295 lines
32 KiB
XML
295 lines
32 KiB
XML
<document id="FD5AEA66F9BFB75D323B8566CAFC2CCB" ID-DOI="10.1080/00222930310001613575" ID-ISSN="1464-5262" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629850176377" checkinUser="felipe" docAuthor="Clausen, Martha Weis" docDate="2004" docId="038587CDFF919A19CFB6FE7140AFFD75" docLanguage="en" docName="JNatHist.38.15.1851-1880.pdf" docOrigin="Journal of Natural History 38 (15)" docSource="http://dx.doi.org/10.1080/00222930310001613575" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Libyodrilus maryae Clausen, 2004, n. sp." docType="treatment" docVersion="2" lastPageNumber="1864" masterDocId="FFBCFFB5FF9D9A17CE4FFFAF4372FFC9" masterDocTitle="Description of seven new species of Libyodrilus Beddard from Cameroon, with a key to the species of the genus (Oligochaeta: Eudrilidae)" masterLastPageNumber="1880" masterPageNumber="1851" pageNumber="1862" updateTime="1699120809012" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="98398B1D7737D3B973EF28BF62D472F0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BA9CF58F3155FF0F7582FD60893BA6E0">
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<mods:title id="FE63E3227D874A70CE56C22F2B34569A">Description of seven new species of Libyodrilus Beddard from Cameroon, with a key to the species of the genus (Oligochaeta: Eudrilidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="6A04932DCCC7F3EDA4BCF2123D8292B9" type="personal">
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<mods:role id="4A61C291998F419C6438610905BA257D">
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<mods:roleTerm id="EE866437C3C9C4562497C3942ECDFDBC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="27DEE3C6745A07FC019FD7609816F787">Clausen, Martha Weis</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E404A568924113927E961E3EEB73AF31">text</mods:typeOfResource>
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<mods:titleInfo id="452D6A75EC71D7FD40EA4837A7F1F5DC">
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<mods:title id="74684D63AA536E6D332AA00E4D6C0B81">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="B2DDBDBE74A9A122D8C0F981246EA377">
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<mods:date id="84D24CD201C87A381ADE3483788E8D8D">2004</mods:date>
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<mods:detail id="E7843F138CCD91040E8F37E5F422F01D" type="pubDate">
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<mods:number id="6C6876E2DADC1F42228197E9D75AE407">2004-08-31</mods:number>
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</mods:detail>
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<mods:detail id="B9BFE4546FC0AEBBFC7D9D99B0E40224" type="volume">
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<mods:number id="CBB95D6D10E29CA57B9C613289AA354C">38</mods:number>
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</mods:detail>
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<mods:detail id="38EAA347343088CA9D00CCC7B983B0E1" type="issue">
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<mods:number id="A00BD29A5468E4F1745A24ACD9E8203C">15</mods:number>
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</mods:detail>
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<mods:extent id="E16AC31AE2B01A5A849905EF1F3F1C8A" unit="page">
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<mods:start id="F7DAFEBC9748AB277F3DD210DE269A23">1851</mods:start>
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||
<mods:end id="35850D5CD8667075453CDF34C376A3E9">1880</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="37F2D3DAAA388C2AAAC3473957D0BDC2">
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<mods:url id="3A4C1DAF8B7BAC76240F11B1B5977162">http://dx.doi.org/10.1080/00222930310001613575</mods:url>
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</mods:location>
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<mods:classification id="0B306A52D3C57CB9C2E6034E7C3EFFAC">journal article</mods:classification>
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<mods:identifier id="847A782CC933B92D1EF83BEF402E4CA5" type="DOI">10.1080/00222930310001613575</mods:identifier>
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<mods:identifier id="1BCABC2FD7ECFBA4D805539286AF6022" type="ISSN">1464-5262</mods:identifier>
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</mods:mods>
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<treatment id="038587CDFF919A19CFB6FE7140AFFD75" LSID="urn:lsid:plazi:treatment:038587CDFF919A19CFB6FE7140AFFD75" httpUri="http://treatment.plazi.org/id/038587CDFF919A19CFB6FE7140AFFD75" lastPageId="14" lastPageNumber="1864" pageId="12" pageNumber="1862">
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<subSubSection id="C3366550FF919A1BCFB6FE714075FDDF" pageId="12" pageNumber="1862" type="nomenclature">
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<paragraph id="8B9336DBFF919A1BCFB6FE714052FE3F" blockId="12.[505,800,478,502]" box="[505,800,478,502]" pageId="12" pageNumber="1862">
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<heading id="D0DB81B7FF919A1BCFB6FE714052FE3F" box="[505,800,478,502]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="1862" reason="5">
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<taxonomicName id="4C2C4D58FF919A1BCFB6FE7141A6FE3F" ID-CoL="8LDDW" authorityName="Clausen" authorityYear="2004" box="[505,724,478,502]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="1862" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
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<emphasis id="B958EAC9FF919A1BCFB6FE7141A6FE3F" bold="true" box="[505,724,478,502]" italics="true" pageId="12" pageNumber="1862">Libyodrilus maryae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26B57B2FF919A1BCC90FE714052FE3F" box="[735,800,478,502]" pageId="12" pageNumber="1862" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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<paragraph id="8B9336DBFF919A1BCC5EFE514075FDDF" blockId="12.[529,775,510,534]" box="[529,775,510,534]" pageId="12" pageNumber="1862">(figures 6B, 8A, 10D)</paragraph>
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</subSubSection>
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||
<subSubSection id="C3366550FF919A19CED5FDE5407BFECC" lastPageId="14" lastPageNumber="1864" pageId="12" pageNumber="1862" type="materials_examined">
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<paragraph id="8B9336DBFF919A1BCED5FDE54203FDAB" blockId="12.[154,1111,586,740]" box="[154,369,586,610]" pageId="12" pageNumber="1862">
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||
<emphasis id="B958EAC9FF919A1BCED5FDE54203FDAB" box="[154,369,586,610]" italics="true" pageId="12" pageNumber="1862">Material examined</emphasis>
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</paragraph>
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<paragraph id="8B9336DBFF919A1BCE8DFDC542ABFD2D" blockId="12.[154,1111,586,740]" pageId="12" pageNumber="1862">
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<emphasis id="B958EAC9FF919A1BCE8DFDC54202FD4B" box="[194,368,618,642]" italics="true" pageId="12" pageNumber="1862">Type material.</emphasis>
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<materialsCitation id="3B443C86FF919A1BCFE5FDC542A7FD2D" collectingDate="1966-01-12" collectionCode="ZMUC" collectorName="M. W. Clausen" country="Cameroon" location="Biwoume River" pageId="12" pageNumber="1862" specimenCode="OLI-5, OLI-6" specimenCount="2" typeStatus="holotype">
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<collectingCountry id="F33B764BFF919A1BCFE5FDC54157FD4B" box="[426,549,618,642]" name="Cameroon" pageId="12" pageNumber="1862">Cameroon</collectingCountry>
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, just
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<locationDeviation id="9A212FF9FF919A1BCC38FDC5407EFD4B" box="[631,780,618,642]" location="Akom" pageId="12" pageNumber="1862">E of Akom</locationDeviation>
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II, 2
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<emphasis id="B958EAC9FF919A1BCD16FDC54011FD4B" bold="true" box="[857,867,618,642]" pageId="12" pageNumber="1862">‡</emphasis>
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51
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<emphasis id="B958EAC9FF919A1BCD30FDC640F7FD4B" bold="true" box="[895,901,617,642]" pageId="12" pageNumber="1862">’</emphasis>
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N, 10
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<emphasis id="B958EAC9FF919A1BCD81FDC540AAFD4B" bold="true" box="[974,984,618,642]" pageId="12" pageNumber="1862">‡</emphasis>
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54
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<emphasis id="B958EAC9FF919A1BCDBBFDC64088FD4B" bold="true" box="[1012,1018,617,642]" pageId="12" pageNumber="1862">’</emphasis>
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E, near
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<location id="8EF36000FF919A1BCED5FD254227FD6B" LSID="urn:lsid:plazi:treatment:038587CDFF919A19CFB6FE7140AFFD75:8EF36000FF919A1BCED5FD254227FD6B" box="[154,341,650,674]" country="Cameroon" name="Biwoume River" pageId="12" pageNumber="1862">Biwoume River</location>
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, clayey to sandy soil in slough,
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<date id="FF92101BFF919A1BCCA4FD2540C7FD6B" box="[747,949,650,674]" pageId="12" pageNumber="1862" value="1966-01-12">
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<collectingDate id="EFD6E9F3FF919A1BCCA4FD2540C7FD6B" box="[747,949,650,674]" pageId="12" pageNumber="1862" value="1966-01-12">12 January 1966</collectingDate>
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</date>
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, coll.
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<collectorName id="26D9530DFF919A1BCA4AFD254387FD0A" pageId="12" pageNumber="1862">M. W. Clausen</collectorName>
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(
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<typeStatus id="54978879FF919A1BCF47FD044203FD0A" box="[264,369,683,707]" pageId="12" pageNumber="1862" type="holotype">holotype</typeStatus>
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||
: one clitellate,
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<collectionCode id="ED3DAE1EFF919A1BCC6BFD04410BFD0A" box="[548,633,683,707]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="12" pageNumber="1862">ZMUC</collectionCode>
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<specimenCode id="DB8A9EA0FF919A1BCCCBFD0441BCFD0A" box="[644,718,683,707]" collectionCode="OLI-" pageId="12" pageNumber="1862">OLI-5</specimenCode>
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;
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<typeStatus id="54978879FF919A1BCC92FD04403DFD0A" box="[733,847,683,707]" pageId="12" pageNumber="1862" type="paratype">paratypes</typeStatus>
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: three clitellates, three aclitellates,
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<collectionCode id="ED3DAE1EFF919A1BCF6DFD634205FD2D" box="[290,375,716,740]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="12" pageNumber="1862">ZMUC</collectionCode>
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<specimenCode id="DB8A9EA0FF919A1BCFCDFD6342BFFD2D" box="[386,461,716,740]" collectionCode="OLI-" pageId="12" pageNumber="1862">OLI-6</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B9336DBFF919A1BCED5FC884289FCF6" blockId="12.[154,1111,807,1458]" box="[154,507,807,831]" pageId="12" pageNumber="1862">
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<emphasis id="B958EAC9FF919A1BCED5FC88420AFCF6" box="[154,376,807,831]" italics="true" pageId="12" pageNumber="1862">External characters</emphasis>
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(figure 8A)
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</paragraph>
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<paragraph id="8B9336DBFF919A1BCE8DFCE841D1FB07" blockId="12.[154,1111,807,1458]" pageId="12" pageNumber="1862">
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<typeStatus id="54978879FF919A1BCE8DFCE84248FC94" box="[194,314,839,863]" pageId="12" pageNumber="1862" type="holotype">HOLOTYPE</typeStatus>
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: length
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<quantity id="4CD49B3EFF919A1BCFEAFCE8428FFC96" box="[421,509,839,863]" metricMagnitude="-1" metricUnit="m" metricValue="1.64" pageId="12" pageNumber="1862" unit="mm" value="164.0">164 mm</quantity>
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(variation in two undamaged clitellate
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<typeStatus id="54978879FF919A1BCDA7FCE84724FC96" box="[1000,1110,839,863]" pageId="12" pageNumber="1862" type="paratype">paratypes</typeStatus>
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||
<quantity id="4CD49B3EFF919A1BCED5FCC7425EFC49" box="[154,300,872,896]" metricMagnitude="-1" metricUnit="m" metricValue="1.525" metricValueMax="1.7" metricValueMin="1.35" pageId="12" pageNumber="1862" unit="mm" value="152.5" valueMax="170.0" valueMin="135.0">135–170 mm</quantity>
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), maximum diameter
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<quantity id="4CD49B3EFF919A1BCC04FCC741EFFC49" box="[587,669,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="12" pageNumber="1862" unit="mm" value="3.6">3.6 mm</quantity>
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(2.5–3.6), 174 segments, probably regeneration of hind end, last 13 segments small (the two undamaged clitellate adults with 200 and 205 segments). Segments 8–13 tetra- to multi-annulate, annulation in clitellum almost indistinguishable, behind clitellum multi-annulate, gradually becoming less annulated and disappearing posteriad. Prostomium prolobous. Clitellum orange, rest of body without pigment (fixed specimens). Setae uniform. Setal formula:
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<emphasis id="B958EAC9FF919A1BCFB8FB9E4160FB80" box="[503,530,1073,1097]" italics="true" pageId="12" pageNumber="1862">aa</emphasis>
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||
:
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<emphasis id="B958EAC9FF919A1BCC56FB9E4146FB80" box="[537,564,1073,1097]" italics="true" pageId="12" pageNumber="1862">ab</emphasis>
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:
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<emphasis id="B958EAC9FF919A1BCC74FB9E4126FB80" box="[571,596,1073,1097]" italics="true" pageId="12" pageNumber="1862">bc</emphasis>
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:
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<emphasis id="B958EAC9FF919A1BCC14FB9E4106FB80" box="[603,628,1073,1097]" italics="true" pageId="12" pageNumber="1862">cd</emphasis>
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<emphasis id="B958EAC9FF919A1BCC3BFB9E41FDFB80" bold="true" box="[628,655,1073,1097]" pageId="12" pageNumber="1862">~</emphasis>
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11.7:1.3:8.3:1.0,
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<emphasis id="B958EAC9FF919A1BCD06FB9E4016FB80" box="[841,868,1073,1097]" italics="true" pageId="12" pageNumber="1862">dd</emphasis>
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<emphasis id="B958EAC9FF919A1BCD2CFB9E400CFB80" bold="true" box="[867,894,1073,1097]" pageId="12" pageNumber="1862">~</emphasis>
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two-thirds of body circumference at segment 10; 8.0:1.0:4.4:1.0,
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<emphasis id="B958EAC9FF919A1BCCEFFBFD41C9FBA3" box="[672,699,1106,1130]" italics="true" pageId="12" pageNumber="1862">dd</emphasis>
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<emphasis id="B958EAC9FF919A1BCCF4FBFD41A4FBA3" bold="true" box="[699,726,1106,1130]" pageId="12" pageNumber="1862">~</emphasis>
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two-thirds of body circumference at segment 30. Setae not visible in clitellar region except for penial setae in 17. Clitellum annular, with up to five segments in 14–3/4 18, 18, replaced by genital swelling ventrally in segment 17 (see below).
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</paragraph>
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<paragraph id="8B9336DBFF919A1BCE8DFB794073FB27" blockId="12.[154,1111,807,1458]" box="[194,769,1238,1262]" pageId="12" pageNumber="1862">No dorsal pores. Nephridiopores not discernible.</paragraph>
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<paragraph id="8B9336DBFF919A1BCE8DFB59404EFAE6" blockId="12.[154,1111,807,1458]" pageId="12" pageNumber="1862">
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Spermarial pore unpaired, minute opening m/v in setal zone in 13. Opening surrounded by circular, whitish area (diameter about
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<emphasis id="B958EAC9FF919A1BCD5DFAB7405FFAF9" box="[786,813,1304,1328]" italics="true" pageId="12" pageNumber="1862">ab</emphasis>
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).
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</paragraph>
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<paragraph id="8B9336DBFF919A1BCE8DFA984250FAB8" blockId="12.[154,1111,807,1458]" pageId="12" pageNumber="1862">
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Female pores paired, minute openings in segment 15 near 14/15, 1
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<emphasis id="B958EAC9FF919A1BCDB3FA984767FA86" box="[1020,1045,1335,1359]" italics="true" pageId="12" pageNumber="1862">cd</emphasis>
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–4
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<emphasis id="B958EAC9FF919A1BCA72FA984724FA86" box="[1085,1110,1335,1359]" italics="true" pageId="12" pageNumber="1862">cd</emphasis>
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dorsal to
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<emphasis id="B958EAC9FF919A1BCF42FAF64268FAB8" box="[269,282,1369,1393]" italics="true" pageId="12" pageNumber="1862">d</emphasis>
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.
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</paragraph>
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<paragraph id="8B9336DBFF919A1BCE8DFAD740D0FA7B" blockId="12.[154,1111,807,1458]" pageId="12" pageNumber="1862">
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Male pore unpaired, m/v in
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<quantity id="4CD49B3EFF919A1BCC4CFAD74149FA59" box="[515,571,1400,1424]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="12" pageNumber="1862" unit="in" value="17.0">17 in</quantity>
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setal zone on small oval swelling covering area between setal lines
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<emphasis id="B958EAC9FF919A1BCF32FA3542F8FA7B" box="[381,394,1434,1458]" italics="true" pageId="12" pageNumber="1862">a</emphasis>
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or
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<emphasis id="B958EAC9FF919A1BCFF7FA3542B7FA7B" box="[440,453,1434,1458]" italics="true" pageId="12" pageNumber="1862">b</emphasis>
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and posterior two-thirds of segment 17.
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</paragraph>
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<paragraph id="8B9336DBFF919A1BCED5FA49421DFA37" blockId="12.[154,1111,1510,1732]" box="[154,367,1510,1534]" pageId="12" pageNumber="1862">
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||
<emphasis id="B958EAC9FF919A1BCED5FA49421DFA37" box="[154,367,1510,1534]" italics="true" pageId="12" pageNumber="1862">Internal characters</emphasis>
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||
</paragraph>
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<paragraph id="8B9336DBFF919A1ACE8DF9AA477FFF2F" blockId="12.[154,1111,1510,1732]" lastBlockId="13.[159,1116,173,1356]" lastPageId="13" lastPageNumber="1863" pageId="12" pageNumber="1862">
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First septum 4/5. Septa 4/5–11/12 with thick muscular layer, all succeeding septa thin. Pharynx extending posteriorly to 4/5. Oesophageal gizzards, ventral oesophageal sacs and calciferous glands absent. Intestine commencing in posterior half of 19. Three intestinal gizzards, one in each of segments 23, 24 and 25. Anterior to gizzards thin-walled sections of gut with four pouches (two dorsally and two ventrally) next to septa. Median typhlosole present from segment 20 (
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||
<specimenCount id="9D2AFD52FF919A1BCDFBF903473DF90D" box="[948,1103,1708,1732]" pageId="12" pageNumber="1862" type="generic">one specimen</specimenCount>
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||
) or 26 (
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<specimenCount id="9D2AFD52FF909A1ACEBCFF0242EFFF0C" box="[243,413,173,197]" pageId="13" pageNumber="1863" type="generic">two specimens</specimenCount>
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||
) becoming gradually less distinct towards hind end (about segment
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<quantity id="4CD49B3EFF909A1ACF49FF61423DFF2F" box="[262,335,206,230]" metricMagnitude="0" metricUnit="m" metricValue="3.429" pageId="13" pageNumber="1863" unit="in" value="135.0">135 in</quantity>
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||
<typeStatus id="54978879FF909A1ACF16FF6142FCFF2F" box="[345,398,206,230]" pageId="13" pageNumber="1863">type</typeStatus>
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). Lateral typhlosoles present in segments 26–35 or 36.
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</paragraph>
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<paragraph id="8B9336DBFF909A1ACE88FF424396FE4F" blockId="13.[159,1116,173,1356]" pageId="13" pageNumber="1863">Dorsal blood vessel bifurcating segmentally, but reuniting passing through septa in 13–20, 21 or 22. Vessel traceable to posterior face of 5/6. Supra-oesophageal trunk visible in 6–12, larger in posterior segments than in anterior ones. Hearts in 6–12. Subneural trunk dividing into two extra-oesophageal vessels just in front of 15/16.</paragraph>
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<paragraph id="8B9336DBFF909A1ACE88FE21415AFE6F" blockId="13.[159,1116,173,1356]" box="[199,552,398,422]" pageId="13" pageNumber="1863">Excretory system holonephric.</paragraph>
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<paragraph id="8B9336DBFF909A1ACE88FE01419BFD8E" blockId="13.[159,1116,173,1356]" pageId="13" pageNumber="1863">
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Holandric. Testes well developed, free, one pair in each of segments 10 and 11. Funnels paired in 10 and 11, large with numerous plications and with white iridescence. Seminal vesicles paired in 11 and 12. Anterior pair restricted to segment 11, posterior one extending to about 19/20. Vasa deferentia entering lateral sides of euprostates approximately
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<quantity id="4CD49B3EFF909A1ACF95FD80415EFD8E" box="[474,556,559,583]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="1863" unit="mm" value="1.2">1.2 mm</quantity>
|
||
from ectal end.
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF909A1ACE88FDE04269FD61" blockId="13.[159,1116,173,1356]" pageId="13" pageNumber="1863">Euprostates. Two almost regular tubular organs, lying ventro-lateral to the intestine in 18–20. They bend mediad, gradually narrowing before penetrating body wall in 17.</paragraph>
|
||
<paragraph id="8B9336DBFF909A1ACE88FD00439DFC81" blockId="13.[159,1116,173,1356]" pageId="13" pageNumber="1863">
|
||
Follicles of penial setae situated just in front of and mediad to euprostates. One seta in each sac. Penial setae (figure 6B) about
|
||
<quantity id="4CD49B3EFF909A1ACC9CFD7F4057FD21" box="[723,805,720,744]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="1863" unit="mm" value="1.1">1.1 mm</quantity>
|
||
long, and with maximum width of
|
||
<quantity id="4CD49B3EFF909A1ACF46FD5F421BFCC1" box="[265,361,752,776]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="13" pageNumber="1863" unit="mm" value="0.07">0.07 mm</quantity>
|
||
(at base). Shape almost cylindrical, only slowly tapering towards ectal end, ending in a small hook. Setae slightly curved. Growth striation hardly visible.
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF909A1ACE88FCFF4399FA85" blockId="13.[159,1116,173,1356]" pageId="13" pageNumber="1863">
|
||
Spermarium (figure 10D) consisting of dorsal sac with two branches emerging from anterior corners of sac and forming two rings in segment 13: one around oesophagus and one around nerve cord with a tube between them. Under nerve cord spermarium is in connection with spermarial pore through narrow tube. Perioesophageal ring continuing dorsally in tubular sac, extending posteriorly to 16/17 or 17/18. Sac with three pairs of diverticula (in
|
||
<specimenCount id="9D2AFD52FF909A1ACD0CFC5D4090FBC3" box="[835,994,1010,1034]" pageId="13" pageNumber="1863" type="generic">one specimen</specimenCount>
|
||
only two diverticula in right side). Branches of each side meeting latero-ventrally in common tube or ending freely. In
|
||
<typeStatus id="54978879FF909A1ACFF3FB9D429DFB83" box="[444,495,1074,1098]" pageId="13" pageNumber="1863">type</typeStatus>
|
||
, left branch of first pair merging with perioesophageal ring, in right side ending freely. Second and third pairs connecting with each other in left side, but not with first diverticle or perioesophageal ring. In right side they end freely. Ovisacs situated on posterior face of 13/14, opening into ental part of oviducts in segment 13. Each oviduct overgrowing funnel in 13, and after penetrating 13/14 traverses 14, where it attaches to 14/15, disappearing into body wall at level of female pore in 15. Receptaculum ovorum in connection with perioesophageal ring by short duct near m/v. No ovaries observed (all specimens adult).
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF909A1ACED0FA2F4270FA51" blockId="13.[159,1116,1408,1625]" box="[159,258,1408,1432]" pageId="13" pageNumber="1863">
|
||
<emphasis id="B958EAC9FF909A1ACED0FA2F4270FA51" box="[159,258,1408,1432]" italics="true" pageId="13" pageNumber="1863">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF909A1ACE88FA0F419FF990" blockId="13.[159,1116,1408,1625]" pageId="13" pageNumber="1863">
|
||
The spermarium of
|
||
<taxonomicName id="4C2C4D58FF909A1ACFF5FA0F41E5FA71" authorityName="Clausen" authorityYear="2004" box="[442,663,1440,1464]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="1863" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
|
||
<emphasis id="B958EAC9FF909A1ACFF5FA0F41E5FA71" box="[442,663,1440,1464]" italics="true" pageId="13" pageNumber="1863">Libyodrilus maryae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57B2FF909A1ACCEAFA0F419AFA71" box="[677,744,1440,1464]" pageId="13" pageNumber="1863" rank="species">n. sp.</taxonomicNameLabel>
|
||
belongs both to group II and group III because latero-ventral connections are in some cases lacking between some of the diverticula. From all other species in group II and
|
||
<taxonomicName id="4C2C4D58FF909A1ACDD8FA4F4748FA31" authorityName="Clausen" authorityYear="2004" box="[919,1082,1504,1528]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="1863" phylum="Annelida" rank="species" species="bipindiensis">
|
||
<emphasis id="B958EAC9FF909A1ACDD8FA4F4748FA31" box="[919,1082,1504,1528]" italics="true" pageId="13" pageNumber="1863">L. bipindiensis</emphasis>
|
||
</taxonomicName>
|
||
in group III (figures 6F, I, J and 5A),
|
||
<taxonomicName id="4C2C4D58FF909A1ACC7EF9AE41D6F9D0" authorityName="Clausen" authorityYear="2004" box="[561,676,1537,1561]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="1863" phylum="Annelida" rank="species" species="maryae">
|
||
<emphasis id="B958EAC9FF909A1ACC7EF9AE41D6F9D0" box="[561,676,1537,1561]" italics="true" pageId="13" pageNumber="1863">L. maryae</emphasis>
|
||
</taxonomicName>
|
||
(figure 6B) differs by the more slender shape of the penial setae. From
|
||
<taxonomicName id="4C2C4D58FF909A1ACC7EF98E419EF9F0" authorityName="Clausen" authorityYear="2004" box="[561,748,1569,1593]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="1863" phylum="Annelida" rank="species" species="choristoporus">
|
||
<emphasis id="B958EAC9FF909A1ACC7EF98E419EF9F0" box="[561,748,1569,1593]" italics="true" pageId="13" pageNumber="1863">L. choristoporus</emphasis>
|
||
</taxonomicName>
|
||
in group III it differs by the position of the genital pores (cf. figure 8A and B).
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF909A1ACED0F922472EF90D" blockId="13.[159,1116,1677,1732]" box="[159,1116,1677,1732]" pageId="13" pageNumber="1863">
|
||
<emphasis id="B958EAC9FF909A1ACED0F9224256F96C" box="[159,292,1677,1701]" italics="true" pageId="13" pageNumber="1863">Distribution</emphasis>
|
||
Only known from the
|
||
<typeStatus id="54978879FF909A1ACF9FF9034172F90D" box="[464,512,1708,1732]" pageId="13" pageNumber="1863">type</typeStatus>
|
||
locality in southern
|
||
<collectingCountry id="F33B764BFF909A1ACCBAF9034002F90D" box="[757,880,1708,1732]" name="Cameroon" pageId="13" pageNumber="1863">Cameroon</collectingCountry>
|
||
, just E of Akom II.
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF939A19CED5FF014261FF0F" blockId="14.[154,1110,174,261]" box="[154,275,174,198]" pageId="14" pageNumber="1864">
|
||
<emphasis id="B958EAC9FF939A19CED5FF014261FF0F" box="[154,275,174,198]" italics="true" pageId="14" pageNumber="1864">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF939A19CE8DFF62407BFECC" blockId="14.[154,1110,174,261]" pageId="14" pageNumber="1864">
|
||
The species
|
||
<taxonomicName id="4C2C4D58FF939A19CF01FF6142B1FF2F" authorityName="Clausen" authorityYear="2004" box="[334,451,206,230]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="1864" phylum="Annelida" rank="species" species="maryae">
|
||
<emphasis id="B958EAC9FF939A19CF01FF6142B1FF2F" box="[334,451,206,230]" italics="true" pageId="14" pageNumber="1864">L. maryae</emphasis>
|
||
</taxonomicName>
|
||
is named after my colleague Mary E. Petersen, who has been an invaluable help in preparing the manuscript.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366550FF939A19CFF6FE8240C2FE4C" pageId="14" pageNumber="1864" type="reference_group">
|
||
<paragraph id="8B9336DBFF939A19CFF6FE82402DFE8C" blockId="14.[154,944,301,389]" box="[441,863,301,325]" pageId="14" pageNumber="1864">
|
||
<taxonomicName id="4C2C4D58FF939A19CFF6FE82402DFE8C" ID-CoL="8LDDX" authority="Clausen, 1965" authorityName="Clausen" authorityYear="1965" box="[441,863,301,325]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="1864" phylum="Annelida" rank="species" species="mekoensis">
|
||
<emphasis id="B958EAC9FF939A19CFF6FE8241C1FE8C" bold="true" box="[441,691,301,325]" italics="true" pageId="14" pageNumber="1864">Libyodrilus mekoensis</emphasis>
|
||
<bibRefCitation id="EFBD4B2AFF939A19CCF2FE82402DFE8C" author="Clausen" box="[701,863,301,325]" firstAuthor="Clausen" pageId="14" pageNumber="1864" pagination="273 - 292" refId="ref12294" refString="CLAUSEN, M. W., 1965, Description of two new species of Libyodrilus Beddard, 1891, with remarks on the genus and on the female system in the family Eudrilidae, Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 128, 273 - 292." type="journal article" year="1965">Clausen, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF939A19CFB2FEE24068FEAC" blockId="14.[154,944,301,389]" box="[509,794,333,357]" pageId="14" pageNumber="1864">(figures 6C, D, 8D, 11C)</paragraph>
|
||
<paragraph id="8B9336DBFF939A19CED5FEC040C2FE4C" blockId="14.[154,944,301,389]" box="[154,944,367,389]" pageId="14" pageNumber="1864">
|
||
<treatmentCitationGroup id="AB3C11F5FF939A19CED5FEC040C2FE4C" box="[154,944,367,389]" pageId="14" pageNumber="1864">
|
||
<treatmentCitation id="0A8D10CAFF939A19CED5FEC0410EFE4C" author="CLAUSEN, M. W." box="[154,636,367,389]" page="286" pageId="14" pageNumber="1864" year="1965">
|
||
<taxonomicName id="4C2C4D58FF939A19CED5FEC0410EFE4C" ID-CoL="8LDDX" authority="Clausen, 1965: 279 - 286" authorityName="Clausen" authorityPageNumber="279 - 286" authorityYear="1965" box="[154,636,367,389]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="1864" phylum="Annelida" rank="species" species="mekoensis">
|
||
<emphasis id="B958EAC9FF939A19CED5FEC04208FE4D" box="[154,378,367,388]" italics="true" pageId="14" pageNumber="1864">Libyodrilus mekoensis</emphasis>
|
||
<bibRefCitation id="EFBD4B2AFF939A19CFCCFEC0410EFE4C" author="Clausen" box="[387,636,367,389]" firstAuthor="Clausen" page="279 - 286" pageId="14" pageNumber="1864" pagination="273 - 292" refId="ref12294" refString="CLAUSEN, M. W., 1965, Description of two new species of Libyodrilus Beddard, 1891, with remarks on the genus and on the female system in the family Eudrilidae, Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 128, 273 - 292." type="journal article" year="1965">Clausen, 1965: 279–286</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, figures 1–3;
|
||
<treatmentCitation id="0A8D10CAFF939A19CD4BFEC040D9FE4C" author="SIMS, R. W." box="[772,939,367,389]" page="364" pageId="14" pageNumber="1864" year="1987">
|
||
<bibRefCitation id="EFBD4B2AFF939A19CD4BFEC040D9FE4C" author="Sims" box="[772,939,367,389]" firstAuthor="Sims" page="364" pageId="14" pageNumber="1864" pagination="359 - 388" refId="ref12684" refString="SIMS, R. W., 1987, Review of the Central African earthworm family Eudrilidae (Oligochaeta), in A. M. B. Pagliai and P. Omodeo (eds) Proceedings of the International Symposium on Earthworms, Bologna (Modena, Italy: Mucchi), pp. 359 - 388." type="proceedings paper" year="1987">Sims, 1987: 364</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366550FF939A19CED5FE1640AFFD75" pageId="14" pageNumber="1864" type="discussion">
|
||
<paragraph id="8B9336DBFF939A19CED5FE16438FFE18" blockId="14.[154,1110,441,593]" box="[154,253,441,465]" pageId="14" pageNumber="1864">
|
||
<emphasis id="B958EAC9FF939A19CED5FE16438FFE18" box="[154,253,441,465]" italics="true" pageId="14" pageNumber="1864">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF939A19CE8DFE7641FEFDF8" blockId="14.[154,1110,441,593]" pageId="14" pageNumber="1864">
|
||
<taxonomicName id="4C2C4D58FF939A19CE8DFE7642C8FE38" authorityName="Clausen" authorityYear="1965" box="[194,442,473,497]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="1864" phylum="Annelida" rank="species" species="mekoensis">
|
||
<emphasis id="B958EAC9FF939A19CE8DFE7642C8FE38" box="[194,442,473,497]" italics="true" pageId="14" pageNumber="1864">Libyodrilus mekoensis</emphasis>
|
||
</taxonomicName>
|
||
belongs to spermarial group IV (table 1; figure 11C). The species differs from all other species of
|
||
<taxonomicName id="4C2C4D58FF939A19CC2AFE564191FDD8" authorityName="Beddard" authorityYear="1891" box="[613,739,505,529]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="1864" phylum="Annelida" rank="genus">
|
||
<emphasis id="B958EAC9FF939A19CC2AFE564191FDD8" box="[613,739,505,529]" italics="true" pageId="14" pageNumber="1864">Libyodrilus</emphasis>
|
||
</taxonomicName>
|
||
in the shape of the genital field (figure 8D) and penial setae (figure 6C, D).
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF939A19CE8DFD964750FD98" blockId="14.[154,1110,441,593]" box="[194,1058,569,593]" pageId="14" pageNumber="1864">
|
||
Relationships with
|
||
<taxonomicName id="4C2C4D58FF939A19CFEBFD964159FD98" authorityName="Beddard" authorityYear="1891" box="[420,555,569,593]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="1864" phylum="Annelida" rank="species" species="violaceus">
|
||
<emphasis id="B958EAC9FF939A19CFEBFD964159FD98" box="[420,555,569,593]" italics="true" pageId="14" pageNumber="1864">L. violaceus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2C4D58FF939A19CC24FD9641B4FD98" authorityName="Clausen" authorityYear="1965" box="[619,710,569,593]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="1864" phylum="Annelida" rank="species" species="simsi">
|
||
<emphasis id="B958EAC9FF939A19CC24FD9641B4FD98" box="[619,710,569,593]" italics="true" pageId="14" pageNumber="1864">L. simsi</emphasis>
|
||
</taxonomicName>
|
||
are discussed under
|
||
<taxonomicName id="4C2C4D58FF939A19CD8FFD964768FD98" authorityName="Clausen" authorityYear="1965" box="[960,1050,569,593]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="1864" phylum="Annelida" rank="species" species="simsi">
|
||
<emphasis id="B958EAC9FF939A19CD8FFD964768FD98" box="[960,1050,569,593]" italics="true" pageId="14" pageNumber="1864">L. simsi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336DBFF939A19CED5FD2A40AFFD75" blockId="14.[154,989,645,700]" box="[154,989,645,700]" pageId="14" pageNumber="1864">
|
||
<emphasis id="B958EAC9FF939A19CED5FD2A4252FD54" box="[154,288,645,669]" italics="true" pageId="14" pageNumber="1864">Distribution</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54978879FF939A19CF81FD0B428CFD75" box="[462,510,676,700]" pageId="14" pageNumber="1864">type</typeStatus>
|
||
locality: Meko, 7
|
||
<emphasis id="B958EAC9FF939A19CC9EFD0B41A9FD75" bold="true" box="[721,731,676,700]" pageId="14" pageNumber="1864">‡</emphasis>
|
||
28
|
||
<emphasis id="B958EAC9FF939A19CCB9FD0C418EFD75" bold="true" box="[758,764,675,700]" pageId="14" pageNumber="1864">’</emphasis>
|
||
N, 2
|
||
<emphasis id="B958EAC9FF939A19CD7EFD0B4049FD75" bold="true" box="[817,827,676,700]" pageId="14" pageNumber="1864">‡</emphasis>
|
||
55
|
||
<emphasis id="B958EAC9FF939A19CD19FD0C402EFD75" bold="true" box="[854,860,675,700]" pageId="14" pageNumber="1864">’</emphasis>
|
||
E,
|
||
<collectingCountry id="F33B764BFF939A19CDCFFD0B40ABFD75" box="[896,985,676,700]" name="Nigeria" pageId="14" pageNumber="1864">Nigeria</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |