303 lines
46 KiB
XML
303 lines
46 KiB
XML
<document id="55F653BDA8D10CC841BC3AA4B6F42A74" ID-CLB-Dataset="5866" ID-DOI="10.5852/ejt.2021.757.1415" ID-GBIF-Dataset="9950dbcc-5627-479c-b9e7-abe6b8a0ddd1" ID-ISSN="2118-9773" ID-Zenodo-Dep="5047683" ID-ZooBank="F95D404B-0A7C-429F-BC4F-AAEDF9DC870C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625079024571" checkinUser="felipe" docAuthor="Bordera, Santiago & Broad, Gavin R." docDate="2021" docId="03858793FFD2FFA4DA08FEB67926FE75" docLanguage="en" docName="ejt-757_80-101.pdf" docOrigin="European Journal of Taxonomy 757" docSource="http://dx.doi.org/10.5852/ejt.2021.757.1415" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Handaoia cuscoensis Bordera 2021, sp. nov." docType="treatment" docUuid="49A19681-3B7F-4E6F-B95B-4E6847D1C5AF" docUuidSource="ZooBank" docVersion="8" lastPageNumber="85" masterDocId="FFBCFFEBFFD1FFA1D83CFFBE781FFF84" masterDocTitle="Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World" masterLastPageNumber="101" masterPageNumber="80" pageNumber="83" updateTime="1699021642505" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ACE6CCEA7340DEC7B9A20E497A051EFA">Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World</mods:title>
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<mods:name id="F8C0DD8F82DA3BA335B2CB3BB1460993" type="personal">
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<mods:roleTerm id="8763D4E69AACDDF5EDF84A75A1957E87">Author</mods:roleTerm>
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<mods:namePart id="3D485BB6FFD50C8715520D449F84196F">Bordera, Santiago</mods:namePart>
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<mods:affiliation id="67809925385B3438E3A05C7AD006E951">Departamento de Ciencias Ambientales y Recursos Naturales, University of Alicante, San Vicente del Raspeig, E- 03690 Alicante, Spain.</mods:affiliation>
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<mods:nameIdentifier id="CFEAB7346103B247988E97AE6B01005A" type="email">s.bordera@ua.es</mods:nameIdentifier>
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</mods:name>
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<mods:namePart id="50532CD6B0896A9468460C3654D12F54">Broad, Gavin R.</mods:namePart>
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<mods:affiliation id="AE9D4E85E4989E5018C50251D350DAE1">Department of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
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<mods:nameIdentifier id="1DEFCABA0DA6E732E56C2B44C83844A5" type="email">g.broad@nhm.ac.uk</mods:nameIdentifier>
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<mods:typeOfResource id="6F67F90927613307C624EA4728104370">text</mods:typeOfResource>
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<mods:title id="5ED00E08256A718FA6E5D7A5E9B08C85">European Journal of Taxonomy</mods:title>
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<mods:part id="E7725EA751893543BE0585A86548C4FD">
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<mods:date id="6616C4E592DE8A487A59620624D0DF7C">2021</mods:date>
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<mods:number id="C0CF4481690BBC1BA5B311327C4846E6">2021-06-30</mods:number>
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<mods:url id="65A4F5BA0268DF5D4BD7F566341E2D93">http://dx.doi.org/10.5852/ejt.2021.757.1415</mods:url>
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<treatment id="03858793FFD2FFA4DA08FEB67926FE75" ID-DOI="http://doi.org/10.5281/zenodo.5056066" ID-GBIF-Taxon="183231746" ID-Zenodo-Dep="5056066" LSID="urn:lsid:zoobank.org:act:49A19681-3B7F-4E6F-B95B-4E6847D1C5AF" httpUri="http://treatment.plazi.org/id/03858793FFD2FFA4DA08FEB67926FE75" lastPageId="5" lastPageNumber="85" pageId="3" pageNumber="83">
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<subSubSection id="C336650EFFD2FFA2DA08FEB67C1FFEA7" box="[564,1024,264,291]" pageId="3" pageNumber="83" type="nomenclature">
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<paragraph id="8B933685FFD2FFA2DA08FEB67C1FFEA7" blockId="3.[367,1220,264,361]" box="[564,1024,264,291]" pageId="3" pageNumber="83">
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<heading id="D0DB81E9FFD2FFA2DA08FEB67C1FFEA7" box="[564,1024,264,291]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="83" reason="2">
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<taxonomicName id="4C2C4D06FFD2FFA2DA08FEB67B82FEA6" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[564,925,264,291]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="83" phylum="Arthropoda" rank="species" species="cuscoensis" status="sp. nov.">
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<emphasis id="B958EA97FFD2FFA2DA08FEB67B27FEA6" bold="true" box="[564,824,264,291]" italics="true" pageId="3" pageNumber="83">Handaoia cuscoensis</emphasis>
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Bordera
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</taxonomicName>
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<taxonomicNameLabel id="A26B57ECFFD2FFA2DB99FEB77C1FFEA7" box="[933,1024,265,291]" pageId="3" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C336650EFFD2FFA2D953FE957B6FFEED" pageId="3" pageNumber="83" type="description">
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<paragraph id="8B933685FFD2FFA2D953FE957CDBFEC2" blockId="3.[367,1220,264,361]" box="[367,1220,299,326]" pageId="3" pageNumber="83">
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<uri id="FFBD3A87FFD2FFA2D953FE957CDBFEC2" box="[367,1220,299,326]" pageId="3" pageNumber="83">
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urn:lsid:zoobank.org:act:
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<uuid id="FF8A0C50FFD2FFA2DAAEFE957CDBFEC2" box="[658,1220,299,326]" pageId="3" pageNumber="83">49A19681-3B7F-4E6F-B95B-4E6847D1C5AF</uuid>
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</uri>
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</paragraph>
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<paragraph id="8B933685FFD2FFA2DAF8FEF17B6FFEED" blockId="3.[367,1220,264,361]" box="[708,880,335,361]" pageId="3" pageNumber="83">
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<figureCitation id="13172A00FFD2FFA2DAF8FEF17B3BFEED" box="[708,804,335,361]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="3" pageNumber="83">Figs 1A</figureCitation>
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,
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<figureCitation id="13172A00FFD2FFA2DB0CFEF17B5EFEED" box="[816,833,335,361]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">2</figureCitation>
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,
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<figureCitation id="13172A00FFD2FFA2DB71FEF17B6FFEED" box="[845,880,335,361]" captionStart="Fig" captionStartId="19.[186,229,1963,1989]" captionTargetBox="[189,1398,1195,1923]" captionTargetId="figure-309@19.[189,1398,1195,1923]" captionTargetPageId="19" captionText="Fig. 9. Distribution of New World species of Handaoia Seyrig, 1952. A. North and Central America. B. South America." figureDoi="http://doi.org/10.5281/zenodo.5047703" httpUri="https://zenodo.org/record/5047703/files/figure.png" pageId="3" pageNumber="83">9B</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C336650EFFD2FFA2D881FE2E7B6EFDC7" pageId="3" pageNumber="83" type="diagnosis">
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<paragraph id="8B933685FFD2FFA2D881FE2E7928FE2E" blockId="3.[189,311,400,426]" box="[189,311,400,426]" pageId="3" pageNumber="83">
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<heading id="D0DB81E9FFD2FFA2D881FE2E7928FE2E" bold="true" box="[189,311,400,426]" fontSize="11" level="3" pageId="3" pageNumber="83" reason="3">
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<emphasis id="B958EA97FFD2FFA2D881FE2E7928FE2E" bold="true" box="[189,311,400,426]" pageId="3" pageNumber="83">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B933685FFD2FFA2D881FE017B6EFDC7" blockId="3.[189,1399,447,579]" pageId="3" pageNumber="83">
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<taxonomicName id="4C2C4D06FFD2FFA2D881FE017A36FE5D" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[189,553,447,473]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="83" phylum="Arthropoda" rank="species" species="cuscoensis" status="sp. nov.">
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<emphasis id="B958EA97FFD2FFA2D881FE0179A0FE5D" box="[189,447,447,473]" italics="true" pageId="3" pageNumber="83">Handaoia cuscoensis</emphasis>
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Bordera
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</taxonomicName>
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<taxonomicNameLabel id="A26B57ECFFD2FFA2DA0AFE017A89FE5D" box="[566,662,447,473]" pageId="3" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
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can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum high, subtriangular with blunt tip (
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<figureCitation id="13172A00FFD2FFA2D8D0FDBB795CFD9B" box="[236,323,517,543]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2F</figureCitation>
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, arrow); juxtacoxal carina absent (
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<figureCitation id="13172A00FFD2FFA2DAE6FDBB7B28FD9B" box="[730,823,517,543]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2C</figureCitation>
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); area externa of propodeum confluent with area dentipara (
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<figureCitation id="13172A00FFD2FFA2D906FD967988FDC7" box="[314,407,552,579]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="3" pageNumber="83">Fig. 1A</figureCitation>
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); hind coxa tinged with white (
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<figureCitation id="13172A00FFD2FFA2DB39FD967B7BFDC7" box="[773,868,552,579]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2A</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C336650EFFD2FFA2D881FDD47CFAFD36" box="[189,1253,618,691]" pageId="3" pageNumber="83" type="etymology">
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<paragraph id="8B933685FFD2FFA2D881FDD4795AFD00" blockId="3.[189,325,618,644]" box="[189,325,618,644]" pageId="3" pageNumber="83">
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||
<heading id="D0DB81E9FFD2FFA2D881FDD4795AFD00" bold="true" box="[189,325,618,644]" fontSize="11" level="3" pageId="3" pageNumber="83" reason="3">
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||
<emphasis id="B958EA97FFD2FFA2D881FDD4795AFD00" bold="true" box="[189,325,618,644]" pageId="3" pageNumber="83">Etymology</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="8B933685FFD2FFA2D881FD267CFAFD36" blockId="3.[189,1253,664,691]" box="[189,1253,664,691]" pageId="3" pageNumber="83">
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The name of the species refers to the Department of
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<collectingRegion id="49E8F867FFD2FFA2DB19FD277B71FD37" box="[805,878,665,691]" country="Peru" name="Cusco" pageId="3" pageNumber="83">Cusco</collectingRegion>
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(
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<collectingCountry id="F33B7615FFD2FFA2DB42FD277BA6FD37" box="[894,953,665,691]" name="Peru" pageId="3" pageNumber="83">Peru</collectingCountry>
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), the
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<typeStatus id="54978827FFD2FFA2DBC5FD277C33FD37" box="[1017,1068,665,691]" pageId="3" pageNumber="83">type</typeStatus>
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locality region.
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</paragraph>
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</subSubSection>
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<subSubSection id="C336650EFFD2FFA2D881FD647CB5FC03" pageId="3" pageNumber="83" type="materials_examined">
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<paragraph id="8B933685FFD2FFA2D881FD6479B2FD70" blockId="3.[189,429,730,756]" box="[189,429,730,756]" pageId="3" pageNumber="83">
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<heading id="D0DB81E9FFD2FFA2D881FD6479B2FD70" bold="true" box="[189,429,730,756]" fontSize="11" level="3" pageId="3" pageNumber="83" reason="3">
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||
<emphasis id="B958EA97FFD2FFA2D881FD6479B2FD70" bold="true" box="[189,429,730,756]" pageId="3" pageNumber="83">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B933685FFD2FFA2D881FCB67A98FCA6" blockId="3.[189,647,776,802]" box="[189,647,776,802]" pageId="3" pageNumber="83">
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Only known from the
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<typeStatus id="54978827FFD2FFA2D9FEFCB67A36FCA6" box="[450,553,776,802]" pageId="3" pageNumber="83" type="holotype">holotype</typeStatus>
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female.
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</paragraph>
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<paragraph id="8B933685FFD2FFA2D8D6FCF47CB5FC03" blockId="3.[189,1194,842,904]" pageId="3" pageNumber="83">
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<materialsCitation id="3B443CD8FFD2FFA2D8D6FCF47CB5FC03" ID-GBIF-Occurrence="3312836310" collectingDate="1962-11-10" collectingDateMax="1962-11-15" collectingDateMin="1962-11-10" collectionCode="AEIC" collectorName="Marcapata & Luis Pena" country="Peru" elevation="750" location="Quincemil" pageId="3" pageNumber="83" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<emphasis id="B958EA97FFD2FFA2D8D6FCF47942FCE0" bold="true" box="[234,349,842,868]" pageId="3" pageNumber="83">
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||
<typeStatus id="54978827FFD2FFA2D8D6FCF47942FCE0" box="[234,349,842,868]" pageId="3" pageNumber="83" type="holotype">Holotype</typeStatus>
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</emphasis>
|
||
<collectingCountry id="F33B7615FFD2FFA2D881FCD07917FC0C" box="[189,264,878,904]" name="Peru" pageId="3" pageNumber="83">PERU</collectingCountry>
|
||
•
|
||
<specimenCount id="9D2AFD0CFFD2FFA2D91DFCD07927FC0C" box="[289,312,878,904]" pageId="3" pageNumber="83" type="female">♀</specimenCount>
|
||
;
|
||
<location id="8EF3605EFFD2FFA2D97AFCD379DAFC03" LSID="urn:lsid:plazi:treatment:03858793FFD2FFA4DA08FEB67926FE75:8EF3605EFFD2FFA2D97AFCD379DAFC03" box="[326,453,877,903]" country="Peru" name="Quincemil" pageId="3" pageNumber="83">Quincemil</location>
|
||
,
|
||
<quantity id="4CD49B60FFD2FFA2D9ECFCD37A05FC0C" box="[464,538,877,904]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="83" unit="m" value="750.0">
|
||
<elevation id="0001D1B6FFD2FFA2D9ECFCD37A05FC0C" box="[464,538,877,904]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="83" unit="m" value="750.0">750 m</elevation>
|
||
</quantity>
|
||
nr.
|
||
<collectorName id="26D95353FFD2FFA2DA7BFCD07AD8FC0C" box="[583,711,878,904]" pageId="3" pageNumber="83">Marcapata</collectorName>
|
||
;
|
||
<date id="FF921045FFD2FFA2DAEEFCD37B82FC03" box="[722,925,877,904]" pageId="3" pageNumber="83" value="1962-11-10" valueMax="1962-11-15" valueMin="1962-11-10">
|
||
<collectingDate id="EFD6E9ADFFD2FFA2DAEEFCD37B82FC03" box="[722,925,877,904]" pageId="3" pageNumber="83" value="1962-11-10" valueMax="1962-11-15" valueMin="1962-11-10">10–15 Nov. 1962</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D95353FFD2FFA2DB95FCD37C01FC0C" box="[937,1054,877,904]" pageId="3" pageNumber="83">Luis Peña</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED3DAE40FFD2FFA2DC61FCD37CBAFC03" box="[1117,1189,877,903]" collectionName="USA, Florida, Gainesville, American Entomological Institute" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1008" lsid="urn:lsid:biocol.org:col:1008" name="American Entomological Institute" pageId="3" pageNumber="83">AEIC</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336650EFFD2FFA4D881FC107926FE75" lastPageId="5" lastPageNumber="85" pageId="3" pageNumber="83" type="description">
|
||
<paragraph id="8B933685FFD2FFA2D881FC107950FC4C" blockId="3.[189,335,942,968]" box="[189,335,942,968]" pageId="3" pageNumber="83">
|
||
<heading id="D0DB81E9FFD2FFA2D881FC107950FC4C" bold="true" box="[189,335,942,968]" fontSize="11" level="3" pageId="3" pageNumber="83" reason="3">
|
||
<emphasis id="B958EA97FFD2FFA2D881FC107950FC4C" bold="true" box="[189,335,942,968]" pageId="3" pageNumber="83">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD2FFA2D8D6FC637959FC73" blockId="3.[189,1164,989,1051]" box="[234,326,989,1015]" pageId="3" pageNumber="83">
|
||
<emphasis id="B958EA97FFD2FFA2D8D6FC637959FC73" bold="true" box="[234,326,989,1015]" pageId="3" pageNumber="83">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD2FFA2D881FBBF7C93FB9E" blockId="3.[189,1164,989,1051]" box="[189,1164,1024,1051]" pageId="3" pageNumber="83">
|
||
MEASUREMENTS. Body length (without ovipositor)
|
||
<quantity id="4CD49B60FFD2FFA2DB3BFBBE7B7BFB9F" box="[775,868,1024,1051]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="83" unit="mm" value="3.8">3.8 mm</quantity>
|
||
. Fore wing
|
||
<quantity id="4CD49B60FFD2FFA2DBD3FBBE7C56FB9F" box="[1007,1097,1024,1051]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="83" unit="mm" value="2.9">2.9 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD2FFA2D881FBFC7B91FA1D" blockId="3.[189,1399,1090,1433]" pageId="3" pageNumber="83">
|
||
HEAD. Very finely and densely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (
|
||
<figureCitation id="13172A00FFD2FFA2D9D3FBDB7A51FBFB" box="[495,590,1125,1151]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2E</figureCitation>
|
||
); gena about 0.15 × length of eye in dorsal view; posterior ocellus separated from eye by 1.43 × its own maximum diameter; distance between posterior ocelli 1.14 × maximum ocellar diameter (
|
||
<figureCitation id="13172A00FFD2FFA2DA28FB157A6CFB41" box="[532,627,1195,1221]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2E</figureCitation>
|
||
); occipital carina gently V-shaped medially; malar space 1.33 × basal width of mandible; face moderately swollen medially (
|
||
<figureCitation id="13172A00FFD2FFA2DBA3FB707BE1FB6D" box="[927,1022,1230,1257]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2B</figureCitation>
|
||
); clypeus very weakly convex, almost flat, finely and slightly granulate with scattered punctures, its apex weakly rounded, about 2.3× as wide as high; mandible slightly tapered to apex, ventral tooth of mandible a little shorter than upper tooth; maxillary palp reaching at most ¾ length of mesosternum; flagellum widened in middle, tapered towards apex, flagellomeres 8–19 flattened ventrally (
|
||
<figureCitation id="13172A00FFD2FFA2DB74FAE57BB6FAF1" box="[840,937,1371,1397]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="3" pageNumber="83">Fig. 1A</figureCitation>
|
||
); flagellum with 20 segments, lengththickness ratios: 1
|
||
<superScript id="7C599BCDFFD2FFA2D9ADFAC07983FA08" attach="left" box="[401,412,1406,1420]" fontSize="6" pageId="3" pageNumber="83">st</superScript>
|
||
segment = 5.64, 2
|
||
<superScript id="7C599BCDFFD2FFA2DA4FFAC07A9BFA08" attach="left" box="[627,644,1406,1420]" fontSize="6" pageId="3" pageNumber="83">nd</superScript>
|
||
= 6.55 and 10
|
||
<superScript id="7C599BCDFFD2FFA2DB16FAC07B28FA08" attach="left" box="[810,823,1406,1420]" fontSize="6" pageId="3" pageNumber="83">th</superScript>
|
||
= 1.08.
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD2FFA2D881FA7E79ACF893" blockId="3.[187,1400,1472,1815]" pageId="3" pageNumber="83">
|
||
MESOSOMA. Very finely and densely granulate, matt (
|
||
<figureCitation id="13172A00FFD2FFA2DB75FA7E7BB3FA5E" box="[841,940,1472,1498]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2D</figureCitation>
|
||
); mesoscutum conspicuously convex, notauli absent (
|
||
<figureCitation id="13172A00FFD2FFA2D94BFA5D79CAFA79" box="[375,469,1507,1533]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2E</figureCitation>
|
||
); scutellum moderately convex, without lateral carinae; sternaulus very short and weak, as a wide V-shaped depression anteriorly, reaching at most 0.3× length of mesopleuron (
|
||
<figureCitation id="13172A00FFD2FFA2D8FAF997793DF9C0" box="[198,290,1577,1604]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2D</figureCitation>
|
||
); juxtacoxal carina barely indicated anteriorly (
|
||
<figureCitation id="13172A00FFD2FFA2DB7DF9977B83F9C0" box="[833,924,1577,1604]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2C</figureCitation>
|
||
); propodeum (
|
||
<figureCitation id="13172A00FFD2FFA2DC7FF9977C83F9C0" box="[1091,1180,1577,1604]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="3" pageNumber="83">Fig. 1A</figureCitation>
|
||
) with mostly weak carinae; area superomedia confluent with area basalis and separated from area petiolaris; area externa and area dentipara confluent; area spiracularis confluent with area lateralis; area dentipara section of lateral longitudinal carina very weak, partly absent; posterior transverse carina strong, forming elevate apophysis joining lateral longitudinal carina, apophysis subtriangular, with blunt tip (
|
||
<figureCitation id="13172A00FFD2FFA2DCA2F9087D08F955" box="[1182,1303,1718,1745]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2D, F</figureCitation>
|
||
, arrow). Length of hind femur about 4.4 × its height. Hind wing with vein
|
||
<emphasis id="B958EA97FFD2FFA2DBF0F9647C1EF977" box="[972,1025,1754,1779]" italics="true" pageId="3" pageNumber="83">cu-a</emphasis>
|
||
0.73 as long as abscissa of
|
||
<emphasis id="B958EA97FFD2FFA2DD73F9647D67F970" box="[1359,1400,1754,1780]" italics="true" pageId="3" pageNumber="83">CU</emphasis>
|
||
between
|
||
<emphasis id="B958EA97FFD2FFA2D91AF9437921F892" box="[294,318,1789,1814]" italics="true" pageId="3" pageNumber="83">M</emphasis>
|
||
and
|
||
<emphasis id="B958EA97FFD2FFA2D944F94379B2F892" box="[376,429,1789,1814]" italics="true" pageId="3" pageNumber="83">cu-a</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD2FFA2D881F8817D21F846" blockId="3.[189,1399,1854,1986]" pageId="3" pageNumber="83">
|
||
METASOMA. Tergite I (
|
||
<figureCitation id="13172A00FFD2FFA2D98BF8807A08F8DC" box="[439,535,1854,1880]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="3" pageNumber="83">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="13172A00FFD2FFA2DA1EF8807A58F8DC" box="[546,583,1854,1880]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">2G</figureCitation>
|
||
) finely granulate and very weakly strigose laterally, matt, 2.5× as long as its maximum width; lateromedian longitudinal carina absent; lateral longitudinal carina complete; tergite II (
|
||
<figureCitation id="13172A00FFD2FFA2D90FF83A7990F81B" box="[307,399,1924,1951]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2G</figureCitation>
|
||
) shallowly granulate, shiny, glabrous; remaining tergites (
|
||
<figureCitation id="13172A00FFD2FFA2DC07F83A7C89F81B" box="[1083,1174,1924,1951]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="3" pageNumber="83">Fig. 2G</figureCitation>
|
||
) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath about 0.4× as long as hind tibia.
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD5FFA4D881FEB77930FE08" blockId="4.[189,1399,265,327]" lastBlockId="5.[189,1400,265,396]" lastPageId="5" lastPageNumber="85" pageId="4" pageNumber="84">
|
||
COLOUR. Head (
|
||
<figureCitation id="13172A00FFD5FFA5D944FEB77A20FEA7" box="[376,575,265,291]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="4" pageNumber="84">Fig. 2A–B, D–E</figureCitation>
|
||
) light orange; antenna with scape and pedicel light brown, flagellum dark brown, distal part of flagellomere 4 to flagellomere 7 dorsally white (
|
||
<figureCitation id="13172A00FFD5FFA5DC2DFE927C71FEC2" box="[1041,1134,300,326]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="4" pageNumber="84">Fig. 2A</figureCitation>
|
||
). Mesosoma (
|
||
<figureCitation id="13172A00FFD5FFA4DD33FE9278CFFEA7" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" lastPageId="5" lastPageNumber="85" pageId="4" pageNumber="84">Fig. 2D– E</figureCitation>
|
||
) light orange. Metasoma (
|
||
<figureCitation id="13172A00FFD4FFA4DA32FEB77A73FEA7" box="[526,620,265,291]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="5" pageNumber="85">Fig. 2G</figureCitation>
|
||
) mostly brown, tergites I–II and anterior part of IV light orange. Legs (
|
||
<figureCitation id="13172A00FFD4FFA4D934FE92797AFEC2" box="[264,357,300,326]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="5" pageNumber="85">Fig. 2A</figureCitation>
|
||
) mostly light orange, distal tarsomeres, dark brown, fore and mid coxae, trochanters and trochantelli white, hind coxa white proximally, dark brown distally. Wings (
|
||
<figureCitation id="13172A00FFD4FFA4DC68FEF17CABFEED" box="[1108,1204,335,361]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="5" pageNumber="85">Fig. 2A</figureCitation>
|
||
) slightly evenly infuscate.
|
||
</paragraph>
|
||
<caption id="DF53660DFFD5FFA5D881F8DD7BB6F863" ID-DOI="http://doi.org/10.5281/zenodo.5047685" ID-Zenodo-Dep="5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="4" pageNumber="84" startId="4.[189,232,1891,1917]" targetBox="[189,1398,400,1853]" targetPageId="4">
|
||
<paragraph id="8B933685FFD5FFA5D881F8DD7BB6F863" blockId="4.[186,1399,1891,2023]" pageId="4" pageNumber="84">
|
||
<emphasis id="B958EA97FFD5FFA5D881F8DD7911F8F9" bold="true" box="[189,270,1891,1917]" pageId="4" pageNumber="84">Fig. 1.</emphasis>
|
||
Propodeum and tergite I of
|
||
<taxonomicName id="4C2C4D06FFD5FFA5DA55F8DD7B05F8FA" box="[617,794,1891,1918]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="84" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B958EA97FFD5FFA5DA55F8DD7AC0F8F9" box="[617,735,1891,1917]" italics="true" pageId="4" pageNumber="84">Handaoia</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, ♀♀.
|
||
<emphasis id="B958EA97FFD5FFA5DB59F8DD7B65F8F9" bold="true" box="[869,890,1891,1917]" pageId="4" pageNumber="84">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C2C4D06FFD5FFA5DBB0F8DA7C87F8F9" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[908,1176,1891,1917]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="84" phylum="Arthropoda" rank="species" species="cuscoensis" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD5FFA5DBB0F8DA7C30F8F9" box="[908,1071,1892,1917]" italics="true" pageId="4" pageNumber="84">H. cuscoensis</emphasis>
|
||
Bordera
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD5FFA5DC9EF8DA7D1EF8FA" box="[1186,1281,1892,1918]" pageId="4" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (AEIC).
|
||
<emphasis id="B958EA97FFD5FFA5D91DF839792AF825" bold="true" box="[289,309,1927,1953]" pageId="4" pageNumber="84">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C2C4D06FFD5FFA5D97CF8397985F825" authority="Bordera & Broad, 2021" authorityName="Bordera & Broad" authorityYear="2021" box="[320,410,1927,1953]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="84" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD5FFA5D97CF8397985F825" box="[320,410,1927,1953]" italics="true" pageId="4" pageNumber="84">H. fritzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD5FFA5D99CF83979E6F825" box="[416,505,1927,1953]" pageId="4" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (NHMUK).
|
||
<emphasis id="B958EA97FFD5FFA5DAC1F8387B0DF824" bold="true" box="[765,786,1926,1952]" pageId="4" pageNumber="84">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C2C4D06FFD5FFA5DB1DF8397C2AF824" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[801,1077,1926,1952]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="84" phylum="Arthropoda" rank="species" species="mercedensis" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD5FFA5DB1DF8397BCBF824" box="[801,980,1926,1952]" italics="true" pageId="4" pageNumber="84">H. mercedensis</emphasis>
|
||
Bordera
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD5FFA5DC07F8397C8AF825" box="[1083,1173,1927,1953]" pageId="4" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (CEUA).
|
||
<emphasis id="B958EA97FFD5FFA5D886F81478D0F840" bold="true" box="[186,207,1962,1988]" pageId="4" pageNumber="84">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C2C4D06FFD5FFA5D8DCF815799CF840" authority="Bordera & Broad, 2021" authorityName="Bordera & Broad" authorityYear="2021" box="[224,387,1962,1988]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="84" phylum="Arthropoda" rank="species" species="plaumanni" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD5FFA5D8DCF815799CF840" box="[224,387,1962,1988]" italics="true" pageId="4" pageNumber="84">H. plaumanni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD5FFA5D9B6F81479FAF840" box="[394,485,1962,1988]" pageId="4" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (NHMUK).
|
||
<emphasis id="B958EA97FFD5FFA5DACCF8147B1BF840" bold="true" box="[752,772,1962,1988]" pageId="4" pageNumber="84">E</emphasis>
|
||
.
|
||
<taxonomicName id="4C2C4D06FFD5FFA5DB2FF8157C30F840" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[787,1071,1962,1988]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="84" phylum="Arthropoda" rank="species" species="ruizcancinoi" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD5FFA5DB2FF8157BD5F840" box="[787,970,1963,1988]" italics="true" pageId="4" pageNumber="84">H. ruizcancinoi</emphasis>
|
||
Bordera
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD5FFA5DC0AF8147C8CF840" box="[1078,1171,1962,1988]" pageId="4" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (CEUA).
|
||
<emphasis id="B958EA97FFD5FFA5D880F87378D1F863" bold="true" box="[188,206,1997,2023]" pageId="4" pageNumber="84">F</emphasis>
|
||
.
|
||
<taxonomicName id="4C2C4D06FFD5FFA5D8E2F870794FF863" authority="Bordera & Broad, 2021" authorityName="Bordera & Broad" authorityYear="2021" box="[222,336,1998,2023]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="84" phylum="Arthropoda" rank="species" species="urceus" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD5FFA5D8E2F870794FF863" box="[222,336,1998,2023]" italics="true" pageId="4" pageNumber="84">H. urceus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD5FFA5D96BF87379ACF863" box="[343,435,1997,2023]" pageId="4" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (NHMUK). Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B933685FFD4FFA4D8D6FE0A7935FE4A" blockId="5.[189,313,436,497]" box="[234,298,436,462]" pageId="5" pageNumber="85">
|
||
<emphasis id="B958EA97FFD4FFA4D8D6FE0A7935FE4A" bold="true" box="[234,298,436,462]" pageId="5" pageNumber="85">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD4FFA4D881FE697926FE75" blockId="5.[189,313,436,497]" box="[189,313,471,497]" pageId="5" pageNumber="85">Unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |