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<document id="C6F633AAD010DB76E8A8379191530CD2" ID-CLB-Dataset="8231" ID-DOI="10.11646/zootaxa.4860.4.1" ID-GBIF-Dataset="1f5ee6c2-2635-44a8-8469-49319b7a8c62" ID-ISSN="1175-5326" ID-Zenodo-Dep="4414137" ID-ZooBank="876F1085-5296-4340-A951-41420C011917" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609705627587" checkinUser="plazi" docAuthor="Zhadan, Anna" docDate="2020" docId="03A787FE3B59087DABBFF9D9FEDC40AD" docLanguage="en" docName="zootaxa.4860.4.1.pdf" docOrigin="Zootaxa 4860 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scoloplos simplex" docType="treatment" docVersion="10" lastPageNumber="464" masterDocId="FF9EFF863B500870AB28FF93FFAA4260" masterDocTitle="Review of Orbiniidae (Annelida, Sedentaria) from Australia" masterLastPageNumber="502" masterPageNumber="451" pageNumber="460" updateTime="1698831485818" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="19923795AB054335AE93E2B8286BC492">Review of Orbiniidae (Annelida, Sedentaria) from Australia</mods:title>
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<mods:namePart id="306A224E016A79B2B53A91863F86958D">Zhadan, Anna</mods:namePart>
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<mods:date id="66479FBB3769F58925B9385FE2139AAC">2020</mods:date>
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<treatment id="03A787FE3B59087DABBFF9D9FEDC40AD" ID-DOI="http://doi.org/10.5281/zenodo.4538363" ID-GBIF-Taxon="177324640" ID-Zenodo-Dep="4538363" LSID="urn:lsid:plazi:treatment:03A787FE3B59087DABBFF9D9FEDC40AD" httpUri="http://treatment.plazi.org/id/03A787FE3B59087DABBFF9D9FEDC40AD" lastPageId="13" lastPageNumber="464" pageId="9" pageNumber="460">
<subSubSection id="C31465633B590879ABBFF9D9FDE74404" box="[151,589,1610,1637]" pageId="9" pageNumber="460" type="nomenclature">
<paragraph id="8BB136E83B590879ABBFF9D9FDE74404" blockId="9.[151,589,1610,1673]" box="[151,589,1610,1637]" pageId="9" pageNumber="460">
<heading id="D0F981843B590879ABBFF9D9FDE74404" bold="true" box="[151,589,1610,1637]" fontSize="11" level="1" pageId="9" pageNumber="460" reason="1">
<taxonomicName id="4C0E4D6B3B590879ABBFF9D9FDE74404" ID-CoL="4VSLZ" authority="(Hutchings 1974)" baseAuthorityName="Hutchings" baseAuthorityYear="1974" box="[151,589,1610,1637]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="9" pageNumber="460" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B590879ABBFF9D9FDE74404" bold="true" box="[151,589,1610,1637]" pageId="9" pageNumber="460">
<emphasis id="B97AEAFA3B590879ABBFF9D9FEC54405" bold="true" box="[151,367,1610,1637]" italics="true" pageId="9" pageNumber="460">Scoloplos simplex</emphasis>
(
<bibRefCitation id="EF9F4B193B590879AA56F9D9FDEE4404" author="Hutchings, P. A." box="[382,580,1610,1636]" pageId="9" pageNumber="460" pagination="175 - 195" refId="ref34514" refString="Hutchings, P. A. (1974) Polychaeta of Wallis Lake, New South Wales. Proceedings of the Linnean Society of New South Wales, 98, 175 - 195." type="journal article" year="1974">Hutchings 1974</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31465633B590879ABBFF9FCFEB644E9" box="[151,284,1646,1673]" pageId="9" pageNumber="460" type="description">
<paragraph id="8BB136E83B590879ABBFF9FCFEB644E9" blockId="9.[151,589,1610,1673]" box="[151,284,1646,1673]" pageId="9" pageNumber="460">
<figureCitation id="13352A6D3B590879ABBFF9FCFEA844E9" box="[151,258,1646,1673]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[151,1437,525,1879]" captionTargetId="figure-184@11.[151,1437,525,1881]" captionTargetPageId="11" captionText="FIGURE 5. Scoloplos simplex, AD: AM W.5246 holotype, methylene blue staining; EH:AM W.6038 holotype of S. difficilis, methylene blue staining; A. Thorax, dorsal view; B. Thorax-abdomen transition, dorso-lateral view; C. abdomen, dorsal view; D. Posterior end of the body with pygidium, dorso-lateral view; E. Thorax, dorsal view; F. Same, lateral view; G. Thorax-abdomen transition, dorso-lateral view; H. Abdomen, dorsal view. Arrows indicate the appearing of the first branchia." figureDoi="http://doi.org/10.5281/zenodo.4414149" httpUri="https://zenodo.org/record/4414149/files/figure.png" pageId="9" pageNumber="460">Figures 5</figureCitation>
,
<figureCitation id="13352A6D3B590879AA26F9FCFEB644E9" box="[270,284,1647,1673]" captionStart="FIGURE 6" captionStartId="12.[151,250,1676,1700]" captionTargetBox="[151,1436,181,1652]" captionTargetId="figure-28@12.[151,1436,181,1652]" captionTargetPageId="12" captionText="FIGURE 6. Scoloplos simplex, AF: AM W.195999, stereomicroscope, methylene blue staining; GJ: AM W.198694, compound microscope. A. General lateral view; B. Thorax-abdomen transition, lateral view; C. Thorax, lateral view; D. Thoraxabdomen transition, lateral view; E. Anterior end, lateral view; F. Thorax, dorsal view; arrow indicates first branchia; G. Parapodium of chaetiger 2; H. Close up of neuropodia of chaetiger 2; I. Parapodium of chaetiger 27; J. Posterior parapodium. br, branchia; cc, crenulated capillaries; ne, neuropodium; no, notopodium; u, uncini." figureDoi="http://doi.org/10.5281/zenodo.4414155" httpUri="https://zenodo.org/record/4414155/files/figure.png" pageId="9" pageNumber="460">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31465633B590879ABBFF920FDD04547" pageId="9" pageNumber="460" type="reference_group">
<paragraph id="8BB136E83B590879ABBFF920FD7844AB" blockId="9.[151,1054,1715,1831]" box="[151,722,1715,1739]" pageId="9" pageNumber="460">
<taxonomicName id="4C0E4D6B3B590879ABBFF920FDCC44AB" ID-CoL="6LD9H" authority="Hutchings, 1974: 118" authorityName="Hutchings" authorityPageNumber="118" authorityYear="1974" box="[151,614,1715,1739]" class="Polychaeta" family="Orbiniidae" genus="Haploscoloplos" kingdom="Animalia" pageId="9" pageNumber="460" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B590879ABBFF920FE2044AB" box="[151,394,1715,1739]" italics="true" pageId="9" pageNumber="460">Haploscoloplos simplex</emphasis>
<bibRefCitation id="EF9F4B193B590879AAB8F920FDCC44AB" author="Hutchings, P. A." box="[400,614,1715,1739]" pageId="9" pageNumber="460" pagination="175 - 195" refId="ref34514" refString="Hutchings, P. A. (1974) Polychaeta of Wallis Lake, New South Wales. Proceedings of the Linnean Society of New South Wales, 98, 175 - 195." type="journal article" year="1974">Hutchings, 1974:118</bibRefCitation>
</taxonomicName>
, fig. 2ad.
</paragraph>
<paragraph id="8BB136E83B590879ABBFF941FBB4448A" blockId="9.[151,1054,1715,1831]" box="[151,1054,1746,1770]" pageId="9" pageNumber="460">
<treatmentCitationGroup id="AB1E11C63B590879ABBFF941FBB4448A" box="[151,1054,1746,1770]" pageId="9" pageNumber="460">
<treatmentCitation id="0AAF10F93B590879ABBFF941FD0B448A" author="Day, J. H." box="[151,673,1746,1770]" page="229" pageId="9" pageNumber="460" year="1977">
<taxonomicName id="4C0E4D6B3B590879ABBFF941FD0B448A" ID-CoL="4VSLZ" authority=": Day 1977: 228 - 229" authorityName="Day" authorityPageNumber="228 - 229" authorityYear="1977" box="[151,673,1746,1770]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="9" pageNumber="460" phylum="Annelida" rank="species" species="simplex" subGenus="Scoloplos">
<emphasis id="B97AEAFA3B590879ABBFF941FE6D448A" box="[151,455,1746,1770]" italics="true" pageId="9" pageNumber="460">Scoloplos (Scoloplos) simplex</emphasis>
:
<bibRefCitation id="EF9F4B193B590879AAFDF941FD0B448A" author="Day, J. H." box="[469,673,1746,1770]" pageId="9" pageNumber="460" pagination="217 - 243" refId="ref33341" refString="Day, J. H. (1977) A review of the Australian and New Zealand Orbiniidae (Annelida: Polychaeta). In: Reish, K. &amp; Fauchald, D. J., (Ed.), Essays on Polychaetous Annelids in Memory of Dr. Olga Hartman. Allan Hancock Foundation, Los Angeles, pp. 217 - 243." type="book chapter" year="1977">Day 1977: 228229</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AAF10F93B590879A983F941FBB3448A" author="Hutchings, P. &amp; Rainer, S." box="[683,1049,1746,1770]" page="761" pageId="9" pageNumber="460" year="1979">
<bibRefCitation id="EF9F4B193B590879A983F941FBB3448A" author="Hutchings, P. &amp; Rainer, S." box="[683,1049,1746,1770]" pageId="9" pageNumber="460" pagination="745 - 796" refId="ref34466" refString="Hutchings, P. &amp; Rainer, S. (1979) The polychaete fauna of Careel Bay, Pittwater, New South Wales, Australia. Journal of Natural History, 13, 745 - 796. https: // doi. org / 10.1080 / 00222937900770561" type="journal article" year="1979">Hutchings &amp; Rainer 1979: 761762</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB136E83B590879ABBFF963FCA34569" blockId="9.[151,1054,1715,1831]" box="[151,777,1776,1801]" pageId="9" pageNumber="460">
<treatmentCitationGroup id="AB1E11C63B590879ABBFF963FCA34569" box="[151,777,1776,1801]" pageId="9" pageNumber="460">
<treatmentCitation id="0AAF10F93B590879ABBFF963FD354569" author="Day, J. H." box="[151,671,1776,1801]" page="230" pageId="9" pageNumber="460" year="1977">
<taxonomicName id="4C0E4D6B3B590879ABBFF963FD354569" ID-CoL="4VSKF" authority="Day 1977: 229 - 230" authorityName="Day" authorityPageNumber="229 - 230" authorityYear="1977" box="[151,671,1776,1801]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="9" pageNumber="460" phylum="Annelida" rank="species" species="difficilis" subGenus="Scoloplos">
<emphasis id="B97AEAFA3B590879ABBFF963FE674568" box="[151,461,1776,1800]" italics="true" pageId="9" pageNumber="460">Scoloplos (Scoloplos) difficilis</emphasis>
<bibRefCitation id="EF9F4B193B590879AAFBF962FD354569" author="Day, J. H." box="[467,671,1777,1801]" pageId="9" pageNumber="460" pagination="217 - 243" refId="ref33341" refString="Day, J. H. (1977) A review of the Australian and New Zealand Orbiniidae (Annelida: Polychaeta). In: Reish, K. &amp; Fauchald, D. J., (Ed.), Essays on Polychaetous Annelids in Memory of Dr. Olga Hartman. Allan Hancock Foundation, Los Angeles, pp. 217 - 243." type="book chapter" year="1977">Day 1977: 229230</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 2ae.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB136E83B590879ABBFF89CFDD04547" blockId="9.[151,1054,1715,1831]" box="[151,634,1807,1831]" pageId="9" pageNumber="460">
<treatmentCitationGroup id="AB1E11C63B590879ABBFF89CFDD04547" box="[151,634,1807,1831]" pageId="9" pageNumber="460">
<treatmentCitation id="0AAF10F93B590879ABBFF89CFDA44547" author="Brown, B." box="[151,526,1807,1831]" page="762" pageId="9" pageNumber="460" year="1979">
<taxonomicName id="4C0E4D6B3B590879ABBFF89CFDA44547" ID-CoL="4VSLZ" authority=": Brown 1979: 762" authorityName=": Brown" authorityPageNumber="762" authorityYear="1979" box="[151,526,1807,1831]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="9" pageNumber="460" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B590879ABBFF89CFEE74547" box="[151,333,1807,1831]" italics="true" pageId="9" pageNumber="460">Scoloplos simplex</emphasis>
:
<bibRefCitation id="EF9F4B193B590879AA73F883FDA44547" author="Brown, B." box="[347,526,1807,1831]" pageId="9" pageNumber="460" pagination="762 - 768" refId="ref32972" refString="Brown, B. (1979) A redescription of Scoloplos simplex (Hutchings, 1974) (Polychaeta: Orbiniidae) from Australia. Proceedings of The Biological Society of Washington, 92, 762 - 768." type="journal article" year="1979">Brown 1979: 762</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 1ad.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31465633B59087AABBFF8C6FD654321" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460" type="materials_examined">
<paragraph id="8BB136E83B59087AABBFF8C6FF7042B5" blockId="9.[151,1436,1877,2012]" lastBlockId="10.[151,1436,151,321]" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460">
<emphasis id="B97AEAFA3B590879ABBFF8C6FE2B450F" bold="true" box="[151,385,1877,1903]" pageId="9" pageNumber="460">Material examined.</emphasis>
<materialsCitation id="3B663CB53B59087AAAA1F8C6FF7042B5" ID-GBIF-Occurrence="3034643326" collectingDate="1970-12" collectingDateMax="1996-08-04" collectingDateMin="1970-12" collectionCode="AM" collectorName="P. Dixon &amp; Pittwater &amp; D. Dexter &amp; P. A. Hutchings &amp; Tamar &amp; D. Henderson" county="Wallis Lake" lastPageId="10" lastPageNumber="461" latitude="-41.1" location="Towlers Bay" longLatPrecision="1158" longitude="146.81667" municipality="O'Gower" pageId="9" pageNumber="460" specimenCount="35" stateProvince="New South Wales" typeStatus="holotype">
<emphasis id="B97AEAFA3B590879AAA1F8C6FDF1450F" bold="true" box="[393,603,1877,1903]" pageId="9" pageNumber="460">
<collectingRegion id="49CAF80A3B590879AAA1F8C6FDF1450F" box="[393,603,1877,1903]" country="Australia" name="New South Wales" pageId="9" pageNumber="460">New South Wales</collectingRegion>
</emphasis>
: entrance to the
<collectingCounty id="62D04E643B590879A831F8C6FC08450F" box="[793,930,1877,1903]" pageId="9" pageNumber="460">Wallis Lake</collectingCounty>
,
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,
<geoCoordinate id="EE3A502F3B590879AF31F8C6FB2D4510" box="[1049,1159,1877,1904]" degrees="152" direction="east" minutes="29" orientation="longitude" pageId="9" pageNumber="460" precision="925" value="152.48334">152°29E</geoCoordinate>
,
<date id="FFB010283B590879AFBCF8C6FAE0450F" box="[1172,1354,1877,1903]" pageId="9" pageNumber="460" value="1970-12">
<collectingDate id="EFF4E9C03B590879AFBCF8C6FAE0450F" box="[1172,1354,1877,1903]" pageId="9" pageNumber="460" value="1970-12">December 1970</collectingDate>
</date>
, sandy, fast flowing water, coll.
<collectorName id="26FB533E3B590879AA8AF8E9FDAF45F3" box="[418,517,1913,1940]" pageId="9" pageNumber="460">P. Dixon</collectorName>
,
<collectingMunicipality id="6BD5AC923B590879A926F8EAFDD345F3" box="[526,633,1913,1939]" pageId="9" pageNumber="460">OGower</collectingMunicipality>
,
<collectionCode id="ED1FAE2D3B590879A9AAF8E9FD0545F4" box="[642,687,1914,1940]" country="Australia" name="Australian Museum" pageId="9" pageNumber="460" type="Museum">AM</collectionCode>
W.5246,
<typeStatus id="54B5884A3B590879A830F8EAFC2E45F3" box="[792,900,1913,1939]" pageId="9" pageNumber="460" type="holotype">Holotype</typeStatus>
;
<location id="8ED160333B590879A8A5F8EAFBB045F4" LSID="urn:lsid:plazi:treatment:03A787FE3B59087DABBFF9D9FEDC40AD:8ED160333B590879A8A5F8EAFBB045F4" box="[909,1050,1913,1940]" county="Wallis Lake" latitude="-41.1" longLatPrecision="1158" longitude="146.81667" municipality="O'Gower" name="Towlers Bay" pageId="9" pageNumber="460" stateProvince="New South Wales">Towlers Bay</location>
,
<collectorName id="26FB533E3B590879AF0CF8EAFB2145F3" box="[1060,1163,1913,1939]" pageId="9" pageNumber="460">Pittwater</collectorName>
,
<geoCoordinate id="EE3A502F3B590879AFBDF8EAFAB545F3" box="[1173,1311,1913,1939]" degrees="33" direction="south" minutes="37" orientation="latitude" pageId="9" pageNumber="460" precision="15" seconds="30" value="-33.625">33°37 30”S</geoCoordinate>
,
<geoCoordinate id="EE3A502F3B590879AE02F8EAFA3D45F4" box="[1322,1431,1913,1940]" degrees="151" direction="east" minutes="17" orientation="longitude" pageId="9" pageNumber="460" precision="925" value="151.28334">151°17E</geoCoordinate>
,
<date id="FFB010283B590879ABBFF80EFEB245D7" box="[151,280,1949,1975]" pageId="9" pageNumber="460" value="1980-12-19">
<collectingDate id="EFF4E9C03B590879ABBFF80EFEB245D7" box="[151,280,1949,1975]" pageId="9" pageNumber="460" value="1980-12-19">19.12.1980</collectingDate>
</date>
, intertidal, sand, coll.
<collectorName id="26FB533E3B590879A92EF80DFDDE45D8" box="[518,628,1950,1976]" pageId="9" pageNumber="460">D. Dexter</collectorName>
,
<collectionCode id="ED1FAE2D3B590879A954F80DFD0345D8" box="[636,681,1950,1976]" country="Australia" name="Australian Museum" pageId="9" pageNumber="460" type="Museum">AM</collectionCode>
W.195999,
<specimenCount id="9D08FD613B590879A803F80EFC6E45D7" box="[811,964,1949,1975]" pageId="9" pageNumber="460" type="generic">21 specimens</specimenCount>
;
<location id="8ED160333B590879A8E5F80EFBE345D8" LSID="urn:lsid:plazi:treatment:03A787FE3B59087DABBFF9D9FEDC40AD:8ED160333B590879A8E5F80EFBE345D8" box="[973,1097,1949,1976]" county="Wallis Lake" latitude="-41.1" longLatPrecision="1158" longitude="146.81667" municipality="O'Gower" name="Careel Bay" pageId="9" pageNumber="460" stateProvince="New South Wales">Careel Bay</location>
,
<collectorName id="26FB533E3B590879AF7AF80EFB1345D7" box="[1106,1209,1949,1975]" pageId="9" pageNumber="460">Pittwater</collectorName>
,
<geoCoordinate id="EE3A502F3B590879AFEAF80EFA8A45D7" box="[1218,1312,1949,1975]" degrees="33" direction="south" minutes="37" orientation="latitude" pageId="9" pageNumber="460" precision="925" value="-33.616665">33°37S</geoCoordinate>
,
<geoCoordinate id="EE3A502F3B590879AE02F80EFA3D45D8" box="[1322,1431,1949,1976]" degrees="151" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="460" precision="925" value="151.31667">151°19E</geoCoordinate>
,
<date id="FFB010283B590879ABBFF852FEB245BB" box="[151,280,1985,2011]" pageId="9" pageNumber="460" value="1996-08-04">
<collectingDate id="EFF4E9C03B590879ABBFF852FEB245BB" box="[151,280,1985,2011]" pageId="9" pageNumber="460" value="1996-08-04">08.04.1996</collectingDate>
</date>
, intertidal,
<taxonomicName id="4C0E4D6B3B590879AAB5F851FE5945BB" box="[413,499,1986,2011]" class="Liliopsida" family="Zosteraceae" genus="Zostera" kingdom="Plantae" order="Alismatales" pageId="9" pageNumber="460" phylum="Tracheophyta" rank="genus">
<emphasis id="B97AEAFA3B590879AAB5F851FE5945BB" box="[413,499,1986,2011]" italics="true" pageId="9" pageNumber="460">Zostera</emphasis>
</taxonomicName>
, coll.
<collectorName id="26FB533E3B590879A916F851FD4745BB" box="[574,749,1985,2012]" pageId="9" pageNumber="460">P.A. Hutchings</collectorName>
,
<collectionCode id="ED1FAE2D3B590879A9D2F851FC8D45BC" box="[762,807,1986,2012]" country="Australia" name="Australian Museum" pageId="9" pageNumber="460" type="Museum">AM</collectionCode>
W.23424,
<specimenCount id="9D08FD613B590879A88FF852FB9D45BB" box="[935,1079,1985,2011]" pageId="9" pageNumber="460" type="generic">3 specimens</specimenCount>
.
<emphasis id="B97AEAFA3B590879AF12F852FB0445BB" bold="true" box="[1082,1198,1985,2011]" pageId="9" pageNumber="460">
<collectingRegion id="49CAF80A3B590879AF12F852FB0445BB" box="[1082,1198,1985,2011]" country="Australia" name="Tasmania" pageId="9" pageNumber="460">Tasmania</collectingRegion>
</emphasis>
: near
<location id="8ED160333B590879AFD2F852FA3245BC" LSID="urn:lsid:plazi:treatment:03A787FE3B59087DABBFF9D9FEDC40AD:8ED160333B590879AFD2F852FA3245BC" box="[1274,1432,1985,2012]" county="Wallis Lake" latitude="-41.1" longLatPrecision="1158" longitude="146.81667" municipality="O'Gower" name="George Town" pageId="9" pageNumber="460" stateProvince="New South Wales">George Town</location>
,
<collectorName id="26FB533E3B5A087AABBFFF04FF7442D1" box="[151,222,151,177]" pageId="10" pageNumber="461">Tamar</collectorName>
estuary,
<geoCoordinate id="EE3A502F3B5A087AAA6FFF04FE0F42D1" box="[327,421,151,177]" degrees="41" direction="south" minutes="06" orientation="latitude" pageId="10" pageNumber="461" precision="925" value="-41.1">41°06S</geoCoordinate>
,
<geoCoordinate id="EE3A502F3B5A087AAA9BFF04FD8B42D1" box="[435,545,151,177]" degrees="146" direction="east" minutes="49" orientation="longitude" pageId="10" pageNumber="461" precision="925" value="146.81667">146°49E</geoCoordinate>
,
<date id="FFB010283B5A087AA907FF04FD7242D1" box="[559,728,151,177]" pageId="10" pageNumber="461" value="1984-02">
<collectingDate id="EFF4E9C03B5A087AA907FF04FD7242D1" box="[559,728,151,177]" pageId="10" pageNumber="461" value="1984-02">February 1984</collectingDate>
</date>
, mud and sand, coll.
<collectorName id="26FB533E3B5A087AA8F8FF04FBDA42D1" box="[976,1136,151,177]" pageId="10" pageNumber="461">D. Henderson</collectorName>
,
<collectionCode id="ED1FAE2D3B5A087AAF54FF04FB0342D1" box="[1148,1193,151,177]" country="Australia" name="Australian Museum" pageId="10" pageNumber="461" type="Museum">AM</collectionCode>
W.198694,
<specimenCount id="9D08FD613B5A087AAE1DFF04FF7C42B5" pageId="10" pageNumber="461" type="generic">11 specimens</specimenCount>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB136E83B5A087AABEFFF4DFB1C437D" blockId="10.[151,1436,151,321]" pageId="10" pageNumber="461">
<emphasis id="B97AEAFA3B5A087AABEFFF4DFDFE4298" bold="true" box="[199,596,222,248]" pageId="10" pageNumber="461">Comparative material examined.</emphasis>
<materialsCitation id="3B663CB53B5A087AA974FF4DFB1C437D" ID-GBIF-Occurrence="3034643338" collectingDate="1973-12-07" collectionCode="AM" collectorName="P. A. Hutchings &amp; Hutchings, C" latitude="-27.35" location="Moreton Bay" longLatPrecision="1237" longitude="153.1" municipality="Jackson Creek" pageId="10" pageNumber="461" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="B97AEAFA3B5A087AA974FF4DFD404298" bold="true" box="[604,746,222,248]" pageId="10" pageNumber="461">
<collectingRegion id="49CAF80A3B5A087AA974FF4DFD404298" box="[604,746,222,248]" country="Australia" name="Queensland" pageId="10" pageNumber="461">Queensland</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6BD5AC923B5A087AA9D1FF4CFC0A4299" box="[761,928,223,249]" pageId="10" pageNumber="461">Jackson Creek</collectingMunicipality>
, mouth of
<location id="8ED160333B5A087AAF33FF4CFB184299" LSID="urn:lsid:plazi:treatment:03A787FE3B59087DABBFF9D9FEDC40AD:8ED160333B5A087AAF33FF4CFB184299" box="[1051,1202,223,249]" latitude="-27.35" longLatPrecision="1237" longitude="153.1" municipality="Jackson Creek" name="Moreton Bay" pageId="10" pageNumber="461" stateProvince="Queensland">Moreton Bay</location>
,
<geoCoordinate id="EE3A502F3B5A087AAF96FF4CFAB64299" box="[1214,1308,223,249]" degrees="27" direction="south" minutes="21" orientation="latitude" pageId="10" pageNumber="461" precision="925" value="-27.35">27°21S</geoCoordinate>
,
<geoCoordinate id="EE3A502F3B5A087AAE02FF4DFA324299" box="[1322,1432,222,249]" degrees="153" direction="east" minutes="06" orientation="longitude" pageId="10" pageNumber="461" precision="925" value="153.1">153°06E</geoCoordinate>
,
<date id="FFB010283B5A087AABBFFE90FEBD437D" box="[151,279,259,285]" pageId="10" pageNumber="461" value="1973-12-07">
<collectingDate id="EFF4E9C03B5A087AABBFFE90FEBD437D" box="[151,279,259,285]" pageId="10" pageNumber="461" value="1973-12-07">12.07.1973</collectingDate>
</date>
, sand flats, coll.
<collectorName id="26FB533E3B5A087AAAFCFE90FDD5437D" box="[468,639,259,285]" pageId="10" pageNumber="461">P.A. Hutchings</collectorName>
, C. Wallace,
<collectionCode id="ED1FAE2D3B5A087AA839FE90FC94437D" box="[785,830,259,285]" country="Australia" name="Australian Museum" pageId="10" pageNumber="461" type="Museum">AM</collectionCode>
W.6038,
<typeStatus id="54B5884A3B5A087AA881FE90FBBB437D" box="[937,1041,259,285]" pageId="10" pageNumber="461" type="holotype">Holotype</typeStatus>
of
<emphasis id="B97AEAFA3B5A087AAF1FFE90FB1C437D" box="[1079,1206,259,285]" italics="true" pageId="10" pageNumber="461">
<taxonomicName id="4C0E4D6B3B5A087AAF1FFE90FB19437D" authorityName="Day" authorityYear="1977" box="[1079,1203,259,285]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="10" pageNumber="461" phylum="Annelida" rank="species" species="difficilis">S. difficilis</taxonomicName>
.
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8BB136E83B5A087AABEFFEB4FD654321" blockId="10.[151,1436,151,321]" box="[199,719,294,321]" pageId="10" pageNumber="461">
<materialsCitation id="3B663CB53B5A087AABEFFEB4FD614321" ID-GBIF-Occurrence="3034643335" box="[199,715,294,321]" country="United Kingdom" county="South Wales" location="Lake" municipality="New South Wales" pageId="10" pageNumber="461" specimenCount="1" typeStatus="holotype">
<emphasis id="B97AEAFA3B5A087AABEFFEB4FECC4320" bold="true" box="[199,358,294,321]" pageId="10" pageNumber="461">
<typeStatus id="54B5884A3B5A087AABEFFEB4FEA84321" box="[199,258,295,321]" pageId="10" pageNumber="461">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31976783B5A087AAA44FEB4FE194321" box="[364,435,295,321]" name="United Kingdom" pageId="10" pageNumber="461">Wallis</collectingCountry>
<location id="8ED160333B5A087AAA92FEB4FE5F4321" LSID="urn:lsid:plazi:treatment:03A787FE3B59087DABBFF9D9FEDC40AD:8ED160333B5A087AAA92FEB4FE5F4321" box="[442,501,295,321]" country="United Kingdom" county="South Wales" municipality="New South Wales" name="Lake" pageId="10" pageNumber="461">Lake</location>
,
<collectingMunicipality id="6BD5AC923B5A087AA928FEB4FD614321" box="[512,715,295,321]" pageId="10" pageNumber="461">
New
<collectingCounty id="62D04E643B5A087AA914FEB4FD614321" box="[572,715,295,321]" pageId="10" pageNumber="461">South Wales</collectingCounty>
</collectingMunicipality>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF7166603B5A087AABBFF956FB8C452A" ID-DOI="http://doi.org/10.5281/zenodo.4414147" ID-Zenodo-Dep="4414147" httpUri="https://zenodo.org/record/4414147/files/figure.png" pageId="10" pageNumber="461" startId="10.[151,250,1733,1757]" targetBox="[151,1436,345,1710]" targetPageId="10">
<paragraph id="8BB136E83B5A087AABBFF956FB8C452A" blockId="10.[151,1436,1733,1866]" pageId="10" pageNumber="461">
<emphasis id="B97AEAFA3B5A087AABBFF956FEB944BE" bold="true" box="[151,275,1733,1758]" pageId="10" pageNumber="461">FIGURE 4.</emphasis>
<taxonomicName id="4C0E4D6B3B5A087AAA31F955FDD644BE" authority=", AM G." authorityName="AM G." box="[281,636,1734,1758]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="10" pageNumber="461" phylum="Annelida" rank="species" species="novaehollandiae">
<emphasis id="B97AEAFA3B5A087AAA31F955FD8144BE" box="[281,555,1734,1758]" italics="true" pageId="10" pageNumber="461">Scoloplos novaehollandiae</emphasis>
, AM G.
</taxonomicName>
11238. AE: stereomicroscope, CE: methylene blue staining; F, G: compound microscope. A. General view; B. Anterior end, dorsal view; C. Parapodia of chaetiger 26, lateral view; D. Thorax-abdomen transition, lateral view; E. Posterior part of abdomen, lateral view; F. Neuropodium of chaetiger 12 showing uncini (u); G. Parapodium of posterior abdominal chaetiger. br, branchia; ne, neuropodium; no, notopodium.
</paragraph>
</caption>
<subSubSection id="C31465633B5A087BABEFF8EBFAFF4395" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461" type="description">
<paragraph id="8BB136E83B5A087BABEFF8EBFAFF4395" blockId="10.[151,1437,1912,2011]" lastBlockId="11.[151,1436,150,501]" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461">
<emphasis id="B97AEAFA3B5A087AABEFF8EBFEFB45F2" bold="true" box="[199,337,1912,1938]" pageId="10" pageNumber="461">Description</emphasis>
.
<typeStatus id="54B5884A3B5A087AAA77F8EBFE6D45F2" box="[351,455,1912,1938]" pageId="10" pageNumber="461" type="holotype">Holotype</typeStatus>
complete, with thoracic width 2.3 mm, other specimens 0.5
<quantity id="4CF69B0D3B5A087AAF47F8EBFB0745F2" box="[1135,1197,1912,1939]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="461" unit="mm" value="3.0">3 mm</quantity>
wide. Thoracic chaetiger numbering
<quantity id="4CF69B0D3B5A087AAA7FF80FFE3845D6" box="[343,402,1948,1974]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="10" pageNumber="461" unit="in" value="16.0">16 in</quantity>
<typeStatus id="54B5884A3B5A087AAAB3F80FFDAA45D6" box="[411,512,1948,1974]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
,
<quantity id="4CF69B0D3B5A087AA926F80FFDD945D6" box="[526,627,1948,1974]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="4.064" metricValueMin="3.048" pageId="10" pageNumber="461" unit="in" value="14.0" valueMax="16.0" valueMin="12.0">1216 in</quantity>
other specimens. Thorax cylindrical, occasionally slightly flattened or swollen anteriorly (
<figureCitation id="13352A6D3B5A087AAA5CF853FDBA45BA" box="[372,528,1984,2011]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[151,1437,525,1879]" captionTargetId="figure-184@11.[151,1437,525,1881]" captionTargetPageId="11" captionText="FIGURE 5. Scoloplos simplex, AD: AM W.5246 holotype, methylene blue staining; EH:AM W.6038 holotype of S. difficilis, methylene blue staining; A. Thorax, dorsal view; B. Thorax-abdomen transition, dorso-lateral view; C. abdomen, dorsal view; D. Posterior end of the body with pygidium, dorso-lateral view; E. Thorax, dorsal view; F. Same, lateral view; G. Thorax-abdomen transition, dorso-lateral view; H. Abdomen, dorsal view. Arrows indicate the appearing of the first branchia." figureDoi="http://doi.org/10.5281/zenodo.4414149" httpUri="https://zenodo.org/record/4414149/files/figure.png" pageId="10" pageNumber="461">Figs 5A, E, F</figureCitation>
,
<figureCitation id="13352A6D3B5A087AA934F853FD2B45BA" box="[540,641,1984,2010]" captionStart="FIGURE 6" captionStartId="12.[151,250,1676,1700]" captionTargetBox="[151,1436,181,1652]" captionTargetId="figure-28@12.[151,1436,181,1652]" captionTargetPageId="12" captionText="FIGURE 6. Scoloplos simplex, AF: AM W.195999, stereomicroscope, methylene blue staining; GJ: AM W.198694, compound microscope. A. General lateral view; B. Thorax-abdomen transition, lateral view; C. Thorax, lateral view; D. Thoraxabdomen transition, lateral view; E. Anterior end, lateral view; F. Thorax, dorsal view; arrow indicates first branchia; G. Parapodium of chaetiger 2; H. Close up of neuropodia of chaetiger 2; I. Parapodium of chaetiger 27; J. Posterior parapodium. br, branchia; cc, crenulated capillaries; ne, neuropodium; no, notopodium; u, uncini." figureDoi="http://doi.org/10.5281/zenodo.4414155" httpUri="https://zenodo.org/record/4414155/files/figure.png" pageId="10" pageNumber="461">6A, C, F</figureCitation>
). Branchiae from chaetiger 916, as small papillae; in abdomen bran- chiae rapidly increasing in size, becoming longer, narrow triangular (
<figureCitation id="13352A6D3B5B087BA8B7FF04FC4F42D1" box="[927,997,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[151,1437,525,1879]" captionTargetId="figure-184@11.[151,1437,525,1881]" captionTargetPageId="11" captionText="FIGURE 5. Scoloplos simplex, AD: AM W.5246 holotype, methylene blue staining; EH:AM W.6038 holotype of S. difficilis, methylene blue staining; A. Thorax, dorsal view; B. Thorax-abdomen transition, dorso-lateral view; C. abdomen, dorsal view; D. Posterior end of the body with pygidium, dorso-lateral view; E. Thorax, dorsal view; F. Same, lateral view; G. Thorax-abdomen transition, dorso-lateral view; H. Abdomen, dorsal view. Arrows indicate the appearing of the first branchia." figureDoi="http://doi.org/10.5281/zenodo.4414149" httpUri="https://zenodo.org/record/4414149/files/figure.png" pageId="11" pageNumber="462">Figs 5</figureCitation>
BD, H, 6D, F, J). Thoracic notopodial postchaetal lobes absent or very small on anterior chaetigers, then increasing in size, becoming triangular in shape and-digitiform in posterior thoracic segments (
<figureCitation id="13352A6D3B5B087BA998FF4CFD5E4299" box="[688,756,223,249]" captionStart="FIGURE 6" captionStartId="12.[151,250,1676,1700]" captionTargetBox="[151,1436,181,1652]" captionTargetId="figure-28@12.[151,1436,181,1652]" captionTargetPageId="12" captionText="FIGURE 6. Scoloplos simplex, AF: AM W.195999, stereomicroscope, methylene blue staining; GJ: AM W.198694, compound microscope. A. General lateral view; B. Thorax-abdomen transition, lateral view; C. Thorax, lateral view; D. Thoraxabdomen transition, lateral view; E. Anterior end, lateral view; F. Thorax, dorsal view; arrow indicates first branchia; G. Parapodium of chaetiger 2; H. Close up of neuropodia of chaetiger 2; I. Parapodium of chaetiger 27; J. Posterior parapodium. br, branchia; cc, crenulated capillaries; ne, neuropodium; no, notopodium; u, uncini." figureDoi="http://doi.org/10.5281/zenodo.4414155" httpUri="https://zenodo.org/record/4414155/files/figure.png" pageId="11" pageNumber="462">Fig. 6</figureCitation>
BE, G). Thoracic neuropodial postchaetal lobes absent on anterior chaetigers, then arising as small, papilliform to triangular lobes, slightly increasing in size along thorax, all simple (
<figureCitation id="13352A6D3B5B087BAA3FFEB4FEF34321" box="[279,345,295,321]" captionStart="FIGURE 6" captionStartId="12.[151,250,1676,1700]" captionTargetBox="[151,1436,181,1652]" captionTargetId="figure-28@12.[151,1436,181,1652]" captionTargetPageId="12" captionText="FIGURE 6. Scoloplos simplex, AF: AM W.195999, stereomicroscope, methylene blue staining; GJ: AM W.198694, compound microscope. A. General lateral view; B. Thorax-abdomen transition, lateral view; C. Thorax, lateral view; D. Thoraxabdomen transition, lateral view; E. Anterior end, lateral view; F. Thorax, dorsal view; arrow indicates first branchia; G. Parapodium of chaetiger 2; H. Close up of neuropodia of chaetiger 2; I. Parapodium of chaetiger 27; J. Posterior parapodium. br, branchia; cc, crenulated capillaries; ne, neuropodium; no, notopodium; u, uncini." figureDoi="http://doi.org/10.5281/zenodo.4414155" httpUri="https://zenodo.org/record/4414155/files/figure.png" pageId="11" pageNumber="462">Fig. 6</figureCitation>
BE, G). Abdominal notopodial lobes narrow, from foliaceous to digitate, longer than branchiae (
<figureCitation id="13352A6D3B5B087BABB7FED8FEC04305" box="[159,362,330,357]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[151,1437,525,1879]" captionTargetId="figure-184@11.[151,1437,525,1881]" captionTargetPageId="11" captionText="FIGURE 5. Scoloplos simplex, AD: AM W.5246 holotype, methylene blue staining; EH:AM W.6038 holotype of S. difficilis, methylene blue staining; A. Thorax, dorsal view; B. Thorax-abdomen transition, dorso-lateral view; C. abdomen, dorsal view; D. Posterior end of the body with pygidium, dorso-lateral view; E. Thorax, dorsal view; F. Same, lateral view; G. Thorax-abdomen transition, dorso-lateral view; H. Abdomen, dorsal view. Arrows indicate the appearing of the first branchia." figureDoi="http://doi.org/10.5281/zenodo.4414149" httpUri="https://zenodo.org/record/4414149/files/figure.png" pageId="11" pageNumber="462">Figs 5C, D, G, H</figureCitation>
,
<figureCitation id="13352A6D3B5B087BAA5FFED8FE644305" box="[375,462,331,357]" captionStart="FIGURE 6" captionStartId="12.[151,250,1676,1700]" captionTargetBox="[151,1436,181,1652]" captionTargetId="figure-28@12.[151,1436,181,1652]" captionTargetPageId="12" captionText="FIGURE 6. Scoloplos simplex, AF: AM W.195999, stereomicroscope, methylene blue staining; GJ: AM W.198694, compound microscope. A. General lateral view; B. Thorax-abdomen transition, lateral view; C. Thorax, lateral view; D. Thoraxabdomen transition, lateral view; E. Anterior end, lateral view; F. Thorax, dorsal view; arrow indicates first branchia; G. Parapodium of chaetiger 2; H. Close up of neuropodia of chaetiger 2; I. Parapodium of chaetiger 27; J. Posterior parapodium. br, branchia; cc, crenulated capillaries; ne, neuropodium; no, notopodium; u, uncini." figureDoi="http://doi.org/10.5281/zenodo.4414155" httpUri="https://zenodo.org/record/4414155/files/figure.png" pageId="11" pageNumber="462">6B, I, J</figureCitation>
). Abdominal neuropodia bilobed, with smaller outer lobe; in posterior segments with subpodal notch and narrow subpodal flange (
<figureCitation id="13352A6D3B5B087BA9A2FEFCFD6543E9" box="[650,719,366,393]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[151,1437,525,1879]" captionTargetId="figure-184@11.[151,1437,525,1881]" captionTargetPageId="11" captionText="FIGURE 5. Scoloplos simplex, AD: AM W.5246 holotype, methylene blue staining; EH:AM W.6038 holotype of S. difficilis, methylene blue staining; A. Thorax, dorsal view; B. Thorax-abdomen transition, dorso-lateral view; C. abdomen, dorsal view; D. Posterior end of the body with pygidium, dorso-lateral view; E. Thorax, dorsal view; F. Same, lateral view; G. Thorax-abdomen transition, dorso-lateral view; H. Abdomen, dorsal view. Arrows indicate the appearing of the first branchia." figureDoi="http://doi.org/10.5281/zenodo.4414149" httpUri="https://zenodo.org/record/4414149/files/figure.png" pageId="11" pageNumber="462">Figs 5</figureCitation>
BD, G, H, 6B, I, J). Thoracic notochaetae all crenulate capillaries; thoracic neurochaetae divided into upper and lower fascicles; 23 rows of crenulate capillaries in both fascicles and slender slightly curved finely serrated uncini forming anterior row in upper fascicle (
<figureCitation id="13352A6D3B5B087BAFA4FE24FAA143B1" box="[1164,1291,439,465]" captionStart="FIGURE 6" captionStartId="12.[151,250,1676,1700]" captionTargetBox="[151,1436,181,1652]" captionTargetId="figure-28@12.[151,1436,181,1652]" captionTargetPageId="12" captionText="FIGURE 6. Scoloplos simplex, AF: AM W.195999, stereomicroscope, methylene blue staining; GJ: AM W.198694, compound microscope. A. General lateral view; B. Thorax-abdomen transition, lateral view; C. Thorax, lateral view; D. Thoraxabdomen transition, lateral view; E. Anterior end, lateral view; F. Thorax, dorsal view; arrow indicates first branchia; G. Parapodium of chaetiger 2; H. Close up of neuropodia of chaetiger 2; I. Parapodium of chaetiger 27; J. Posterior parapodium. br, branchia; cc, crenulated capillaries; ne, neuropodium; no, notopodium; u, uncini." figureDoi="http://doi.org/10.5281/zenodo.4414155" httpUri="https://zenodo.org/record/4414155/files/figure.png" pageId="11" pageNumber="462">Fig. 6G, H</figureCitation>
). Abdominal chaetae all thin capillaries, forked chaetae not found. Pygidium with two short cirri, anus terminal (
<figureCitation id="13352A6D3B5B087BAFC6FE48FAE24395" box="[1262,1352,474,501]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[151,1437,525,1879]" captionTargetId="figure-184@11.[151,1437,525,1881]" captionTargetPageId="11" captionText="FIGURE 5. Scoloplos simplex, AD: AM W.5246 holotype, methylene blue staining; EH:AM W.6038 holotype of S. difficilis, methylene blue staining; A. Thorax, dorsal view; B. Thorax-abdomen transition, dorso-lateral view; C. abdomen, dorsal view; D. Posterior end of the body with pygidium, dorso-lateral view; E. Thorax, dorsal view; F. Same, lateral view; G. Thorax-abdomen transition, dorso-lateral view; H. Abdomen, dorsal view. Arrows indicate the appearing of the first branchia." figureDoi="http://doi.org/10.5281/zenodo.4414149" httpUri="https://zenodo.org/record/4414149/files/figure.png" pageId="11" pageNumber="462">Fig. 5D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF7166603B5B087BABBFF8E3FAA54594" ID-DOI="http://doi.org/10.5281/zenodo.4414149" ID-Zenodo-Dep="4414149" httpUri="https://zenodo.org/record/4414149/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,250,1904,1928]" targetBox="[151,1437,525,1879]" targetPageId="11">
<paragraph id="8BB136E83B5B087BABBFF8E3FAA54594" blockId="11.[151,1436,1904,2036]" pageId="11" pageNumber="462">
<emphasis id="B97AEAFA3B5B087BABBFF8E3FEB845E8" bold="true" box="[151,274,1904,1928]" pageId="11" pageNumber="462">FIGURE 5.</emphasis>
<taxonomicName id="4C0E4D6B3B5B087BAA30F8E3FE6745E8" authorityName=": Brown" authorityYear="1979" box="[280,461,1904,1928]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="11" pageNumber="462" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B5B087BAA30F8E3FE6745E8" box="[280,461,1904,1928]" italics="true" pageId="11" pageNumber="462">Scoloplos simplex</emphasis>
</taxonomicName>
, AD: AM W.5246 holotype, methylene blue staining; EH: AM W.6038 holotype of
<taxonomicName id="4C0E4D6B3B5B087BAE02F8E3FA3C45E8" authorityName="Day" authorityYear="1977" box="[1322,1430,1904,1928]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="11" pageNumber="462" phylum="Annelida" rank="species" species="difficilis">
<emphasis id="B97AEAFA3B5B087BAE02F8E3FA3C45E8" box="[1322,1430,1904,1928]" italics="true" pageId="11" pageNumber="462">S. difficilis</emphasis>
</taxonomicName>
, methylene blue staining; A. Thorax, dorsal view; B. Thorax-abdomen transition, dorso-lateral view; C. abdomen, dorsal view; D. Posterior end of the body with pygidium, dorso-lateral view; E. Thorax, dorsal view; F. Same, lateral view; G. Thorax-abdomen transition, dorso-lateral view; H. Abdomen, dorsal view. Arrows indicate the appearing of the first branchia.
</paragraph>
</caption>
<caption id="DF7166603B5C087CABBFF91FFC614554" ID-DOI="http://doi.org/10.5281/zenodo.4414155" ID-Zenodo-Dep="4414155" httpUri="https://zenodo.org/record/4414155/files/figure.png" pageId="12" pageNumber="463" startId="12.[151,250,1676,1700]" targetBox="[151,1436,181,1652]" targetPageId="12">
<paragraph id="8BB136E83B5C087CABBFF91FFC614554" blockId="12.[151,1437,1676,1844]" pageId="12" pageNumber="463">
<emphasis id="B97AEAFA3B5C087CABBFF91FFEBF44C4" bold="true" box="[151,277,1676,1700]" pageId="12" pageNumber="463">FIGURE 6.</emphasis>
<taxonomicName id="4C0E4D6B3B5C087CAA35F91FFE7F44C4" authorityName=": Brown" authorityYear="1979" box="[285,469,1676,1700]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="12" pageNumber="463" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B5C087CAA35F91FFE7F44C4" box="[285,469,1676,1700]" italics="true" pageId="12" pageNumber="463">Scoloplos simplex</emphasis>
</taxonomicName>
, AF: AM W.195999, stereomicroscope, methylene blue staining; GJ: AM W.198694, compound microscope. A. General lateral view; B. Thorax-abdomen transition, lateral view; C. Thorax, lateral view; D. Thoraxabdomen transition, lateral view; E. Anterior end, lateral view; F. Thorax, dorsal view; arrow indicates first branchia; G. Parapodium of chaetiger 2; H. Close up of neuropodia of chaetiger 2; I. Parapodium of chaetiger 27; J. Posterior parapodium. br, branchia; cc, crenulated capillaries; ne, neuropodium; no, notopodium; u, uncini.
</paragraph>
</caption>
<subSubSection id="C31465633B5C087CABEFF8F1FC1245A5" pageId="12" pageNumber="463" type="distribution">
<paragraph id="8BB136E83B5C087CABEFF8F1FF1145C0" blockId="12.[151,1436,1890,2025]" pageId="12" pageNumber="463">
<emphasis id="B97AEAFA3B5C087CABEFF8F1FEF3451C" bold="true" box="[199,345,1890,1916]" pageId="12" pageNumber="463">Distribution</emphasis>
.
<collectingCountry id="F31976783B5C087CAA4FF8F1FE7B451C" box="[359,465,1890,1916]" name="Australia" pageId="12" pageNumber="463">Australia</collectingCountry>
,
<collectingRegion id="49CAF80A3B5C087CAAF4F8F1FDC9451C" box="[476,611,1890,1916]" country="Australia" name="Queensland" pageId="12" pageNumber="463">Queensland</collectingRegion>
,
<collectingRegion id="49CAF80A3B5C087CA958F8F0FC96451C" box="[624,828,1890,1917]" country="Australia" name="New South Wales" pageId="12" pageNumber="463">New South Wales</collectingRegion>
,
<collectingRegion id="49CAF80A3B5C087CA860F8F1FC0F451C" box="[840,933,1890,1916]" country="Australia" name="Victoria" pageId="12" pageNumber="463">Victoria</collectingRegion>
,
<collectingRegion id="49CAF80A3B5C087CA898F8F1FBB4451C" box="[944,1054,1890,1916]" country="Australia" name="Tasmania" pageId="12" pageNumber="463">Tasmania</collectingRegion>
,
<collectingRegion id="49CAF80A3B5C087CAF02F8F1FB76451C" box="[1066,1244,1890,1916]" country="Australia" name="South Australia" pageId="12" pageNumber="463">South Australia</collectingRegion>
,
<collectingRegion id="49CAF80A3B5C087CAFCFF8F0FF1C45C0" country="Australia" name="Western Australia" pageId="12" pageNumber="463">Western Australia</collectingRegion>
.
</paragraph>
<paragraph id="8BB136E83B5C087CABEFF839FC1245A5" blockId="12.[151,1436,1890,2025]" box="[199,952,1962,1989]" pageId="12" pageNumber="463">
<emphasis id="B97AEAFA3B5C087CABEFF839FE8045A4" bold="true" box="[199,298,1962,1988]" pageId="12" pageNumber="463">Habitat.</emphasis>
Intertidal and upper subtidal, sand, muddy sand, seagrass.
</paragraph>
</subSubSection>
<subSubSection id="C31465633B5C087DABEFF85DFEDC40AD" lastPageId="13" lastPageNumber="464" pageId="12" pageNumber="463" type="discussion">
<paragraph id="8BB136E83B5C087CABEFF85DFA364588" blockId="12.[151,1436,1890,2025]" box="[199,1436,1998,2025]" pageId="12" pageNumber="463">
<emphasis id="B97AEAFA3B5C087CABEFF85DFE9E4588" bold="true" box="[199,308,1998,2024]" pageId="12" pageNumber="463">Remarks</emphasis>
.
<taxonomicName id="4C0E4D6B3B5C087CAA6DF85DFDB84588" authorityName=": Brown" authorityYear="1979" box="[325,530,1998,2024]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="12" pageNumber="463" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B5C087CAA6DF85DFDB84588" box="[325,530,1998,2024]" italics="true" pageId="12" pageNumber="463">Scoloplos simplex</emphasis>
</taxonomicName>
was originally described as a part of the genus
<taxonomicName id="4C0E4D6B3B5C087CAF6EF85DFA324588" authority="Monro, 1933" authorityName="Monro" authorityYear="1933" box="[1094,1432,1998,2025]" class="Polychaeta" family="Orbiniidae" genus="Haploscoloplos" kingdom="Animalia" pageId="12" pageNumber="463" phylum="Annelida" rank="genus">
<emphasis id="B97AEAFA3B5C087CAF6EF85DFB5C4588" box="[1094,1270,1998,2024]" italics="true" pageId="12" pageNumber="463">Haploscoloplos</emphasis>
<bibRefCitation id="EF9F4B193B5C087CAE28F85CFA324588" author="Monro, C. C. A." box="[1280,1432,1998,2025]" pageId="12" pageNumber="463" pagination="244 - 269" refId="ref34967" refString="Monro, C. C. A. (1933) On a collection of Polychaeta from Dry Tortugas, Florida. Annals and Magazine of Natural History, Series 10, 12 (69), 244 - 269. https: // doi. org / 10.1080 / 00222933308655413" type="journal article" year="1933">Monro, 1933</bibRefCitation>
</taxonomicName>
,
</paragraph>
<paragraph id="8BB136E83B5D087DABBFFF04FEDC40AD" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="464">
although the presence of uncini in the thoracic neuropodia was noted (
<bibRefCitation id="EF9F4B193B5D087DA88CFF04FBF142D1" author="Hutchings, P. A." box="[932,1115,151,177]" pageId="13" pageNumber="464" pagination="175 - 195" refId="ref34514" refString="Hutchings, P. A. (1974) Polychaeta of Wallis Lake, New South Wales. Proceedings of the Linnean Society of New South Wales, 98, 175 - 195." type="journal article" year="1974">Hutchings 1974</bibRefCitation>
).
<bibRefCitation id="EF9F4B193B5D087DAF46FF04FB4742D1" author="Day, J. H." box="[1134,1261,151,177]" pageId="13" pageNumber="464" pagination="217 - 243" refId="ref33341" refString="Day, J. H. (1977) A review of the Australian and New Zealand Orbiniidae (Annelida: Polychaeta). In: Reish, K. &amp; Fauchald, D. J., (Ed.), Essays on Polychaetous Annelids in Memory of Dr. Olga Hartman. Allan Hancock Foundation, Los Angeles, pp. 217 - 243." type="book chapter" year="1977">Day (1977)</bibRefCitation>
transferred this species to the genus
<emphasis id="B97AEAFA3B5D087DAAAEFF28FE5042B5" box="[390,506,187,213]" italics="true" pageId="13" pageNumber="464">
<taxonomicName id="4C0E4D6B3B5D087DAAAEFF28FE5C42B5" authorityName="Blainville" authorityYear="1828" box="[390,502,187,213]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="genus">Scoloplos</taxonomicName>
,
</emphasis>
and described a similar species,
<taxonomicName id="4C0E4D6B3B5D087DA850FF28FB6242B5" authority="Day, 1977" authorityName="Day" authorityYear="1977" box="[888,1224,187,213]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="difficilis">
<emphasis id="B97AEAFA3B5D087DA850FF28FBE042B5" box="[888,1098,187,213]" italics="true" pageId="13" pageNumber="464">Scoloplos difficilis</emphasis>
<bibRefCitation id="EF9F4B193B5D087DAF7BFF28FB6242B5" author="Day, J. H." box="[1107,1224,187,213]" pageId="13" pageNumber="464" pagination="217 - 243" refId="ref33341" refString="Day, J. H. (1977) A review of the Australian and New Zealand Orbiniidae (Annelida: Polychaeta). In: Reish, K. &amp; Fauchald, D. J., (Ed.), Essays on Polychaetous Annelids in Memory of Dr. Olga Hartman. Allan Hancock Foundation, Los Angeles, pp. 217 - 243." type="book chapter" year="1977">Day, 1977</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49CAF80A3B5D087DAE39FF28FA3342B5" box="[1297,1433,187,213]" country="Australia" name="Queensland" pageId="13" pageNumber="464">Queensland</collectingRegion>
. The only difference between these two species was the starting-point of the branchiae segment (
<quantity id="4CF69B0D3B5D087DAFE6FF4CFA894299" box="[1230,1315,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="13" pageNumber="464" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">911 in</quantity>
<taxonomicName id="4C0E4D6B3B5D087DAE02FF4CFA374299" authorityName=": Brown" authorityYear="1979" box="[1322,1437,223,249]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B5D087DAE02FF4CFA374299" box="[1322,1437,223,249]" italics="true" pageId="13" pageNumber="464">S. simplex</emphasis>
</taxonomicName>
vs
<quantity id="4CF69B0D3B5D087DAB90FE91FEB7437D" box="[184,285,258,285]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="3.81" metricValueMin="3.556" pageId="13" pageNumber="464" unit="in" value="14.5" valueMax="15.0" valueMin="14.0">1415 in</quantity>
<taxonomicName id="4C0E4D6B3B5D087DAA0DFE90FE34437D" authorityName="Day" authorityYear="1977" box="[293,414,259,285]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="difficilis">
<emphasis id="B97AEAFA3B5D087DAA0DFE90FE34437D" box="[293,414,259,285]" italics="true" pageId="13" pageNumber="464">S. difficilis</emphasis>
</taxonomicName>
).
<bibRefCitation id="EF9F4B193B5D087DAA9EFE90FD47437D" author="Hutchings, P. &amp; Rainer, S." box="[438,749,259,285]" pageId="13" pageNumber="464" pagination="745 - 796" refId="ref34466" refString="Hutchings, P. &amp; Rainer, S. (1979) The polychaete fauna of Careel Bay, Pittwater, New South Wales, Australia. Journal of Natural History, 13, 745 - 796. https: // doi. org / 10.1080 / 00222937900770561" type="journal article" year="1979">Hutchings &amp; Rainer (1979)</bibRefCitation>
synonymised
<taxonomicName id="4C0E4D6B3B5D087DA8B9FE90FBA0437D" authorityName="Day" authorityYear="1977" box="[913,1034,259,285]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="difficilis">
<emphasis id="B97AEAFA3B5D087DA8B9FE90FBA0437D" box="[913,1034,259,285]" italics="true" pageId="13" pageNumber="464">S. difficilis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C0E4D6B3B5D087DAF64FE90FB6A437D" authorityName=": Brown" authorityYear="1979" box="[1100,1216,259,285]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B5D087DAF64FE90FB6A437D" box="[1100,1216,259,285]" italics="true" pageId="13" pageNumber="464">S. simplex</emphasis>
</taxonomicName>
as the branchiae in the
<typeStatus id="54B5884A3B5D087DABE8FEB4FE884321" box="[192,290,295,321]" pageId="13" pageNumber="464" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0E4D6B3B5D087DAA60FEB4FE6A4321" authorityName="Day" authorityYear="1977" box="[328,448,295,321]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="difficilis">
<emphasis id="B97AEAFA3B5D087DAA60FEB4FE6A4321" box="[328,448,295,321]" italics="true" pageId="13" pageNumber="464">S. difficilis</emphasis>
</taxonomicName>
were found on chaetiger 11. In addition, they noticed the presence of flail chaetae in the abdominal parapodia in larger specimens of both species. The flail chaetae have not been found in the present study because only chaetae of small specimens were investigated.
<taxonomicName id="4C0E4D6B3B5D087DA866FEFCFBB043E9" authorityName=": Brown" authorityYear="1979" box="[846,1050,367,393]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B5D087DA866FEFCFBB043E9" box="[846,1050,367,393]" italics="true" pageId="13" pageNumber="464">Scoloplos simplex</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E4D6B3B5D087DAF7DFEFCFB6543E9" authorityName="Day" authorityYear="1977" box="[1109,1231,367,393]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="difficilis">
<emphasis id="B97AEAFA3B5D087DAF7DFEFCFB6543E9" box="[1109,1231,367,393]" italics="true" pageId="13" pageNumber="464">S. difficilis</emphasis>
</taxonomicName>
have overlapping distribution and same habitat, occurring intertidally and subtidally in sheltered estuarine area. A re-investigation of the
<typeStatus id="54B5884A3B5D087DABCAFE24FEE543B1" box="[226,335,439,465]" pageId="13" pageNumber="464" type="holotype">holotypes</typeStatus>
of both species showed that in the
<typeStatus id="54B5884A3B5D087DA9CBFE24FCEF43B1" box="[739,837,439,465]" pageId="13" pageNumber="464" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0E4D6B3B5D087DA845FE24FC4B43B1" authorityName=": Brown" authorityYear="1979" box="[877,993,439,465]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="simplex">
<emphasis id="B97AEAFA3B5D087DA845FE24FC4B43B1" box="[877,993,439,465]" italics="true" pageId="13" pageNumber="464">S. simplex</emphasis>
</taxonomicName>
the branchiae were found on chaetiger 15 (
<figureCitation id="13352A6D3B5D087DABEBFE48FE964395" box="[195,316,474,501]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[151,1437,525,1879]" captionTargetId="figure-184@11.[151,1437,525,1881]" captionTargetPageId="11" captionText="FIGURE 5. Scoloplos simplex, AD: AM W.5246 holotype, methylene blue staining; EH:AM W.6038 holotype of S. difficilis, methylene blue staining; A. Thorax, dorsal view; B. Thorax-abdomen transition, dorso-lateral view; C. abdomen, dorsal view; D. Posterior end of the body with pygidium, dorso-lateral view; E. Thorax, dorsal view; F. Same, lateral view; G. Thorax-abdomen transition, dorso-lateral view; H. Abdomen, dorsal view. Arrows indicate the appearing of the first branchia." figureDoi="http://doi.org/10.5281/zenodo.4414149" httpUri="https://zenodo.org/record/4414149/files/figure.png" pageId="13" pageNumber="464">Fig. 5A, B</figureCitation>
), whereas the
<typeStatus id="54B5884A3B5D087DAAF6FE48FDEA4395" box="[478,576,475,501]" pageId="13" pageNumber="464" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0E4D6B3B5D087DA94DFE48FD744395" authorityName="Day" authorityYear="1977" box="[613,734,475,501]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="difficilis">
<emphasis id="B97AEAFA3B5D087DA94DFE48FD744395" box="[613,734,475,501]" italics="true" pageId="13" pageNumber="464">S. difficilis</emphasis>
</taxonomicName>
was poorly preserved, so the position of the first branchia was difficult to determine, but presumably the branchiae start on chaetiger 10 (
<figureCitation id="13352A6D3B5D087DA8FDFE6CFBE34079" box="[981,1097,510,537]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[151,1437,525,1879]" captionTargetId="figure-184@11.[151,1437,525,1881]" captionTargetPageId="11" captionText="FIGURE 5. Scoloplos simplex, AD: AM W.5246 holotype, methylene blue staining; EH:AM W.6038 holotype of S. difficilis, methylene blue staining; A. Thorax, dorsal view; B. Thorax-abdomen transition, dorso-lateral view; C. abdomen, dorsal view; D. Posterior end of the body with pygidium, dorso-lateral view; E. Thorax, dorsal view; F. Same, lateral view; G. Thorax-abdomen transition, dorso-lateral view; H. Abdomen, dorsal view. Arrows indicate the appearing of the first branchia." figureDoi="http://doi.org/10.5281/zenodo.4414149" httpUri="https://zenodo.org/record/4414149/files/figure.png" pageId="13" pageNumber="464">Fig. 5E, F</figureCitation>
). Examination of the
<typeStatus id="54B5884A3B5D087DAE12FE6CFA364079" box="[1338,1436,511,537]" pageId="13" pageNumber="464" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0E4D6B3B5D087DAB9FFDB0FE9A405D" authorityName="Day" authorityYear="1977" box="[183,304,547,573]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="difficilis">
<emphasis id="B97AEAFA3B5D087DAB9FFDB0FE9A405D" box="[183,304,547,573]" italics="true" pageId="13" pageNumber="464">S. difficilis</emphasis>
</taxonomicName>
reveals that the branchiae occur earlier than in
<emphasis id="B97AEAFA3B5D087DA867FDB0FC62405D" box="[847,968,547,573]" italics="true" pageId="13" pageNumber="464">
<taxonomicName id="4C0E4D6B3B5D087DA867FDB0FC6E405D" authorityName=": Brown" authorityYear="1979" box="[847,964,547,573]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="13" pageNumber="464" phylum="Annelida" rank="species" species="simplex">S. simplex</taxonomicName>
,
</emphasis>
which is the reverse of what
<bibRefCitation id="EF9F4B193B5D087DAE33FDB0FA36405D" author="Day, J. H." box="[1307,1436,547,573]" pageId="13" pageNumber="464" pagination="217 - 243" refId="ref33341" refString="Day, J. H. (1977) A review of the Australian and New Zealand Orbiniidae (Annelida: Polychaeta). In: Reish, K. &amp; Fauchald, D. J., (Ed.), Essays on Polychaetous Annelids in Memory of Dr. Olga Hartman. Allan Hancock Foundation, Los Angeles, pp. 217 - 243." type="book chapter" year="1977">Day (1977)</bibRefCitation>
states. The present study confirmed the synonymy of these two species as the starting segment of branchiae varies from chaetiger 9 to 16 even within one sample. Therefore, it cannot be used as evidence for species recognition. Moreover, the few first pairs of branchia are so small that they can only be seen in well-preserved species and with the help of staining.
</paragraph>
</subSubSection>
</treatment>
</document>