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<document id="C4B59C899A0AC6FCB70C54E088904C77" ID-DOI="10.5281/zenodo.282239" ID-GBIF-Dataset="b8acb8ee-6490-4aa4-a002-52468cc2587d" ID-ISSN="1175-5326" ID-Zenodo-Dep="282239" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460523611886" checkinUser="plazi" docAuthor="Zhou, Yu-Lingzi &amp; Zhou, Hong-Zhang" docDate="2012" docId="0385D33C593A8C5AC5BAEC77FC3AF975" docLanguage="en" docName="zt03478p191.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Medhiama xiaolongmenensis Zhou &amp; Zhou, sp. n." docType="treatment" docVersion="9" lastPageNumber="175" masterDocId="FFBCAB44593E8C5CC52DE84DFFD4FFC4" masterDocTitle="Taxonomy of the genus Medhiama Bordoni, 2002 (Coleoptera: Staphylinidae, Staphylininae, Xantholinini) with descriptions of three new species" masterLastPageNumber="191" masterPageNumber="169" pageNumber="173" updateTime="1698314837013" updateUser="plazi">
<mods:mods id="D48BADE9F86E9B4122B54E71182BEE82" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DCF319A447C6767B611FE163AF1866A3">
<mods:title id="85638190F54F36383009F59C9204946B">Taxonomy of the genus Medhiama Bordoni, 2002 (Coleoptera: Staphylinidae, Staphylininae, Xantholinini) with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name id="3660D8B82AD3582A7A8789D90AFB0372" type="personal">
<mods:role id="8A7137DB10E3FE1B099002F476CA096D">
<mods:roleTerm id="6BF7489DDB7957D7CA45F95C94A44824">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3AFC6D6C029CC4705CC4B9E7AA5A58D0">Zhou, Yu-Lingzi</mods:namePart>
</mods:name>
<mods:name id="DF63BC76D3F2067F7B74636A4EB8D496" type="personal">
<mods:role id="1D039C417D6DF943100886E292BCE42E">
<mods:roleTerm id="D88D50ABCC7B4EB878B173FAD1CAFEEE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D5A0F77206A011F953DE848A81387C6">Zhou, Hong-Zhang</mods:namePart>
</mods:name>
<mods:typeOfResource id="DD7743A879D74C77B116AB926DC484E0">text</mods:typeOfResource>
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<mods:title id="8397EE47EBA35AA1EB6635F812C2C3E6">Zootaxa</mods:title>
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<mods:date id="FB8AA0FDD8136DFDFFC696A213273D3A">2012</mods:date>
<mods:detail id="44CFF8C9F3C0F966A076989E059A43BF" type="volume">
<mods:number id="92D748A20E153FFB1FA0D2FEBCE79D0C">3478</mods:number>
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<mods:classification id="1D9CCAD31E5CB3839BB2E2A178776B18">journal article</mods:classification>
<mods:identifier id="723C80A5A243ADC9CA5C17979E0ECEFB" type="DOI">10.5281/zenodo.282239</mods:identifier>
<mods:identifier id="2F3F2A4B6E8E418C1DE13F3D430AA0C8" type="GBIF-Dataset">b8acb8ee-6490-4aa4-a002-52468cc2587d</mods:identifier>
<mods:identifier id="BAC87C23E39CE1B14CA124F52CD6299C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="582754FCBCCCD67D72C514DAD865B1AD" type="Zenodo-Dep">282239</mods:identifier>
</mods:mods>
<treatment id="0385D33C593A8C5AC5BAEC77FC3AF975" ID-DOI="http://doi.org/10.5281/zenodo.6173768" ID-GBIF-Taxon="119526003" ID-Zenodo-Dep="6173768" LSID="urn:lsid:zoobank.org:act:1854EEB5-1041-418F-8C20-84187527388B" httpUri="http://treatment.plazi.org/id/0385D33C593A8C5AC5BAEC77FC3AF975" lastPageId="6" lastPageNumber="175" pageId="4" pageNumber="173">
<subSubSection id="C33631A1593A8C58C5BAEC77FCD4FB97" box="[151,768,1082,1107]" pageId="4" pageNumber="173" type="nomenclature">
<paragraph id="8B93622A593A8C58C5BAEC77FCD4FB97" blockId="4.[151,768,1082,1142]" box="[151,768,1082,1107]" pageId="4" pageNumber="173">
<heading id="D0DBD546593A8C58C5BAEC77FCD4FB97" bold="true" box="[151,768,1082,1107]" fontSize="10" level="2" pageId="4" pageNumber="173" reason="2">
<emphasis id="B958BE38593A8C58C5BAEC77FCD4FB97" bold="true" box="[151,768,1082,1107]" pageId="4" pageNumber="173">
1.
<taxonomicName id="4C2C19A9593A8C58C59EEC77FD60FB97" ID-CoL="3YNFP" authority="Zhou &amp; Zhou" authorityName="Zhou &amp; Zhou" box="[179,692,1082,1107]" class="Insecta" family="Staphylinidae" genus="Medhiama" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="xiaolongmenensis" status="sp. nov.">
<emphasis id="B958BE38593A8C58C59EEC77FDD3FB97" bold="true" box="[179,519,1082,1107]" italics="true" pageId="4" pageNumber="173">Medhiama xiaolongmenensis</emphasis>
Zhou &amp; Zhou
</taxonomicName>
,
<taxonomicNameLabel id="A26B0343593A8C58C7ECEC76FCD4FB97" box="[705,768,1083,1107]" pageId="4" pageNumber="173" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33631A1593A8C58C5BAEC13FEF5FBB2" box="[151,289,1118,1142]" pageId="4" pageNumber="173" type="description">
<paragraph id="8B93622A593A8C58C5BAEC13FEF5FBB2" blockId="4.[151,768,1082,1142]" box="[151,289,1118,1142]" pageId="4" pageNumber="173">
(
<figureCitation id="13177EAF593A8C58C5B2EC13FF34FBB2" box="[159,224,1118,1142]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="4" pageNumber="173">Fig. 3</figureCitation>
A–H)
</paragraph>
</subSubSection>
<subSubSection id="C33631A1593A8C58C5BAECEFFDE5FA8D" pageId="4" pageNumber="173" type="materials_examined">
<paragraph id="8B93622A593A8C58C5BAECEFFDE5FA8D" blockId="4.[151,1436,1185,2037]" pageId="4" pageNumber="173">
<emphasis id="B958BE38593A8C58C5BAECEFFE96FB7E" bold="true" box="[151,322,1185,1210]" pageId="4" pageNumber="173">
<typeStatus id="5497DC88593A8C58C5BAECEFFF04FB7E" box="[151,208,1186,1210]" pageId="4" pageNumber="173">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5497DC88593A8C58C466ECECFE6CFB7D" box="[331,440,1185,1209]" pageId="4" pageNumber="173" type="holotype">Holotype</typeStatus>
: male,
<emphasis id="B958BE38593A8C58C728ECECFD05FB7E" bold="true" box="[517,721,1185,1210]" pageId="4" pageNumber="173">
<collectingCountry id="F33B22BA593A8C58C728ECECFDB2FB7E" box="[517,614,1185,1210]" name="China" pageId="4" pageNumber="173">CHINA</collectingCountry>
: Beijing:
</emphasis>
Mentougou District: Xiaolongmen,
<quantity id="4CD4CFCF593A8C58C145ECECFB6BFB7D" box="[1128,1215,1185,1209]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="173" unit="m" value="1600.0">1600 m</quantity>
,
<date id="FF9244EA593A8C58C1E7ECECFA82FB7D" box="[1226,1366,1185,1209]" pageId="4" pageNumber="173" value="1998-07-03">03.VII.1998</date>
, Zhou Haisheng collected (IZ-CAS);
<typeStatus id="5497DC88593A8C58C4D1EC8BFDB9FB1A" box="[508,621,1222,1246]" pageId="4" pageNumber="173" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D2AA9A3593A8C58C751EC88FD0EFB19" box="[636,730,1221,1245]" pageId="4" pageNumber="173" type="male">2 males</specimenCount>
,
<specimenCount id="9D2AA9A3593A8C58C7C7EC88FCBEFB19" box="[746,874,1221,1245]" pageId="4" pageNumber="173" type="female">11 females</specimenCount>
, same data as
<typeStatus id="5497DC88593A8C58C136EC88FB55FB19" box="[1051,1153,1221,1245]" pageId="4" pageNumber="173" type="holotype">holotype</typeStatus>
;
<emphasis id="B958BE38593A8C58C1BDEC88FA40FB19" bold="true" box="[1168,1428,1221,1246]" pageId="4" pageNumber="173">Sichuan: Baoxing co.</emphasis>
: Longdong town, Ganyang Gou,
<specimenCount id="9D2AA9A3593A8C58C739ECA4FDB3FAC5" box="[532,615,1257,1281]" pageId="4" pageNumber="173" type="male">1 male</specimenCount>
,
<specimenCount id="9D2AA9A3593A8C58C75AECA4FD3FFAC5" box="[631,747,1257,1281]" pageId="4" pageNumber="173" type="female">5 females</specimenCount>
,
<quantity id="4CD4CFCF593A8C58C7D6ECA4FC83FAC5" box="[763,855,1257,1281]" metricMagnitude="3" metricUnit="m" metricValue="1.796" pageId="4" pageNumber="173" unit="m" value="1796.0">1796 m</quantity>
,
<date id="FF9244EA593A8C58C645ECA4FC29FAC5" box="[872,1021,1257,1281]" pageId="4" pageNumber="173" value="2003-08-12">12.VIII.2003</date>
, Wu Jie collected; same locality,
<specimenCount id="9D2AA9A3593A8C58C0A3ECA4FF32FAE1" pageId="4" pageNumber="173" type="female">1 female</specimenCount>
,
<quantity id="4CD4CFCF593A8C58C5DFED40FE9EFAE1" box="[242,330,1293,1317]" metricMagnitude="3" metricUnit="m" metricValue="1.795" pageId="4" pageNumber="173" unit="m" value="1795.0">1795 m</quantity>
,
<date id="FF9244EA593A8C58C47AED40FE38FAE1" box="[343,492,1293,1317]" pageId="4" pageNumber="173" value="2003-08-12">12.VIII.2003</date>
, Yu Xiongdong collected; Puji Gou,
<specimenCount id="9D2AA9A3593A8C58C6A2ED40FC20FAE1" box="[911,1012,1293,1317]" pageId="4" pageNumber="173" type="female">1 female</specimenCount>
,
<quantity id="4CD4CFCF593A8C58C12DED40FB8CFAE1" box="[1024,1112,1293,1317]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="4" pageNumber="173" unit="m" value="2180.0">2180 m</quantity>
,
<date id="FF9244EA593A8C58C148ED40FB2EFAE1" box="[1125,1274,1293,1317]" pageId="4" pageNumber="173" value="2003-08-14">14.VIII.2003</date>
, Yu Xiaodong &amp; Peng Xuewei collected (IZ-CAS).
</paragraph>
</subSubSection>
<subSubSection id="C33631A1593A8C5AC5E8ED18FE68FA97" lastPageId="6" lastPageNumber="175" pageId="4" pageNumber="173" type="description">
<paragraph id="8B93622A593A8C58C5E8ED18FD5AFA55" blockId="4.[151,1436,1185,2037]" pageId="4" pageNumber="173">
<emphasis id="B958BE38593A8C58C5E8ED18FDC4FAA9" bold="true" box="[197,528,1365,1390]" pageId="4" pageNumber="173">Description. Measurement.</emphasis>
BL=
<quantity id="4CD4CFCF593A8C58C77FED18FD14FAA9" box="[594,704,1365,1389]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" pageId="4" pageNumber="173" unit="mm" value="6.75">6.75 mm</quantity>
, FL=
<quantity id="4CD4CFCF593A8C58C62EED18FCA5FAA9" box="[771,881,1365,1389]" metricMagnitude="-3" metricUnit="m" metricValue="3.52" pageId="4" pageNumber="173" unit="mm" value="3.52">3.52 mm</quantity>
, HL=
<quantity id="4CD4CFCF593A8C58C697ED18FBF3FAA9" box="[954,1063,1365,1389]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="4" pageNumber="173" unit="mm" value="1.16">1.16 mm</quantity>
, HW=
<quantity id="4CD4CFCF593A8C58C154ED18FB32FAA9" box="[1145,1254,1365,1389]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="4" pageNumber="173" unit="mm" value="0.86">0.86 mm</quantity>
, PL=
<quantity id="4CD4CFCF593A8C58C006ED18FA43FAA9" box="[1323,1431,1365,1389]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="173" unit="mm" value="1.2">1.20 mm</quantity>
, PW=
<quantity id="4CD4CFCF593A8C58C5FFED34FEEDFA55" box="[210,313,1401,1425]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="4" pageNumber="173" unit="mm" value="0.74">0.74 mm</quantity>
, EL=
<quantity id="4CD4CFCF593A8C58C45AED34FE08FA55" box="[375,476,1401,1425]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="173" unit="mm" value="1.05">1.05 mm</quantity>
, EW=
<quantity id="4CD4CFCF593A8C58C708ED34FD5EFA55" box="[549,650,1401,1425]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="173" unit="mm" value="1.05">1.05 mm</quantity>
.
</paragraph>
<paragraph id="8B93622A593A8C58C5E8EDD0FC34FA1D" blockId="4.[151,1436,1185,2037]" pageId="4" pageNumber="173">Body nearly cylindrical and medium sized. Entirely dark brown. Legs brown. Antennae dark brown, apical half of last antennomere lighter in color. Maxillary and labial palpi dark brown.</paragraph>
<paragraph id="8B93622A593A8C58C5E8EDA8FE6BF8A1" blockId="4.[151,1436,1185,2037]" pageId="4" pageNumber="173">
<emphasis id="B958BE38593A8C58C5E8EDA8FED1FA39" bold="true" box="[197,261,1509,1533]" pageId="4" pageNumber="173">Head</emphasis>
. Oblong (HL to HW ratio 1.36), tempora widened posteriorly, posterior angles widely rounded. Dorsal integument entirely bearing microsculpture composed of distinct polygonal reticulum, and extensively distributed deep and medium sized punctures, puncture intervals rather narrow, about 1/2 puncture diameter. Median longitudinal region impunctate, width equals to 2–3 puncture diameters; frontal region with distinct transverse microstriae, and 3 pairs of tiny punctures. Each side of cranium without anterolateral puncture, but with midlateral puncture far from dorsal margin of eye (5–6 puncture diameters to eye), temporal puncture rather near lateral margin (at lateral 1/6) and occipital puncture rather near posterior margin (at posterior 1/6). Frontal furrows deep and long, convergent backwards; anteocular furrows indistinct, scarcely observable. Eye of small size and flat, diameter nearly 1/4 of temporal length (eye: tempora = 0.19:
<quantity id="4CD4CFCF593A8C58C661EF48FC6CF8D9" box="[844,952,1797,1821]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="4" pageNumber="173" unit="mm" value="0.76">0.76 mm</quantity>
). Epistoma relatively wide, subrectangular and flat, with a pair of tiny punctures. Distance between antennal insertions
<quantity id="4CD4CFCF593A8C58C12CEF64FBBDF885" box="[1025,1129,1833,1857]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="173" unit="mm" value="0.23">0.23 mm</quantity>
, subequal to distance from antenna to eye (
<quantity id="4CD4CFCF593A8C58C464EF00FE67F8A1" box="[329,435,1869,1893]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="173" unit="mm" value="0.21">0.21 mm</quantity>
).
</paragraph>
<paragraph id="8B93622A593A8C58C5E8EF3CFE65F831" blockId="4.[151,1436,1185,2037]" pageId="4" pageNumber="173">
<emphasis id="B958BE38593A8C58C5E8EF3CFEECF84E" bold="true" box="[197,312,1905,1930]" pageId="4" pageNumber="173">Antennae</emphasis>
. Scape stout, thickened apically, much longer than three subsequent antennomeres combined,
<quantity id="4CD4CFCF593A8C58C046EF3CFF1CF869" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="173" unit="mm" value="0.44">0.44 mm</quantity>
; antennomere II slightly elongate or subglobular,
<quantity id="4CD4CFCF593A8C58C7D0EFD8FCB1F869" box="[765,869,1941,1965]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="173" unit="mm" value="0.1">0.10 mm</quantity>
; III elongate, slightly longer than II,
<quantity id="4CD4CFCF593A8C58C027EFD8FAA6F869" box="[1290,1394,1941,1965]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="173" unit="mm" value="0.11">0.11 mm</quantity>
; IV and V subequal in length,
<quantity id="4CD4CFCF593A8C58C4E2EFF4FDEDF815" box="[463,569,1977,2001]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="173" unit="mm" value="0.09">0.09 mm</quantity>
; last antennomere of medium length,
<quantity id="4CD4CFCF593A8C58C6DEEFF4FB88F815" box="[1011,1116,1977,2001]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="173" unit="mm" value="0.19">0.19 mm</quantity>
, subequal to two preceding antennomeres combined.
</paragraph>
<caption id="DF5332A2593B8C59C5BAEF18FF11F86D" httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="5" pageNumber="174" targetBox="[170,1412,208,1842]" targetPageId="5">
<paragraph id="8B93622A593B8C59C5BAEF18FF11F86D" blockId="5.[151,1436,1877,1961]" pageId="5" pageNumber="174">
<emphasis id="B958BE38593B8C59C5BAEF18FEC6F8AF" bold="true" box="[151,274,1877,1900]" pageId="5" pageNumber="174">FIGURE 3.</emphasis>
<taxonomicName id="4C2C19A9593B8C59C434EF1BFD94F8A8" box="[281,576,1878,1900]" class="Insecta" family="Staphylinidae" genus="Medhiama" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="174" phylum="Arthropoda" rank="species" species="xiaolongmenensis" status="sp. nov.">
<emphasis id="B958BE38593B8C59C434EF1BFD94F8A8" box="[281,576,1878,1900]" italics="true" pageId="5" pageNumber="174">Medhiama xiaolongmenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B0343593B8C59C76AEF1BFDABF8AF" box="[583,639,1878,1899]" pageId="5" pageNumber="174" rank="species">
<emphasis id="B958BE38593B8C59C76AEF1BFDABF8AF" bold="true" box="[583,639,1878,1899]" pageId="5" pageNumber="174">sp. n.</emphasis>
</taxonomicNameLabel>
: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm.
</paragraph>
</caption>
<paragraph id="8B93622A59388C5AC5E8E8D5FD84FF32" blockId="6.[153,1437,151,1713]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8E8D5FE80FF74" bold="true" box="[197,340,152,176]" pageId="6" pageNumber="175">Mouthparts</emphasis>
. Labrum bilobed and relatively long, with a median longitudinal groove. Maxillary palpus elongate, segment III longest, last segment slender and obconical and shorter than the penultimate. Labial palpus distinctly slender, last segment longest.
</paragraph>
<paragraph id="8B93622A59388C5AC5E8E94CFA46FEDC" blockId="6.[153,1437,151,1713]" box="[197,1426,256,281]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8E94CFED5FEDD" bold="true" box="[197,257,257,281]" pageId="6" pageNumber="175">Neck</emphasis>
. Medium width (
<quantity id="4CD4CFCF59388C5AC4E9E94DFDF9FEDC" box="[452,557,256,280]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="175" unit="mm" value="0.3">0.30 mm</quantity>
), nearly 1/3 of head width, with a transverse substraight ridge on anterior 1/3.
</paragraph>
<paragraph id="8B93622A59388C5AC5E8E969FEE0FE03" blockId="6.[153,1437,151,1713]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8E969FEEAFEF8" bold="true" box="[197,318,292,316]" pageId="6" pageNumber="175">Pronotum</emphasis>
. Distinctly elongate (PL to PW ratio 1.62), of same length as head, but narrower. Widest at anterior 1/3, narrowest at posterior 1/3. Anterior angles widely rounded, not protruding, lateral margins sinuate from middle, posterior angles rounded. Integument bearing distinct transverse microstriae, and a pair of admedian row of 12–14 punctures, smaller than those on head. Areas outside admedian rows with additional, irregular, sparsely set punctures.
</paragraph>
<paragraph id="8B93622A59388C5AC5E8E99EFF10FDC9" blockId="6.[153,1437,151,1713]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8E99EFEABFE2F" bold="true" box="[197,383,467,491]" pageId="6" pageNumber="175">Mesoscutellum.</emphasis>
Shiny, extensively bearing polygonal reticulum and with a pair of small punctures on apical 1/4.
</paragraph>
<paragraph id="8B93622A59388C5AC5E8EA55FC95FD5D" blockId="6.[153,1437,151,1713]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8EA55FEC5FDF4" bold="true" box="[197,273,536,560]" pageId="6" pageNumber="175">Elytra</emphasis>
. Subquadrate (EL to EW ratio 1.0), shorter but distinctly wider than pronotum. Humeri well developed, lateral margins widened posteriorly, hind margin rounded. Integument slightly wrinkled, without microsculpture; each side symmetrically with regular rows of punctures, interspaces between rows 1–2 puncture diameters; deflexed portion of each elytron with 3–4 rows of punctures.
</paragraph>
<paragraph id="8B93622A59388C5AC5E8EAE9FD62FCC5" blockId="6.[153,1437,151,1713]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8EAE9FF29FD78" bold="true" box="[197,253,676,700]" pageId="6" pageNumber="175">Legs</emphasis>
. First four segments stout but not dilated in protarsi and metatarsi, whereas relatively slender in mesotarsi; each last segment as long as the II–IV combined. Protibia with apical ctenidium and subapical ctenidia, meso- and metatibia only with apical ctenidium.
</paragraph>
<paragraph id="8B93622A59388C5AC5E8EB41FCC6FC17" blockId="6.[153,1437,151,1713]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8EB41FE94FCE1" bold="true" box="[197,320,780,805]" pageId="6" pageNumber="175">Abdomen.</emphasis>
Cylindrical, broadest at segment VII. Tergites III–VII shiny, surface entirely covered with observable transverse microstriae, also with denser polygonal reticulum near anterior margin; punctures small, sparsely scattered, interspace between them approximately 3–4 puncture diameters, but much denser on tergites VII and VIII. Each tergite without distinct basal impression near anterior margin. Surface between two basal transverse carinae of tergites III–VII bearing distinct polygonal reticulum. All abdominal sternites shiny, with microstriae and setiferous punctures as those on tergites.
</paragraph>
<paragraph id="8B93622A59388C5AC5E8EB93FED1FB2F" blockId="6.[153,1437,151,1713]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8EB93FED5FC32" bold="true" box="[197,257,990,1014]" pageId="6" pageNumber="175">Male</emphasis>
(
<figureCitation id="13177EAF59388C5AC439EB93FE83FC32" box="[276,343,990,1014]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
A–F). Abdominal segment VIII entirely covered with setiferous punctures, posterior margin of tergite VIII subtruncated, that of sternite VIII sharply protruding posteriorly (
<figureCitation id="13177EAF59388C5AC6D2EC4CFB95FBDD" box="[1023,1089,1025,1049]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
A, B). Tergite IX symmetrical, connected mediobasally. Sternite IX, with long linear-shaped base and sharp apex (
<figureCitation id="13177EAF59388C5AC18AEC69FB27FBF8" box="[1191,1267,1060,1084]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
D). Tergite X symmetrical, broadest at anterior 1/3, lateral margins quite expanded outwards (
<figureCitation id="13177EAF59388C5AC11CEC0AFBA2FB9B" box="[1073,1142,1095,1119]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
C). Aedeagus of irregular shape and small in size (
<figureCitation id="13177EAF59388C5AC481EC27FE3AFB46" box="[428,494,1130,1154]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
E, F), basal bulb
<quantity id="4CD4CFCF59388C5AC782EC27FCC0FB46" box="[687,788,1130,1154]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="175" unit="mm" value="0.48">0.48 mm</quantity>
long. Parameres asymmetrical and foot-shaped, right one obviously larger than left one in dorsal view (
<figureCitation id="13177EAF59388C5AC794ECC0FD2AFB61" box="[697,766,1165,1189]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
F), base distinctly darker; large sized,
<quantity id="4CD4CFCF59388C5AC1E9ECC0FAF8FB61" box="[1220,1324,1165,1189]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="175" unit="mm" value="0.4">0.40 mm</quantity>
long and subequal to 4/5 of basal bulb length. Internal sac located outside basal bulb, brown and soft, without distinct spines (
<figureCitation id="13177EAF59388C5AC58CEC9EFF37FB2F" box="[161,227,1235,1259]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
E).
</paragraph>
<paragraph id="8B93622A59388C5AC5E8ECBBFE68FA97" blockId="6.[153,1437,151,1713]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8ECBBFEC8FACA" bold="true" box="[197,284,1270,1294]" pageId="6" pageNumber="175">Female</emphasis>
(
<figureCitation id="13177EAF59388C5AC400ECBBFEBAFACA" box="[301,366,1270,1294]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
G, H). Sternite VIII not distinctly oblong, and posterior margin protruding medially (
<figureCitation id="13177EAF59388C5AC01EECBBFAA3FACA" box="[1331,1399,1270,1294]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
G). Genital segment with pair of subtriangular supplementary sclerites and broad sternite with distinctly protruding anterior margin (
<figureCitation id="13177EAF59388C5AC478ED76FE43FA97" box="[341,407,1339,1363]" captionStart="FIGURE 3" captionStartId="5.[151,250,1877,1900]" captionTargetBox="[170,1412,208,1842]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Medhiama xiaolongmenensis sp. n.: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, ventral view; F. aedeagus, dorsal view; G. female sternite VIII; H. female genital segment. Scale bars 0.15 mm." httpUri="https://zenodo.org/record/282241/files/figure.png" pageId="6" pageNumber="175">Fig. 3</figureCitation>
H).
</paragraph>
</subSubSection>
<subSubSection id="C33631A159388C5AC5E8ED13FD50FAB2" box="[197,644,1374,1399]" pageId="6" pageNumber="175" type="distribution">
<paragraph id="8B93622A59388C5AC5E8ED13FD50FAB2" blockId="6.[153,1437,151,1713]" box="[197,644,1374,1399]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8ED13FE8AFAB3" bold="true" box="[197,350,1374,1399]" pageId="6" pageNumber="175">Distribution.</emphasis>
<collectingCountry id="F33B22BA59388C5AC448ED13FE7CFAB2" box="[357,424,1374,1398]" name="China" pageId="6" pageNumber="175">China</collectingCountry>
(Beijing, Sichuan).
</paragraph>
</subSubSection>
<subSubSection id="C33631A159388C5AC5E8EDCFFA49FA5D" box="[197,1437,1409,1434]" pageId="6" pageNumber="175" type="etymology">
<paragraph id="8B93622A59388C5AC5E8EDCFFA49FA5D" blockId="6.[153,1437,151,1713]" box="[197,1437,1409,1434]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8EDCFFE90FA5E" bold="true" box="[197,324,1410,1434]" pageId="6" pageNumber="175">Etymology</emphasis>
. The specific epithet is derived from the Chinese name (Pin-yin) of the
<typeStatus id="5497DC8859388C5AC14BEDCFFB42FA5E" box="[1126,1174,1410,1434]" pageId="6" pageNumber="175">type</typeStatus>
locality, Xiaolongmen.
</paragraph>
</subSubSection>
<subSubSection id="C33631A159388C5AC5E8EDE8FC3AF975" pageId="6" pageNumber="175" type="discussion">
<paragraph id="8B93622A59388C5AC5E8EDE8FC3AF975" blockId="6.[153,1437,151,1713]" pageId="6" pageNumber="175">
<emphasis id="B958BE3859388C5AC5E8EDE8FEEDFA79" bold="true" box="[197,313,1445,1469]" pageId="6" pageNumber="175">Remarks.</emphasis>
This new species could be distinguished from its congeners by the following character combination: in male by widened anterior angles of pronotum, protruding posterior margin of male sternite VIII, distinctive shape of sternite IX, tergite X, asymmetrical parameres, unique form of inner sac, and in female by broad sternite of genital segment with a distinctly protruding anterior margin. This is the second species of the genus with asymmetrical parameres (the other being
<taxonomicName id="4C2C19A959388C5AC740EE7CFC51F98C" authority="Bordoni, 2003" authorityName="Bordoni" authorityYear="2003" box="[621,901,1584,1608]" class="Insecta" family="Staphylinidae" genus="Medhiama" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="puetzi">
<emphasis id="B958BE3859388C5AC740EE7CFD02F98C" box="[621,726,1585,1608]" italics="true" pageId="6" pageNumber="175">M. puetzi</emphasis>
Bordoni, 2003
</taxonomicName>
), and the two could be distinguished from each other by different shapes of sternite IX, tergite X and aedeagi. The new species also shared similar foot-shaped parameres with
<taxonomicName id="4C2C19A959388C5AC47DEE3AFDB6F94A" authority="Bordoni" authorityName="Bordoni" box="[336,610,1654,1678]" class="Insecta" family="Staphylinidae" genus="Medhiama" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="schawalleri">
<emphasis id="B958BE3859388C5AC47DEE3AFE2EF94A" box="[336,506,1654,1678]" italics="true" pageId="6" pageNumber="175">M. schawalleri</emphasis>
Bordoni
</taxonomicName>
, but the pronotum widened distinctly near anterior angles, punctures on abdomen more densely distributed, and tergite X more outwardly expanded.
</paragraph>
</subSubSection>
</treatment>
</document>
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