638 lines
109 KiB
XML
638 lines
109 KiB
XML
<document id="8ABEF284F38BE02C8A8413B9231CA8EB" ID-ISSN="1536-9307" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638231627386" checkinUser="felipe" docAuthor="Kovařík, František & Lowe, Graeme" docDate="2012" docId="039B9366F53CFFE2FF54B195EB8CFB5B" docLanguage="en" docName="Euscorpius.138.1-25.pdf" docOrigin="Euscorpius 138" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Neobuthus awashensis Kovarik et Lowe 2012, sp. n." docType="treatment" docVersion="3" lastPageNumber="16" masterDocId="FFA2EB1EF534FFF3FF83B738EA4EFFE8" masterDocTitle="Review of the genus Neobuthus Hirst, 1911 with description of a new species from Ethiopia (Scorpiones: Buthidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="7" updateTime="1699434625454" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="249060BAED2A5551FEEE2D31991A41AD">Review of the genus Neobuthus Hirst, 1911 with description of a new species from Ethiopia (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="B3E545C98DED8938FC9366A706E79A27">Kovařík, František</mods:namePart>
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<mods:namePart id="23AE9A511FC61348B051AC02A3AA6DC2">Lowe, Graeme</mods:namePart>
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<mods:title id="FEA51D66AEA7035B0BC52698A7A752CE">Euscorpius</mods:title>
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<mods:part id="0D4B2A3A33A6BA5D6711ACF5D9096EA5">
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<mods:date id="5D7490E409C5948BAD8EA61CA5EDC05C">2012</mods:date>
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<mods:number id="E74E8EA74C016779935D99222E4D3509">138</mods:number>
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<mods:classification id="19320351A6CD378471B9AC7BFD7011EA">journal article</mods:classification>
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<mods:identifier id="E7FD59703BB59F31837468627BE27D36" type="ISSN">1536-9307</mods:identifier>
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<treatment id="039B9366F53CFFE2FF54B195EB8CFB5B" ID-DOI="http://doi.org/10.5281/zenodo.5738999" ID-Zenodo-Dep="5738999" LSID="urn:lsid:plazi:treatment:039B9366F53CFFE2FF54B195EB8CFB5B" httpUri="http://treatment.plazi.org/id/039B9366F53CFFE2FF54B195EB8CFB5B" lastPageId="17" lastPageNumber="16" pageId="8" pageNumber="7">
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<subSubSection id="C32871FBF53CFFFBFF54B195E944F920" box="[215,778,1709,1737]" pageId="8" pageNumber="7" type="nomenclature">
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<paragraph id="8B8D2270F53CFFFBFF54B195E944F920" blockId="8.[196,797,1709,1797]" box="[215,778,1709,1737]" pageId="8" pageNumber="7">
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<heading id="D0C5951CF53CFFFBFF54B195E944F920" box="[215,778,1709,1737]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="7" reason="2">
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<taxonomicName id="4C3259F3F53CFFFBFF54B195E8F3F921" ID-CoL="46BZV" authority="Kovarik et Lowe" authorityName="Kovarik et Lowe" authorityYear="2012" box="[215,701,1709,1737]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
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<emphasis id="B946FE62F53CFFFBFF54B195EBA2F920" bold="true" box="[215,492,1709,1736]" italics="true" pageId="8" pageNumber="7">Neobuthus awashensis</emphasis>
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Kovařík et Lowe
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</taxonomicName>
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,
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<emphasis id="B946FE62F53CFFFBFD4BB196E944F920" bold="true" box="[712,778,1710,1736]" pageId="8" pageNumber="7">
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<taxonomicNameLabel id="A2754319F53CFFFBFD4BB196E944F920" box="[712,778,1710,1736]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C32871FBF53CFFFBFF47B1F7E876F8ED" pageId="8" pageNumber="7" type="description">
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<paragraph id="8B8D2270F53CFFFBFF47B1F7E876F8ED" blockId="8.[196,797,1709,1797]" pageId="8" pageNumber="7">
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(
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<figureCitation id="13093EF5F53CFFFBFF4EB1F7EB7EF90E" box="[205,304,1743,1766]" captionStart="Figures 1–8" captionStartId="5.[192,278,1733,1757]" captionTargetBox="[192,1440,220,1680]" captionTargetId="figure-6@5.[192,1440,200,1704]" captionTargetPageId="5" captionText="Figures 1–8: Figures 1–7: Fifth (or fourth and fifth) metasomal segment and telson. Figures 1–2: Neobuthus berberensis Hirst, 1911, ♀ (1) and ♂ (2), Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figures 3–4: Neobuthus ferrugineus (Kraepelin, 1898) comb. n., ♀ (3), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP and ♂ (4), Somaliland, 15 km near Sheikh, Goolis mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l., FKCP. Figures 5–6: Neobuthus awashensis sp. n., ♀ allotype (5) and ♂ (6) holotype. Figure 7: Butheolus thalassinus Simon, 1882, ♂, Yemen, Ta’izz government, N of Al Makha by road, 13°23'37"N 43°16'22"E, 5 m a.s.l., FKCP. Figure 8: Carapace, Butheolus thalassinus Simon, 1882, ♂, same specimen locality as Figure 7." pageId="8" pageNumber="7">Figs. 5–6</figureCitation>
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,
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<figureCitation id="13093EF5F53CFFFBFEBDB1F7EBCCF90E" box="[318,386,1743,1766]" captionStart="Figures 14–21" captionStartId="8.[192,278,1360,1384]" captionTargetBox="[227,1416,123,1323]" captionTargetId="figure-13@8.[223,1427,120,1333]" captionTargetPageId="8" captionText="Figures 14–21: Figures 14–17: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figures 14–15: Dorsal and ventral views, ♂ (20 mm). Figures 16–17: Dorsal and ventral views, ♀ (30 mm). Figures 18–21: Neobuthus awashensis sp. n.. Figures 18–19: Dorsal and ventral views, ♂ (20 mm) holotype. Figures 20–21: Dorsal and ventral views, ♀ (29 mm) allotype." pageId="8" pageNumber="7">18–21</figureCitation>
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,
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<figureCitation id="13093EF5F53CFFFBFE0DB1F7EB9DF90E" box="[398,467,1743,1766]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="8" pageNumber="7">34–38</figureCitation>
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,
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<figureCitation id="13093EF5F53CFFFBFE5DB1F7E86DF90E" box="[478,547,1743,1766]" captionStart="Figures 39–47" captionStartId="10.[192,278,1730,1754]" captionTargetBox="[193,1440,205,1693]" captionTargetId="figure-13@10.[192,1440,192,1701]" captionTargetPageId="10" captionText="Figures 39–47: Figures 39–40: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Trochanter and femur of pedipalp (39), and patela and chela of pedipalp (40), ♀ (30 mm). Figures 41–43: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Pedipalp (41), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Pedipalp (42), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Pedipalp (43), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Figures 44–47: Neobuthus awashensis sp. n.. Pedipalp (44), ♂‚ (21 mm) paratype. Chela dorsal (45), tarsomeres of the second leg (46), and trichobothrial pattern (47), ♀ (30 mm) paratype." pageId="8" pageNumber="7">44–47</figureCitation>
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,
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<figureCitation id="13093EF5F53CFFFBFDADB1F7E88EF90E" box="[558,704,1743,1766]" captionStart-0="Figures 67–71" captionStart-1="Figure 72" captionStart-2="Figure 73" captionStart-3="Figures 74–91" captionStartId-0="16.[192,278,1866,1890]" captionStartId-1="18.[192,268,1250,1274]" captionStartId-2="19.[192,268,1330,1354]" captionStartId-3="20.[192,278,1752,1776]" captionTargetBox-0="[227,1398,172,1836]" captionTargetBox-1="[217,1416,217,1207]" captionTargetBox-2="[191,771,198,1302]" captionTargetBox-3="[243,1388,188,1729]" captionTargetId-0="figure-13@16.[228,1398,173,1836]" captionTargetId-1="figure-13@18.[192,1441,192,1232]" captionTargetId-2="figure-6@19.[192,771,199,1301]" captionTargetId-3="figure-13@20.[216,1418,176,1733]" captionTargetPageId-0="16" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="Figures 67–71: Neobuthus awashensis sp. n. Figures 67–68: Females paratypes at the type locality. Figure 69–70. Males paratypes at the type locality. Figure 71: Ethiopia, Awash, Metahara env., 08°54'N 39°54'E, 960-1050 m a.s.l. Type locality of Neobuthus awashensis sp. n. and Buthus awashensis Kovařík, 2011 together with Compsobuthus abyssinicus Birula, 1903 and Parabuthus liosoma (Ehrenberg, 1828)." captionText-1="Figure 72: Distribution map of the three African closely related species of Neobuthus except Chisimaio (= Kismaayo), locality of Neobuthus ferrugineus (Kraepelin, 1898) in the southern part of Somalia." captionText-2="Figure 73: Neobuthus awashensis sp. n. Right hemispermatophore of paratype ♂, viewed from convex side. The structure has been compressed under a cover slip to reveal flat profiles of the basal lobes. Scale bar: 0.2 mm." captionText-3="Figures 74–91: Prosomal and mesosomal characters in Neobuthus and Butheolus. Figures 74–76: Carapace and tergite I of ♂ Neobuthus awashensis sp. n. (74), ♂ Butheolus thalassinus (75), and ♂ B. gallagheri (76), showing granulation, carination, and margins. Figures 77–85: Carapace anterior margin of 9 ♀♀ B. gallagheri. Figures 86–88: Pectines and sternites of ♂ Neobuthus awashensis sp. n. (86), ♂ Butheolus thalassinus (87), and ♂ B. gallagheri (88). Figures 89–91: Higher magnification views of sternite III posterior margins of ♂ Neobuthus awashensis sp. n. (89), ♂ Butheolus thalassinus (90), and ♂ B. gallagheri (91), showing differences in denticle armature. Scale bars: 1 mm in 74–76, 86–88; 0.5 mm in 77–85 (bar in 77 applies to 77–85), 89– 91. All photographs acquired by UV fluorescence imaging. Comparative material: Butheolus gallagheri, ♂, ♀, Oman, wadi above Khor Rhori Beach, E of Taqah, 17°3.22'N 54°25.33'E 18 Oct 1993, leg. G. Lowe, GL; Butheolus thalassinus, ♂ Yemen, Ta'izz, under rock on Euphorbia-Aloe hill 3800 ft, 10 Jan 1951, leg. H. Hoogstraal, WDS." pageId="8" pageNumber="7">67–74, 86, 89</figureCitation>
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,
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<figureCitation id="13093EF5F53CFFFBFD4EB1F7E8A6F90E" box="[717,744,1743,1766]" captionStart="Figures 92–94" captionStartId="21.[192,278,906,930]" captionTargetBox="[201,1429,178,869]" captionTargetId="figure-6@21.[192,1439,168,877]" captionTargetPageId="21" captionText="Figures 92–94: Ventral aspect of chelicerae of Neobuthus and Butheolus. Figure 92: Neobuthus awashensis sp. n., paratype ♂, showing single denticle (d) on fixed finger. Figures 93-94: ♂ Butheolus gallagheri (93) and ♂ Butheolus anthracinus (94), showing two denticles, proximal (d1) and distal (d2), on fixed finger. We also examined the cheliceral fixed finger of Butheolus thalassinus and confirmed the presence of two ventral denticles (photo not shown). Scale bars: 0.5 mm. All photographs acquired by UV fluorescence imaging. Comparative material: Butheolus gallagheri, see data under caption for Figs. 74–91. Butheolus anthracinus, ♂, Oman, Nejd Desert, SW of Thumrait, 17°30.77N 54°2.28'E, 600 m a.s.l., 19 Oct 1993, leg. G. Lowe, NMBS." pageId="8" pageNumber="7">92</figureCitation>
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,
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<figureCitation id="13093EF5F53CFFFBFD76B1F7EB8BF8ED" captionStart="Figures 95–99" captionStartId="22.[192,278,1384,1408]" captionTargetBox="[214,1386,223,1323]" captionTargetId="figure-13@22.[192,1440,192,1355]" captionTargetPageId="22" captionText="Figures 95–99: Ventral aspect of metasoma and telson in Neobuthus and Butheolus. Figures 95–96: Neobuthus awashensis sp. n., paratype ♂ (95) and ♀ (96). Figures 97–98: Butheolus gallagheri, ♂ (97) and ♀ (98). Figure 99: Butheolus thalassinus, ♂. Scale bars: 1 mm. All photographs acquired by UV fluorescence imaging. Comparative material: see data under caption for Figs. 74–91." pageId="8" pageNumber="7">95– 96</figureCitation>
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,
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<figureCitation id="13093EF5F53CFFFBFE51B1D6E861F8ED" box="[466,559,1774,1797]" captionStart="Figures 100–104" captionStartId="23.[192,278,1630,1654]" captionTargetBox="[201,1409,236,1545]" captionTargetId="figure-6@23.[192,1440,192,1601]" captionTargetPageId="23" captionText="Figures 100–104: Lateral aspect of metasoma and telson in Neobuthus and Butheolus. Figures 100–101: Neobuthus awashensis sp. n., paratype ♂ (100) and ♀ (101). Figures 102–103: Butheolus gallagheri, ♂ (102) and ♀ (103). Figure 104: Butheolus thalassinus, ♂. Scale bars: 1 mm. All photographs acquired by UV fluorescence imaging. Comparative material: see data under caption for Figs. 74–91." pageId="8" pageNumber="7">100–101</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32871FBF53CFFFBFF43B01EEE58F90C" pageId="8" pageNumber="7" type="materials_examined">
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<paragraph id="8B8D2270F53CFFFBFF43B01EE8FBF893" blockId="8.[192,809,1830,1915]" pageId="8" pageNumber="7">
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<materialsCitation id="3B5A282DF53CFFFBFF43B01EE8FFF893" collectionCode="FKCP" country="Ethiopia" elevation="1005" latitude="8.9" location="Awash" longLatPrecision="1300" longitude="39.9" municipality="Metahara" pageId="8" pageNumber="7" specimenCount="1">
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TYPE LOCALITY AND
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<typeStatus id="54899CD2F53CFFFBFE51B012E849F8D5" box="[466,519,1834,1853]" pageId="8" pageNumber="7">TYPE</typeStatus>
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DEPOSITORY.
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<emphasis id="B946FE62F53CFFFBFD43B01EE96CF8D6" bold="true" box="[704,802,1830,1854]" pageId="8" pageNumber="7">
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<collectingCountry id="F32562E0F53CFFFBFD43B01EE96CF8D6" box="[704,802,1830,1854]" name="Ethiopia" pageId="8" pageNumber="7">Ethiopia</collectingCountry>
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</emphasis>
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,
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<location id="8EED74ABF53CFFFBFF43B07DEB44F8B4" LSID="urn:lsid:plazi:treatment:039B9366F53CFFE2FF54B195EB8CFB5B:8EED74ABF53CFFFBFF43B07DEB44F8B4" box="[192,266,1861,1884]" country="Ethiopia" latitude="8.9" longLatPrecision="1300" longitude="39.9" municipality="Metahara" name="Awash" pageId="8" pageNumber="7">Awash</location>
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,
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<collectingMunicipality id="6BE9B80AF53CFFFBFE9EB07DEBCCF8B4" box="[285,386,1861,1884]" pageId="8" pageNumber="7">Metahara</collectingMunicipality>
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env.,
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<geoCoordinate id="EE0644B7F53CFFFBFE4DB07DE868F8B4" box="[462,550,1861,1884]" degrees="08" direction="north" minutes="54" orientation="latitude" pageId="8" pageNumber="7" precision="925" value="8.9">08°54'N</geoCoordinate>
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<geoCoordinate id="EE0644B7F53CFFFBFDB0B07DE8C4F8B4" box="[563,650,1861,1884]" degrees="39" direction="east" minutes="54" orientation="longitude" pageId="8" pageNumber="7" precision="925" value="39.9">39°54'E</geoCoordinate>
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,
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<quantity id="4CCA8F95F53CFFFBFD19B07DE96FF8B4" box="[666,801,1861,1884]" metricMagnitude="3" metricUnit="m" metricValue="1.005" metricValueMax="1.05" metricValueMin="0.96" pageId="8" pageNumber="7" unit="m" value="1005.0" valueMax="1050.0" valueMin="960.0">
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<elevation id="001FC543F53CFFFBFD19B07DE96FF8B4" box="[666,801,1861,1884]" metricMagnitude="3" metricUnit="m" metricValue="1.005" metricValueMax="1.05" metricValueMin="0.96" pageId="8" pageNumber="7" unit="m" value="1005.0" valueMax="1050.0" valueMin="960.0">960-1050 m</elevation>
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</quantity>
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. a.s.l. (
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<figureCitation id="13093EF5F53CFFFBFE83B05CEB03F893" box="[256,333,1892,1915]" captionStart="Figures 67–71" captionStartId="16.[192,278,1866,1890]" captionTargetBox="[227,1398,172,1836]" captionTargetId="figure-13@16.[228,1398,173,1836]" captionTargetPageId="16" captionText="Figures 67–71: Neobuthus awashensis sp. n. Figures 67–68: Females paratypes at the type locality. Figure 69–70. Males paratypes at the type locality. Figure 71: Ethiopia, Awash, Metahara env., 08°54'N 39°54'E, 960-1050 m a.s.l. Type locality of Neobuthus awashensis sp. n. and Buthus awashensis Kovařík, 2011 together with Compsobuthus abyssinicus Birula, 1903 and Parabuthus liosoma (Ehrenberg, 1828)." pageId="8" pageNumber="7">Fig. 71</figureCitation>
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); the authorś collection (
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<collectionCode id="ED23BAB5F53CFFFBFDE2B05CE8E4F893" box="[609,682,1892,1915]" pageId="8" pageNumber="7">FKCP</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8D2270F53CFFFBFCBDB2EFEE58F90C" blockId="8.[830,1442,1495,1764]" pageId="8" pageNumber="7">
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||
<materialsCitation id="3B5A282DF53CFFFBFCBDB2EFEE41F90C" collectingDate="2010-11" collectingDateMax="2011-07-22" collectingDateMin="2010-11" collectionCode="FKCP, GL" collectorName="V. Trailin & T. Mazuch & P. Novak & F. Kovarik. Most" country="Ethiopia" elevation="1005" latitude="8.9" location="Awash" longLatPrecision="1300" longitude="39.9" municipality="Metahara" pageId="8" pageNumber="7" specimenCount="73" specimenCount-female="34" specimenCount-male="38" typeStatus="paratype">
|
||
TYPE MATERIAL.
|
||
<collectingCountry id="F32562E0F53CFFFBFB96B2EFEE3CFA06" box="[1045,1138,1495,1518]" name="Ethiopia" pageId="8" pageNumber="7">Ethiopia</collectingCountry>
|
||
,
|
||
<location id="8EED74ABF53CFFFBFB0FB2EFEE98FA06" LSID="urn:lsid:plazi:treatment:039B9366F53CFFE2FF54B195EB8CFB5B:8EED74ABF53CFFFBFB0FB2EFEE98FA06" box="[1164,1238,1495,1518]" country="Ethiopia" latitude="8.9" longLatPrecision="1300" longitude="39.9" municipality="Metahara" name="Awash" pageId="8" pageNumber="7">Awash</location>
|
||
,
|
||
<collectingMunicipality id="6BE9B80AF53CFFFBFB71B2EFEF19FA06" box="[1266,1367,1495,1518]" pageId="8" pageNumber="7">Metahara</collectingMunicipality>
|
||
env.,
|
||
<geoCoordinate id="EE0644B7F53CFFFBFCBDB2CEE9D8F9E5" box="[830,918,1526,1549]" degrees="08" direction="north" minutes="54" orientation="latitude" pageId="8" pageNumber="7" precision="925" value="8.9">08°54'N</geoCoordinate>
|
||
<geoCoordinate id="EE0644B7F53CFFFBFC2AB2CEEE4FF9E5" box="[937,1025,1526,1549]" degrees="39" direction="east" minutes="54" orientation="longitude" pageId="8" pageNumber="7" precision="925" value="39.9">39°54'E</geoCoordinate>
|
||
,
|
||
<elevation id="001FC543F53CFFFBFB9BB2CEEEA7F9E5" box="[1048,1257,1526,1549]" metricMagnitude="3" metricUnit="m" metricValue="1.005" metricValueMax="1.05" metricValueMin="0.96" pageId="8" pageNumber="7" unit="m" value="1005.0" valueMax="1050.0" valueMin="960.0">
|
||
<quantity id="4CCA8F95F53CFFFBFB9BB2CEEEE8F9E5" box="[1048,1190,1526,1549]" metricMagnitude="3" metricUnit="m" metricValue="1.005" metricValueMax="1.05" metricValueMin="0.96" pageId="8" pageNumber="7" unit="m" value="1005.0" valueMax="1050.0" valueMin="960.0">960-1050 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(
|
||
<figureCitation id="13093EF5F53CFFFBFA86B2CEEF13F9E5" box="[1285,1373,1526,1549]" captionStart="Figures 67–71" captionStartId="16.[192,278,1866,1890]" captionTargetBox="[227,1398,172,1836]" captionTargetId="figure-13@16.[228,1398,173,1836]" captionTargetPageId="16" captionText="Figures 67–71: Neobuthus awashensis sp. n. Figures 67–68: Females paratypes at the type locality. Figure 69–70. Males paratypes at the type locality. Figure 71: Ethiopia, Awash, Metahara env., 08°54'N 39°54'E, 960-1050 m a.s.l. Type locality of Neobuthus awashensis sp. n. and Buthus awashensis Kovařík, 2011 together with Compsobuthus abyssinicus Birula, 1903 and Parabuthus liosoma (Ehrenberg, 1828)." pageId="8" pageNumber="7">Fig. 71</figureCitation>
|
||
),
|
||
<specimenCount id="9D34E9F9F53CFFFBFAFCB2CEEFEEF9E5" box="[1407,1440,1526,1549]" pageId="8" pageNumber="7" type="female">1♀</specimenCount>
|
||
(
|
||
<typeStatus id="54899CD2F53CFFFBFCC5B12DE9E6F9C4" box="[838,936,1557,1580]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
|
||
), 2008, leg.
|
||
<collectorName id="26C747A6F53CFFFBFBAEB12DEED6F9C4" box="[1069,1176,1557,1580]" pageId="8" pageNumber="7">V. Trailin</collectorName>
|
||
,
|
||
<specimenCount id="9D34E9F9F53CFFFBFB25B12DEEA6F9C4" box="[1190,1256,1557,1580]" pageId="8" pageNumber="7" type="female">2♀1♀</specimenCount>
|
||
im. (
|
||
<typeStatus id="54899CD2F53CFFFBFA99B12DEF3CF9C4" box="[1306,1394,1557,1580]" pageId="8" pageNumber="7" type="allotype">allotype</typeStatus>
|
||
and
|
||
<typeStatus id="54899CD2F53CFFFBFCC3B10BE9EBF9A2" box="[832,933,1587,1610]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
),
|
||
<date id="FF8C04B0F53CFFFBFC3FB10BEE5AF9A2" box="[956,1044,1587,1610]" pageId="8" pageNumber="7" value="2010-11">
|
||
<collectingDate id="EFC8FD58F53CFFFBFC3FB10BEE5AF9A2" box="[956,1044,1587,1610]" pageId="8" pageNumber="7" value="2010-11">XI.2010</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26C747A6F53CFFFBFBD0B10BEE86F9A2" box="[1107,1224,1587,1610]" pageId="8" pageNumber="7">T. Mazuch</collectorName>
|
||
and
|
||
<collectorName id="26C747A6F53CFFFBFB7DB10BEF2DF9A2" box="[1278,1379,1587,1610]" pageId="8" pageNumber="7">P. Novák</collectorName>
|
||
,
|
||
<specimenCount id="9D34E9F9F53CFFFBFAF1B10BEFEEF9A2" box="[1394,1440,1587,1610]" pageId="8" pageNumber="7" type="male">32♂</specimenCount>
|
||
(
|
||
<typeStatus id="54899CD2F53CFFFBFCC5B16AE9EBF981" box="[838,933,1618,1641]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54899CD2F53CFFFBFC70B16AEE17F981" box="[1011,1113,1618,1641]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
)
|
||
<specimenCount id="9D34E9F9F53CFFFBFBF5B16AEEEBF981" box="[1142,1189,1618,1641]" pageId="8" pageNumber="7" type="female">18♀</specimenCount>
|
||
(
|
||
<typeStatus id="54899CD2F53CFFFBFB42B16AEF67F981" box="[1217,1321,1618,1641]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
)
|
||
<specimenCount id="9D34E9F9F53CFFFBFAC7B16AEF3DF981" box="[1348,1395,1618,1641]" pageId="8" pageNumber="7" type="female">11♀</specimenCount>
|
||
ims.
|
||
<specimenCount id="9D34E9F9F53CFFFBFCBDB149E92EF960" box="[830,864,1649,1672]" pageId="8" pageNumber="7" type="male">5♂</specimenCount>
|
||
ims. (
|
||
<typeStatus id="54899CD2F53CFFFBFC27B149EE42F960" box="[932,1036,1649,1672]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
),
|
||
<date id="FF8C04B0F53CFFFBFBABB149EE9BF960" box="[1064,1237,1649,1672]" pageId="8" pageNumber="7" value="2011-07-19" valueMax="2011-07-22" valueMin="2011-07-19">
|
||
<collectingDate id="EFC8FD58F53CFFFBFBABB149EE9BF960" box="[1064,1237,1649,1672]" pageId="8" pageNumber="7" value="2011-07-19" valueMax="2011-07-22" valueMin="2011-07-19">19.-22.VII.2011</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26C747A6F53CFFFBFA9DB149E93BF94E" pageId="8" pageNumber="7">F. Kovařík. Most</collectorName>
|
||
types are in the collection of the first author (
|
||
<collectionCode id="ED23BAB5F53CFFFBFCC5B196E9C0F92D" box="[838,910,1710,1733]" pageId="8" pageNumber="7">FKCP</collectionCode>
|
||
),
|
||
<specimenCount id="9D34E9F9F53CFFFBFC20B196EE76F92D" box="[931,1080,1710,1733]" pageId="8" pageNumber="7" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
(
|
||
<specimenCount id="9D34E9F9F53CFFFBFBCAB196EE2FF92D" box="[1097,1121,1710,1733]" pageId="8" pageNumber="7" type="male">♂</specimenCount>
|
||
<specimenCount id="9D34E9F9F53CFFFBFBE2B196EE3FF92D" box="[1121,1137,1710,1733]" pageId="8" pageNumber="7" type="female">♀</specimenCount>
|
||
) are in the collection of the second author (
|
||
<collectionCode id="ED23BAB5F53CFFFBFC62B1F5EE48F90C" box="[993,1030,1741,1764]" pageId="8" pageNumber="7">GL</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32871FBF53CFFFBFCBDB032EF5BF8C9" box="[830,1301,1802,1825]" pageId="8" pageNumber="7" type="etymology">
|
||
<paragraph id="8B8D2270F53CFFFBFCBDB032EF5BF8C9" blockId="8.[830,1301,1802,1825]" box="[830,1301,1802,1825]" pageId="8" pageNumber="7">
|
||
ETYMOLOGY. Named after the
|
||
<typeStatus id="54899CD2F53CFFFBFB0BB032EEF8F8C9" box="[1160,1206,1802,1825]" pageId="8" pageNumber="7">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32871FBF53CFFF8FCBDB07FEB36F890" lastPageId="11" lastPageNumber="10" pageId="8" pageNumber="7" type="diagnosis">
|
||
<paragraph id="8B8D2270F53CFFF8FCBDB07FEB36F890" blockId="8.[830,1440,1863,1917]" lastBlockId="11.[192,802,1735,1912]" lastPageId="11" lastPageNumber="10" pageId="8" pageNumber="7">
|
||
DIAGNOSIS. Total length
|
||
<quantity id="4CCA8F95F53CFFFBFBD7B07FEE84F8B6" box="[1108,1226,1863,1886]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="8" pageNumber="7" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">20–21 mm</quantity>
|
||
(males) and
|
||
<quantity id="4CCA8F95F53CFFFBFADEB07FE926F895" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="8" pageNumber="7" unit="mm" value="28.5" valueMax="30.0" valueMin="27.0">27–30 mm</quantity>
|
||
(females); carapace densely granulated with only anterior median carinae developed, area between these carinae fuscous; anterior margin of carapace straight or slightly convex; pectine teeth
|
||
<quantity id="4CCA8F95F53DFFFAFD8CB07DE83FF8B4" box="[527,625,1861,1884]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.572" metricValueMin="3.81" pageId="9" pageNumber="8" unit="in" value="16.5" valueMax="18.0" valueMin="15.0">15–18 in</quantity>
|
||
females,
|
||
<quantity id="4CCA8F95F53DFFFAFD5CB07DEA9BF893" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.334" metricValueMin="4.318" pageId="9" pageNumber="8" unit="in" value="19.0" valueMax="21.0" valueMin="17.0">17–21 in</quantity>
|
||
males; sternites III–VI smooth, sternite VII with 4 weakly granulated or obsolete carinae; pedipalps including trochanter, metasoma and telson with sparse dark, long thin setae in females and shorter spiniform setae in males; metasomal segment V with ca. 35–40 long setae in females, shorter spiniform setae in males; legs I–III with bristle combs; movable finger of pedipalp bears 5–6 rows of denticles, with external and internal accessory denticles and three distal denticles; pedipalp femur length to width ratio
|
||
<quantity id="4CCA8F95F53FFFF8FDD0B07AE88EF8B1" box="[595,704,1858,1881]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="6.604" metricValueMin="6.096" pageId="11" pageNumber="10" unit="in" value="2.5" valueMax="2.6" valueMin="2.4">2.4–2.6 in</quantity>
|
||
females,
|
||
<quantity id="4CCA8F95F53FFFF8FF41B059EB62F890" box="[194,300,1889,1912]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" metricValueMax="6.858" metricValueMin="6.35" pageId="11" pageNumber="10" unit="in" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 in</quantity>
|
||
males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4D72F8F53DFFFAFF43B2D3EB3CF90A" pageId="9" pageNumber="8" startId="9.[192,278,1515,1539]" targetBox="[192,1440,192,1484]" targetPageId="9">
|
||
<paragraph id="8B8D2270F53DFFFAFF43B2D3EB3CF90A" blockId="9.[192,1442,1515,1762]" pageId="9" pageNumber="8">
|
||
<emphasis id="B946FE62F53DFFFAFF43B2D3E840F9EA" bold="true" box="[192,526,1515,1539]" pageId="9" pageNumber="8">Figures 22–38: Figures 22–27:</emphasis>
|
||
<taxonomicName id="4C3259F3F53DFFFAFD9BB2D6E93CF9EB" authority="Hirst, 1911" authorityName="Hirst" authorityYear="1911" box="[536,882,1518,1539]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="berberensis">
|
||
<emphasis id="B946FE62F53DFFFAFD9BB2D6E8B6F9EB" box="[536,760,1518,1539]" italics="true" pageId="9" pageNumber="8">Neobuthus berberensis</emphasis>
|
||
Hirst, 1911
|
||
</taxonomicName>
|
||
, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (
|
||
<emphasis id="B946FE62F53DFFFAFEDDB13EEB38F9F3" bold="true" box="[350,374,1542,1563]" pageId="9" pageNumber="8">22</emphasis>
|
||
), trochanter and femur of pedipalp (
|
||
<emphasis id="B946FE62F53DFFFAFD63B13EE8B6F9F3" bold="true" box="[736,760,1542,1563]" pageId="9" pageNumber="8">23</emphasis>
|
||
), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (
|
||
<emphasis id="B946FE62F53DFFFAFAF9B13EEFDCF9F3" bold="true" box="[1402,1426,1542,1563]" pageId="9" pageNumber="8">26</emphasis>
|
||
), and trochanter and femur of pedipalp (
|
||
<emphasis id="B946FE62F53DFFFAFDCBB126E82EF9DB" bold="true" box="[584,608,1566,1587]" pageId="9" pageNumber="8">27</emphasis>
|
||
), ♂ (20 mm).
|
||
<emphasis id="B946FE62F53DFFFAFD79B126E9DAF9DB" bold="true" box="[762,916,1566,1588]" pageId="9" pageNumber="8">Figures 28–33:</emphasis>
|
||
<taxonomicName id="4C3259F3F53DFFFAFC1DB127EF7BF9DC" authority="(Kraepelin, 1898)" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" box="[926,1333,1567,1588]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F53DFFFAFC1DB127EE30F9DC" box="[926,1150,1567,1588]" italics="true" pageId="9" pageNumber="8">Neobuthus ferrugineus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA35F81F53DFFFAFB0DB127EF60F9DC" author="KRAEPELIN" box="[1166,1326,1567,1588]" pageId="9" pageNumber="8" pagination="39 - 44" refId="ref13141" refString="KRAEPELIN, K. 1898. Neue Pedipalpen und Scorpione des Hamburger Museums. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 15: 39 - 44." type="journal article" year="1898">Kraepelin, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F53DFFFAFABCB127EFD4F9DC" bold="true" box="[1343,1434,1567,1588]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="A2754319F53DFFFAFABCB127EFD4F9DC" box="[1343,1434,1567,1588]" pageId="9" pageNumber="8" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. Carapace (
|
||
<emphasis id="B946FE62F53DFFFAFEABB10FEB0EF9A4" bold="true" box="[296,320,1591,1612]" pageId="9" pageNumber="8">28</emphasis>
|
||
) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (
|
||
<emphasis id="B946FE62F53DFFFAFD2DB168E888F98D" bold="true" box="[686,710,1616,1637]" pageId="9" pageNumber="8">29</emphasis>
|
||
), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (
|
||
<emphasis id="B946FE62F53DFFFAFEFDB151EBD8F996" bold="true" box="[382,406,1641,1662]" pageId="9" pageNumber="8">31</emphasis>
|
||
) and trochanter and femur of pedipalp (
|
||
<emphasis id="B946FE62F53DFFFAFCCBB151E92EF996" bold="true" box="[840,864,1641,1662]" pageId="9" pageNumber="8">33</emphasis>
|
||
), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (
|
||
<emphasis id="B946FE62F53DFFFAFCF0B1BAE9C5F97F" bold="true" box="[883,907,1666,1687]" pageId="9" pageNumber="8">32</emphasis>
|
||
), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP.
|
||
<emphasis id="B946FE62F53DFFFAFD2BB1A3E90CF958" bold="true" box="[680,834,1691,1713]" pageId="9" pageNumber="8">Figures 34–38:</emphasis>
|
||
<taxonomicName id="4C3259F3F53DFFFAFCCFB1A3EE66F958" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" box="[844,1064,1691,1712]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F53DFFFAFCCFB1A3EE66F958" box="[844,1064,1691,1712]" italics="true" pageId="9" pageNumber="8">Neobuthus awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F53DFFFAFBB1B1A3EE25F958" bold="true" box="[1074,1131,1691,1712]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="A2754319F53DFFFAFBB1B1A3EE25F958" box="[1074,1131,1691,1712]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. Carapace (
|
||
<emphasis id="B946FE62F53DFFFAFB67B1A3EEB2F958" bold="true" box="[1252,1276,1691,1712]" pageId="9" pageNumber="8">34</emphasis>
|
||
), trochanter and femur of pedipalp (
|
||
<emphasis id="B946FE62F53DFFFAFEFDB18BEBD8F920" bold="true" box="[382,406,1715,1736]" pageId="9" pageNumber="8">35</emphasis>
|
||
), and lateral eyes (
|
||
<emphasis id="B946FE62F53DFFFAFDD0B18BE825F920" bold="true" box="[595,619,1715,1736]" pageId="9" pageNumber="8">36</emphasis>
|
||
), ♀ (30 mm) paratype. Carapace (
|
||
<emphasis id="B946FE62F53DFFFAFC3DB18BE998F920" bold="true" box="[958,982,1715,1736]" pageId="9" pageNumber="8">37</emphasis>
|
||
) and trochanter and femur of pedipalp (
|
||
<emphasis id="B946FE62F53DFFFAFAE2B18BEF37F920" bold="true" box="[1377,1401,1715,1736]" pageId="9" pageNumber="8">38</emphasis>
|
||
), ♂ (21 mm) paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D72F8F53EFFF9FF43B1FAE8DEF887" pageId="10" pageNumber="9" startId="10.[192,278,1730,1754]" targetBox="[193,1440,205,1693]" targetPageId="10">
|
||
<paragraph id="8B8D2270F53EFFF9FF43B1FAE8DEF887" blockId="10.[192,1441,1730,1903]" pageId="10" pageNumber="9">
|
||
<emphasis id="B946FE62F53EFFF9FF43B1FAE840F931" bold="true" box="[192,526,1730,1754]" pageId="10" pageNumber="9">Figures 39–47: Figures 39–40:</emphasis>
|
||
<taxonomicName id="4C3259F3F53EFFF9FD9BB1FDE93CF932" authority="Hirst, 1911" authorityName="Hirst" authorityYear="1911" box="[536,882,1733,1754]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="berberensis">
|
||
<emphasis id="B946FE62F53EFFF9FD9BB1FDE8B6F932" box="[536,760,1733,1754]" italics="true" pageId="10" pageNumber="9">Neobuthus berberensis</emphasis>
|
||
Hirst, 1911
|
||
</taxonomicName>
|
||
, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Trochanter and femur of pedipalp (
|
||
<emphasis id="B946FE62F53EFFF9FDD9B1E5E83CF91A" bold="true" box="[602,626,1757,1778]" pageId="10" pageNumber="9">39</emphasis>
|
||
), and patela and chela of pedipalp (
|
||
<emphasis id="B946FE62F53EFFF9FC60B1E5E9B5F91A" bold="true" box="[995,1019,1757,1778]" pageId="10" pageNumber="9">40</emphasis>
|
||
), ♀ (30 mm).
|
||
<emphasis id="B946FE62F53EFFF9FB16B1E5EF7EF91A" bold="true" box="[1173,1328,1757,1779]" pageId="10" pageNumber="9">Figures 41–43:</emphasis>
|
||
<taxonomicName id="4C3259F3F53EFFF9FABAB1E6EBAFF8E3" authority="(Kraepelin, 1898)" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F53EFFF9FABAB1E6EB61F8E3" italics="true" pageId="10" pageNumber="9">Neobuthus ferrugineus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA35F81F53EFFF9FEBFB1CEEB97F8E3" author="KRAEPELIN" box="[316,473,1782,1803]" pageId="10" pageNumber="9" pagination="39 - 44" refId="ref13141" refString="KRAEPELIN, K. 1898. Neue Pedipalpen und Scorpione des Hamburger Museums. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 15: 39 - 44." type="journal article" year="1898">Kraepelin, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F53EFFF9FE64B1CEE870F8E3" bold="true" box="[487,574,1782,1803]" pageId="10" pageNumber="9">
|
||
<taxonomicNameLabel id="A2754319F53EFFF9FE64B1CEE870F8E3" box="[487,574,1782,1803]" pageId="10" pageNumber="9" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. Pedipalp (
|
||
<emphasis id="B946FE62F53EFFF9FD2EB1CEE88BF8E3" bold="true" box="[685,709,1782,1803]" pageId="10" pageNumber="9">41</emphasis>
|
||
), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Pedipalp (
|
||
<emphasis id="B946FE62F53EFFF9FEF1B037EBC4F8CC" bold="true" box="[370,394,1807,1828]" pageId="10" pageNumber="9">42</emphasis>
|
||
), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Pedipalp (
|
||
<emphasis id="B946FE62F53EFFF9FEAAB01FEB0FF8D4" bold="true" box="[297,321,1831,1852]" pageId="10" pageNumber="9">43</emphasis>
|
||
), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP.
|
||
<emphasis id="B946FE62F53EFFF9FA81B01FEFEEF8D4" bold="true" box="[1282,1440,1831,1853]" pageId="10" pageNumber="9">Figures 44–47:</emphasis>
|
||
<taxonomicName id="4C3259F3F53EFFF9FF43B079EBD3F8BE" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" box="[192,413,1857,1878]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F53EFFF9FF43B079EBD3F8BE" box="[192,413,1857,1878]" italics="true" pageId="10" pageNumber="9">Neobuthus awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F53EFFF9FE25B079EB91F8BE" bold="true" box="[422,479,1857,1878]" pageId="10" pageNumber="9">
|
||
<taxonomicNameLabel id="A2754319F53EFFF9FE25B079EB91F8BE" box="[422,479,1857,1878]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. Pedipalp (
|
||
<emphasis id="B946FE62F53EFFF9FDD0B078E825F8BD" bold="true" box="[595,619,1856,1877]" pageId="10" pageNumber="9">44</emphasis>
|
||
), ♂‚ (21 mm) paratype. Chela dorsal (
|
||
<emphasis id="B946FE62F53EFFF9FC71B078EE44F8BD" bold="true" box="[1010,1034,1856,1877]" pageId="10" pageNumber="9">45</emphasis>
|
||
), tarsomeres of the second leg (
|
||
<emphasis id="B946FE62F53EFFF9FACCB078EF29F8BD" bold="true" box="[1359,1383,1856,1877]" pageId="10" pageNumber="9">46</emphasis>
|
||
), and trichobothrial pattern (
|
||
<emphasis id="B946FE62F53EFFF9FE1AB061EBFFF886" bold="true" box="[409,433,1881,1902]" pageId="10" pageNumber="9">47</emphasis>
|
||
), ♀ (30 mm) paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D72F8F53FFFF8FF43B2DCE8A2F949" pageId="11" pageNumber="10" startId="11.[192,278,1508,1532]" targetBox="[200,1435,152,1462]" targetPageId="11">
|
||
<paragraph id="8B8D2270F53FFFF8FF43B2DCE8A2F949" blockId="11.[192,1441,1508,1697]" pageId="11" pageNumber="10">
|
||
<emphasis id="B946FE62F53FFFF8FF43B2DCE976FA13" bold="true" box="[192,824,1508,1532]" pageId="11" pageNumber="10">Figures 48–55: Figures 48–49: Movable finger. Figure 48:</emphasis>
|
||
<taxonomicName id="4C3259F3F53FFFF8FCC2B2DEEED3FA13" authority="Hirst, 1911" authorityName="Hirst" authorityYear="1911" box="[833,1181,1510,1531]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="berberensis">
|
||
<emphasis id="B946FE62F53FFFF8FCC2B2DEEE6FFA13" box="[833,1057,1510,1531]" italics="true" pageId="11" pageNumber="10">Neobuthus berberensis</emphasis>
|
||
Hirst, 1911
|
||
</taxonomicName>
|
||
, ♂ (20 mm), Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP.
|
||
<emphasis id="B946FE62F53FFFF8FD71B139E92EF9FE" bold="true" box="[754,864,1537,1559]" pageId="11" pageNumber="10">Figure 49:</emphasis>
|
||
<taxonomicName id="4C3259F3F53FFFF8FCE5B13AEEB7F9FF" authority="(Kraepelin, 1898)" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" box="[870,1273,1538,1559]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F53FFFF8FCE5B13AEE0DF9FF" box="[870,1091,1538,1559]" italics="true" pageId="11" pageNumber="10">Neobuthus ferrugineus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA35F81F53FFFF8FBD0B13AEEBCF9FF" author="KRAEPELIN" box="[1107,1266,1538,1559]" pageId="11" pageNumber="10" pagination="39 - 44" refId="ref13141" refString="KRAEPELIN, K. 1898. Neue Pedipalpen und Scorpione des Hamburger Museums. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 15: 39 - 44." type="journal article" year="1898">Kraepelin, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F53FFFF8FA82B13AEF16F9FF" bold="true" box="[1281,1368,1538,1559]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="A2754319F53FFFF8FA82B13AEF16F9FF" box="[1281,1368,1538,1559]" pageId="11" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP.
|
||
<emphasis id="B946FE62F53FFFF8FB8FB125EB9EF9A6" bold="true" pageId="11" pageNumber="10">
|
||
Figures 50–55:
|
||
<taxonomicName id="4C3259F3F53FFFF8FB37B125EB3DF9A6" authority="(Kraepelin, 1898)" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F53FFFF8FB37B125EFEEF9DB" bold="true" box="[1204,1440,1565,1587]" italics="true" pageId="11" pageNumber="10">Neobuthus ferrugineus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA35F81F53FFFF8FF45B100EB22F9A6" author="KRAEPELIN" box="[198,364,1592,1614]" pageId="11" pageNumber="10" pagination="39 - 44" refId="ref13141" refString="KRAEPELIN, K. 1898. Neue Pedipalpen und Scorpione des Hamburger Museums. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 15: 39 - 44." type="journal article" year="1898">Kraepelin, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2754319F53FFFF8FEFAB101EB9EF9A6" box="[377,464,1593,1614]" pageId="11" pageNumber="10" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B946FE62F53FFFF8FE5EB101E83AF9A6" bold="true" box="[477,628,1593,1615]" pageId="11" pageNumber="10">Figures 50–51:</emphasis>
|
||
Dorsal and ventral views, ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP.
|
||
<emphasis id="B946FE62F53FFFF8FDF3B16CE945F981" bold="true" box="[624,779,1620,1642]" pageId="11" pageNumber="10">Figures 52–53:</emphasis>
|
||
Dorsal and ventral views, ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP.
|
||
<emphasis id="B946FE62F53FFFF8FD3AB148E91BF96D" bold="true" box="[697,853,1648,1670]" pageId="11" pageNumber="10">Figures 54–55:</emphasis>
|
||
Dorsal and ventral views, ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32871FBF53FFFE2FCBDB1FFEB8CFB5B" lastPageId="17" lastPageNumber="16" pageId="11" pageNumber="10" type="description">
|
||
<paragraph id="8B8D2270F53FFFFFFCBDB1FFEB47FD63" blockId="11.[830,1440,1735,1912]" lastBlockId="12.[192,802,199,651]" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10">
|
||
DESCRIPTION. Adult males are
|
||
<quantity id="4CCA8F95F53FFFF8FB0FB1FFEEB1F936" box="[1164,1279,1735,1758]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="11" pageNumber="10" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">20–21 mm</quantity>
|
||
long and adult females are
|
||
<quantity id="4CCA8F95F53FFFF8FC4AB1DEEE0FF915" box="[969,1089,1766,1789]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="11" pageNumber="10" unit="mm" value="28.5" valueMax="30.0" valueMin="27.0">27–30 mm</quantity>
|
||
long. The habitus is shown in
|
||
<figureCitation id="13093EF5F53FFFF8FCBDB03DE988F8F4" box="[830,966,1797,1820]" captionStart="Figures 14–21" captionStartId="8.[192,278,1360,1384]" captionTargetBox="[227,1416,123,1323]" captionTargetId="figure-13@8.[223,1427,120,1333]" captionTargetPageId="8" captionText="Figures 14–21: Figures 14–17: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figures 14–15: Dorsal and ventral views, ♂ (20 mm). Figures 16–17: Dorsal and ventral views, ♀ (30 mm). Figures 18–21: Neobuthus awashensis sp. n.. Figures 18–19: Dorsal and ventral views, ♂ (20 mm) holotype. Figures 20–21: Dorsal and ventral views, ♀ (29 mm) allotype." pageId="11" pageNumber="10">Figs. 18–19</figureCitation>
|
||
(male) and
|
||
<figureCitation id="13093EF5F53FFFF8FBE5B03DEEA3F8F4" box="[1126,1261,1797,1820]" captionStart="Figures 14–21" captionStartId="8.[192,278,1360,1384]" captionTargetBox="[227,1416,123,1323]" captionTargetId="figure-13@8.[223,1427,120,1333]" captionTargetPageId="8" captionText="Figures 14–21: Figures 14–17: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figures 14–15: Dorsal and ventral views, ♂ (20 mm). Figures 16–17: Dorsal and ventral views, ♀ (30 mm). Figures 18–21: Neobuthus awashensis sp. n.. Figures 18–19: Dorsal and ventral views, ♂ (20 mm) holotype. Figures 20–21: Dorsal and ventral views, ♀ (29 mm) allotype." pageId="11" pageNumber="10">Figs. 20–21</figureCitation>
|
||
(females). For position and distribution of trichobothria of pedipalps see
|
||
<figureCitation id="13093EF5F53FFFF8FCE4B07AE9FDF8B1" box="[871,947,1858,1881]" captionStart="Figures 39–47" captionStartId="10.[192,278,1730,1754]" captionTargetBox="[193,1440,205,1693]" captionTargetId="figure-13@10.[192,1440,192,1701]" captionTargetPageId="10" captionText="Figures 39–47: Figures 39–40: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Trochanter and femur of pedipalp (39), and patela and chela of pedipalp (40), ♀ (30 mm). Figures 41–43: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Pedipalp (41), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Pedipalp (42), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Pedipalp (43), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Figures 44–47: Neobuthus awashensis sp. n.. Pedipalp (44), ♂‚ (21 mm) paratype. Chela dorsal (45), tarsomeres of the second leg (46), and trichobothrial pattern (47), ♀ (30 mm) paratype." pageId="11" pageNumber="10">Fig. 47</figureCitation>
|
||
. Trichobothrium
|
||
<emphasis id="B946FE62F53FFFF8FBF3B07AEE33F8B1" box="[1136,1149,1858,1881]" italics="true" pageId="11" pageNumber="10">d</emphasis>
|
||
<subScript id="17B62035F53FFFF8FBFEB074EEC8F8B3" attach="left" box="[1149,1158,1868,1883]" fontSize="6" pageId="11" pageNumber="10">2</subScript>
|
||
of pedipalp femur present on dorsal surface but often smaller than other trichobothria or absent. Trichobothrium
|
||
<emphasis id="B946FE62F538FFFFFDB9B7FFE80BFF36" box="[570,581,199,222]" italics="true" pageId="12" pageNumber="11">d</emphasis>
|
||
<subScript id="17B62035F538FFFFFDC5B7E9E801FF08" attach="left" box="[582,591,209,224]" fontSize="6" pageId="12" pageNumber="11">2</subScript>
|
||
of pedipalp patella either present but often smaller than other trichobothria, or absent. Sexual dimorphism is noticeable. Males are substantially smaller with bristle combs composed of long spiniform setae, whereas females have bristle combs composed of long thin setae. All segments of metasoma and pedipalps are sparsely hirsute, with dark long setae in females (
|
||
<figureCitation id="13093EF5F538FFFFFE37B6A6EBB1FE5D" box="[436,511,414,437]" captionStart="Figures 1–8" captionStartId="5.[192,278,1733,1757]" captionTargetBox="[192,1440,220,1680]" captionTargetId="figure-6@5.[192,1440,200,1704]" captionTargetPageId="5" captionText="Figures 1–8: Figures 1–7: Fifth (or fourth and fifth) metasomal segment and telson. Figures 1–2: Neobuthus berberensis Hirst, 1911, ♀ (1) and ♂ (2), Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figures 3–4: Neobuthus ferrugineus (Kraepelin, 1898) comb. n., ♀ (3), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP and ♂ (4), Somaliland, 15 km near Sheikh, Goolis mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l., FKCP. Figures 5–6: Neobuthus awashensis sp. n., ♀ allotype (5) and ♂ (6) holotype. Figure 7: Butheolus thalassinus Simon, 1882, ♂, Yemen, Ta’izz government, N of Al Makha by road, 13°23'37"N 43°16'22"E, 5 m a.s.l., FKCP. Figure 8: Carapace, Butheolus thalassinus Simon, 1882, ♂, same specimen locality as Figure 7." pageId="12" pageNumber="11">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13093EF5F538FFFFFD8CB6A6E866FE5D" box="[527,552,414,437]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="12" pageNumber="11">35</figureCitation>
|
||
) and shorter spiniform setae in males (
|
||
<figureCitation id="13093EF5F538FFFFFEF6B684EB9FFE3B" box="[373,465,444,467]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="12" pageNumber="11">Figs. 38</figureCitation>
|
||
,
|
||
<figureCitation id="13093EF5F538FFFFFE67B684E84DFE3B" box="[484,515,444,467]" captionStart="Figures 39–47" captionStartId="10.[192,278,1730,1754]" captionTargetBox="[193,1440,205,1693]" captionTargetId="figure-13@10.[192,1440,192,1701]" captionTargetPageId="10" captionText="Figures 39–47: Figures 39–40: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Trochanter and femur of pedipalp (39), and patela and chela of pedipalp (40), ♀ (30 mm). Figures 41–43: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Pedipalp (41), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Pedipalp (42), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Pedipalp (43), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Figures 44–47: Neobuthus awashensis sp. n.. Pedipalp (44), ♂‚ (21 mm) paratype. Chela dorsal (45), tarsomeres of the second leg (46), and trichobothrial pattern (47), ♀ (30 mm) paratype." pageId="12" pageNumber="11">44</figureCitation>
|
||
). Males also have longer pectines than females and are more granulated, with e.g. the chela of pedipalp granulate with carinae in males, and smooth without granules in females. Males are matte, females and juveniles are glossy. Other differences such as metasomal carination are described below.
|
||
</paragraph>
|
||
<paragraph id="8B8D2270F538FFFFFF43B58DE89CFB70" blockId="12.[192,809,693,1176]" pageId="12" pageNumber="11">
|
||
COLORATION. The mesosoma and carapace are dark (brown to black) with orange to yellow spots, but proportions of the colors are quite variable so that some specimens may be described as orange to yellow with brown to black spots. However, the space between the anterior median carinae on the carapace is always dark, and does not appear as an orange to yellow spot (
|
||
<figureCitation id="13093EF5F538FFFFFD7EB455EA90FC4A" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="12" pageNumber="11">Fig. 34</figureCitation>
|
||
). The mesosoma bears a median black stripe and two orange stripes or only a stripe with orange spots surrounded by black. The base color of the metasoma, pedipalps and legs is yellow or orange with dark spots or reticulations. Carinae on the metasoma and on the pedipalps are dark. The fifth metasomal segment is darker than others, with the contrast more profound in males. The chelicerae are yellow with reticulation only in anterior part, and with reddish denticles (
|
||
<figureCitation id="13093EF5F538FFFFFDF5B3B9E88DFB70" box="[630,707,1153,1176]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="12" pageNumber="11">Fig. 34</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8D2270F538FFFFFF43B3F9EB4BFA78" blockId="12.[192,802,1217,1424]" pageId="12" pageNumber="11">
|
||
CARAPACE (
|
||
<figureCitation id="13093EF5F538FFFFFECAB3F9EB9BFB30" box="[329,469,1217,1240]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="12" pageNumber="11">Figs. 34, 37</figureCitation>
|
||
,
|
||
<figureCitation id="13093EF5F538FFFFFE6AB3F9E84AFB30" box="[489,516,1217,1240]" captionStart="Figures 74–91" captionStartId="20.[192,278,1752,1776]" captionTargetBox="[243,1388,188,1729]" captionTargetId="figure-13@20.[216,1418,176,1733]" captionTargetPageId="20" captionText="Figures 74–91: Prosomal and mesosomal characters in Neobuthus and Butheolus. Figures 74–76: Carapace and tergite I of ♂ Neobuthus awashensis sp. n. (74), ♂ Butheolus thalassinus (75), and ♂ B. gallagheri (76), showing granulation, carination, and margins. Figures 77–85: Carapace anterior margin of 9 ♀♀ B. gallagheri. Figures 86–88: Pectines and sternites of ♂ Neobuthus awashensis sp. n. (86), ♂ Butheolus thalassinus (87), and ♂ B. gallagheri (88). Figures 89–91: Higher magnification views of sternite III posterior margins of ♂ Neobuthus awashensis sp. n. (89), ♂ Butheolus thalassinus (90), and ♂ B. gallagheri (91), showing differences in denticle armature. Scale bars: 1 mm in 74–76, 86–88; 0.5 mm in 77–85 (bar in 77 applies to 77–85), 89– 91. All photographs acquired by UV fluorescence imaging. Comparative material: Butheolus gallagheri, ♂, ♀, Oman, wadi above Khor Rhori Beach, E of Taqah, 17°3.22'N 54°25.33'E 18 Oct 1993, leg. G. Lowe, GL; Butheolus thalassinus, ♂ Yemen, Ta'izz, under rock on Euphorbia-Aloe hill 3800 ft, 10 Jan 1951, leg. H. Hoogstraal, WDS." pageId="12" pageNumber="11">74</figureCitation>
|
||
). The surface is densely granulated. Carinae are absent or inconspicuous, only anterior median carinae are well developed. The anterior margin of the carapace is straight, but in males may be slightly convex; it bears 8–11 macrosetae, which are long setae in females and shorter spiniform setae in males.
|
||
</paragraph>
|
||
<paragraph id="8B8D2270F538FFFFFF43B283E9EDFED2" blockId="12.[192,802,1467,1919]" lastBlockId="12.[830,1441,199,314]" pageId="12" pageNumber="11">
|
||
MESOSOMA. The mesosoma bears three carinae, of which the lateral pair are incomplete or absent; in males even the median carina may be less conspicuous. The pectinal tooth count is 15–18 (9x15, 23x16, 14x17, 2x18) in females and 17–21 (1x17, 6x18, 28x19, 27x20, 6x21) in males. The marginal tips of the pectines extend to the proximal half of sternite IV in females, and to the proximal half of sternite V in males. The pectines have three marginal lamellae and seven or eight middle lamellae. The lamellae bear numerous dark setae, three to six on each fulcrum; the setae are shorter in males. Sternites III–VI without carinae, but may be weakly granulated, mainly in males. Sternite VII is more strongly granulated, with granulation apparent even in females; there may be a weakly granulated pair of median carinae, and obsolete lateral carinae indicated by granules. Hemispermatophore with three short, spatulate or laminate basal lobes, and prominent pointed hook (
|
||
<figureCitation id="13093EF5F538FFFFFCC5B61BE9DDFED2" box="[838,915,291,314]" captionStart="Figure 73" captionStartId="19.[192,268,1330,1354]" captionTargetBox="[191,771,198,1302]" captionTargetId="figure-6@19.[192,771,199,1301]" captionTargetPageId="19" captionText="Figure 73: Neobuthus awashensis sp. n. Right hemispermatophore of paratype ♂, viewed from convex side. The structure has been compressed under a cover slip to reveal flat profiles of the basal lobes. Scale bar: 0.2 mm." pageId="12" pageNumber="11">Fig. 73</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8D2270F538FFFFFCBDB662EF57FB66" blockId="12.[830,1441,346,1166]" pageId="12" pageNumber="11">
|
||
METASOMA AND TELSON (
|
||
<figureCitation id="13093EF5F538FFFFFBE6B662EE82FE99" box="[1125,1228,346,369]" captionStart="Figures 1–8" captionStartId="5.[192,278,1733,1757]" captionTargetBox="[192,1440,220,1680]" captionTargetId="figure-6@5.[192,1440,200,1704]" captionTargetPageId="5" captionText="Figures 1–8: Figures 1–7: Fifth (or fourth and fifth) metasomal segment and telson. Figures 1–2: Neobuthus berberensis Hirst, 1911, ♀ (1) and ♂ (2), Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figures 3–4: Neobuthus ferrugineus (Kraepelin, 1898) comb. n., ♀ (3), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP and ♂ (4), Somaliland, 15 km near Sheikh, Goolis mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l., FKCP. Figures 5–6: Neobuthus awashensis sp. n., ♀ allotype (5) and ♂ (6) holotype. Figure 7: Butheolus thalassinus Simon, 1882, ♂, Yemen, Ta’izz government, N of Al Makha by road, 13°23'37"N 43°16'22"E, 5 m a.s.l., FKCP. Figure 8: Carapace, Butheolus thalassinus Simon, 1882, ♂, same specimen locality as Figure 7." pageId="12" pageNumber="11">Figs. 5–6</figureCitation>
|
||
,
|
||
<figureCitation id="13093EF5F538FFFFFB5DB662EF6FFE99" box="[1246,1313,346,369]" captionStart="Figures 95–99" captionStartId="22.[192,278,1384,1408]" captionTargetBox="[214,1386,223,1323]" captionTargetId="figure-13@22.[192,1440,192,1355]" captionTargetPageId="22" captionText="Figures 95–99: Ventral aspect of metasoma and telson in Neobuthus and Butheolus. Figures 95–96: Neobuthus awashensis sp. n., paratype ♂ (95) and ♀ (96). Figures 97–98: Butheolus gallagheri, ♂ (97) and ♀ (98). Figure 99: Butheolus thalassinus, ♂. Scale bars: 1 mm. All photographs acquired by UV fluorescence imaging. Comparative material: see data under caption for Figs. 74–91." pageId="12" pageNumber="11">95–96</figureCitation>
|
||
,
|
||
<figureCitation id="13093EF5F538FFFFFAB0B662EFDEFE99" box="[1331,1424,346,369]" captionStart="Figures 100–104" captionStartId="23.[192,278,1630,1654]" captionTargetBox="[201,1409,236,1545]" captionTargetId="figure-6@23.[192,1440,192,1601]" captionTargetPageId="23" captionText="Figures 100–104: Lateral aspect of metasoma and telson in Neobuthus and Butheolus. Figures 100–101: Neobuthus awashensis sp. n., paratype ♂ (100) and ♀ (101). Figures 102–103: Butheolus gallagheri, ♂ (102) and ♀ (103). Figure 104: Butheolus thalassinus, ♂. Scale bars: 1 mm. All photographs acquired by UV fluorescence imaging. Comparative material: see data under caption for Figs. 74–91." pageId="12" pageNumber="11">100–101</figureCitation>
|
||
). Metasomal segments I–III bear eight or ten carinae; segment IV bears eight carinae that are more complete in males, but in females only the ventrolateral carinae may be complete. Other carinae in females may be indicated only by several granules. Metasomal segment V of both sexes has only ventrolateral carinae, which in their posterior halves bear several lobate granules usually more conspicuous in females. Granules on the ventral surface of metasomal segment V may form an irregular median carina in both sexes. Metasomal segments I–III are densely granulated, with granules of approximately equal size. Granules on segments IV–V are of unequal sizes and unevenly distributed, especially in females. Granulation in females is sparse on all surfaces of segments I–III, and lateral and dorsal surfaces of segment IV (these surfaces more densely granulated in males). The anal arch consists of two lobes in females, three or four lobes in males. All segments are sparsely setose in females or spinose in males; on metasomal segment V there are ca. 35–40 long setae in females or shorter spiniform setae in males. The telson is rather elongate in both sexes. The aculeus is slightly shorter than the vesicle in both sexes. The surface of the telson is bumpy, sparsely hirsute and without a subaculear tubercle, although the angular shape of the vesicle may appear as a protuberance (
|
||
<figureCitation id="13093EF5F538FFFFFBD4B34FEEEFFB66" box="[1111,1185,1143,1166]" captionStart="Figures 1–8" captionStartId="5.[192,278,1733,1757]" captionTargetBox="[192,1440,220,1680]" captionTargetId="figure-6@5.[192,1440,200,1704]" captionTargetPageId="5" captionText="Figures 1–8: Figures 1–7: Fifth (or fourth and fifth) metasomal segment and telson. Figures 1–2: Neobuthus berberensis Hirst, 1911, ♀ (1) and ♂ (2), Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figures 3–4: Neobuthus ferrugineus (Kraepelin, 1898) comb. n., ♀ (3), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP and ♂ (4), Somaliland, 15 km near Sheikh, Goolis mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l., FKCP. Figures 5–6: Neobuthus awashensis sp. n., ♀ allotype (5) and ♂ (6) holotype. Figure 7: Butheolus thalassinus Simon, 1882, ♂, Yemen, Ta’izz government, N of Al Makha by road, 13°23'37"N 43°16'22"E, 5 m a.s.l., FKCP. Figure 8: Carapace, Butheolus thalassinus Simon, 1882, ♂, same specimen locality as Figure 7." pageId="12" pageNumber="11">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13093EF5F538FFFFFB2FB34FEF47FB66" box="[1196,1289,1143,1166]" captionStart="Figures 100–104" captionStartId="23.[192,278,1630,1654]" captionTargetBox="[201,1409,236,1545]" captionTargetId="figure-6@23.[192,1440,192,1601]" captionTargetPageId="23" captionText="Figures 100–104: Lateral aspect of metasoma and telson in Neobuthus and Butheolus. Figures 100–101: Neobuthus awashensis sp. n., paratype ♂ (100) and ♀ (101). Figures 102–103: Butheolus gallagheri, ♂ (102) and ♀ (103). Figure 104: Butheolus thalassinus, ♂. Scale bars: 1 mm. All photographs acquired by UV fluorescence imaging. Comparative material: see data under caption for Figs. 74–91." pageId="12" pageNumber="11">100–101</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8D2270F538FFFFFCBDB397EF28FAB7" blockId="12.[830,1448,1199,1375]" pageId="12" pageNumber="11">LEGS. The tarsomeres bear two rows of macrosetae on the ventral surface and numerous macrosetae on the other surfaces, which on legs I–III form bristle combs. The macrosetae are longer in females. The femur and patella may bear four to six carinae, which however may be obsolete. The femur bears only solitary macrosetae.</paragraph>
|
||
<paragraph id="8B8D2270F538FFFFFCBDB247E9EDF8CD" blockId="12.[830,1447,1407,1829]" pageId="12" pageNumber="11">
|
||
PEDIPALPS. The femur is granulated and bears three to five carinae; the ventroexternal carina is incomplete or absent, the other carinae are granular. The patella is granular, with seven coarsely granular carinae in males, and smooth, without carinae or obsolete carinae in females. The chela is granulate with five carinae, which may be weak and incomplete in males and smooth, without granules and carinae in females. All pedipalps including the trochanter are sparsely hirsute, with dark long setae in females (
|
||
<figureCitation id="13093EF5F538FFFFFBCAB1ABEED3F942" box="[1097,1181,1683,1706]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="12" pageNumber="11">Fig. 35</figureCitation>
|
||
) and shorter spiniform setae in males (
|
||
<figureCitation id="13093EF5F538FFFFFB89B18AEE2BF921" box="[1034,1125,1714,1737]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="12" pageNumber="11">Fig. 38</figureCitation>
|
||
). The movable and fixed fingers bear five to six rows of denticles, with external and internal accessory denticles and three distal denticles (
|
||
<figureCitation id="13093EF5F538FFFFFCC4B036E9DDF8CD" box="[839,915,1806,1829]" captionStart="Figures 48–55" captionStartId="11.[192,278,1508,1532]" captionTargetBox="[200,1435,152,1462]" captionTargetId="figure-6@11.[192,1439,144,1476]" captionTargetPageId="11" captionText="Figures 48–55: Figures 48–49: Movable finger. Figure 48: Neobuthus berberensis Hirst, 1911, ♂ (20 mm), Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figure 49: Neobuthus ferrugineus (Kraepelin, 1898) comb. n., ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Figures 50–55: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Figures 50–51: Dorsal and ventral views, ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 52–53: Dorsal and ventral views, ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Figures 54–55: Dorsal and ventral views, ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP." pageId="12" pageNumber="11">Fig. 49</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8D2270F538FFFCFCBDB070EE4AF8FB" blockId="12.[830,1440,1864,1918]" lastBlockId="15.[830,1441,1727,1903]" lastPageId="15" lastPageNumber="14" pageId="12" pageNumber="11">
|
||
MEASUREMENTS IN MM. Male
|
||
<typeStatus id="54899CD2F538FFFFFB0BB070EEABF8B7" box="[1160,1253,1864,1887]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
. Total length 21; carapace length 2.28, width 2.65; metasoma and telson length 12.4; first metasomal segment length 1.45, width 1.57; second metasomal segment length 1.72, width 1.45; third metasomal segment length 1.80, width 1.45; fourth metasomal segment length 2.12, width 1.40; fifth metasomal segment length 2.78, width 1.35; telson length 2.36; telson width 0.85; pedipalp femur length 1.57, width 0.62; pedipalp patella length 2.15, width 0.85; chela length 2.60; manus width 0.55; movable finger length 1.60.
|
||
</paragraph>
|
||
<caption id="DF4D72F8F539FFFEFF43B1D8E927F8A2" pageId="13" pageNumber="12" startId="13.[192,278,1760,1784]" targetBox="[191,1441,214,1729]" targetPageId="13">
|
||
<paragraph id="8B8D2270F539FFFEFF43B1D8E927F8A2" blockId="13.[192,1441,1760,1867]" pageId="13" pageNumber="12">
|
||
<emphasis id="B946FE62F539FFFEFF43B1D8EB24F910" bold="true" box="[192,362,1760,1784]" pageId="13" pageNumber="12">Figures 56–61:</emphasis>
|
||
<taxonomicName id="4C3259F3F539FFFEFEF2B1DAE94AF91F" authority="(Kraepelin, 1898)" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" box="[369,772,1762,1783]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F539FFFEFEF2B1DAE800F91F" box="[369,590,1762,1783]" italics="true" pageId="13" pageNumber="12">Neobuthus ferrugineus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA35F81F539FFFEFDDDB1DAE8B2F91F" author="KRAEPELIN" box="[606,764,1762,1783]" pageId="13" pageNumber="12" pagination="39 - 44" refId="ref13141" refString="KRAEPELIN, K. 1898. Neue Pedipalpen und Scorpione des Hamburger Museums. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 15: 39 - 44." type="journal article" year="1898">Kraepelin, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F539FFFEFC88B1DAE92CF91F" bold="true" box="[779,866,1762,1783]" pageId="13" pageNumber="12">
|
||
<taxonomicNameLabel id="A2754319F539FFFEFC88B1DAE92CF91F" box="[779,866,1762,1783]" pageId="13" pageNumber="12" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B946FE62F539FFFEFCF3B1DAE992F91F" bold="true" box="[880,988,1762,1784]" pageId="13" pageNumber="12">Figure 56:</emphasis>
|
||
Juvenile at the locality in Fig. 59.
|
||
<emphasis id="B946FE62F539FFFEFAB6B1DAEFEEF91F" bold="true" box="[1333,1440,1762,1784]" pageId="13" pageNumber="12">Figure 57:</emphasis>
|
||
Male at the locality in Fig. 59.
|
||
<emphasis id="B946FE62F539FFFEFE72B1C5E812F8FA" bold="true" box="[497,604,1789,1811]" pageId="13" pageNumber="12">Figure 58:</emphasis>
|
||
Female at the locality in Fig. 59.
|
||
<emphasis id="B946FE62F539FFFEFC24B1C5EE5CF8FA" bold="true" box="[935,1042,1789,1811]" pageId="13" pageNumber="12">Figure 59:</emphasis>
|
||
Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l.
|
||
<emphasis id="B946FE62F539FFFEFE57B021E80DF8C6" bold="true" box="[468,579,1817,1839]" pageId="13" pageNumber="12">Figure 60:</emphasis>
|
||
Female at the locality in Djibouti, near Goubetto, ca. 25 km S of Gulf of Tadjura, 10°23.5'N 42°55.5'E.
|
||
<emphasis id="B946FE62F539FFFEFE11B00DEBB2F8A2" bold="true" box="[402,508,1845,1867]" pageId="13" pageNumber="12">Figure 61:</emphasis>
|
||
Male at the same locality in Djibouti.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D72F8F53AFFFDFF43B00BEE41F88B" pageId="14" pageNumber="13" startId="14.[192,278,1843,1867]" targetBox="[213,1417,191,1814]" targetPageId="14">
|
||
<paragraph id="8B8D2270F53AFFFDFF43B00BEE41F88B" blockId="14.[192,1440,1843,1891]" pageId="14" pageNumber="13">
|
||
<emphasis id="B946FE62F53AFFFDFF43B00BEB24F8A3" bold="true" box="[192,362,1843,1867]" pageId="14" pageNumber="13">Figures 62–64:</emphasis>
|
||
<taxonomicName id="4C3259F3F53AFFFDFEF0B00EE948F8A3" authority="(Kraepelin, 1898)" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" box="[371,774,1846,1867]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F53AFFFDFEF0B00EE81EF8A3" box="[371,592,1846,1867]" italics="true" pageId="14" pageNumber="13">Neobuthus ferrugineus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA35F81F53AFFFDFDDCB00EE8B1F8A3" author="KRAEPELIN" box="[607,767,1846,1867]" pageId="14" pageNumber="13" pagination="39 - 44" refId="ref13141" refString="KRAEPELIN, K. 1898. Neue Pedipalpen und Scorpione des Hamburger Museums. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 15: 39 - 44." type="journal article" year="1898">Kraepelin, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F53AFFFDFC8CB00DE926F8A2" bold="true" box="[783,872,1845,1866]" pageId="14" pageNumber="13">
|
||
<taxonomicNameLabel id="A2754319F53AFFFDFC8CB00DE926F8A2" box="[783,872,1845,1866]" pageId="14" pageNumber="13" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B946FE62F53AFFFDFCF4B00DE9ADF8A2" bold="true" box="[887,995,1845,1867]" pageId="14" pageNumber="13">Figure 62:</emphasis>
|
||
Male at the locality in Figs. 63–64.
|
||
<emphasis id="B946FE62F53AFFFDFAD0B00DEB4AF88B" bold="true" pageId="14" pageNumber="13">Figures 63–64:</emphasis>
|
||
Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D72F8F53BFFFCFF43B168E82CF972" pageId="15" pageNumber="14" startId="15.[192,278,1616,1640]" targetBox="[191,1441,191,1588]" targetPageId="15">
|
||
<paragraph id="8B8D2270F53BFFFCFF43B168E82CF972" blockId="15.[192,1440,1616,1690]" pageId="15" pageNumber="14">
|
||
<emphasis id="B946FE62F53BFFFCFF43B168EB26F980" bold="true" box="[192,360,1616,1640]" pageId="15" pageNumber="14">Figures 65–66:</emphasis>
|
||
<taxonomicName id="4C3259F3F53BFFFCFEECB16BE8B2F980" authority="(Kraepelin, 1898)" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" box="[367,764,1619,1640]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F53BFFFCFEECB16BE804F980" box="[367,586,1619,1640]" italics="true" pageId="15" pageNumber="14">Neobuthus ferrugineus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA35F81F53BFFFCFDD4B16BE8BBF980" author="KRAEPELIN" box="[599,757,1619,1640]" pageId="15" pageNumber="14" pagination="39 - 44" refId="ref13141" refString="KRAEPELIN, K. 1898. Neue Pedipalpen und Scorpione des Hamburger Museums. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 15: 39 - 44." type="journal article" year="1898">Kraepelin, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F53BFFFCFC81B16AE917F98F" bold="true" box="[770,857,1618,1639]" pageId="15" pageNumber="14">
|
||
<taxonomicNameLabel id="A2754319F53BFFFCFC81B16AE917F98F" box="[770,857,1618,1639]" pageId="15" pageNumber="14" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B946FE62F53BFFFCFCE5B16AE99EF98F" bold="true" box="[870,976,1618,1640]" pageId="15" pageNumber="14">Figure 65:</emphasis>
|
||
Female from the locality in Fig. 66. The female was not photographed directly at the place of capture, the substrate is sand from another locality.
|
||
<emphasis id="B946FE62F53BFFFCFB0AB153EEB8F968" bold="true" box="[1161,1270,1643,1665]" pageId="15" pageNumber="14">Figure 66:</emphasis>
|
||
Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l..
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D2270F53BFFE2FCBDB023EAB8FE5D" blockId="15.[830,1441,1727,1903]" lastBlockId="17.[192,802,199,437]" lastPageId="17" lastPageNumber="16" pageId="15" pageNumber="14">
|
||
Female
|
||
<typeStatus id="54899CD2F53BFFFCFC14B023E9BEF8DA" box="[919,1008,1819,1842]" pageId="15" pageNumber="14" type="allotype">allotype</typeStatus>
|
||
. Total length 29; carapace length 3.05, width 3.65; metasoma and telson length 16.6; first metasomal segment length 1.97, width 2.17; second metasomal segment length 2.28, width 1.95; third metasomal segment length 2.42, width 1.92; fourth metasomal segment length 2.95, width 1.85; fifth metasomal segment length 3.67, width 1.85; telson length 3.30; telson width 1.15; pedipalp femur length 2.00, width 0.83; pedipalp patella length 2.65, width 1.15; chela length 3.37; manus width 0.80; movable finger length 2.33.
|
||
</paragraph>
|
||
<caption id="DF4D72F8F524FFE3FF43B072E871F844" pageId="16" pageNumber="15" startId="16.[192,278,1866,1890]" targetBox="[227,1398,172,1836]" targetPageId="16">
|
||
<paragraph id="8B8D2270F524FFE3FF43B072E871F844" blockId="16.[192,1440,1866,1964]" pageId="16" pageNumber="15">
|
||
<emphasis id="B946FE62F524FFE3FF43B072EB25F88A" bold="true" box="[192,363,1866,1890]" pageId="16" pageNumber="15">Figures 67–71:</emphasis>
|
||
<taxonomicName id="4C3259F3F524FFE3FEF6B074E81CF889" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" box="[373,594,1868,1889]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F524FFE3FEF6B074E81CF889" box="[373,594,1868,1889]" italics="true" pageId="16" pageNumber="15">Neobuthus awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F524FFE3FDDFB074E974F888" bold="true" box="[604,826,1867,1889]" pageId="16" pageNumber="15">
|
||
<taxonomicNameLabel id="A2754319F524FFE3FDDFB074E8DBF889" box="[604,661,1868,1889]" pageId="16" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
|
||
Figures 67–68:
|
||
</emphasis>
|
||
Females paratypes at the type locality.
|
||
<emphasis id="B946FE62F524FFE3FB4EB073EF12F888" bold="true" box="[1229,1372,1867,1889]" pageId="16" pageNumber="15">Figure 69–70.</emphasis>
|
||
Males paratypes at the type locality.
|
||
<emphasis id="B946FE62F524FFE3FE6AB05CE81AF891" bold="true" box="[489,596,1892,1914]" pageId="16" pageNumber="15">Figure 71:</emphasis>
|
||
Ethiopia, Awash, Metahara env., 08°54'N 39°54'E, 960-1050 m a.s.l. Type locality of
|
||
<taxonomicName id="4C3259F3F524FFE3FF41B045EBD3F87A" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" box="[194,413,1917,1938]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F524FFE3FF41B045EBD3F87A" box="[194,413,1917,1938]" italics="true" pageId="16" pageNumber="15">Neobuthus awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F524FFE3FE27B046EB93F87B" bold="true" box="[420,477,1918,1939]" pageId="16" pageNumber="15">
|
||
<taxonomicNameLabel id="A2754319F524FFE3FE27B046EB93F87B" box="[420,477,1918,1939]" pageId="16" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3259F3F524FFE3FD92B045E912F87B" authority="Kovarik, 2011" authorityName="Kovarik" authorityYear="2011" box="[529,860,1917,1939]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="awashensis">
|
||
<emphasis id="B946FE62F524FFE3FD92B045E889F87A" box="[529,711,1917,1938]" italics="true" pageId="16" pageNumber="15">Buthus awashensis</emphasis>
|
||
Kovařík, 2011
|
||
</taxonomicName>
|
||
together with
|
||
<taxonomicName id="4C3259F3F524FFE3FC6DB045EF3BF87B" authority="Birula, 1903" authorityName="Birula" authorityYear="1903" box="[1006,1397,1917,1939]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="abyssinicus">
|
||
<emphasis id="B946FE62F524FFE3FC6DB045EEBDF87A" box="[1006,1267,1917,1938]" italics="true" pageId="16" pageNumber="15">Compsobuthus abyssinicus</emphasis>
|
||
Birula, 1903
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3259F3F524FFE3FF43B0AEE875F844" authority="(Ehrenberg, 1828)" baseAuthorityName="Ehrenberg" baseAuthorityYear="1828" box="[192,571,1942,1964]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="liosoma">
|
||
<emphasis id="B946FE62F524FFE3FF43B0AEEBCCF843" box="[192,386,1942,1963]" italics="true" pageId="16" pageNumber="15">Parabuthus liosoma</emphasis>
|
||
(Ehrenberg, 1828)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D2270F525FFE2FF43B6E3EB8CFB5B" blockId="17.[192,803,475,1204]" pageId="17" pageNumber="16">
|
||
AFFINITIES. The described features distinguish
|
||
<taxonomicName id="4C3259F3F525FFE2FD50B6E3EB3DFDF9" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F525FFE2FD50B6E3EB3DFDF9" italics="true" pageId="17" pageNumber="16">Neobuthus awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FE01B6C1EB8BFDF9" bold="true" box="[386,453,505,529]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FE01B6C1EB8BFDF9" box="[386,453,505,529]" pageId="17" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from all other species of the genus. In contrast to
|
||
<taxonomicName id="4C3259F3F525FFE2FE2AB520E80BFDC7" authorityName="Hirst" authorityYear="1911" box="[425,581,536,559]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="berberensis">
|
||
<emphasis id="B946FE62F525FFE2FE2AB520E80BFDC7" box="[425,581,536,559]" italics="true" pageId="17" pageNumber="16">N. berberensis</emphasis>
|
||
</taxonomicName>
|
||
the entire pedipalps are sparsely hirsute with dark setae (females,
|
||
<figureCitation id="13093EF5F525FFE2FD2DB50FE8B4FDA6" box="[686,762,567,590]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="17" pageNumber="16">Fig. 35</figureCitation>
|
||
) or spiniform setae (males,
|
||
<figureCitation id="13093EF5F525FFE2FE53B56EE86DFD85" box="[464,547,598,621]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="17" pageNumber="16">Fig. 38</figureCitation>
|
||
) and there are one or more dark spots (
|
||
<figureCitation id="13093EF5F525FFE2FE07B54CE846FD63" box="[388,520,628,651]" captionStart="Figures 39–47" captionStartId="10.[192,278,1730,1754]" captionTargetBox="[193,1440,205,1693]" captionTargetId="figure-13@10.[192,1440,192,1701]" captionTargetPageId="10" captionText="Figures 39–47: Figures 39–40: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Trochanter and femur of pedipalp (39), and patela and chela of pedipalp (40), ♀ (30 mm). Figures 41–43: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Pedipalp (41), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Pedipalp (42), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Pedipalp (43), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Figures 44–47: Neobuthus awashensis sp. n.. Pedipalp (44), ♂‚ (21 mm) paratype. Chela dorsal (45), tarsomeres of the second leg (46), and trichobothrial pattern (47), ♀ (30 mm) paratype." pageId="17" pageNumber="16">Figs. 44, 47</figureCitation>
|
||
), whereas
|
||
<taxonomicName id="4C3259F3F525FFE2FD00B54CE96FFD63" authorityName="Hirst" authorityYear="1911" box="[643,801,628,651]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="berberensis">
|
||
<emphasis id="B946FE62F525FFE2FD00B54CE96FFD63" box="[643,801,628,651]" italics="true" pageId="17" pageNumber="16">N. berberensis</emphasis>
|
||
</taxonomicName>
|
||
has the entire femur of pedipalp yellow without dark spots and with only several white setae (
|
||
<figureCitation id="13093EF5F525FFE2FD0CB58AE958FD21" box="[655,790,690,713]" captionStart="Figures 39–47" captionStartId="10.[192,278,1730,1754]" captionTargetBox="[193,1440,205,1693]" captionTargetId="figure-13@10.[192,1440,192,1701]" captionTargetPageId="10" captionText="Figures 39–47: Figures 39–40: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Trochanter and femur of pedipalp (39), and patela and chela of pedipalp (40), ♀ (30 mm). Figures 41–43: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Pedipalp (41), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Pedipalp (42), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Pedipalp (43), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Figures 44–47: Neobuthus awashensis sp. n.. Pedipalp (44), ♂‚ (21 mm) paratype. Chela dorsal (45), tarsomeres of the second leg (46), and trichobothrial pattern (47), ♀ (30 mm) paratype." pageId="17" pageNumber="16">Figs. 39–40</figureCitation>
|
||
). The pubescence on trochanter and femur of pedipalps is the same in
|
||
<taxonomicName id="4C3259F3F525FFE2FECAB5D7EBADFCEE" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" box="[329,483,751,774]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F525FFE2FECAB5D7EBADFCEE" box="[329,483,751,774]" italics="true" pageId="17" pageNumber="16">N. ferrugineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FE6DB5D6E801FCEE" bold="true" box="[494,591,750,774]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FE6DB5D6E801FCEE" box="[494,591,750,774]" pageId="17" pageNumber="16" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3259F3F525FFE2FD04B5D7E96CFCEE" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" box="[647,802,751,774]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F525FFE2FD04B5D7E96CFCEE" box="[647,802,751,774]" italics="true" pageId="17" pageNumber="16">N. awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FF41B435EB4DFCCD" bold="true" box="[194,259,781,805]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FF41B435EB4DFCCD" box="[194,259,781,805]" pageId="17" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but the latter species is darker, which is best apparent from the coloration of the femur of pedipalp and the carapace; in
|
||
<taxonomicName id="4C3259F3F525FFE2FE2BB473E80BFC8A" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" box="[424,581,843,866]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F525FFE2FE2BB473E80BFC8A" box="[424,581,843,866]" italics="true" pageId="17" pageNumber="16">N. ferrugineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FDCDB472E8FDFC8A" bold="true" box="[590,691,842,866]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FDCDB472E8FDFC8A" box="[590,691,842,866]" pageId="17" pageNumber="16" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the space between the anterior median carinae on the carapace always has an orange to yellow spot (
|
||
<figureCitation id="13093EF5F525FFE2FDDDB4B0E8E0FC77" box="[606,686,904,927]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="17" pageNumber="16">Fig. 28</figureCitation>
|
||
), which is absent in
|
||
<taxonomicName id="4C3259F3F525FFE2FEB3B49FEB80FC56" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" box="[304,462,935,958]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F525FFE2FEB3B49FEB80FC56" box="[304,462,935,958]" italics="true" pageId="17" pageNumber="16">N. awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FE59B49EE855FC56" bold="true" box="[474,539,934,958]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FE59B49EE855FC56" box="[474,539,934,958]" pageId="17" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13093EF5F525FFE2FDADB49FE8CEFC56" box="[558,640,935,958]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="17" pageNumber="16">Fig. 34</figureCitation>
|
||
). Males of
|
||
<taxonomicName id="4C3259F3F525FFE2FC8AB49FEB75FC35" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F525FFE2FC8AB49FEB75FC35" italics="true" pageId="17" pageNumber="16">N. ferrugineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FEC4B4FDEBE0FC35" bold="true" box="[327,430,965,989]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FEC4B4FDEBE0FC35" box="[327,430,965,989]" pageId="17" pageNumber="16" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
have spiniform setae/macrosetae shorter than
|
||
<taxonomicName id="4C3259F3F525FFE2FECCB4DCEBA3FC13" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" box="[335,493,996,1019]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F525FFE2FECCB4DCEBA3FC13" box="[335,493,996,1019]" italics="true" pageId="17" pageNumber="16">N. awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FE7BB4DCE877FC14" bold="true" box="[504,569,996,1020]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FE7BB4DCE877FC14" box="[504,569,996,1020]" pageId="17" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13093EF5F525FFE2FDCFB4DCE8D0FC13" box="[588,670,996,1019]" captionStart="Figures 22–38" captionStartId="9.[192,278,1515,1539]" captionTargetBox="[192,1440,192,1484]" captionTargetId="figure-6@9.[192,1440,192,1486]" captionTargetPageId="9" captionText="Figures 22–38: Figures 22–27: Neobuthus berberensis Hirst, 1911, Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l. Carapace (22), trochanter and femur of pedipalp (23), and lateral eyes (24), ♀ (30 mm). Lateral eyes (25), carapace (26), and trochanter and femur of pedipalp (27), ♂ (20 mm). Figures 28–33: Neobuthus ferrugineus (Kraepelin, 1898) comb. n.. Carapace (28) and trochanter and femur of pedipalp (30), ♀ (29 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (29), ♀ (28 mm), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP. Carapace (31) and trochanter and femur of pedipalp (33), ♂ (21 mm), Somaliland, near Berbera, 10°15'30.5"N 45°06'04.2"E, 376 m a.s.l., FKCP. Trochanter and femur of pedipalp (32), ♂ (21 mm), Somaliland, 15 km near Sheikh, Goolis Mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l. FKCP. Figures 34–38: Neobuthus awashensis sp. n.. Carapace (34), trochanter and femur of pedipalp (35), and lateral eyes (36), ♀ (30 mm) paratype. Carapace (37) and trochanter and femur of pedipalp (38), ♂ (21 mm) paratype." pageId="17" pageNumber="16">Fig. 33</figureCitation>
|
||
vs. 38).
|
||
<taxonomicName id="4C3259F3F525FFE2FC8AB4DCEB74FBF2" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F525FFE2FC8AB4DCEB74FBF2" italics="true" pageId="17" pageNumber="16">N. awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FEC3B33AEB32FBF2" bold="true" box="[320,380,1026,1050]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FEC3B33AEB32FBF2" box="[320,380,1026,1050]" pageId="17" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also differs from
|
||
<taxonomicName id="4C3259F3F525FFE2FDBFB33BE898FBF2" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" box="[572,726,1027,1050]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F525FFE2FDBFB33BE898FBF2" box="[572,726,1027,1050]" italics="true" pageId="17" pageNumber="16">N. ferrugineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FD5EB33AEA9BFBD1" bold="true" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FD5EB33AEA9BFBD1" pageId="17" pageNumber="16" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
morphometrically, in having a longer and narrower femur of pedipalp. The telson of females of
|
||
<taxonomicName id="4C3259F3F525FFE2FC8AB378EB75FB9E" authorityName="Kovařík & Lowe" authorityYear="2012" baseAuthorityName="Kraepelin" baseAuthorityYear="1898" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="ferrugineus" status="comb. nov.">
|
||
<emphasis id="B946FE62F525FFE2FC8AB378EB75FB9E" italics="true" pageId="17" pageNumber="16">N. ferrugineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FEC4B366EBE2FB9E" bold="true" box="[327,428,1118,1142]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FEC4B366EBE2FB9E" box="[327,428,1118,1142]" pageId="17" pageNumber="16" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3259F3F525FFE2FE69B367E8C4FB9E" authorityName="Hirst" authorityYear="1911" box="[490,650,1119,1142]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="berberensis">
|
||
<emphasis id="B946FE62F525FFE2FE69B367E8C4FB9E" box="[490,650,1119,1142]" italics="true" pageId="17" pageNumber="16">N. berberensis</emphasis>
|
||
</taxonomicName>
|
||
have a more bulky vesicle compared to females of
|
||
<taxonomicName id="4C3259F3F525FFE2FDE2B346E8B4FB7D" authority="Kovařík & Lowe, 2012" authorityName="Kovařík & Lowe" authorityYear="2012" box="[609,762,1150,1173]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="awashensis" status="sp. nov.">
|
||
<emphasis id="B946FE62F525FFE2FDE2B346E8B4FB7D" box="[609,762,1150,1173]" italics="true" pageId="17" pageNumber="16">N. awashensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946FE62F525FFE2FC80B345EA9BFB5C" bold="true" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="A2754319F525FFE2FC80B345EA9BFB5C" pageId="17" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13093EF5F525FFE2FF66B3A4EB06FB5B" box="[229,328,1180,1203]" captionStart="Figures 1–8" captionStartId="5.[192,278,1733,1757]" captionTargetBox="[192,1440,220,1680]" captionTargetId="figure-6@5.[192,1440,200,1704]" captionTargetPageId="5" captionText="Figures 1–8: Figures 1–7: Fifth (or fourth and fifth) metasomal segment and telson. Figures 1–2: Neobuthus berberensis Hirst, 1911, ♀ (1) and ♂ (2), Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figures 3–4: Neobuthus ferrugineus (Kraepelin, 1898) comb. n., ♀ (3), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP and ♂ (4), Somaliland, 15 km near Sheikh, Goolis mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l., FKCP. Figures 5–6: Neobuthus awashensis sp. n., ♀ allotype (5) and ♂ (6) holotype. Figure 7: Butheolus thalassinus Simon, 1882, ♂, Yemen, Ta’izz government, N of Al Makha by road, 13°23'37"N 43°16'22"E, 5 m a.s.l., FKCP. Figure 8: Carapace, Butheolus thalassinus Simon, 1882, ♂, same specimen locality as Figure 7." pageId="17" pageNumber="16">Figs. 1, 3</figureCitation>
|
||
vs.
|
||
<figureCitation id="13093EF5F525FFE2FEF7B3A4EBFDFB5B" box="[372,435,1180,1203]" captionStart="Figures 1–8" captionStartId="5.[192,278,1733,1757]" captionTargetBox="[192,1440,220,1680]" captionTargetId="figure-6@5.[192,1440,200,1704]" captionTargetPageId="5" captionText="Figures 1–8: Figures 1–7: Fifth (or fourth and fifth) metasomal segment and telson. Figures 1–2: Neobuthus berberensis Hirst, 1911, ♀ (1) and ♂ (2), Somaliland, Berbera env., 10°22.8'N 45°02.2'E, 107 m a.s.l., FKCP. Figures 3–4: Neobuthus ferrugineus (Kraepelin, 1898) comb. n., ♀ (3), Somaliland, near Sheikh, 09°46.1'N 45°17.5'E, 1329 m a.s.l., FKCP and ♂ (4), Somaliland, 15 km near Sheikh, Goolis mts., 09°58.9'N 45°10.3'E, 1247 m a.s.l., FKCP. Figures 5–6: Neobuthus awashensis sp. n., ♀ allotype (5) and ♂ (6) holotype. Figure 7: Butheolus thalassinus Simon, 1882, ♂, Yemen, Ta’izz government, N of Al Makha by road, 13°23'37"N 43°16'22"E, 5 m a.s.l., FKCP. Figure 8: Carapace, Butheolus thalassinus Simon, 1882, ♂, same specimen locality as Figure 7." pageId="17" pageNumber="16">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |