136 lines
16 KiB
XML
136 lines
16 KiB
XML
<document id="F68CDC4B3089EE2F6D322A2070A405C7" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE3C1FF56FF75FB91BAB9FE70" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trinomadraena clusa Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="200" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="199" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:mods id="1509707865CC4C206E3806D5E350859C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4946B4FCDECFF11157D4D1E5BACC24BD">
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<mods:title id="1D86046021E356EF10F49763B1A38C6F">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="7CF65208606737E7ABD59BB33EE755B7" type="personal">
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<mods:role id="B982FD5000C0B59C9E9E72152775DAD4">
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<mods:roleTerm id="45CB71F815240310D5DBBF6A08478FEE">Author</mods:roleTerm>
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<mods:namePart id="80452E29ABCE450DA9D74B2E9FB7C957">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="E67ED37C09AE4C141E13A19CC3B7791F">text</mods:typeOfResource>
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<mods:title id="6B3C299860B953C8069D3712DC793FB9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B2FCCFED13E1EDBF1B96A5FDCADE0BAF">
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<mods:date id="6AD353B91C9A67FB500E6483CD7509EF">2017</mods:date>
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<mods:detail id="762627AD14EC61FB2060206B0D52277F" type="pubDate">
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<mods:number id="F44D689047A0F9532E2B993A84996FE9">2017-11-03</mods:number>
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<mods:detail id="9462D1B26E3CF344D410E1F78516F6DE" type="volume">
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<mods:number id="86BCE9EEB932D187DE33704E45B50A61">4342</mods:number>
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</mods:detail>
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<mods:detail id="E1B8EB3F15A4E5B6B52060BEEF6C2F8D" type="issue">
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<mods:number id="A1A86C12E0839E387F36F09A625EC779">1</mods:number>
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<mods:start id="70D351CFFFD96CF3193A572CBA004649">1</mods:start>
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<mods:end id="5507150F70197C0C32D17B9904089C57">264</mods:end>
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<mods:classification id="37E814BC04596361FE889B089AB47DE9">journal article</mods:classification>
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<mods:identifier id="BD74FF227B1DB2C65ADD1346E93F806F" type="CLB-Dataset">31639</mods:identifier>
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<mods:identifier id="3F064683234F28AD581EA73EBBCE74EC" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="F341EA3098FEB065E44022D06A529E01" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
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<mods:identifier id="4137B83DA603B521EE3F88574B410E4E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B2F406413C72EDC9554AF3FA913549EB" type="Zenodo-Dep">1041066</mods:identifier>
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<mods:identifier id="9DC474E28D127CF17E310A1246B830D8" type="ZooBank">2ACD54D2-3487-432D-9323-EEC131FE2E64</mods:identifier>
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</mods:mods>
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<treatment id="038587BBE3C1FF56FF75FB91BAB9FE70" ID-DOI="http://doi.org/10.5281/zenodo.5323756" ID-GBIF-Taxon="137233374" ID-Zenodo-Dep="5323756" LSID="urn:lsid:plazi:treatment:038587BBE3C1FF56FF75FB91BAB9FE70" httpUri="http://treatment.plazi.org/id/038587BBE3C1FF56FF75FB91BAB9FE70" lastPageId="199" lastPageNumber="200" pageId="198" pageNumber="199">
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<subSubSection id="C3366526E3C1FF57FF75FB91BD2AFBEA" pageId="198" pageNumber="199" type="nomenclature">
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<paragraph id="8B9336ADE3C1FF57FF75FB91BB06FBCB" blockId="198.[151,1046,1076,1135]" box="[151,570,1076,1102]" pageId="198" pageNumber="199">
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<heading id="D0DB81C1E3C1FF57FF75FB91BB06FBCB" bold="true" box="[151,570,1076,1102]" fontSize="11" level="1" pageId="198" pageNumber="199" reason="1">
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<emphasis id="B958EABFE3C1FF57FF75FB91BB06FBCB" bold="true" box="[151,570,1076,1102]" pageId="198" pageNumber="199">
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<taxonomicName id="4C2C4D2EE3C1FF57FF75FB91B8A7FBCB" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,411,1076,1102]" class="Insecta" family="Hydraenidae" genus="Trinomadraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="clusa" status="sp. nov.">
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<emphasis id="B958EABFE3C1FF57FF75FB91B8A7FBCB" bold="true" box="[151,411,1076,1102]" italics="true" pageId="198" pageNumber="199">Trinomadraena clusa</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26B57C4E3C1FF57FE48FB91BB06FBCB" box="[426,570,1076,1102]" pageId="198" pageNumber="199" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336ADE3C1FF57FF75FBF3BD2AFBEA" blockId="198.[151,1046,1076,1135]" box="[151,1046,1110,1135]" pageId="198" pageNumber="199">
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<figureCitation id="13172A28E3C1FF57FF75FBF3B9C3FBEA" box="[151,255,1110,1135]" captionStart="FIGURE 191" captionStartId="199.[151,250,1741,1763]" captionText="FIGURE 191. Trinomadraena clusa, holotype, habitus dorsal, lateral, and ventral." pageId="198" pageNumber="199">Figs. 191</figureCitation>
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(habitus), 192 (aedeagus), 219 (antenna, maxillary palpus), 300 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3C1FF57FF75FB3BB8C4FB5E" pageId="198" pageNumber="199" type="materials_examined">
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<paragraph id="8B9336ADE3C1FF57FF75FB3BB8C4FB5E" blockId="198.[151,1437,1182,2035]" pageId="198" pageNumber="199">
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<materialsCitation id="3B443CF0E3C1FF57FF75FB3BB8C4FB5E" ID-GBIF-Occurrence="1697157773" collectingDate="1970-10-16" collectionCode="BMNH" collectorName="P. Hammond" latitude="-12.6" location="Mt. d'Ambre" longLatPrecision="1293" longitude="49.166668" pageId="198" pageNumber="199" specimenCount="2" stateProvince="Antsiranana" typeStatus="holotype">
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<emphasis id="B958EABFE3C1FF57FF75FB3BB887FB32" bold="true" box="[151,443,1182,1207]" pageId="198" pageNumber="199">
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<typeStatus id="5497880FE3C1FF57FF75FB3BB9EEFB32" box="[151,210,1182,1207]" pageId="198" pageNumber="199">Type</typeStatus>
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Material.
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<typeStatus id="5497880FE3C1FF57FEACFB3BB887FB32" box="[334,443,1182,1207]" pageId="198" pageNumber="199" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<collectingRegion id="49E8F84FE3C1FF57FDFBFB3BBB92FB32" box="[537,686,1182,1207]" country="Madagascar" name="Antsiranana" pageId="198" pageNumber="199">
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<emphasis id="B958EABFE3C1FF57FDFBFB3BBB92FB32" bold="true" box="[537,686,1182,1207]" pageId="198" pageNumber="199">Antsiranana</emphasis>
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</collectingRegion>
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,
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<location id="8EF36076E3C1FF57FD5EFB3ABA6EFB32" LSID="urn:lsid:plazi:treatment:038587BBE3C1FF56FF75FB91BAB9FE70:8EF36076E3C1FF57FD5EFB3ABA6EFB32" box="[700,850,1182,1207]" latitude="-12.6" longLatPrecision="1293" longitude="49.166668" name="Mt. d'Ambre" pageId="198" pageNumber="199" stateProvince="Antsiranana">Mt. d’Ambre</location>
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, wet moss by petite cascade,
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<geoCoordinate id="EE18506AE3C1FF57FB7BFB3ABC3CFB32" box="[1177,1280,1182,1207]" direction="south" orientation="latitude" pageId="198" pageNumber="199" precision="925" value="-12.6">12° 36' S</geoCoordinate>
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,
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<geoCoordinate id="EE18506AE3C1FF57FAEEFB3ABC49FB33" box="[1292,1397,1183,1207]" direction="east" orientation="longitude" pageId="198" pageNumber="199" precision="925" value="49.166668">49° 10' E</geoCoordinate>
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,
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<collectingDate id="EFD6E985E3C1FF57FA62FB3BB9DBFB5E" pageId="198" pageNumber="199" value="1970-10-16">
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<date id="FF92106DE3C1FF57FA62FB3BB9DBFB5E" pageId="198" pageNumber="199" value="1970-10-16">16 x 1970</date>
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</collectingDate>
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,
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<collectorName id="26D9537BE3C1FF57FF10FB66B8BEFB5E" box="[242,386,1218,1243]" pageId="198" pageNumber="199">P. Hammond</collectorName>
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(
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<collectionCode id="ED3DAE68E3C1FF57FE73FB66B8D7FB5E" box="[401,491,1219,1243]" pageId="198" pageNumber="199">BMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3C1FF57FF25FB43BA39FA52" pageId="198" pageNumber="199" type="diagnosis">
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<paragraph id="8B9336ADE3C1FF57FF25FB43BA39FA52" blockId="198.[151,1437,1182,2035]" pageId="198" pageNumber="199">
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<emphasis id="B958EABFE3C1FF57FF25FB43B8EEFB7A" bold="true" box="[199,466,1254,1279]" pageId="198" pageNumber="199">Differential Diagnosis.</emphasis>
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Dorsal habitus as in
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<figureCitation id="13172A28E3C1FF57FD58FB43BA26FB7A" box="[698,794,1254,1279]" captionStart="FIGURE 191" captionStartId="199.[151,250,1741,1763]" captionText="FIGURE 191. Trinomadraena clusa, holotype, habitus dorsal, lateral, and ventral." pageId="198" pageNumber="199">Fig. 191</figureCitation>
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; a moderately sized species with strong elytral costae, the first of which has an interrupted section. Differentiated from all
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<taxonomicName id="4C2C4D2EE3C1FF57FC92FAAFBD38FAA6" authorityName="Perkins" authorityYear="1997" box="[880,1028,1290,1315]" class="Insecta" family="Hydraenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="tribe" tribe="Madagastrini">Madagastrini</taxonomicName>
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by the hydrofuge pubescence being restricted to the first three ventrites plus the anterior part of the fourth; this vestiture is very dense and very short, perhaps plastron forming.
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<taxonomicName id="4C2C4D2EE3C1FF57FE2AFAF1BB1CFAEE" authorityName="Perkins" authorityYear="2017" box="[456,544,1362,1387]" class="Insecta" family="Hydraenidae" genus="Trinomadraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="clusa">
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<emphasis id="B958EABFE3C1FF57FE2AFAF1BB1CFAEE" box="[456,544,1362,1387]" italics="true" pageId="198" pageNumber="199">T. clusa</emphasis>
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</taxonomicName>
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also differs from other members of the tribe in having the procoxal cavities closed behind: the prosternal process is slightly widened behind the coxae, and is contacted on each side by the postcoxal processes of the hypomeron. The aedeagus differs in basic plan from that of other
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<taxonomicName id="4C2C4D2EE3C1FF57FB2BFA3FBC61FA36" authorityName="Perkins" authorityYear="1997" box="[1225,1373,1434,1459]" class="Insecta" family="Hydraenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="tribe" tribe="Madagastrini">Madagastrini</taxonomicName>
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(
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<figureCitation id="13172A28E3C1FF57FA8DFA3FB9F8FA52" captionStart="FIGURE 192" captionStartId="200.[151,250,1281,1303]" captionText="FIGURE 192. Trinomadraena clusa, aedeagus of holotype." pageId="198" pageNumber="199">Fig. 192</figureCitation>
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), as do the antennae and maxillary palpi (
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<figureCitation id="13172A28E3C1FF57FD74FA1BBBC5FA52" box="[662,761,1470,1495]" captionStart="FIGURES 219 – 223" captionStartId="225.[151,264,1807,1830]" captionTargetBox="[270,1302,212,1759]" captionTargetId="figure@225.[264,1323,193,1786]" captionTargetPageId="225" captionText="FIGURES 219 – 223. Antennae, maxillary palpi, and spermathecae." httpUri="https://zenodo.org/record/1041146/files/figure.png" pageId="198" pageNumber="199">Fig. 219</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3C1FF56FF25FA47BAC8FE4A" lastPageId="199" lastPageNumber="200" pageId="198" pageNumber="199" type="description">
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<paragraph id="8B9336ADE3C1FF57FF25FA47BA37F99A" blockId="198.[151,1437,1182,2035]" pageId="198" pageNumber="199">
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<emphasis id="B958EABFE3C1FF57FF25FA47B86DFA7E" bold="true" box="[199,337,1506,1531]" pageId="198" pageNumber="199">Description</emphasis>
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. Size:
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<typeStatus id="5497880FE3C1FF57FE41FA47BB39FA7E" box="[419,517,1506,1531]" pageId="198" pageNumber="199" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.42/1.00; head width 0.60; pronotum 0.60/0.88, PA 0.70, PB 0.67; elytra 1.45/1.00.
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</paragraph>
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<paragraph id="8B9336ADE3C1FF57FF75F98EBAE8F9C6" blockId="198.[151,1437,1182,2035]" box="[151,980,1578,1603]" pageId="198" pageNumber="199">Dorsum and venter dark brown to piceous, legs brown, ocelli light brown.</paragraph>
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<paragraph id="8B9336ADE3C1FF57FF25F9EABB95F972" blockId="198.[151,1437,1182,2035]" pageId="198" pageNumber="199">Dorsum of head and pronotum dull, with very small, closely spaced granules, each with a short recumbent seta. Labrum smooth, anterior margin emarginate, setae very sparse. Labrum originating at anterior of non-overhanging margin of clypeus. Labroclypeal suture deep. Small impression in front of each ocellus. Frons with concavity between ocelli. Maxillary palpi shorter than antenna, ca. as long as non-club articles together. Antenna with penultimate antennomere shorter than ultimate.</paragraph>
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<paragraph id="8B9336ADE3C1FF57FF25F8A6BC28F8E6" blockId="198.[151,1437,1182,2035]" pageId="198" pageNumber="199">Pronotum with two narrowly separated midlongitudinal fovea, anterior ca. twice as long as posterior; each side with two admedian and two sublateral foveae; posteroadmedian fovea oblique, deep; sublateral foveae very deep and very large; transverse fovea along base; lateral margin rounded; postocular emarginations shallow.</paragraph>
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<paragraph id="8B9336ADE3C1FF57FF25F8CBBC69F877" blockId="198.[151,1437,1182,2035]" pageId="198" pageNumber="199">Elytral suture costate in posterior. Each elytron with four costae; first costa (from suture) interrupted for short distance at ca. middle of length. Each costa with unilinear row of granules. Areas between costae punctate, each puncture with minute setigerous granule at anterior margin. Areas between costae with three irregular rows of punctures, except part of area between costae two and three with four irregular rows. Elytral apices truncate.</paragraph>
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<paragraph id="8B9336ADE3C0FF56FF25FF3DBB27FF7D" blockId="199.[151,1437,151,501]" pageId="199" pageNumber="200">Mentum and postmentum micropunctulate, mentum pubescent, especially at anterior; Genae shining, concave behind cardo, with very low posterior ridge. Subocular groove for antenna moderately deep and wide. Posterior margin of eye with low raised rim.</paragraph>
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<paragraph id="8B9336ADE3C0FF56FF25FEA0BB8DFE98" blockId="199.[151,1437,151,501]" box="[199,689,260,285]" pageId="199" pageNumber="200">Prosternum without midlongitudinal carina.</paragraph>
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<paragraph id="8B9336ADE3C0FF56FF25FE82BAC8FE4A" blockId="199.[151,1437,151,501]" pageId="199" pageNumber="200">
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Mesoventrite without tubercle on intercoxal process. Metaventrite densely micropunctulate and with very short and very dense vestiture; median depression oval, moderately deep (
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<figureCitation id="13172A28E3C0FF56FC79FEE9BAC2FEE1" box="[923,1022,332,357]" captionStart="FIGURE 191" captionStartId="199.[151,250,1741,1763]" captionText="FIGURE 191. Trinomadraena clusa, holotype, habitus dorsal, lateral, and ventral." pageId="199" pageNumber="200">Fig. 191</figureCitation>
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). First three abdominal ventrites and anterior of fourth with very dense, short setae (plastron?). Last ventrite oval. Protibia widened in distal 2/3. protrochanter with small, sharply pointed tubercle. Mesotibia inner margin very slightly emarginate. Metafemur rather large. Metatibia slender, bisinuate, inner margin with recumbent setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3C0FF56FF25FE79BAB9FE70" box="[199,901,476,501]" pageId="199" pageNumber="200" type="etymology">
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<paragraph id="8B9336ADE3C0FF56FF25FE79BAB9FE70" blockId="199.[151,1437,151,501]" box="[199,901,476,501]" pageId="199" pageNumber="200">
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<emphasis id="B958EABFE3C0FF56FF25FE79B872FE70" bold="true" box="[199,334,476,501]" pageId="199" pageNumber="200">Etymology.</emphasis>
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Name in reference to the closed procoxal cavities.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |