box="[1245,1423,482,500]" gridcol="3" gridrow="5" pageId="4" pageNumber="728">Di1, De1</td>
</tr>
<tr id="352834ACFFD3000D2C07BECAB6DCFD96" box="[96,1423,510,528]" gridrow="6" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BECAB226FD96" box="[96,373,510,528]" gridcol="0" gridrow="6" pageId="4" pageNumber="728">(DL + L + So)</th>
<td id="76F95DD0FFD3000D2E70BECAB1FAFD96" box="[535,681,510,528]" gridcol="1" gridrow="6" pageId="4" pageNumber="728">11</td>
<td id="76F95DD0FFD3000D2FE5BECAB750FD96" box="[898,1027,510,528]" gridcol="2" gridrow="6" pageId="4" pageNumber="728">Ml, Mc, me</td>
<td id="76F95DD0FFD3000D28BABECAB6DCFD96" box="[1245,1423,510,528]" gridcol="3" gridrow="6" pageId="4" pageNumber="728">not recognizable</td>
</tr>
<tr id="352834ACFFD3000D2C07BD11B6DCFDB1" box="[96,1423,549,567]" gridrow="7" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BD11B6DCFDB1" box="[96,1423,549,567]" colspan="4" colspanRight="3" gridcol="0" gridrow="7" pageId="4" pageNumber="728">Cephalic chaetotaxy–ventral side</th>
</tr>
<tr id="352834ACFFD3000D2C07BD7FB6DCFDDB" box="[96,1423,587,605]" gridrow="8" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BD7FB226FDDB" box="[96,373,587,605]" gridcol="0" gridrow="8" pageId="4" pageNumber="728">Vi</th>
<td id="76F95DD0FFD3000D2E70BD7FB1FAFDDB" box="[535,681,587,605]" gridcol="1" gridrow="8" pageId="4" pageNumber="728">5</td>
<td id="76F95DD0FFD3000D2FE5BD7FB750FDDB" box="[898,1027,587,605]" gridcol="2" gridrow="8" pageId="4" pageNumber="728">—</td>
<td id="76F95DD0FFD3000D28BABD7FB6DCFDDB" box="[1245,1423,587,605]" gridcol="3" gridrow="8" pageId="4" pageNumber="728">—</td>
</tr>
<tr id="352834ACFFD3000D2C07BD52B6DCFDFE" box="[96,1423,614,632]" gridrow="9" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BD52B226FDFE" box="[96,373,614,632]" gridcol="0" gridrow="9" pageId="4" pageNumber="728">Vea</th>
<td id="76F95DD0FFD3000D2E70BD52B1FAFDFE" box="[535,681,614,632]" gridcol="1" gridrow="9" pageId="4" pageNumber="728">3</td>
<td id="76F95DD0FFD3000D2FE5BD52B750FDFE" box="[898,1027,614,632]" gridcol="2" gridrow="9" pageId="4" pageNumber="728">—</td>
<td id="76F95DD0FFD3000D28BABD52B6DCFDFE" box="[1245,1423,614,632]" gridcol="3" gridrow="9" pageId="4" pageNumber="728">—</td>
</tr>
<tr id="352834ACFFD3000D2C07BDB6B6DCFD12" box="[96,1423,642,660]" gridrow="10" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BDB6B226FD12" box="[96,373,642,660]" gridcol="0" gridrow="10" pageId="4" pageNumber="728">Vem</th>
<td id="76F95DD0FFD3000D2E70BDB6B1FAFD12" box="[535,681,642,660]" gridcol="1" gridrow="10" pageId="4" pageNumber="728">2-3</td>
<td id="76F95DD0FFD3000D2FE5BDB6B750FD12" box="[898,1027,642,660]" gridcol="2" gridrow="10" pageId="4" pageNumber="728">—</td>
<td id="76F95DD0FFD3000D28BABDB6B6DCFD12" box="[1245,1423,642,660]" gridcol="3" gridrow="10" pageId="4" pageNumber="728">—</td>
</tr>
<tr id="352834ACFFD3000D2C07BDABB6DCFD37" box="[96,1423,671,689]" gridrow="11" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BDABB226FD37" box="[96,373,671,689]" gridcol="0" gridrow="11" pageId="4" pageNumber="728">Vep</th>
<td id="76F95DD0FFD3000D2E70BDABB1FAFD37" box="[535,681,671,689]" gridcol="1" gridrow="11" pageId="4" pageNumber="728">3</td>
<td id="76F95DD0FFD3000D2FE5BDABB750FD37" box="[898,1027,671,689]" gridcol="2" gridrow="11" pageId="4" pageNumber="728">—</td>
<td id="76F95DD0FFD3000D28BABDABB6DCFD37" box="[1245,1423,671,689]" gridcol="3" gridrow="11" pageId="4" pageNumber="728">—</td>
</tr>
<tr id="352834ACFFD3000D2C07BD8FB6DCFD4B" box="[96,1423,699,717]" gridrow="12" pageId="4" pageNumber="728">
<th id="76F95DD0FFD3000D2C07BD8FB226FD4B" box="[96,373,699,717]" gridcol="0" gridrow="12" pageId="4" pageNumber="728">Labium</th>
<td id="76F95DD0FFD3000D2E70BD8FB1FAFD4B" box="[535,681,699,717]" gridcol="1" gridrow="12" pageId="4" pageNumber="728">6, 0x</td>
<td id="76F95DD0FFD3000D2FE5BD8FB750FD4B" box="[898,1027,699,717]" gridcol="2" gridrow="12" pageId="4" pageNumber="728">—</td>
<td id="76F95DD0FFD3000D28BABD8FB6DCFD4B" box="[1245,1423,699,717]" gridcol="3" gridrow="12" pageId="4" pageNumber="728">—</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA736EEFFD3FFF62F2BBCA8B020FB8E" blockId="4.[808,1489,777,1090]" pageId="4" pageNumber="728">
<emphasis id="B96CEAFCFFD3FFF62F2BBCA8B0E1FC36" box="[844,946,924,944]" italics="true" pageId="4" pageNumber="728">Etymology</emphasis>
. The name of the new genus is derived from the Latin combining form 
<emphasis id="B96CEAFCFFD3FFF62FA1BC8DB741FC4B" box="[966,1042,953,973]" italics="true" pageId="4" pageNumber="728">intricat-</emphasis>
(entangled), referring to its intricate combination of characters and the common root – 
<emphasis id="B96CEAFCFFD3FFF628DEBCE2B7B5FC6C" box="[1209,1254,982,1002]" italics="true" pageId="4" pageNumber="728">nura</emphasis>
used for many neanurid genera.
</paragraph>
<paragraph id="8BA736EEFFD3FFF62F2BBB25B033FBC4" blockId="4.[808,1489,777,1090]" pageId="4" pageNumber="728">
Revised key to 
<taxonomicName id="4C184D6DFFD3FFF62F8DBB25B719FBA3" authorityName="Salmon" authorityYear="1951" box="[1002,1098,1041,1061]" class="Collembola" family="Neanuridae" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="tribe" tribe="Neanurini">Neanurini</taxonomicName>
genera (modified from 
<bibRefCitation id="EF894B1FFFD3FFF62923BB25B008FBC4" author="Mayvan MM &amp; Shayanmehr M &amp; Smolis A &amp; Skarzynski D." pageId="4" pageNumber="728" pagination="552 - 558" refId="ref4669" refString="Mayvan MM, Shayanmehr M, Smolis A, Skarzynski D. 2015. Persanura hyrca- nica, a new genus and species of Neanurinae (Collembola: Neanuridae) from Iran, with a key to genera of the tribe Neanurini. Zootaxa 3918: 552 - 558." type="journal article" year="2015">Mayvan et al. 2015</bibRefCitation>
)
</paragraph>
<paragraph id="8BA736EEFFD3FFF62CD4BB55B62CFBF3" blockId="4.[179,1407,1121,1141]" box="[179,1407,1121,1141]" pageId="4" pageNumber="728">
In order to assign the new genus within the tribe 
<taxonomicName id="4C184D6DFFD3FFF62EF5BB55B1A5FBF3" authorityName="Salmon" authorityYear="1951" box="[658,758,1121,1141]" class="Collembola" family="Neanuridae" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="tribe" tribe="Neanurini">Neanurini</taxonomicName>
, we modified the key published by 
<bibRefCitation id="EF894B1FFFD3FFF6282DBB55B642FBF3" author="Mayvan MM &amp; Shayanmehr M &amp; Smolis A &amp; Skarzynski D." box="[1098,1297,1121,1141]" pageId="4" pageNumber="728" pagination="552 - 558" refId="ref4669" refString="Mayvan MM, Shayanmehr M, Smolis A, Skarzynski D. 2015. Persanura hyrca- nica, a new genus and species of Neanurinae (Collembola: Neanuridae) from Iran, with a key to genera of the tribe Neanurini. Zootaxa 3918: 552 - 558." type="journal article" year="2015">Mayvan et al. (2015)</bibRefCitation>
as follows:
</paragraph>
</subSubSection>
<subSubSection id="C3026565FFD3FFF62C07BBA5B682FA12" pageId="4" pageNumber="728" type="key">
<key id="FF8961DFFFD3FFF62C07BBA5B682FA12" pageId="4" pageNumber="728">
<keyStep id="30EC2E4BFFD3FFF62C07BBA5B683FB52" pageId="4" pageNumber="728">
<paragraph id="8BA736EEFFD3FFF62C07BBA5B683FB23" blockId="4.[96,1488,1169,1189]" box="[96,1488,1169,1189]" pageId="4" pageNumber="728">
<keyLead id="30E995DBFFD3FFF62C07BBA5B683FB23" box="[96,1488,1169,1189]" pageId="4" pageNumber="728">1. — Head with complete fusion of lateral tubercles DL, L and So............................................................... 2</keyLead>
</paragraph>
<paragraph id="8BA736EEFFD3FFF62C07BBF4B683FB52" blockId="4.[96,1488,1216,1236]" box="[96,1488,1216,1236]" pageId="4" pageNumber="728">
<keyLead id="30E995DBFFD3FFF62C07BBF4B683FB52" box="[96,1488,1216,1236]" pageId="4" pageNumber="728">1’. — Head with separation or incomplete fusion of lateral tubercles DL, L and So (e.g., only tubercles DL and L fused)................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E4BFFD3FFF62C07BBC4B683FAB2" pageId="4" pageNumber="728">
<paragraph id="8BA736EEFFD3FFF62C07BBC4B683FA82" blockId="4.[96,1488,1264,1284]" box="[96,1488,1264,1284]" pageId="4" pageNumber="728">
<keyLead id="30E995DBFFD3FFF62C07BBC4B683FA82" box="[96,1488,1264,1284]" pageId="4" pageNumber="728">
2. (1) Abdomen IV with tubercles Di fused............................ 
<taxonomicName id="4C184D6DFFD3FFF62F36BBC4B775FA82" ID-CoL="63BFG" authority="Cassagnau" authorityName="Cassagnau" authorityYear="1979" box="[849,1062,1264,1284]" class="Collembola" family="Neanuridae" genus="Monobella" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAFCFFD3FFF62F36BBC4B0E9FA82" box="[849,954,1264,1284]" italics="true" pageId="4" pageNumber="728">Monobella</emphasis>
Cassagnau
</taxonomicName>
(partim: 
<taxonomicName id="4C184D6DFFD3FFF62818BBC4B698FA82" ID-CoL="444WD" authority="Deharveng" authorityName="Deharveng" box="[1151,1483,1264,1284]" class="Collembola" family="Neanuridae" genus="Monobella" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="species" species="cassagnaui">
<emphasis id="B96CEAFCFFD3FFF62818BBC4B60AFA82" box="[1151,1369,1264,1284]" italics="true" pageId="4" pageNumber="728">Monobella cassagnaui</emphasis>
Deharveng
</taxonomicName>
)
</keyLead>
</paragraph>
<paragraph id="8BA736EEFFD3FFF62C07BA14B683FAB2" blockId="4.[96,1488,1312,1332]" box="[96,1488,1312,1332]" pageId="4" pageNumber="728">
<keyLead id="30E995DBFFD3FFF62C07BA14B683FAB2" box="[96,1488,1312,1332]" pageId="4" pageNumber="728">2’. — Abdomen IV with tubercles separate.................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E4BFFD3FFF62C07BA64B682FA12" pageId="4" pageNumber="728">
<paragraph id="8BA736EEFFD3FFF62C07BA64B683FAE2" blockId="4.[96,1488,1360,1380]" box="[96,1488,1360,1380]" pageId="4" pageNumber="728">
<keyLead id="30E995DBFFD3FFF62C07BA64B683FAE2" box="[96,1488,1360,1380]" pageId="4" pageNumber="728">
3. (2) Head with fusion of tubercles Af and Oc; Abdomen V with 3 tubercles: (Di + Di) and 2 (De + DL).......... 
<taxonomicName id="4C184D6DFFD3FFF628D1BA64B683FAE2" ID-CoL="86QT" authority="Deharveng &amp; Bedos" authorityName="Deharveng &amp; Bedos" authorityYear="2000" box="[1206,1488,1360,1380]" class="Collembola" family="Neanuridae" genus="Vietnura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAFCFFD3FFF628D1BA64B659FAE2" box="[1206,1290,1360,1380]" italics="true" pageId="4" pageNumber="728">Vietnura</emphasis>
Deharveng &amp; Bedos
</taxonomicName>
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</paragraph>
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<keyLead id="30E995DBFFD3FFF62C07BAB4B682FA12" box="[96,1489,1408,1428]" pageId="4" pageNumber="728">
3’. — Head with separation of tubercles Af and Oc; Abdomen V with 2 tubercles: 2 (Di + De + DL).................. 
<taxonomicName id="4C184D6DFFD3FFF62895BAB4B622FA12" ID-CoL="8ZNRP" authorityName="Smolis &amp; Bernard" authorityYear="2017" box="[1266,1393,1408,1428]" class="Collembola" family="Neanuridae" genus="Intricatonura" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="728" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAFCFFD3FFF62895BAB4B622FA12" box="[1266,1393,1408,1428]" italics="true" pageId="4" pageNumber="728">Intricatonura</emphasis>
</taxonomicName>
<emphasis id="B96CEAFCFFD3FFF62911BAB4B682FA12" bold="true" box="[1398,1489,1408,1428]" pageId="4" pageNumber="728">
<taxonomicNameLabel id="A25F5787FFD3FFF62911BAB4B682FA12" box="[1398,1489,1408,1428]" pageId="4" pageNumber="728" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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