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<document id="42C38A5A62C4328301FA3472F3744D96" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE32FFFBBFF75FAA5B812FF50" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena bergsteni Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="41" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:titleInfo id="30D7FA9E9EEF3C8E345DDD72CC4B9EF3">
<mods:title id="B7E5B9BBE7E68619785A65B68A51D544">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
</mods:titleInfo>
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<mods:namePart id="33503F7C8351B17018B188713AA120A1">Perkins, Philip D.</mods:namePart>
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<mods:title id="A82FA3066CE4D63FC99EC3A8C38BD67F">Zootaxa</mods:title>
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<mods:part id="95B66632DD39F2816252EAB00C4D6948">
<mods:date id="3E99E09BD16CF65FBC1149B553AFC390">2017</mods:date>
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<mods:number id="8A54BFD7D0469E6EAD7396A9302E79C5">2017-11-03</mods:number>
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<mods:number id="7920252F6CE4E1E53E2EF45E8D2624C1">4342</mods:number>
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<mods:identifier id="C662F1E5C2E16FECAC78492FC9018CB8" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<treatment id="038587BBE32FFFBBFF75FAA5B812FF50" ID-DOI="http://doi.org/10.5281/zenodo.5323467" ID-GBIF-Taxon="137233458" ID-Zenodo-Dep="5323467" LSID="urn:lsid:plazi:treatment:038587BBE32FFFBBFF75FAA5B812FF50" httpUri="http://treatment.plazi.org/id/038587BBE32FFFBBFF75FAA5B812FF50" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection id="C3366526E32FFFB9FF75FAA5BBB4FABE" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8B9336ADE32FFFB9FF75FAA5BB18FA9F" blockId="40.[151,648,1280,1339]" box="[151,548,1280,1306]" pageId="40" pageNumber="41">
<heading id="D0DB81C1E32FFFB9FF75FAA5BB18FA9F" bold="true" box="[151,548,1280,1306]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis id="B958EABFE32FFFB9FF75FAA5BB18FA9F" bold="true" box="[151,548,1280,1306]" pageId="40" pageNumber="41">
<taxonomicName id="4C2C4D2EE32FFFB9FF75FAA5B8B9FA9F" ID-CoL="3N3LH" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,389,1280,1306]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="bergsteni" status="sp. nov.">
<emphasis id="B958EABFE32FFFB9FF75FAA5B8B9FA9F" bold="true" box="[151,389,1280,1306]" italics="true" pageId="40" pageNumber="41">Hydraena bergsteni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B57C4E32FFFB9FE76FAA5BB18FA9F" box="[404,548,1280,1306]" pageId="40" pageNumber="41" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336ADE32FFFB9FF75FA87BBB4FABE" blockId="40.[151,648,1280,1339]" box="[151,648,1314,1339]" pageId="40" pageNumber="41">
<figureCitation id="13172A28E32FFFB9FF75FA87B9CDFABE" box="[151,241,1314,1339]" captionStart="FIGURE 30" captionStartId="41.[151,250,1506,1528]" captionText="FIGURE 30. Hydraena bergsteni, holotype, habitus dorsal, lateral, and ventral." pageId="40" pageNumber="41">Figs. 30</figureCitation>
(habitus), 32 (aedeagus), 241 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E32FFFB9FF75FAC0B8A3F9A1" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8B9336ADE32FFFB9FF75FAC0B8A3F9A1" blockId="40.[151,1437,1381,2006]" pageId="40" pageNumber="41">
<materialsCitation id="3B443CF0E32FFFB9FF75FAC0BB98FA45" ID-GBIF-Occurrence="1697157520" collectingDate="2012-11-22" collectionCode="NHRS" collectorName="J. Bergsten &amp; R. Bukontaite &amp; T. Ranarilalatiana &amp; J. H. Randriamihaja" county="Diana" elevation="328" latitude="-14.02944" location="East" longLatPrecision="1" longitude="48.27491" municipality="Beraty" pageId="40" pageNumber="41" specimenCode="MAD12-22" specimenCount="2" stateProvince="Antsiranana" typeStatus="holotype">
<emphasis id="B958EABFE32FFFB9FF75FAC0B8F0FAFB" bold="true" box="[151,460,1381,1406]" pageId="40" pageNumber="41">
<typeStatus id="5497880FE32FFFB9FF75FAC0B9EEFAFB" box="[151,210,1381,1406]" pageId="40" pageNumber="41">Type</typeStatus>
Material.
<typeStatus id="5497880FE32FFFB9FEBDFAC0B8F0FAFB" box="[351,460,1381,1406]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<collectingRegion id="49E8F84FE32FFFB9FDD9FAC0BBECFAFB" box="[571,720,1381,1406]" country="Madagascar" name="Antsiranana" pageId="40" pageNumber="41">
<emphasis id="B958EABFE32FFFB9FDD9FAC0BBECFAFB" bold="true" box="[571,720,1381,1406]" pageId="40" pageNumber="41">Antsiranana</emphasis>
</collectingRegion>
,
<collectingCounty id="62F24E21E32FFFB9FD04FAC0BA12FAFB" box="[742,814,1381,1406]" pageId="40" pageNumber="41">Diana</collectingCounty>
:
<collectingMunicipality id="6BF7ACD7E32FFFB9FCA2FAC3BAB3FAFB" box="[832,911,1382,1406]" pageId="40" pageNumber="41">Beraty</collectingMunicipality>
:
<quantity id="4CD49B48E32FFFB9FC43FAC0BAEFFAFB" box="[929,979,1381,1406]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="41" unit="km" value="2.0">2km</quantity>
<location id="8EF36076E32FFFB9FC00FAC3BD2EFAF8" LSID="urn:lsid:plazi:treatment:038587BBE32FFFBBFF75FAA5B812FF50:8EF36076E32FFFB9FC00FAC3BD2EFAF8" box="[994,1042,1382,1405]" county="Diana" latitude="-14.02944" longLatPrecision="1" longitude="48.27491" municipality="Beraty" name="East" pageId="40" pageNumber="41" stateProvince="Antsiranana">East</location>
from
<location id="8EF36076E32FFFB9FB85FAC3BC2CFAFB" LSID="urn:lsid:plazi:treatment:038587BBE32FFFBBFF75FAA5B812FF50:8EF36076E32FFFB9FB85FAC3BC2CFAFB" box="[1127,1296,1381,1406]" county="Diana" latitude="-14.02944" longLatPrecision="1" longitude="48.27491" municipality="Beraty" name="Beraty village" pageId="40" pageNumber="41" stateProvince="Antsiranana">Beraty village</location>
;
<geoCoordinate id="EE18506AE32FFFB9FAC1FAC3BCA0FAFB" box="[1315,1436,1382,1406]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-14.02944">14.02944S</geoCoordinate>
<geoCoordinate id="EE18506AE32FFFB9FF75FA22B828FA1A" box="[151,276,1415,1439]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="48.27491">48.27491E</geoCoordinate>
, big forest stream with big rocks, elev.
<elevation id="0001D19EE32FFFB9FD25FA22BA33FA1B" box="[711,783,1415,1439]" metricMagnitude="2" metricUnit="m" metricValue="3.28" pageId="40" pageNumber="41" unit="m" value="328.0">
<quantity id="4CD49B48E32FFFB9FD25FA22BA33FA1B" box="[711,783,1415,1439]" metricMagnitude="2" metricUnit="m" metricValue="3.28" pageId="40" pageNumber="41" unit="m" value="328.0">328 m</quantity>
</elevation>
,
<collectingDate id="EFD6E985E32FFFB9FCF8FA22BAAFFA1A" box="[794,915,1414,1439]" pageId="40" pageNumber="41" value="2012-11-22">
<date id="FF92106DE32FFFB9FCF8FA22BAAFFA1A" box="[794,915,1414,1439]" pageId="40" pageNumber="41" value="2012-11-22">22 xi 2012</date>
</collectingDate>
,
<collectorName id="26D9537BE32FFFB9FC7CFA22BD20FA1A" box="[926,1052,1415,1439]" pageId="40" pageNumber="41">J. Bergsten</collectorName>
,
<collectorName id="26D9537BE32FFFB9FBC5FA22BDF9FA1A" box="[1063,1221,1414,1439]" pageId="40" pageNumber="41">R. Bukontaite</collectorName>
,
<collectorName id="26D9537BE32FFFB9FB2DFA22BCA4FA1A" box="[1231,1432,1414,1439]" pageId="40" pageNumber="41">T. Ranarilalatiana</collectorName>
, &amp;
<collectorName id="26D9537BE32FFFB9FF5AFA0DB8AAFA45" box="[184,406,1447,1472]" pageId="40" pageNumber="41">J.H. Randriamihaja</collectorName>
(
<specimenCode id="DB8A9ED6E32FFFB9FE48FA0DBB13FA45" box="[426,559,1448,1472]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="40" pageNumber="41">MAD12-22</specimenCode>
) (
<collectionCode id="ED3DAE68E32FFFB9FDA9FA0DBBA0FA45" box="[587,668,1448,1472]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="40" pageNumber="41">NHRS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B443CF0E32FFFB9FD56FA02B821F987" ID-GBIF-Occurrence="1697157517" collectionCode="NHRS" county="Same" elevation="173" location="Maromiandra" longitude="14.02948" municipality="Sofia" pageId="40" pageNumber="41" specimenCount="2" stateProvince="Mahajanga" typeStatus="paratype">
<typeStatus id="5497880FE32FFFB9FD56FA02BA10FA45" box="[692,812,1447,1472]" pageId="40" pageNumber="41" type="paratype">
<emphasis id="B958EABFE32FFFB9FD56FA02BA10FA45" bold="true" box="[692,812,1447,1472]" pageId="40" pageNumber="41">Paratypes</emphasis>
</typeStatus>
(3):
<collectingCounty id="62F24E21E32FFFB9FC89FA0DBA95FA45" box="[875,937,1448,1472]" pageId="40" pageNumber="41">Same</collectingCounty>
data as
<typeStatus id="5497880FE32FFFB9FBF1FA02BD49FA45" box="[1043,1141,1447,1472]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
(2
<collectionCode id="ED3DAE68E32FFFB9FB47FA0DBDC8FA45" box="[1189,1268,1448,1472]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="40" pageNumber="41">NHRS</collectionCode>
);
<collectingRegion id="49E8F84FE32FFFB9FAEEFA02BCA9FA45" box="[1292,1429,1447,1472]" country="Madagascar" name="Mahajanga" pageId="40" pageNumber="41">
<emphasis id="B958EABFE32FFFB9FAEEFA02BCA9FA45" bold="true" box="[1292,1429,1447,1472]" pageId="40" pageNumber="41">Mahajanga</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6BF7ACD7E32FFFB9FF75FA6CB9EAFA67" box="[151,214,1481,1506]" pageId="40" pageNumber="41">Sofia</collectingMunicipality>
:
<location id="8EF36076E32FFFB9FF06FA6CB843FA67" LSID="urn:lsid:plazi:treatment:038587BBE32FFFBBFF75FAA5B812FF50:8EF36076E32FFFB9FF06FA6CB843FA67" box="[228,383,1481,1506]" county="Same" longitude="14.02948" municipality="Sofia" name="Maromiandra" pageId="40" pageNumber="41" stateProvince="Mahajanga">Maromiandra</location>
, forest stream, GB nets and sieves; stream with stones and gravels, S
<geoCoordinate id="EE18506AE32FFFB9FB44FA6FBC10FA67" box="[1190,1324,1482,1506]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="14.02948">14.02948 E</geoCoordinate>
48.22443, elev.
<elevation id="0001D19EE32FFFB9FF31FA4EB821F987" box="[211,285,1515,1539]" metricMagnitude="2" metricUnit="m" metricValue="1.73" pageId="40" pageNumber="41" unit="m" value="173.0">
<quantity id="4CD49B48E32FFFB9FF31FA4EB821F987" box="[211,285,1515,1539]" metricMagnitude="2" metricUnit="m" metricValue="1.73" pageId="40" pageNumber="41" unit="m" value="173.0">173 m</quantity>
</elevation>
</materialsCitation>
, 14° 2' S, 48° 13' E,
<date id="FF92106DE32FFFB9FDEDFA4EBBB1F986" box="[527,653,1514,1539]" pageId="40" pageNumber="41" value="2012-11-20">20 xi 2012</date>
, J. Bergsten, R. Bukontaite, J.H. Randriamihaja &amp; T. Ranarilalatiana (Mad12-15) (1 NHRS).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E32FFFB9FF25F988BD21F90D" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8B9336ADE32FFFB9FF25F988BD21F90D" blockId="40.[151,1437,1381,2006]" pageId="40" pageNumber="41">
<emphasis id="B958EABFE32FFFB9FF25F988B8EEF9C3" bold="true" box="[199,466,1581,1606]" pageId="40" pageNumber="41">Differential Diagnosis.</emphasis>
Similar in color and size to
<taxonomicName id="4C2C4D2EE32FFFB9FCEEF98BBAB2F9C3" authorityName="Perkins" authorityYear="2017" box="[780,910,1581,1606]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="oscillata">
<emphasis id="B958EABFE32FFFB9FCEEF98BBAB2F9C3" box="[780,910,1581,1606]" italics="true" pageId="40" pageNumber="41">H. oscillata</emphasis>
</taxonomicName>
, but having a differently shaped pronotum that is more transverse and with more distinctly bisinuate lateral margins; also, the elytra are less ovate and the pronotal punctures smaller. The aedeagi of the two species markedly differ (
<figureCitation id="13172A28E32FFFB9FC65F9CABAD9F90D" box="[903,997,1647,1672]" captionStart="FIGURES 31 32" captionStartId="42.[151,264,1951,1974]" captionTargetBox="[366,1228,273,1907]" captionTargetId="figure@42.[290,1297,267,1930]" captionTargetPageId="42" captionText="FIGURES 31 32. Hydraena, aedeagi of H. impressicollis (lectotype) and H. bergsteni (holotype)." httpUri="https://zenodo.org/record/1041094/files/figure.png" pageId="40" pageNumber="41">Figs. 32</figureCitation>
,
<figureCitation id="13172A28E32FFFB9FC12F9CABD2CF90D" box="[1008,1040,1647,1672]" captionStart="FIGURES 35 36" captionStartId="46.[151,264,1842,1865]" captionText="FIGURES 35 36. Hydraena, aedeagi of holotypes." pageId="40" pageNumber="41">36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E32FFFB8FF25F934BC3CF862" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
<paragraph id="8B9336ADE32FFFB9FF25F934BA37F94E" blockId="40.[151,1437,1381,2006]" pageId="40" pageNumber="41">
<emphasis id="B958EABFE32FFFB9FF25F934B86DF92F" bold="true" box="[199,337,1681,1706]" pageId="40" pageNumber="41">Description</emphasis>
. Size:
<typeStatus id="5497880FE32FFFB9FE41F934BB39F92F" box="[419,517,1681,1706]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.29/0.93; head width 0.55; pronotum 0.53/0.68, PA 0.55, PB 0.58; elytra 1.45/0.93.
</paragraph>
<paragraph id="8B9336ADE32FFFB9FF75F970BD94F8AA" blockId="40.[151,1437,1381,2006]" pageId="40" pageNumber="41">Dorsum general coloration dark reddish brown; head darker than pronotum, almost piceous; pronotum narrowly bordered at anterior and posterior with light brown; elytra as dark as pronotal disc, except lighter humeral angles; legs brown to dark brown; maxillary palpi light brown, distal ½ of last palpomere not darker.</paragraph>
<paragraph id="8B9336ADE32FFFB9FF25F892B9C9F853" blockId="40.[151,1437,1381,2006]" pageId="40" pageNumber="41">Head with clypeus finely moderately densely punctate; frons slightly more coarsely punctate laterally than medially where interstices ca. 13xpd, each puncture with short distinctive seta. Pronotum densely coarsely punctate, interstices on disc ca. 0.51xpd, shining. Mentum moderately coarsely, sparsely punctate, interstices weakly shining; postmentum coarsely densely punctate. Genae raised, rugulose, with posterior ridge, interrupted in midline.</paragraph>
<paragraph id="8B9336ADE32EFFB8FF25FF3DBB88FF57" blockId="41.[151,1436,151,210]" pageId="41" pageNumber="42">Pronotum bisinuate laterally; anterior margin straight behind eyes, emarginate behind frons; PF1 and PF2 absent; PF3 moderately deep; PF4 very shallow.</paragraph>
<caption id="DF536625E32EFFB8FF75FA47BA83FA7D" box="[151,959,1506,1528]" pageId="41" pageNumber="42">
<paragraph id="8B9336ADE32EFFB8FF75FA47BA83FA7D" blockId="41.[151,959,1506,1528]" box="[151,959,1506,1528]" pageId="41" pageNumber="42">
<emphasis id="B958EABFE32EFFB8FF75FA47B823FA72" bold="true" box="[151,287,1506,1528]" pageId="41" pageNumber="42">FIGURE 30.</emphasis>
<taxonomicName id="4C2C4D2EE32EFFB8FEC7FA47B8D2FA7D" authorityName="Perkins" authorityYear="2017" box="[293,494,1506,1528]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B958EABFE32EFFB8FEC7FA47B8D2FA7D" box="[293,494,1506,1528]" italics="true" pageId="41" pageNumber="42">Hydraena bergsteni</emphasis>
</taxonomicName>
, holotype, habitus dorsal, lateral, and ventral.
</paragraph>
</caption>
<paragraph id="8B9336ADE32EFFB8FF25F9BABA7DF921" blockId="41.[151,1437,1567,2024]" pageId="41" pageNumber="42">Elytra weakly arcuate laterally; summit of moderately steep posterior declivity at ca. midlength; lateral explanate margins wide; serial punctures large, round, separated by ca. 1xpd, slightly smaller than largest pronotal punctures. Intervals not raised, shining, ca. 0.51xpd. Apices in dorsal aspect rather sharply conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="8B9336ADE32EFFB8FF25F90ABC3CF862" blockId="41.[151,1437,1567,2024]" pageId="41" pageNumber="42">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.75/3/1. P1 as wide as P2; median carina markedly sinuate in profile. P2 l/w ca.2/1, sides parallel, apex blunt and on same level as anteromedian carina of metaventrite. Plaques roundly raised, converging very slightly, at anterior extreme becoming carinate lines and joining anteromedian carina; separated basally by slightly more than plaque width, located at sides of median depression. Metaventrite anteromedian carina shorter than plaques, as 1 to 2, joining carinae of plaques, together forming inverted Y-shape; long and strong, slightly arcuate longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at arcuate posterior margin ca.
<date id="FF92106DE32EFFB8FBD0F822BD72F81A" box="[1074,1102,1927,1951]" pageId="41" pageNumber="42">1x</date>
P2. Protibia straight, slender. Mesotibia very slightly arcuate, middle 1/3 of medial margin with 5 or 6 short sharp spines, surface of tibia notched between spines. Metatibia straight, slender. Metafemur moderately slender. Abdominal apex arcuate.
</paragraph>
</subSubSection>
<subSubSection id="C3366526E32DFFBBFF25FF32B812FF50" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8B9336ADE32DFFBBFF25FF32B812FF50" blockId="42.[151,1436,151,213]" pageId="42" pageNumber="43">
<emphasis id="B958EABFE32DFFBBFF25FF32B872FF35" bold="true" box="[199,334,151,176]" pageId="42" pageNumber="43">Etymology.</emphasis>
Named in honor of Johannes Bergsten, who has collected many new species of Malagasy
<taxonomicName id="4C2C4D2EE32DFFBBFF75FF19B816FF50" authorityName="Mulsant" authorityYear="1844" box="[151,298,188,213]" class="Insecta" family="Hydraenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="family">Hydraenidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>