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<mods:title id="68308A8B3CE360E5BC4683F9085B7C02">On the morphology and classification of larval water mites (Hydrachnidia, Acari) from springs in Luxembourg</mods:title>
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<subSubSection id="C302657DFFD05751CD09F95B0A66CD1B" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BA736F6FFD05751CD09F95B0CE2CCC4" blockId="16.[264,700,1689,1716]" box="[264,700,1689,1716]" pageId="16" pageNumber="17">
<heading id="D0EF819AFFD05751CD09F95B0CE2CCC4" bold="true" box="[264,700,1689,1716]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C184D75FFD05751CD09F95B0CE2CCC4" authority="Kramer, 1879" authorityName="Kramer" authorityYear="1879" box="[264,700,1689,1716]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="squamosus">
<emphasis id="B96CEAE4FFD05751CD09F95B0CE2CCC4" bold="true" box="[264,700,1689,1716]" pageId="16" pageNumber="17">
<emphasis id="B96CEAE4FFD05751CD09F95B0C56CCC3" bold="true" box="[264,520,1690,1716]" italics="true" pageId="16" pageNumber="17">Sperchon squamosus</emphasis>
Kramer, 1879
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA736F6FFD05751CD09F91B0A66CD1B" blockId="16.[264,1321,1754,1900]" pageId="16" pageNumber="17">
Records (
<tableCitation id="C69A034DFFD05751CD7CF91B0F80CC83" box="[381,478,1754,1780]" captionStart="TABLE 2" captionStartId="5.[264,352,283,307]" captionText="TABLE 2. The water mite species known from Luxembourg springs and their presence at different sites. In bold: species and numbers of sampled larvae of species collected in the respective emergence traps at sites E 1 ­ E 7, x = species at the emergence sites collected only in benthos samples by Gerecke et al. (2005) and / or during the present study, o. s. = other spring sites in Luxembourg (see Gerecke et al. 2005), + = species collected at these sites and at the 7 emergence sites, ++ = species only collected in these benthic samples." pageId="16" pageNumber="17">Table 2</tableCitation>
): Benthos samples at ten spring sites in Gutland and in Ösling, 48 individuals (
<bibRefCitation id="EF894B07FFD05751CD9CF8C30CC8CD6B" author="Gerecke" box="[413,662,1794,1820]" pageId="16" pageNumber="43" refString="Gerecke, R., Stoch, F., Meisch, C. &amp; Schrankel, I. (2005) Die Fauna der Quellen und des hyporheischen Interstitials in Luxemburg. Unter besonderer Berucksichtigung der Acari, Ostracoda und Copepoda. Ferrantia - Travaux scientifiques du Musee national d'histoire naturelle, Luxembourg, 41, 1 - 134." type="journal article" year="2005">
Gerecke
<emphasis id="B96CEAE4FFD05751CE0DF8C30C15CD6C" box="[524,587,1794,1819]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2005
</bibRefCitation>
).
<geoCoordinate id="EE2C5031FFD05751CEACF8C30CA9CD6B" box="[685,759,1794,1820]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="2.9666667">
E2
<specimenCount id="9D1EFD7FFFD05751CEDBF8C30CA9CD6B" box="[730,759,1794,1820]" pageId="16" pageNumber="17" type="larva">58</specimenCount>
</geoCoordinate>
larvae,
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E3
<specimenCount id="9D1EFD7FFFD05751CF89F8C30DFBCD6B" box="[904,933,1794,1820]" pageId="16" pageNumber="17" type="larva">93</specimenCount>
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larvae,
<geoCoordinate id="EE2C5031FFD05751C80AF8C30A1BCD6B" box="[1035,1093,1794,1820]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="5.016667">E5 1</geoCoordinate>
larva, parasitic on chironomids. Attribution based on individuals reared from a female from a small lowland stream in the
<collectingCountry id="F30F7666FFD05751CDA6F8930C5BCD1B" box="[423,517,1874,1900]" name="Germany" pageId="16" pageNumber="17">German</collectingCountry>
lowlands (Himmelreichbach, see
<bibRefCitation id="EF894B07FFD05751CF94F8930A75CD1B" author="Martin" box="[917,1067,1874,1900]" pageId="16" pageNumber="43" refString="Martin, P. (1997) Zur Faunistik und Substratpraferenz der Wassermilben (Hydrachnidia, Acari) zweier durch feine mineralische Substrate gepragten Bache in Schleswig-Holstein. Faunistisch-Okologische Mitteilungen, 7, 221 - 237." type="journal article" year="1997">Martin 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C302657DFFD15750CD39FEDD0C0BCD39" pageId="17" pageNumber="18" type="description">
<paragraph id="8BA736F6FFD15750CD39FEDD0A22CB41" blockId="17.[264,1324,284,1870]" box="[312,1148,284,310]" pageId="17" pageNumber="18">
Descriptions of larvae:
<bibRefCitation id="EF894B07FFD15750CE4BFEDD0D4CCB41" author="Lundblad" box="[586,786,284,310]" pageId="17" pageNumber="43" refString="Lundblad, O. (1927) Die Hydracarinen Schwedens. I. Beitrag zur Systematik, Embryologie, Okologie und Verbreitungsgeschichte der schwedischen Arten. Zoologiska bidrag, 11, 185 - 540." type="journal article" year="1927">Lundblad (1927)</bibRefCitation>
,
<bibRefCitation id="EF894B07FFD15750CF1FFEDD0D9ECB41" author="MotaS" box="[798,960,284,310]" pageId="17" pageNumber="43" refString="Motas, C. (1961) Hydrachnellae. The Zoology of Iceland, 3, 1 - 26." type="journal article" year="1961">MotaŞ (1961)</bibRefCitation>
,
<bibRefCitation id="EF894B07FFD15750CFCDFEDD0A26CB41" author="Ullrich" box="[972,1144,284,310]" pageId="17" pageNumber="44" refString="Ullrich, F. (1976) Biologisch-okologische Studien an rheophilen Wassermilben (Hydrachnellae, Acari), unter besonderer Berucksichtigung von Sperchon setiger (Thor 1898). PhD thesis, University of Kiel, 241 pp." type="book" year="1976">Ullrich (1976)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA736F6FFD15750CD39FE850A55CBD9" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">
Description (n = 5, Himmelreichbach): Idiosoma ovate; in non­engorged specimens and to a lesser extent in slightly engorged specimens with distinct distal indentation (
<figureCitation id="13232A73FFD15750C8ECFEAD0F0BCBD9" captionStart="FIGURES 28 30" captionStartId="18.[264,383,1835,1859]" captionTargetBox="[275,1300,534,1803]" captionTargetId="figure@18.[264,1323,524,1809]" captionTargetPageId="18" captionText="FIGURES 28 30. Sperchon squamosus (original). Fig. 28: dorsal idiosoma, Fig. 29: ventral idiosoma, Fig. 30: chelicera. Scale bars: 28, 29: 100 µm, 30 50 µm." httpUri="https://zenodo.org/record/172013/files/figure.png" pageId="17" pageNumber="18">Figs. 28, 29</figureCitation>
). Length/width of idiosoma 234261 (251)/174198 (187).
</paragraph>
<paragraph id="8BA736F6FFD15750CD39FE7D0F1EC961" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">
Dorsal idiosoma (
<figureCitation id="13232A73FFD15750CE08FE7D0C3CCBA1" box="[521,610,444,470]" captionStart="FIGURES 28 30" captionStartId="18.[264,383,1835,1859]" captionTargetBox="[275,1300,534,1803]" captionTargetId="figure@18.[264,1323,524,1809]" captionTargetPageId="18" captionText="FIGURES 28 30. Sperchon squamosus (original). Fig. 28: dorsal idiosoma, Fig. 29: ventral idiosoma, Fig. 30: chelicera. Scale bars: 28, 29: 100 µm, 30 50 µm." httpUri="https://zenodo.org/record/172013/files/figure.png" pageId="17" pageNumber="18">Fig. 28</figureCitation>
): Dp relatively large and ovate, anteriorly and medially with a fine lineation, the lines meeting each other laterally and posteriorly and building almost a rhombic pattern. Length/width Dp 202245 (230)/153168 (161), length eye capsule 36 43 (39), Mp2­Amdp 4957 (53), Mp1­Mp1 5562 (60), Mp2­Mp2 4858 (53), Lp1­Lp1 3440 (37), Lp2­Lp2 100106 (103), Mp1­Lp
<date id="FFA61036FFD15750CF34FD9D0DCEC801" box="[821,912,604,630]" pageId="17" pageNumber="18" value="1914-11-01">1 1114</date>
(12), Mp2­Lp2 3336 (34), Mp1­ Mp2 3740 (39), Lp1­Lp2 4042 (41), Mp
<date id="FFA61036FFD15750CF01FD450D07C8E9" box="[768,857,644,670]" pageId="17" pageNumber="18" value="1932-01-25">1 2532</date>
(28), Mp
<date id="FFA61036FFD15750CFC8FD450A79C8E9" box="[969,1063,644,670]" pageId="17" pageNumber="18" value="1935-02-20">2 2035</date>
(26), Lp1 7296 (87), Lp2 140150 (146), Hu 120140 (134), Mh1 118150 (141), Mh2 105140 (127), Mh3 103113 (109), Mh4 78103 (94), Lh1 128140 (134), Lh2 108118 (114), Lh3 85105 (92).
</paragraph>
<paragraph id="8BA736F6FFD15750CD39FCE50C12CE21" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">
Ventral idiosoma (
<figureCitation id="13232A73FFD15750CE12FCE50C30C949" box="[531,622,804,830]" captionStart="FIGURES 28 30" captionStartId="18.[264,383,1835,1859]" captionTargetBox="[275,1300,534,1803]" captionTargetId="figure@18.[264,1323,524,1809]" captionTargetPageId="18" captionText="FIGURES 28 30. Sperchon squamosus (original). Fig. 28: dorsal idiosoma, Fig. 29: ventral idiosoma, Fig. 30: chelicera. Scale bars: 28, 29: 100 µm, 30 50 µm." httpUri="https://zenodo.org/record/172013/files/figure.png" pageId="17" pageNumber="18">Fig. 29</figureCitation>
): Length/width CX­I 8893 (91)/3845 (42), CX­II 7593 (85)/5865 (61), CX­III 133150 (143)/8898 (93), coxal plates with an alveolar pattern, between the median margins of the coxal plates III, one or two small sclerites, urstigmata rounded, C1­C2 5357 (55), C1­Mmcp 2025 (23), C4­Pmcp 105132 (144), C1­C4 40 55 (48), C1 5565 (60), C2 82105 (90), C3 8089 (83), C4 7494 (81), Expp relative large and pronounced, length/width 2022 (21)/1922 (21), E1­
<geoCoordinate id="EE2C5031FFD15750CFF5FC2D0A77CE71" box="[1012,1065,1004,1030]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="1.1333333">
E
<date id="FFA61036FFD15750C80BFC2D0A77CE71" box="[1034,1065,1004,1030]" pageId="17" pageNumber="18" value="1911-08-01">1 8</date>
</geoCoordinate>
11 (9), E2­
<geoCoordinate id="EE2C5031FFD15750C8B6FC2D0AA2CE71" box="[1207,1276,1004,1030]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="2.25">
E
<date id="FFA61036FFD15750C8CBFC2D0AA2CE71" box="[1226,1276,1004,1030]" pageId="17" pageNumber="18" value="1915-02-19">2 15</date>
</geoCoordinate>
19 (16), E1­
<geoCoordinate id="EE2C5031FFD15750CD6FFBD50FECCE59" box="[366,434,1044,1070]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="2.1833334">
E
<date id="FFA61036FFD15750CD83FBD50FECCE59" box="[386,434,1044,1070]" pageId="17" pageNumber="18" value="1914-11-02">2 11</date>
</geoCoordinate>
14 (13),
<geoCoordinate id="EE2C5031FFD15750CE27FBD50C37CE59" box="[550,617,1044,1070]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="1.25">
E
<date id="FFA61036FFD15750CE3DFBD50C37CE59" box="[572,617,1044,1070]" pageId="17" pageNumber="18" value="1915-01-22">1 15</date>
</geoCoordinate>
22 (18),
<geoCoordinate id="EE2C5031FFD15750CEDCFBD50D41CE59" box="[733,799,1044,1070]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="2.35">
E
<date id="FFA61036FFD15750CEEEFBD50D41CE59" box="[751,799,1044,1070]" pageId="17" pageNumber="18" value="1942-02-21">2 21</date>
</geoCoordinate>
42 (26),
<date id="FFA61036FFD15750CF92FBD50D85CE59" box="[915,987,1044,1070]" pageId="17" pageNumber="18" value="1959-05-01">V1 59</date>
68 (63),
<date id="FFA61036FFD15750C84DFBD50ACACE59" box="[1100,1172,1044,1070]" pageId="17" pageNumber="18" value="1962-05-02">V2 62</date>
76 (68),
<date id="FFA61036FFD15750C907FBD50F7ACE21" pageId="17" pageNumber="18" value="1980-05-03">V3 80</date>
92 (86),
<date id="FFA61036FFD15750CD99FBFD0F81CE21" box="[408,479,1084,1110]" pageId="17" pageNumber="18" value="1985-05-04">V4 85</date>
94 (90).
</paragraph>
<paragraph id="8BA736F6FFD15750CD39FBA50FCFCE81" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">
Gnathosoma: Base (n=4) 9097 (95), length chelicera (
<figureCitation id="13232A73FFD15750CFDEFBA50A64CE09" box="[991,1082,1124,1150]" captionStart="FIGURES 28 30" captionStartId="18.[264,383,1835,1859]" captionTargetBox="[275,1300,534,1803]" captionTargetId="figure@18.[264,1323,524,1809]" captionTargetPageId="18" captionText="FIGURES 28 30. Sperchon squamosus (original). Fig. 28: dorsal idiosoma, Fig. 29: ventral idiosoma, Fig. 30: chelicera. Scale bars: 28, 29: 100 µm, 30 50 µm." httpUri="https://zenodo.org/record/172013/files/figure.png" pageId="17" pageNumber="18">Fig. 30</figureCitation>
) (n=3) 8387 (85), chela 1416 (15), length/width P
<date id="FFA61036FFD15750CE86FB4D0CBBCED1" box="[647,741,1164,1190]" pageId="17" pageNumber="18" value="1940-02-28">2 2840</date>
(36)/2830 (29), P
<date id="FFA61036FFD15750CFC4FB4D0A7CCED1" box="[965,1058,1164,1190]" pageId="17" pageNumber="18" value="1935-03-30">3 3035</date>
(32)/2629 (27), claw (n=4) 1218 (15), palpal seta C3 82105 (98), C4 5772 (63), C5 6779 (73), lateral seta 7990 (86).
</paragraph>
<paragraph id="8BA736F6FFD15750CD39FAC50C1ECF31" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">Legs: First segments with a little pronounced transverse lineation, some of the other segments slightly lineated.</paragraph>
<paragraph id="8BA736F6FFD15750CD39FA950CC2CFC9" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">
Leg I (
<figureCitation id="13232A73FFD15750CD89FA950FBFCF19" box="[392,481,1364,1390]" captionStart="FIGURES 31 33" captionStartId="19.[264,382,1581,1605]" captionTargetBox="[280,1306,334,1560]" captionTargetId="figure@19.[264,1323,329,1593]" captionTargetPageId="19" captionText="FIGURES 31 33. Sperchon squamosus (original). Fig. 31: I­L, Fig. 32: II­L, Fig. 33: III­L. Scale bar 100 µm." httpUri="https://zenodo.org/record/172014/files/figure.png" pageId="17" pageNumber="18">Fig. 31</figureCitation>
): Total length (n=5) 276300 (289), length/height I­L1 3538 (36)/29 30 (30), I­L2 4348 (46)/3031 (31), I­L3 5660 (59)/2528 (26), I­L4 6672 (70)/2425 (25), I­L5 7682 (78)/ 1920 (20).
</paragraph>
<paragraph id="8BA736F6FFD15750CD39FA0D0CF8CC41" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">
Leg II (
<figureCitation id="13232A73FFD15750CD96FA0D0FADCF91" box="[407,499,1484,1510]" captionStart="FIGURES 31 33" captionStartId="19.[264,382,1581,1605]" captionTargetBox="[280,1306,334,1560]" captionTargetId="figure@19.[264,1323,329,1593]" captionTargetPageId="19" captionText="FIGURES 31 33. Sperchon squamosus (original). Fig. 31: I­L, Fig. 32: II­L, Fig. 33: III­L. Scale bar 100 µm." httpUri="https://zenodo.org/record/172014/files/figure.png" pageId="17" pageNumber="18">Fig. 32</figureCitation>
): Total length 311326 (319), length/height II­L1 4446 (45)/2830 (29), II­L2 4951 (50)/2930 (30), II­L3 6163 (62)/2527 (26), II­L4 7582 (79)/2428 (25), II­L5 8284 (83)/ 1920 (20).
</paragraph>
<paragraph id="8BA736F6FFD15750CD39F9850CA7CCD9" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">
Leg III (
<figureCitation id="13232A73FFD15750CD9FF9850FA9CC29" box="[414,503,1604,1630]" captionStart="FIGURES 31 33" captionStartId="19.[264,382,1581,1605]" captionTargetBox="[280,1306,334,1560]" captionTargetId="figure@19.[264,1323,329,1593]" captionTargetPageId="19" captionText="FIGURES 31 33. Sperchon squamosus (original). Fig. 31: I­L, Fig. 32: II­L, Fig. 33: III­L. Scale bar 100 µm." httpUri="https://zenodo.org/record/172014/files/figure.png" pageId="17" pageNumber="18">Fig. 33</figureCitation>
): Total length 376436 (418), length/width III­L1 7275 (74)/2931 (30), III­L2 6368 (66)/2730 (28), III­L3 8591 (89)/2224 (23), III­L4 104108 (106)/ 2224 (23), III­L5 5294 (83)/1718 (18).
</paragraph>
<paragraph id="8BA736F6FFD15750CD39F97D0F8ACC89" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">Diagnostic characters of larvae: Structure of the Dp (laterally and posteriorly with a rhombic pattern).</paragraph>
<paragraph id="8BA736F6FFD15750CD39F8CD0C0BCD39" blockId="17.[264,1324,284,1870]" pageId="17" pageNumber="18">
Remarks: Previous descriptions of
<taxonomicName id="4C184D75FFD15750CED6F8CD0D90CD52" box="[727,974,1804,1829]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="squamosus">
<emphasis id="B96CEAE4FFD15750CED6F8CD0D90CD52" box="[727,974,1804,1829]" italics="true" pageId="17" pageNumber="18">Sperchon squamosus</emphasis>
</taxonomicName>
often lack measurement data and/or accurate illustrations.
</paragraph>
</subSubSection>
</treatment>
</document>