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<document id="04D8E3ADC6E6BBAB474187123FA8E5D0" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629656228228" checkinUser="felipe" docAuthor="Fancello, Luca, Hernando, Carles &amp; Leo, Piero" docDate="2009" docId="03A387F2FFC9FFC2D2E7FD4BFF325803" docLanguage="en" docName="zt02318p338.pdf" docOrigin="Zootaxa 2318" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Entomoculia (Entomoculia) carbonaria Fancello &amp; Hernando &amp; Leo 2009, sp. nov." docType="treatment" docVersion="2" lastPageNumber="322" masterDocId="FF9AFF8AFFCCFFC7D270FFC5FFC75F79" masterDocTitle="The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *" masterLastPageNumber="338" masterPageNumber="317" pageNumber="322" updateTime="1699090143914" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="07F171F2B7603642F61DA57F3109203E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1E51D14FBF1FEBB321971CEDF75B69FE">
<mods:title id="472A83288599FFE1B868052C7CC4D839">The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *</mods:title>
</mods:titleInfo>
<mods:name id="6F3B2ABC764888BF3A9F72E1C66FC0DA" type="personal">
<mods:role id="B2104B3AEE7B92F62B50C60142AF1F03">
<mods:roleTerm id="64C1E64A42C370BDED41E8FE0D0A2313">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A80D45F857D12AA09B0439AE8793311F">Fancello, Luca</mods:namePart>
<mods:affiliation id="6EC1DB576BE252594E2A17665E5B3D16">Via Bainsizza 12, I- 09123 Cagliari, Italy. E-mail: L. fancello @ hotmail. it Museu de Ciènces Naturals de la Ciutadella (Zoologia), Passeig Picasso s / n, E- 08003 Barcelona, Spain. E-mail: c _ hernando @ telefonica. net Via Tola 21, I- 09128 Cagliari, Italy. E-mail: piero. leo @ tiscali. it</mods:affiliation>
</mods:name>
<mods:name id="BCC7415A7715A80754705BD926F2680B" type="personal">
<mods:role id="626221963A3BFA4A051856AB5325FE48">
<mods:roleTerm id="9E62EF08AD0B6A2C43D83DD11B857A25">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E1552908FDC7C3D94209671C5B64FBD2">Hernando, Carles</mods:namePart>
</mods:name>
<mods:name id="63051EFEBDB1ED76B9F209A71544BA8D" type="personal">
<mods:role id="5ACDFB9226E585B8B373B7EA7BFF4FE6">
<mods:roleTerm id="72883459A180AC00DC643DAFEA076215">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B2D5942CF1B9412A25E0F3C25961E8D">Leo, Piero</mods:namePart>
</mods:name>
<mods:typeOfResource id="815BA5486D04F7B786CA015BADB72A31">text</mods:typeOfResource>
<mods:relatedItem id="1F67A0284EC1B0E5997C941E915E28FB" type="host">
<mods:titleInfo id="51A5971E8BC6C72EA11DB4847F1BC2E7">
<mods:title id="2112B0E6BC6E039E3C337B5D51111865">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2F8BE78ECB8DAE4130991A135631ECC8">
<mods:date id="903140D1734670FB329AD83CFEF76955">2009</mods:date>
<mods:detail id="5DB73402F4EC7B74A816BF0C611C3FEF" type="pubDate">
<mods:number id="2FAE4AC5FD15EC2F4F6652183AFD208F">2009-12-22</mods:number>
</mods:detail>
<mods:detail id="E70DB01FEE2F331E3CD27F01BCABE8FC" type="volume">
<mods:number id="498E8F95FD3F9A4854D9BFEA61E6B26D">2318</mods:number>
</mods:detail>
<mods:extent id="8A97DC68A591614E44AE46E69571E518" unit="page">
<mods:start id="2FD25721F986E0E0FD8FEAACAB89C5C7">317</mods:start>
<mods:end id="B6A06E043FCCFB7FE2E6774AFE6B85FF">338</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FF10E3111DA2B90B2E57F7A59D2ABB5B">journal article</mods:classification>
<mods:identifier id="C808F8AE47FD4B06883486F50A49470C" type="ISSN">1175-5326</mods:identifier>
</mods:mods>
<treatment id="03A387F2FFC9FFC2D2E7FD4BFF325803" LSID="urn:lsid:plazi:treatment:03A387F2FFC9FFC2D2E7FD4BFF325803" httpUri="http://treatment.plazi.org/id/03A387F2FFC9FFC2D2E7FD4BFF325803" lastPageNumber="322" pageId="5" pageNumber="322">
<subSubSection id="C310656FFFC9FFC2D2E7FD4BFD1B5DD1" box="[151,732,654,680]" pageId="5" pageNumber="322" type="nomenclature">
<paragraph id="8BB536E4FFC9FFC2D2E7FD4BFD1B5DD1" blockId="5.[151,732,654,715]" box="[151,732,654,680]" pageId="5" pageNumber="322">
<heading id="D0FD8188FFC9FFC2D2E7FD4BFD1B5DD1" bold="true" box="[151,732,654,680]" fontSize="11" level="1" pageId="5" pageNumber="322" reason="1">
<emphasis id="B97EEAF6FFC9FFC2D2E7FD4BFD1B5DD1" bold="true" box="[151,732,654,680]" pageId="5" pageNumber="322">
<taxonomicName id="4C0A4D67FFC9FFC2D2E7FD4BFDB05DD1" ID-CoL="6FGP3" authority="Fancello &amp; Hernando &amp; Leo, 2009" authorityName="Fancello &amp; Hernando &amp; Leo" authorityYear="2009" box="[151,631,654,680]" class="Insecta" family="Staphylinidae" genus="Entomoculia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="carbonaria" status="sp. nov." subGenus="Entomoculia">
<emphasis id="B97EEAF6FFC9FFC2D2E7FD4BFEF45DD1" bold="true" box="[151,307,654,680]" italics="true" pageId="5" pageNumber="322">Entomoculia</emphasis>
(
<emphasis id="B97EEAF6FFC9FFC2D334FD4BFE275DD1" bold="true" box="[324,480,654,680]" italics="true" pageId="5" pageNumber="322">Entomoculia</emphasis>
)
<emphasis id="B97EEAF6FFC9FFC2D382FD4BFDB05DD1" bold="true" box="[498,631,654,680]" italics="true" pageId="5" pageNumber="322">carbonaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D578DFFC9FFC2D00EFD4BFD1B5DD1" box="[638,732,654,680]" pageId="5" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFC9FFC2D2E7FD74FED95DB2" box="[151,286,689,715]" pageId="5" pageNumber="322" type="description">
<paragraph id="8BB536E4FFC9FFC2D2E7FD74FED95DB2" blockId="5.[151,732,654,715]" box="[151,286,689,715]" pageId="5" pageNumber="322">
(
<figureCitation id="13312A61FFC9FFC2D2EFFD74FF2C5DB2" box="[159,235,689,715]" captionStart="FIGURES 7–10" captionStartId="3.[151,273,1925,1949]" captionTargetBox="[188,1388,1080,1907]" captionTargetId="figure-247@3.[186,1391,1074,1912]" captionTargetPageId="3" captionText="FIGURES 7–10. Male copulatory organ of Entomoculia spp. in lateral view, scale bars: 0.05 mm. 7. Entomoculia villascemae sp. nov. (holotype). 8. Entomoculia shardana sp. nov. (paratype). 9. Entomoculia carbonaria sp. nov. (paratype, Gutturu di Monte Nieddu). 10. Entomoculia melonii sp. nov., paratype." pageId="5" pageNumber="322">Figs 9</figureCitation>
,
<figureCitation id="13312A61FFC9FFC2D288FD74FED15DB2" box="[248,278,689,715]" captionStart="FIGURES 11–14" captionStartId="4.[151,269,1709,1733]" captionTargetBox="[283,1301,431,1683]" captionTargetId="figure-207@4.[276,1311,426,1686]" captionTargetPageId="4" captionText="FIGURES 11–14. Female genital armatures of Entomoculia spp. in ventral view, scale bars: 0.05 mm. 11. Entomoculia melonii sp. nov. (paratype). 12. Entomoculia villascemae sp. nov. (paratype, Foresta Demaniale di Montimannu). 13. Entomoculia carbonaria sp. nov. (paratype, Gutturu di Monte Nieddu). 14. Entomoculia shardana sp. nov. (paratype)." pageId="5" pageNumber="322">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFC9FFC2D2E7FD3BFB0A5C1F" pageId="5" pageNumber="322" type="diagnosis">
<paragraph id="8BB536E4FFC9FFC2D2E7FD3BFB0A5C1F" blockId="5.[151,1437,766,1914]" pageId="5" pageNumber="322">
<emphasis id="B97EEAF6FFC9FFC2D2E7FD3BFED05C61" bold="true" box="[151,279,766,792]" pageId="5" pageNumber="322">Diagnosis.</emphasis>
An
<taxonomicName id="4C0A4D67FFC9FFC2D339FD3AFE185C61" authorityName="Croissandeau" authorityYear="1891" box="[329,479,767,792]" class="Insecta" family="Staphylinidae" genus="Entomoculia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAF6FFC9FFC2D339FD3AFE185C61" box="[329,479,767,792]" italics="true" pageId="5" pageNumber="322">Entomoculia</emphasis>
</taxonomicName>
ascribable to the nominal subgenus for its robust build, the presence of chitinized thickenings on the male genital segment and for the characters of the male and female genitalia; welldifferentiated from its congeners by the shape of the aedeagus and female copulatory plates.
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFC9FFC2D2B6FCB7FF0D5B91" pageId="5" pageNumber="322" type="materials_examined">
<paragraph id="8BB536E4FFC9FFC2D2B6FCB7FAD75CF5" blockId="5.[151,1437,766,1914]" box="[198,1296,882,908]" pageId="5" pageNumber="322">
<materialsCitation id="3B623CB9FFC9FFC2D2B6FCB7FACB5CF5" box="[198,1292,882,908]" county="Carbonia-Iglesias prov." location="Gutturu di Monte Nieddu" municipality="Domusnovas" pageId="5" pageNumber="322" specimenCount="1" stateProvince="Sardegna" typeStatus="holotype">
<emphasis id="B97EEAF6FFC9FFC2D2B6FCB7FEAB5CF5" bold="true" box="[198,364,882,908]" pageId="5" pageNumber="322">
<typeStatus id="54B18846FFC9FFC2D2B6FCB7FEC35CF5" box="[198,260,882,908]" pageId="5" pageNumber="322">Type</typeStatus>
locality.
</emphasis>
SW
<collectingRegion id="49CEF806FFC9FFC2D3D6FCB7FDCD5CF5" box="[422,522,882,908]" country="Italy" name="Sardegna" pageId="5" pageNumber="322">Sardinia</collectingRegion>
,
<collectingCounty id="62D44E68FFC9FFC2D066FCB7FCED5CF5" box="[534,810,882,908]" pageId="5" pageNumber="322">Carbonia-Iglesias prov.</collectingCounty>
,
<collectingMunicipality id="6BD1AC9EFFC9FFC2D146FCB7FC155CF5" box="[822,978,882,908]" pageId="5" pageNumber="322">Domusnovas</collectingMunicipality>
,
<location id="8ED5603FFFC9FFC2D1AEFCB7FACB5CF5" LSID="urn:lsid:plazi:treatment:03A387F2FFC9FFC2D2E7FD4BFF325803:8ED5603FFFC9FFC2D1AEFCB7FACB5CF5" box="[990,1292,882,908]" county="Carbonia-Iglesias prov." municipality="Domusnovas" name="Gutturu di Monte Nieddu" pageId="5" pageNumber="322" stateProvince="Sardegna">Gutturu di Monte Nieddu</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB536E4FFC9FFC2D2B6FC5DFF0D5B91" blockId="5.[151,1437,766,1914]" pageId="5" pageNumber="322">
<emphasis id="B97EEAF6FFC9FFC2D2B6FC5DFE9F5CCB" bold="true" box="[198,344,920,946]" pageId="5" pageNumber="322">Type series.</emphasis>
<materialsCitation id="3B623CB9FFC9FFC2D32FFC5CFCD35CA3" collectingDate="1985-10-25" collectionCode="MSNG" collectorName="L. Fancello &amp; P. Leo" county="Domusnovas" elevation="290" location="Gutturu di Monte Nieddu" municipality="Carbonia-Iglesias" pageId="5" pageNumber="322" specimenCount="1" specimenCount-male="1" stateProvince="Sardegna" typeStatus="holotype">
<typeStatus id="54B18846FFC9FFC2D32FFC5CFE0B5CCA" box="[351,460,921,947]" pageId="5" pageNumber="322" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0CFD6DFFC9FFC2D3A3FC5FFE205CC8" box="[467,487,922,945]" pageId="5" pageNumber="322" type="male">♂</specimenCount>
: “SW
<collectingRegion id="49CEF806FFC9FFC2D047FC5CFD5C5CCA" box="[567,667,921,947]" country="Italy" name="Sardegna" pageId="5" pageNumber="322">Sardinia</collectingRegion>
,
<collectingCounty id="62D44E68FFC9FFC2D0D6FC5CFCF85CCA" box="[678,831,921,947]" pageId="5" pageNumber="322">Domusnovas</collectingCounty>
(
<collectingMunicipality id="6BD1AC9EFFC9FFC2D13FFC5CFBE35CCA" box="[847,1060,921,947]" pageId="5" pageNumber="322">Carbonia-Iglesias</collectingMunicipality>
),
<location id="8ED5603FFFC9FFC2D647FC5CFAA35CCA" LSID="urn:lsid:plazi:treatment:03A387F2FFC9FFC2D2E7FD4BFF325803:8ED5603FFFC9FFC2D647FC5CFAA35CCA" box="[1079,1380,921,947]" county="Domusnovas" municipality="Carbonia-Iglesias" name="Gutturu di Monte Nieddu" pageId="5" pageNumber="322" stateProvince="Sardegna">Gutturu di Monte Nieddu</location>
,
<quantity id="4CF29B01FFC9FFC2D700FC5CFF765CA3" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="322" unit="m" value="290.0">
<elevation id="0027D1D7FFC9FFC2D700FC5CFF765CA3" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="322" unit="m" value="290.0">290 m</elevation>
</quantity>
,
<date id="FFB41024FFC9FFC2D2CFFC05FE825CA3" box="[191,325,960,986]" pageId="5" pageNumber="322" value="1985-10-25">
<collectingDate id="EFF0E9CCFFC9FFC2D2CFFC05FE825CA3" box="[191,325,960,986]" pageId="5" pageNumber="322" value="1985-10-25">25.X.1985</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFC9FFC2D3FBFC05FDDE5CA3" box="[395,537,960,986]" pageId="5" pageNumber="322">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFC9FFC2D033FC05FD535CA3" box="[579,660,960,986]" pageId="5" pageNumber="322">P. Leo</collectorName>
” (
<collectionCode id="ED1BAE21FFC9FFC2D0C1FC05FCCB5CA3" box="[689,780,960,986]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="5" pageNumber="322">MSNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B623CB9FFC9FFC2D156FC05FD215B0D" collectingDate="1985-05-22" collectingDateMax="1985-10-25" collectingDateMin="1985-05-22" collectionCode="CLF-PL, CCH, CNBFVR, MSNG" collectorName="L. Fancello &amp; P. Leo" county="Gutturu di Monte Nieddu" elevation="290" location="Domusnovas" municipality="Carbonia-Iglesias prov." pageId="5" pageNumber="322" specimenCount="37" specimenCount-female="15" specimenCount-male="22" stateProvince="Sardegna" typeStatus="paratype">
<typeStatus id="54B18846FFC9FFC2D156FC05FC645CA3" box="[806,931,960,986]" pageId="5" pageNumber="322" type="paratype">Paratypes</typeStatus>
:
<location id="8ED5603FFFC9FFC2D1C1FC05FB965CA3" LSID="urn:lsid:plazi:treatment:03A387F2FFC9FFC2D2E7FD4BFF325803:8ED5603FFFC9FFC2D1C1FC05FB965CA3" box="[945,1105,960,986]" county="Gutturu di Monte Nieddu" municipality="Carbonia-Iglesias prov." name="Domusnovas" pageId="5" pageNumber="322" stateProvince="Sardegna">Domusnovas</location>
(
<collectingMunicipality id="6BD1AC9EFFC9FFC2D614FC05FA495CA3" box="[1124,1422,960,986]" pageId="5" pageNumber="322">Carbonia-Iglesias prov.</collectingMunicipality>
),
<collectingCounty id="62D44E68FFC9FFC2D2E7FC23FE255B79" box="[151,482,998,1024]" pageId="5" pageNumber="322">Gutturu di Monte Nieddu</collectingCounty>
,
<quantity id="4CF29B01FFC9FFC2D382FC23FD825B79" box="[498,581,998,1024]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="322" unit="m" value="290.0">
<elevation id="0027D1D7FFC9FFC2D382FC23FD825B79" box="[498,581,998,1024]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="322" unit="m" value="290.0">290 m</elevation>
</quantity>
,
<date id="FFB41024FFC9FFC2D025FC23FD1E5B79" box="[597,729,998,1024]" pageId="5" pageNumber="322" value="1985-05-22">
<collectingDate id="EFF0E9CCFFC9FFC2D025FC23FD1E5B79" box="[597,729,998,1024]" pageId="5" pageNumber="322" value="1985-05-22">22.V.1985</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFC9FFC2D154FC23FC715B79" box="[804,950,998,1024]" pageId="5" pageNumber="322">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFC9FFC2D192FC23FBF15B79" box="[994,1078,998,1024]" pageId="5" pageNumber="322">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFC9FFC2D635FC23FB4F5C86" box="[1093,1160,998,1024]" pageId="5" pageNumber="322" type="male">4 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFC9FFC2D6B8FC23FB305C86" box="[1224,1271,998,1024]" pageId="5" pageNumber="322" type="female">3 ♀</specimenCount>
(
<collectionCode id="ED1BAE21FFC9FFC2D76EFC23FA485B79" box="[1310,1423,998,1024]" pageId="5" pageNumber="322">CLF-PL</collectionCode>
);
<date id="FFB41024FFC9FFC2D2E7FBC8FEE75B5E" box="[151,288,1037,1063]" pageId="5" pageNumber="322" value="1985-09-18">
<collectingDate id="EFF0E9CCFFC9FFC2D2E7FBC8FEE75B5E" box="[151,288,1037,1063]" pageId="5" pageNumber="322" value="1985-09-18">18.IX.1985</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFC9FFC2D312FBC8FE2E5B5E" box="[354,489,1037,1063]" pageId="5" pageNumber="322">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFC9FFC2D061FBC8FD985B5E" box="[529,607,1037,1063]" pageId="5" pageNumber="322">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFC9FFC2D01BFBC8FD6C5B5F" box="[619,683,1037,1063]" pageId="5" pageNumber="322" type="male">4 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFC9FFC2D096FBC8FCD55B5F" box="[742,786,1037,1063]" pageId="5" pageNumber="322" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED1BAE21FFC9FFC2D152FBC8FC4B5B5E" box="[802,908,1037,1063]" pageId="5" pageNumber="322">CLF-PL</collectionCode>
);
<date id="FFB41024FFC9FFC2D1D1FBC8FBD65B5E" box="[929,1041,1037,1063]" pageId="5" pageNumber="322" value="1985-10-01">
<collectingDate id="EFF0E9CCFFC9FFC2D1D1FBC8FBD65B5E" box="[929,1041,1037,1063]" pageId="5" pageNumber="322" value="1985-10-01">1.X.1985</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFC9FFC2D623FBC8FB1D5B5E" box="[1107,1242,1037,1063]" pageId="5" pageNumber="322">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFC9FFC2D772FBC8FA975B5E" box="[1282,1360,1037,1063]" pageId="5" pageNumber="322">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFC9FFC2D72DFBC8FA5A5B5F" box="[1373,1437,1037,1063]" pageId="5" pageNumber="322" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFC9FFC2D2B9FBF1FF345B35" box="[201,243,1076,1102]" pageId="5" pageNumber="322" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED1BAE21FFC9FFC2D372FBF1FEAD5B37" box="[258,362,1076,1102]" pageId="5" pageNumber="322">CLF-PL</collectionCode>
);
<date id="FFB41024FFC9FFC2D30FFBF1FE3A5B37" box="[383,509,1076,1102]" pageId="5" pageNumber="322" value="1985-10-25">
<collectingDate id="EFF0E9CCFFC9FFC2D30FFBF1FE3A5B37" box="[383,509,1076,1102]" pageId="5" pageNumber="322" value="1985-10-25">25.X.1985</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFC9FFC2D04CFBF1FD065B37" box="[572,705,1076,1102]" pageId="5" pageNumber="322">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFC9FFC2D097FBF1FCF45B37" box="[743,819,1076,1102]" pageId="5" pageNumber="322">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFC9FFC2D130FBF1FCB95B35" box="[832,894,1076,1102]" pageId="5" pageNumber="322" type="male">6 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFC9FFC2D1C7FBF1FC265B35" box="[951,993,1076,1102]" pageId="5" pageNumber="322" type="female">5 ♀</specimenCount>
(
<specimenCount id="9D0CFD6DFFC9FFC2D678FBF1FBF65B35" box="[1032,1073,1076,1102]" pageId="5" pageNumber="322" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFC9FFC2D619FBF1FB535B35" box="[1129,1172,1076,1102]" pageId="5" pageNumber="322" type="female">1 ♀</specimenCount>
<collectionCode id="ED1BAE21FFC9FFC2D6EBFBF1FB1D5B37" box="[1179,1242,1076,1102]" pageId="5" pageNumber="322">CCH</collectionCode>
,
<specimenCount id="9D0CFD6DFFC9FFC2D696FBF1FAE25B35" box="[1254,1317,1076,1102]" pageId="5" pageNumber="322" type="male">4 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFC9FFC2D72EFBF1FA4F5B35" box="[1374,1416,1076,1102]" pageId="5" pageNumber="322" type="female">2 ♀</specimenCount>
CLF-PL,
<specimenCount id="9D0CFD6DFFC9FFC2D362FB9FFE865B0A" box="[274,321,1114,1140]" pageId="5" pageNumber="322" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFC9FFC2D3F1FB9FFE685B0A" box="[385,431,1114,1140]" pageId="5" pageNumber="322" type="female">1 ♀</specimenCount>
<collectionCode id="ED1BAE21FFC9FFC2D3C8FB9FFDF05B0D" box="[440,567,1114,1140]" pageId="5" pageNumber="322">CNBFVR</collectionCode>
,
<specimenCount id="9D0CFD6DFFC9FFC2D036FB9FFDB15B0A" box="[582,630,1114,1140]" pageId="5" pageNumber="322" type="female">1 ♀</specimenCount>
<collectionCode id="ED1BAE21FFC9FFC2D00EFB9FFD1A5B0D" box="[638,733,1114,1140]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="5" pageNumber="322">MSNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B623CB9FFC9FFC2D085FB9FFA8C5BB9" collectingDate="1986-01-10" collectingDateMax="1989-12-18" collectingDateMin="1986-01-10" collectionCode="CLF-PL" collectorName="L. Fancello &amp; P. Leo" county="San Giovanni" elevation="200" location="Domusnovas" municipality="Grotta" pageId="5" pageNumber="322" specimenCount="8" specimenCount-female="5" specimenCount-male="3" stateProvince="Sardegna" typeStatus="paratype">
<date id="FFB41024FFC9FFC2D085FB9FFCB55B0D" box="[757,882,1114,1140]" pageId="5" pageNumber="322" value="1986-01-10">
<collectingDate id="EFF0E9CCFFC9FFC2D085FB9FFCB55B0D" box="[757,882,1114,1140]" pageId="5" pageNumber="322" value="1986-01-10">10.I.1986</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFC9FFC2D1CCFB9FFB895B0D" box="[956,1102,1114,1140]" pageId="5" pageNumber="322">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFC9FFC2D60AFB9FFB0A5B0D" box="[1146,1229,1114,1140]" pageId="5" pageNumber="322">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFC9FFC2D6ADFB9FFACB5B0A" box="[1245,1292,1114,1140]" pageId="5" pageNumber="322" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED1BAE21FFC9FFC2D76EFB9FFA485B0D" box="[1310,1423,1114,1140]" pageId="5" pageNumber="322">CLF-PL</collectionCode>
);
<date id="FFB41024FFC9FFC2D2E7FB44FEEB5BE2" box="[151,300,1153,1179]" pageId="5" pageNumber="322" value="1989-12-18">
<collectingDate id="EFF0E9CCFFC9FFC2D2E7FB44FEEB5BE2" box="[151,300,1153,1179]" pageId="5" pageNumber="322" value="1989-12-18">18.XII.1989</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFC9FFC2D31EFB44FE325BE2" box="[366,501,1153,1179]" pageId="5" pageNumber="322">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFC9FFC2D06EFB44FDAA5BE2" box="[542,621,1153,1179]" pageId="5" pageNumber="322">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFC9FFC2D00AFB44FD7D5BE0" box="[634,698,1153,1179]" pageId="5" pageNumber="322" type="male">3 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFC9FFC2D085FB44FCE65BE0" box="[757,801,1153,1179]" pageId="5" pageNumber="322" type="female">3 ♀</specimenCount>
(
<collectionCode id="ED1BAE21FFC9FFC2D136FB44FC775BE2" box="[838,944,1153,1179]" pageId="5" pageNumber="322">CLF-PL</collectionCode>
).
<location id="8ED5603FFFC9FFC2D1B6FB44FBA65BE2" LSID="urn:lsid:plazi:treatment:03A387F2FFC9FFC2D2E7FD4BFF325803:8ED5603FFFC9FFC2D1B6FB44FBA65BE2" box="[966,1121,1153,1179]" county="San Giovanni" municipality="Grotta" name="Domusnovas" pageId="5" pageNumber="322" stateProvince="Sardegna">Domusnovas</location>
(
<location id="8ED5603FFFC9FFC2D602FB44FA575BE2" LSID="urn:lsid:plazi:treatment:03A387F2FFC9FFC2D2E7FD4BFF325803:8ED5603FFFC9FFC2D602FB44FA575BE2" box="[1138,1424,1153,1179]" county="San Giovanni" municipality="Grotta" name="Carbonia-Iglesias prov." pageId="5" pageNumber="322" stateProvince="Sardegna">Carbonia-Iglesias prov.</location>
),
<collectingMunicipality id="6BD1AC9EFFC9FFC2D2E7FB6DFF255BBB" box="[151,226,1192,1218]" pageId="5" pageNumber="322">Grotta</collectingMunicipality>
[= cave]
<collectingCounty id="62D44E68FFC9FFC2D321FB6DFE335BBB" box="[337,500,1192,1218]" pageId="5" pageNumber="322">San Giovanni</collectingCounty>
, ingresso nord [= northern entrance],
<quantity id="4CF29B01FFC9FFC2D1DCFB6DFC3F5BBB" box="[940,1016,1192,1218]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="322" unit="m" value="200.0">
<elevation id="0027D1D7FFC9FFC2D1DCFB6DFC3F5BBB" box="[940,1016,1192,1218]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="322" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="FFB41024FFC9FFC2D675FB6DFB4C5BBB" box="[1029,1163,1192,1218]" pageId="5" pageNumber="322" value="1987-03-28">
<collectingDate id="EFF0E9CCFFC9FFC2D675FB6DFB4C5BBB" box="[1029,1163,1192,1218]" pageId="5" pageNumber="322" value="1987-03-28">28.III.1987</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFC9FFC2D6BAFB6DFAD05BBB" box="[1226,1303,1192,1218]" pageId="5" pageNumber="322">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFC9FFC2D752FB6DFA8C5BB9" box="[1314,1355,1192,1218]" pageId="5" pageNumber="322" type="female">1 ♀</specimenCount>
</materialsCitation>
(CLF- PL).
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFC9FFC2D2B6FB30FB255855" pageId="5" pageNumber="322" type="description">
<paragraph id="8BB536E4FFC9FFC2D2B6FB30FB145AA9" blockId="5.[151,1437,766,1914]" pageId="5" pageNumber="322">
<emphasis id="B97EEAF6FFC9FFC2D2B6FB30FE995A76" bold="true" box="[198,350,1269,1295]" pageId="5" pageNumber="322">Description.</emphasis>
Total length 1.00–1.10 mm. Head subparallel-sided, with prominant frontal carinas slightly diverging posteriorly; microreticulation obvious and punctuation strong and sparse. Pronotum about as long as wide, shiny, with faint microreticulation and superficial and sparse punctuation, much more sparse than on the head; median longitudinal dimples lacking. Elytra shiny with imperceptible microsculpture and scattered punctuation. Aedeagus as in
<figureCitation id="13312A61FFC9FFC2D39AFA55FDF75AD3" box="[490,560,1424,1450]" captionStart="FIGURES 7–10" captionStartId="3.[151,273,1925,1949]" captionTargetBox="[188,1388,1080,1907]" captionTargetId="figure-247@3.[186,1391,1074,1912]" captionTargetPageId="3" captionText="FIGURES 7–10. Male copulatory organ of Entomoculia spp. in lateral view, scale bars: 0.05 mm. 7. Entomoculia villascemae sp. nov. (holotype). 8. Entomoculia shardana sp. nov. (paratype). 9. Entomoculia carbonaria sp. nov. (paratype, Gutturu di Monte Nieddu). 10. Entomoculia melonii sp. nov., paratype." pageId="5" pageNumber="322">Fig. 9</figureCitation>
, with sternal lamina, in lateral view, long and robust, tapering at apex; left paramere long, thin and flexuous. Female genital armature complex, structured as in
<figureCitation id="13312A61FFC9FFC2D609FA73FB085AA9" box="[1145,1231,1462,1488]" captionStart="FIGURES 11–14" captionStartId="4.[151,269,1709,1733]" captionTargetBox="[283,1301,431,1683]" captionTargetId="figure-207@4.[276,1311,426,1686]" captionTargetPageId="4" captionText="FIGURES 11–14. Female genital armatures of Entomoculia spp. in ventral view, scale bars: 0.05 mm. 11. Entomoculia melonii sp. nov. (paratype). 12. Entomoculia villascemae sp. nov. (paratype, Foresta Demaniale di Montimannu). 13. Entomoculia carbonaria sp. nov. (paratype, Gutturu di Monte Nieddu). 14. Entomoculia shardana sp. nov. (paratype)." pageId="5" pageNumber="322">Fig. 13</figureCitation>
.
</paragraph>
<paragraph id="8BB536E4FFC9FFC2D2B6FA18FF335912" blockId="5.[151,1437,766,1914]" pageId="5" pageNumber="322">
<emphasis id="B97EEAF6FFC9FFC2D2B6FA18FEB45A8E" bold="true" box="[198,371,1501,1527]" pageId="5" pageNumber="322">Observations.</emphasis>
The new species is well-differentiated from the already-known ones and from those described in the present paper; some similarity in the structure of the female genital plates can be found in
<taxonomicName id="4C0A4D67FFC9FFC2D7F3F9C1FEE2593D" authorityName="Pace" authorityYear="1978" class="Insecta" family="Staphylinidae" genus="Entomoculia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="sassariensis">
<emphasis id="B97EEAF6FFC9FFC2D7F3F9C1FA525964" box="[1411,1429,1540,1565]" italics="true" pageId="5" pageNumber="322">E</emphasis>
.
<emphasis id="B97EEAF6FFC9FFC2D2E7F9EEFEE2593D" box="[151,293,1579,1604]" italics="true" pageId="5" pageNumber="322">sassariensis</emphasis>
</taxonomicName>
from northern
<collectingRegion id="49CEF806FFC9FFC2D3A6F9EFFDFD593D" box="[470,570,1578,1604]" country="Italy" name="Sardegna" pageId="5" pageNumber="322">Sardinia</collectingRegion>
, but they are easily separable by the shape of the aedeagus (
<emphasis id="B97EEAF6FFC9FFC2D688F9EEFACA593D" box="[1272,1293,1579,1604]" italics="true" pageId="5" pageNumber="322">cf</emphasis>
.
<bibRefCitation id="EF9B4B15FFC9FFC2D76CF9EFFA50593D" author="Pace R. &amp; Fauna d'Italia, XXXIV" box="[1308,1431,1578,1604]" pageId="5" pageNumber="322" refId="ref13459" refString="Pace R., (1996). Fauna d'Italia, XXXIV. Coleoptera Staphylinidae Leptotyphlinae. Calderini, Bologna, 328 pp." type="book" year="1996">Pace 1996</bibRefCitation>
, fig. 87).
</paragraph>
<paragraph id="8BB536E4FFC9FFC2D2B6F9B2FB255855" blockId="5.[151,1437,766,1914]" pageId="5" pageNumber="322">
<emphasis id="B97EEAF6FFC9FFC2D2B6F9B2FE5D59E8" bold="true" box="[198,410,1655,1681]" pageId="5" pageNumber="322">Ecological notes.</emphasis>
This species was collected in woodland in soil samples taken at the base of an old
<taxonomicName id="4C0A4D67FFC9FFC2D2E7F95AFEE959C1" box="[151,302,1695,1720]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="322" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis id="B97EEAF6FFC9FFC2D2E7F95AFEE959C1" box="[151,302,1695,1720]" italics="true" pageId="5" pageNumber="322">Quercus ilex</emphasis>
</taxonomicName>
and under stone, near a natural cave. Sampling localities are situated on Palaeozoic limestone soil. The following endogean beetles were collected in association with the new species:
<taxonomicName id="4C0A4D67FFC9FFC2D771F900FA5059A6" box="[1281,1431,1733,1759]" class="Insecta" family="Staphylinidae" genus="Mayetia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97EEAF6FFC9FFC2D771F900FAA159A7" box="[1281,1382,1733,1758]" italics="true" pageId="5" pageNumber="322">Mayetia</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="4C0A4D67FFC9FFC2D2E7F929FE70587C" authority="Fancello &amp; Hernando &amp; Leo, 2009" authorityName="Fancello &amp; Hernando &amp; Leo" authorityYear="2009" box="[151,439,1772,1797]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="minator" status="sp. nov.">
<emphasis id="B97EEAF6FFC9FFC2D2E7F929FE70587C" box="[151,439,1772,1797]" italics="true" pageId="5" pageNumber="322">Leptotyphlus minator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D578DFFC9FFC2D3B3F929FDE8587F" box="[451,559,1772,1798]" pageId="5" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C0A4D67FFC9FFC2D030F929FB11587F" authority="Saulcy, 1878" authorityName="Saulcy" authorityYear="1878" box="[576,1238,1772,1798]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="raymondi">
<emphasis id="B97EEAF6FFC9FFC2D030F929FCF8587C" box="[576,831,1772,1797]" italics="true" pageId="5" pageNumber="322">Octavius raymondi</emphasis>
<bibRefCitation id="EF9B4B15FFC9FFC2D13BF929FC3F587F" author="Saulcy, F. H. C." box="[843,1016,1772,1798]" pageId="5" pageNumber="322" pagination="113 - 125" refId="ref13812" refString="Saulcy, F. H. C. (de) (1878) Descriptions de plusieurs especes aveugles de staphylinides. Bulletin de la Societe d'Histoire Naturelle de Metz, 15, 113 - 125." type="journal article" year="1878">Saulcy, 1878</bibRefCitation>
(Staphylinidae)
</taxonomicName>
,
<taxonomicName id="4C0A4D67FFC9FFC2D698F929FE945855" class="Insecta" family="Curculionidae" genus="Torneuma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97EEAF6FFC9FFC2D698F929FAAC587C" box="[1256,1387,1772,1797]" italics="true" pageId="5" pageNumber="322">Torneuma</emphasis>
sp. (Curculionidae)
</taxonomicName>
,
<taxonomicName id="4C0A4D67FFC9FFC2D32FF8D6FD885855" authorityName="Perris" authorityYear="1869" box="[351,591,1811,1836]" class="Insecta" family="Brachyceridae" genus="Alaocyba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="carinulata">
<emphasis id="B97EEAF6FFC9FFC2D32FF8D6FD885855" box="[351,591,1811,1836]" italics="true" pageId="5" pageNumber="322">Alaocyba carinulata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0A4D67FFC9FFC2D02EF8D6FB195855" authority="sardous" authorityName="sardous (Perris" authorityYear="1869" box="[606,1246,1810,1836]" class="Insecta" family="Brachyceridae" genus="Raymondiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="subSpecies" species="sardous" subSpecies="sardous">
<emphasis id="B97EEAF6FFC9FFC2D02EF8D6FC155855" box="[606,978,1811,1836]" italics="true" pageId="5" pageNumber="322">Raymondiellus sardous sardous</emphasis>
(Raymondionymidae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFC9FFC2D2B6F8FDFF325803" pageId="5" pageNumber="322" type="etymology">
<paragraph id="8BB536E4FFC9FFC2D2B6F8FDFF325803" blockId="5.[151,1437,766,1914]" pageId="5" pageNumber="322">
<emphasis id="B97EEAF6FFC9FFC2D2B6F8FDFE94582B" bold="true" box="[198,339,1848,1874]" pageId="5" pageNumber="322">Etymology.</emphasis>
Named after the vegetal coal production that took place in the
<typeStatus id="54B18846FFC9FFC2D613F8FCFB52582A" box="[1123,1173,1849,1875]" pageId="5" pageNumber="322">type</typeStatus>
locality until the last century.
</paragraph>
</subSubSection>
</treatment>
</document>
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