treatments-xml/data/03/85/06/03850669FF85FFD6FE6425543A2AC9B3.xml
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<document id="6C4931DBE926E82BD9BE09E1C17804E5" ID-CLB-Dataset="8176" ID-DOI="10.5852/ejt.2021.734.1233" ID-GBIF-Dataset="fd8f5cb9-13e9-4c90-ac15-502fa7ee8d38" ID-ISSN="2118-9773" ID-Zenodo-Dep="4542107" ID-ZooBank="663A0C2D-E3CF-4B0C-8BA1-566B174829A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613423098920" checkinUser="plazi" docAuthor="Montoya-Molina, Santiago &amp; Vaz-De, Fernando Z." docDate="2021" docId="03850669FF85FFD6FE6425543A2AC9B3" docLanguage="en" docName="ejt-734_1-64.pdf" docOrigin="European Journal of Taxonomy 734" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius (Luederwaldtinia) deyrollei" docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="FFBC7E11FF90FFCEFFAD26453B59CB56" masterDocTitle="Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="22" updateTime="1698929631379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="111D97F2677A052744683170578BA8B5">Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="17DD41FF950FE63DE4134A2004F2C3C0">Montoya-Molina, Santiago</mods:namePart>
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<mods:affiliation id="0FB8E304887B6A970ACC3D18FA1C3105">Federal University of Mato Grosso, Institute of Biosciences, Programa de Pós Graduaç „ o em Ecología e Conservaç „ o da Biodiversidade, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil. Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 129, Praha-Suchdol, 16500, Czech Republic. Federal University of Mato Grosso, Institute of Biosciences, Department of Biology and Zoology, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil.</mods:affiliation>
<mods:nameIdentifier id="C05C6CBC41C8339FA9EACB7DC9B3C6C4" type="email">horomo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="045ECAAA36A5A426D6FC123621F0FCB8">Vaz-De, Fernando Z.</mods:namePart>
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<mods:date id="3B85DAB862052739B2EB8C24C9EA62C4">2021</mods:date>
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<treatment id="03850669FF85FFD6FE6425543A2AC9B3" ID-DOI="http://doi.org/10.5281/zenodo.4545849" ID-GBIF-Taxon="177420817" ID-Zenodo-Dep="4545849" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3" httpUri="http://treatment.plazi.org/id/03850669FF85FFD6FE6425543A2AC9B3" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
<subSubSection id="C336E4F4FF85FFDBFE6425543F33C87B" box="[457,1130,785,813]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8B93B77FFF85FFDBFE6425543F33C87B" blockId="21.[457,1130,785,849]" box="[457,1130,785,813]" pageId="21" pageNumber="22">
<heading id="D0DB0013FF85FFDBFE6425543F33C87B" box="[457,1130,785,813]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="22" reason="2">
<taxonomicName id="4C2CCCFCFF85FFDBFE6425543F33C87B" ID-CoL="35NBJ" authority="(Harold, 1869)" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[457,1130,785,813]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="deyrollei" subGenus="Luederwaldtinia">
<emphasis id="B9586B6DFF85FFDBFE642554393DC87A" bold="true" box="[457,612,785,812]" italics="true" pageId="21" pageNumber="22">Dichotomius</emphasis>
(
<emphasis id="B9586B6DFF85FFDBFDD825543865C87A" bold="true" box="[629,828,785,812]" italics="true" pageId="21" pageNumber="22">Luederwaldtinia</emphasis>
)
<emphasis id="B9586B6DFF85FFDBFCE0255438ECC87A" bold="true" box="[845,949,785,812]" italics="true" pageId="21" pageNumber="22">deyrollei</emphasis>
(
<bibRefCitation id="EFBDCA8EFF85FFDBFC6925573F3BC87B" author="Harold E." box="[964,1122,786,813]" pageId="21" pageNumber="22" pagination="124 - 144" refId="ref29859" refString="Harold E. 1869. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 124 - 144." type="journal article" year="1869">Harold, 1869</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FF85FFDBFD6725733830C807" box="[714,873,822,849]" pageId="21" pageNumber="22" type="description">
<paragraph id="8B93B77FFF85FFDBFD6725733830C807" blockId="21.[457,1130,785,849]" box="[714,873,822,849]" pageId="21" pageNumber="22">
<figureCitation id="1317ABFAFF85FFDBFD6725733848C806" box="[714,785,822,848]" captionStart="Fig" captionStartId="23.[189,232,1927,1953]" captionTargetBox="[210,1378,272,1894]" captionTargetId="figure-59@23.[204,1383,265,1899]" captionTargetPageId="23" captionText="Fig. 7. Dichotomius deyrollei (Harold, 1869). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the head cephalic carina. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Metasternum dense pilosity." figureDoi="http://doi.org/10.5281/zenodo.4542121" httpUri="https://zenodo.org/record/4542121/files/figure.png" pageId="21" pageNumber="22">Fig. 7</figureCitation>
, Map 5
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FF85FFDBFF10253C3E2EC8AF" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="8B93B77FFF85FFDBFF10253C3833C8C5" blockId="21.[189,874,888,915]" box="[189,874,888,915]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB3C9051FF85FFDBFF10253C3833C8C5" box="[189,874,888,915]" pageId="21" pageNumber="22">
<treatmentCitation id="0A8D916EFF85FFDBFF10253C393AC8C4" author="Harold E." box="[189,611,888,914]" page="139" pageId="21" pageNumber="22" year="1869">
<taxonomicName id="4C2CCCFCFF85FFDBFF10253C393AC8C4" ID-CoL="77LBZ" authority="Harold, 1869: 139" authorityName="Harold" authorityPageNumber="139" authorityYear="1869" box="[189,611,888,914]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="B9586B6DFF85FFDBFF10253C3ADCC8C4" box="[189,389,888,914]" italics="true" pageId="21" pageNumber="22">Pinotus deyrollei</emphasis>
<bibRefCitation id="EFBDCA8EFF85FFDBFE21253D393AC8C4" author="Harold E." box="[396,611,888,914]" pageId="21" pageNumber="22" pagination="124 - 144" refId="ref29859" refString="Harold E. 1869. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 124 - 144." type="journal article" year="1869">Harold, 1869: 139</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B93B77FFF85FFDBFF1025FE38BAC882" blockId="21.[189,1399,954,1017]" box="[189,995,954,981]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB3C9051FF85FFDBFF1025FE38BAC882" box="[189,995,954,981]" pageId="21" pageNumber="22">
<taxonomicName id="4C2CCCFCFF85FFDBFF1025FE3ADCC882" ID-CoL="77LBZ" authorityName="Harold" authorityYear="1869" box="[189,389,954,980]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="B9586B6DFF85FFDBFF1025FE3ADCC882" box="[189,389,954,980]" italics="true" pageId="21" pageNumber="22">Pinotus deyrollei</emphasis>
</taxonomicName>
<treatmentCitation id="0A8D916EFF85FFDBFE0F25FF39F0C882" author="Luederwaldt H." box="[418,681,954,980]" page="99" pageId="21" pageNumber="22" year="1929">
<bibRefCitation id="EFBDCA8EFF85FFDBFE0F25FF39F0C882" author="Luederwaldt H." box="[418,681,954,980]" pageId="21" pageNumber="22" pagination="1 - 173" refId="ref30128" refString="Luederwaldt H. 1929. As especies Brasileiras do genero Pinotus. Revista do Museu Paulista 16: 1 - 173." type="journal article" year="1929">Luederwaldt 1929: 99</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0A8D916EFF85FFDBFD7425FF38B9C882" author="Blackwelder R. E." box="[729,992,954,980]" page="69" pageId="21" pageNumber="22" year="1944">
<bibRefCitation id="EFBDCA8EFF85FFDBFD7425FF38B9C882" author="Blackwelder R. E." box="[729,992,954,980]" pageId="21" pageNumber="22" pagination="189 - 341" refId="ref29526" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 69</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B93B77FFF85FFDBFF10259B3E2EC8AF" blockId="21.[189,1399,954,1017]" box="[189,1399,990,1017]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB3C9051FF85FFDBFF10259B3E2EC8AF" box="[189,1399,990,1017]" pageId="21" pageNumber="22">
<taxonomicName id="4C2CCCFCFF85FFDBFF10259B3A99C8AE" ID-CoL="35NBJ" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[189,448,990,1016]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="B9586B6DFF85FFDBFF10259B3A99C8AE" box="[189,448,990,1016]" italics="true" pageId="21" pageNumber="22">Dichotomius deyrollei</emphasis>
</taxonomicName>
<treatmentCitation id="0A8D916EFF85FFDBFE76259A39D1C8AE" author="Roze J. A." box="[475,648,990,1017]" page="44" pageId="21" pageNumber="22" year="1955">
<bibRefCitation id="EFBDCA8EFF85FFDBFE76259A39D1C8AE" author="Roze J. A." box="[475,648,990,1017]" pageId="21" pageNumber="22" pagination="39 - 63" refId="ref30793" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0A8D916EFF85FFDBFD18259B3894C8AE" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[693,973,990,1016]" page="138" pageId="21" pageNumber="22" year="2001">
<bibRefCitation id="EFBDCA8EFF85FFDBFD18259B3894C8AE" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[693,973,990,1016]" pageId="21" pageNumber="22" pagination="131 - 144" refId="ref30377" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="B9586B6DFF85FFDBFCB9259A3817C8AE" box="[788,846,990,1016]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2001: 138
</bibRefCitation>
</treatmentCitation>
. — Sarmiento &amp; Amat 2014: 9395.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FF85FFDBFF1022653F66CE48" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8B93B77FFF85FFDBFF1022653A6FCF6C" blockId="21.[189,310,1056,1082]" box="[189,310,1056,1082]" pageId="21" pageNumber="22">
<heading id="D0DB0013FF85FFDBFF1022653A6FCF6C" bold="true" box="[189,310,1056,1082]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B9586B6DFF85FFDBFF1022653A6FCF6C" bold="true" box="[189,310,1056,1082]" pageId="21" pageNumber="22">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFF85FFDBFF1022153F66CE48" blockId="21.[189,1401,1104,1311]" pageId="21" pageNumber="22">
<taxonomicName id="4C2CCCFCFF85FFDBFF1022153A9DCF3C" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[189,452,1104,1130]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="B9586B6DFF85FFDBFF1022153A9DCF3C" box="[189,452,1104,1130]" italics="true" pageId="21" pageNumber="22">Dichotomius deyrollei</emphasis>
</taxonomicName>
is separated from the other species in the group by the following combination of characters: clypeus with central emargination displaying two rounded, dorsally angled clypeal teeth. Cephalic carina, higher than wide, antero-posteriorly flattened. Median tubercle approximately twice as high as lateral tubercles and gently inclined anteriorly, apically expanded and emarginate. Anterior portion of pronotum displaying sharp declivity, antero-medial area deeper and convex than lateral and where the longer, median tubercle can fit if head capsules is rotated dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FF85FFD8FF1023033E03C96E" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8B93B77FFF85FFDBFF1023033AF3CE36" blockId="21.[189,426,1350,1376]" box="[189,426,1350,1376]" pageId="21" pageNumber="22">
<heading id="D0DB0013FF85FFDBFF1023033AF3CE36" bold="true" box="[189,426,1350,1376]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B9586B6DFF85FFDBFF1023033AF3CE36" bold="true" box="[189,426,1350,1376]" pageId="21" pageNumber="22">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFF85FFDBFF4723333A35CD76" blockId="21.[189,1399,1398,1568]" pageId="21" pageNumber="22">
<materialsCitation id="3B44BD22FF85FFDBFF4723333A35CD76" ID-GBIF-Occurrence="3037418373" collectionCode="MNHN" collectorName="F. Z. Vaz-de-Mello" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B9586B6DFF85FFDBFF4723333A3FCEC6" bold="true" box="[234,358,1398,1424]" pageId="21" pageNumber="22">
<typeStatus id="549709DDFF85FFDBFF4723333A3FCEC6" box="[234,358,1398,1424]" pageId="21" pageNumber="22" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated) COUNTRY UNKNOWN •
<specimenCount id="9D2A7CF6FF85FFDBFDBF23DD3970CEE2" box="[530,553,1432,1460]" pageId="21" pageNumber="22" type="male"></specimenCount>
; “Brasilia. P.
<taxonomicName id="4C2CCCFCFF85FFDBFD7023DF38F2CEE2" authority="Harold" authorityName="Harold" box="[733,939,1434,1460]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="B9586B6DFF85FFDBFD7023DF3812CEE2" box="[733,843,1434,1460]" italics="true" pageId="21" pageNumber="22">Deyrollei</emphasis>
Harold
</taxonomicName>
” [white with red margins, hw]/ “Ex- Musaeo E. Harold” [white with black margins, p]/ “Museum Paris ex Coll. R. Oberthur” [green, p]/ “LECTOTYPE ♂
<taxonomicName id="4C2CCCFCFF85FFDBFE6C23A739BCCEAA" authority="Harold" authorityName="Harold" authorityYear="1869" box="[449,741,1506,1532]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="B9586B6DFF85FFDBFE6C23A739D2CEAA" box="[449,651,1506,1532]" italics="true" pageId="21" pageNumber="22">Pinotus deyrollei</emphasis>
Harold
</taxonomicName>
des.
<collectorName id="26D9D2A9FF85FFDBFC8B23A73F58CEAA" box="[806,1025,1506,1532]" pageId="21" pageNumber="22">F.Z. Vaz-de-Mello</collectorName>
, 2014” [red with black margin, phw];
<collectionCode id="ED3D2FBAFF85FFDBFEA720433A31CD76" box="[266,360,1542,1568]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="22">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B93B77FFF85FFDBFF47200D3ACECD34" blockId="21.[189,1399,1608,1743]" box="[234,407,1608,1634]" pageId="21" pageNumber="22">
<emphasis id="B9586B6DFF85FFDBFF47200D3ACECD34" bold="true" box="[234,407,1608,1634]" pageId="21" pageNumber="22">
<typeStatus id="549709DDFF85FFDBFF47200D3ACECD34" box="[234,407,1608,1634]" pageId="21" pageNumber="22" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B93B77FFF85FFDBFF102029398ACD99" blockId="21.[189,1399,1608,1743]" pageId="21" pageNumber="22">
COUNTRY UNKNOWN •
<materialsCitation id="3B44BD22FF85FFDBFDAC20293989CD99" ID-GBIF-Occurrence="3037418404" collectionCode="MNHN" collectorName="F. Z. Vaz-de-Mello" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="9D2A7CF6FF85FFDBFDAC20293977CDD0" box="[513,558,1644,1670]" pageId="21" pageNumber="22" type="female">1 ♀</specimenCount>
; “Ex-Musaeo E. Harold” [white with black margins, p]/ “Museum Paris ex Coll. R. Oberthur” [green, p]/ “LECTOTYPE ♀
<taxonomicName id="4C2CCCFCFF85FFDBFC8320D43F0ACDFC" authority="Harold" authorityName="Harold" authorityYear="1869" box="[814,1107,1680,1706]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="B9586B6DFF85FFDBFC8320D438A1CDFC" box="[814,1016,1680,1706]" italics="true" pageId="21" pageNumber="22">Pinotus deyrollei</emphasis>
Harold
</taxonomicName>
des.
<collectorName id="26D9D2A9FF85FFDBFB3B20D43E2BCDFC" box="[1174,1394,1680,1707]" pageId="21" pageNumber="22">F.Z. Vaz-de-Mello</collectorName>
, 2014” [red with black margin, phw];
<collectionCode id="ED3D2FBAFF85FFDBFDDF20F03989CD99" box="[626,720,1717,1743]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="22">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B93B77FFF85FFDBFF4720B33AB8CC46" blockId="21.[188,1399,1782,2025]" box="[234,481,1782,1808]" pageId="21" pageNumber="22">
<emphasis id="B9586B6DFF85FFDBFF4720B33AB8CC46" bold="true" box="[234,481,1782,1808]" pageId="21" pageNumber="22">Additional material</emphasis>
</paragraph>
<paragraph id="8B93B77FFF85FFDBFF10215F39E1CCBF" blockId="21.[188,1399,1782,2025]" pageId="21" pageNumber="22">
<materialsCitation id="3B44BD22FF85FFDBFF10215F3AD4CC0E" ID-GBIF-Occurrence="3037418458" collectingDate="1996-03" collectionCode="ECC" collectorName="A. Lopera" country="Colombia" county="Yopal" elevation="600" location="Bosque seco Tropical" municipality="Guarataro" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Casanare">
<collectingCountry id="F33BF7EFFF85FFDBFF10215F3A01CC62" box="[189,344,1818,1844]" name="Colombia" pageId="21" pageNumber="22">COLOMBIA</collectingCountry>
<emphasis id="B9586B6DFF85FFDBFEDE215F3AB0CC62" bold="true" box="[371,489,1818,1844]" pageId="21" pageNumber="22">
<collectingRegion id="49E8799DFF85FFDBFEDE215F3AB0CC62" box="[371,489,1818,1844]" country="Colombia" name="Casanare" pageId="21" pageNumber="22">Casanare</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FF85FFDBFDAC215F390CCC63" box="[513,597,1818,1845]" pageId="21" pageNumber="22" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="62F2CFF3FF85FFDBFDCD215F39F1CC62" box="[608,680,1818,1844]" pageId="21" pageNumber="22">Yopal</collectingCounty>
,
<collectingMunicipality id="6BF72D05FF85FFDBFD1F215F3872CC62" box="[690,811,1818,1844]" pageId="21" pageNumber="22">Guarataro</collectingMunicipality>
;
<elevation id="0001504CFF85FFDBFC9B215F38E6CC62" box="[822,959,1818,1845]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="m" value="600.0">
<quantity id="4CD41A9AFF85FFDBFC9B215F38D9CC63" box="[822,896,1818,1845]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
;
<location id="8EF3E1A4FF85FFDBFC66215E3F93CC62" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF85FFDBFC66215E3F93CC62" box="[971,1226,1818,1845]" country="Colombia" county="Yopal" municipality="Guarataro" name="Bosque seco Tropical" pageId="21" pageNumber="22" stateProvince="Casanare">Bosque seco Tropical</location>
;
<date id="FF9291BFFF85FFDBFB78215E3E09CC62" box="[1237,1360,1818,1845]" pageId="21" pageNumber="22" value="1996-03">
<collectingDate id="EFD66857FF85FFDBFB78215E3E09CC62" box="[1237,1360,1818,1845]" pageId="21" pageNumber="22" value="1996-03">Mar. 1996</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF85FFDBFAF7215F3A56CC0F" pageId="21" pageNumber="22">A. Lopera</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF85FFDBFEFD217B3AD4CC0E" box="[336,397,1854,1880]" pageId="21" pageNumber="22">ECC</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B44BD22FF85FFDBFE00217B3E56CC0E" ID-GBIF-Occurrence="3037418619" box="[429,1295,1854,1881]" collectingDate="1993-03" collectionCode="ECC" collectorName="Ivan Jimenez" country="Colombia" location="Varzea" municipality="Tinigua" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Meta">
<emphasis id="B9586B6DFF85FFDBFE00217B3AB7CC0E" bold="true" box="[429,494,1854,1880]" pageId="21" pageNumber="22">
<collectingRegion id="49E8799DFF85FFDBFE00217B3AB7CC0E" box="[429,494,1854,1880]" country="Colombia" name="Meta" pageId="21" pageNumber="22">Meta</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FF85FFDBFDAA217B3902CC0F" box="[519,603,1854,1881]" pageId="21" pageNumber="22" type="generic">1 spec.</specimenCount>
; PNN
<collectingMunicipality id="6BF72D05FF85FFDBFD05217B385CCC0E" box="[680,773,1854,1880]" pageId="21" pageNumber="22">Tinigua</collectingMunicipality>
,
<location id="8EF3E1A4FF85FFDBFCBD217A383DCC0F" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF85FFDBFCBD217A383DCC0F" box="[784,868,1855,1881]" country="Colombia" municipality="Tinigua" name="Varzea" pageId="21" pageNumber="22" stateProvince="Meta">Varzea</location>
;
<date id="FF9291BFFF85FFDBFCC2217A38B3CC0E" box="[879,1002,1854,1881]" pageId="21" pageNumber="22" value="1993-03">
<collectingDate id="EFD66857FF85FFDBFCC2217A38B3CC0E" box="[879,1002,1854,1881]" pageId="21" pageNumber="22" value="1993-03">Mar. 1993</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF85FFDBFC5B217A3FC8CC0E" box="[1014,1169,1854,1881]" pageId="21" pageNumber="22">Ivan Jiménez</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF85FFDBFB7F217B3E56CC0E" box="[1234,1295,1854,1880]" pageId="21" pageNumber="22">ECC</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B44BD22FF85FFDBFA83217B3AD3CCF6" ID-GBIF-Occurrence="3037418366" collectingDate="1974-05-10" collectingMethod="dung carrion traps" collectionCode="CEMT" collectorName="S. Peck" country="Colombia" elevation="1000" location="Norte de Santander" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Norte de Santander">
<emphasis id="B9586B6DFF85FFDBFA83217B3A3FCC2A" bold="true" pageId="21" pageNumber="22">
<collectingRegion id="49E8799DFF85FFDBFA83217B3A3FCC2A" country="Colombia" name="Norte de Santander" pageId="21" pageNumber="22">Norte de Santander</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FF85FFDBFE2921273AE8CC2A" box="[388,433,1888,1916]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
;
<locationDeviation id="9A21AE5DFF85FFDBFE6F21273993CC2A" box="[450,714,1890,1917]" pageId="21" pageNumber="22">3 Km N of Chinacota</locationDeviation>
;
<elevation id="0001504CFF85FFDBFD7A2127382DCC2A" box="[727,884,1890,1917]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="m" value="1000.0">
<quantity id="4CD41A9AFF85FFDBFD7A2127386BCC2B" box="[727,818,1890,1917]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
;
<collectingMethod id="526DCF68FF85FFDBFC2E21273F06CC2B" box="[899,1119,1890,1917]" pageId="21" pageNumber="22">dung carrion traps</collectingMethod>
, T10;
<date id="FF9291BFFF85FFDBFB0021273E08CC2A" box="[1197,1361,1890,1917]" pageId="21" pageNumber="22" value="1974-05-10">
<collectingDate id="EFD66857FF85FFDBFB0021273E08CC2A" box="[1197,1361,1890,1917]" pageId="21" pageNumber="22" value="1974-05-10">10 May 1974</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF85FFDBFAF221273BAFCCF6" pageId="21" pageNumber="22">S. Peck</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF85FFDBFE9421C33AD3CCF6" box="[313,394,1926,1952]" collectionName="CEMT" pageId="21" pageNumber="22">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FF85FFDBFE0921C33F98CCF6" ID-GBIF-Occurrence="3037418585" box="[420,1217,1926,1953]" collectingDate="2007-02" collectionCode="ECC" collectorName="S. Ardila" country="Colombia" elevation="1200" location="Villa Marina" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Norte de Santander">
<specimenCount id="9D2A7CF6FF85FFDBFE0921C33AA0CCF7" box="[420,505,1926,1953]" pageId="21" pageNumber="22" type="generic">1 spec.</specimenCount>
;
<location id="8EF3E1A4FF85FFDBFDA821C339C6CCF6" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF85FFDBFDA821C339C6CCF6" box="[517,671,1926,1952]" country="Colombia" name="Villa Marina" pageId="21" pageNumber="22" stateProvince="Norte de Santander">Villa Marina</location>
;
<elevation id="0001504CFF85FFDBFD0621C3381DCCF6" box="[683,836,1926,1953]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="22" unit="m" value="1200.0">
<quantity id="4CD41A9AFF85FFDBFD0621C3385DCCF7" box="[683,772,1926,1953]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="22" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
;
<date id="FF9291BFFF85FFDBFCFF21C33892CCF6" box="[850,971,1926,1952]" pageId="21" pageNumber="22" value="2007-02">
<collectingDate id="EFD66857FF85FFDBFCFF21C33892CCF6" box="[850,971,1926,1952]" pageId="21" pageNumber="22" value="2007-02">Feb. 2007</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF85FFDBFC7521C33F18CCF6" box="[984,1089,1926,1952]" pageId="21" pageNumber="22">S. Ardila</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF85FFDBFB2921C33F98CCF6" box="[1156,1217,1926,1952]" pageId="21" pageNumber="22">ECC</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B44BD22FF85FFDBFB4E21C339EACCBF" ID-GBIF-Occurrence="3037418393" collectingDate="2016-04-11" collectionCode="IAvH" collectorName="C. Giraldo &amp; Leon, J" country="Colombia" elevation="1730" latitude="6.5441666" location="Fca. Buenos Aires" longLatPrecision="15" longitude="-73.65192" municipality="Charala Vda. El Salitre" pageId="21" pageNumber="22" specimenCount="10" stateProvince="Santander">
<emphasis id="B9586B6DFF85FFDBFB4E21C33E3CCCF6" bold="true" box="[1251,1381,1926,1952]" pageId="21" pageNumber="22">
<collectingRegion id="49E8799DFF85FFDBFB4E21C33E3CCCF6" box="[1251,1381,1926,1952]" country="Colombia" name="Santander" pageId="21" pageNumber="22">Santander</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FF85FFDBFF1021EF3A7CCC93" box="[189,293,1962,1989]" pageId="21" pageNumber="22">10 specs</specimenCount>
;
<collectingMunicipality id="6BF72D05FF85FFDBFE9F21EF391ECC92" box="[306,583,1962,1988]" pageId="21" pageNumber="22">Charalá Vda. El Salitre</collectingMunicipality>
,
<location id="8EF3E1A4FF85FFDBFDFF21EE3874CC92" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF85FFDBFDFF21EE3874CC92" box="[594,813,1962,1989]" country="Colombia" latitude="6.5441666" longLatPrecision="15" longitude="-73.65192" municipality="Charala Vda. El Salitre" name="Fca. Buenos Aires" pageId="21" pageNumber="22" stateProvince="Santander">Fca. Buenos Aires</location>
;
<elevation id="0001504CFF85FFDBFC9721EF388DCC92" box="[826,980,1962,1989]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="21" pageNumber="22" unit="m" value="1730.0">
<quantity id="4CD41A9AFF85FFDBFC9721EF38CDCC93" box="[826,916,1962,1989]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="21" pageNumber="22" unit="m" value="1730.0">1730 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FF85FFDBFC4F21EC3F3ECC93" box="[994,1127,1961,1989]" degrees="6" direction="north" minutes="32" orientation="latitude" pageId="21" pageNumber="22" precision="15" seconds="39" value="6.5441666">6º3239″ N</geoCoordinate>
,
<geoCoordinate id="EE18D1B8FF85FFDBFBDE21EC3E4DCC93" box="[1139,1300,1961,1989]" degrees="73" direction="west" minutes="39" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="6.9" value="-73.65192">73º396.9″ W</geoCoordinate>
;
<date id="FF9291BFFF85FFDBFA8C21EF3BA2CCBE" pageId="21" pageNumber="22" value="2016-04-11">
<collectingDate id="EFD66857FF85FFDBFA8C21EF3BA2CCBE" pageId="21" pageNumber="22" value="2016-04-11">11 Apr. 2016</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF85FFDBFEAA218B3A25CCBE" box="[263,380,1998,2024]" pageId="21" pageNumber="22">C.Giraldo</collectorName>
and A.
<collectorName id="26D9D2A9FF85FFDBFE7A218B3974CCBF" box="[471,557,1998,2025]" pageId="21" pageNumber="22">León, J</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF85FFDBFDC2218A39EACCBF" box="[623,691,1999,2025]" pageId="21" pageNumber="22">IAvH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF10274C3F81CA75" blockId="22.[189,1240,263,292]" box="[189,1240,263,292]" pageId="22" pageNumber="23">
<materialsCitation id="3B44BD22FF86FFD8FF10274C38D6CA75" ID-GBIF-Occurrence="3037418594" box="[189,911,263,292]" collectingDate="1950-04-30" collectionCode="CEMT" country="Trinidad and Tobago" location="Arima Valley" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Arima">
<collectingCountry id="F33BF7EFFF86FFD8FF10274C3A12CA75" box="[189,331,265,291]" name="Trinidad and Tobago" pageId="22" pageNumber="23">TRINIDAD</collectingCountry>
<specimenCount id="9D2A7CF6FF86FFD8FEC9274C3AC8CA75" box="[356,401,263,291]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
; B.W.I
<location id="8EF3E1A4FF86FFD8FE47274C39D2CA75" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF86FFD8FE47274C39D2CA75" box="[490,651,265,291]" country="Trinidad and Tobago" name="Arima Valley" pageId="22" pageNumber="23" stateProvince="Arima">Arima Valley</location>
;
<date id="FF9291BFFF86FFD8FD3B274C386BCA72" box="[662,818,265,292]" pageId="22" pageNumber="23" value="1950-04-30">
<collectingDate id="EFD66857FF86FFD8FD3B274C386BCA72" box="[662,818,265,292]" pageId="22" pageNumber="23" value="1950-04-30">30 Apr. 1950</collectingDate>
</date>
;
<collectionCode id="ED3D2FBAFF86FFD8FC93274C38D6CA75" box="[830,911,265,291]" collectionName="CEMT" pageId="22" pageNumber="23">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FF86FFD8FC05274C3F8DCA75" ID-GBIF-Occurrence="3037418326" box="[936,1236,265,292]" collectingDate="1953-05-02" collectionCode="CEMT" country="Trinidad and Tobago" pageId="22" pageNumber="23" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D2A7CF6FF86FFD8FC05274C388CCA75" box="[936,981,265,291]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
;
<date id="FF9291BFFF86FFD8FC4E274C3F2DCA72" box="[995,1140,265,292]" pageId="22" pageNumber="23" value="1953-05-02">
<collectingDate id="EFD66857FF86FFD8FC4E274C3F2DCA72" box="[995,1140,265,292]" pageId="22" pageNumber="23" value="1953-05-02">2 May 1953</collectingDate>
</date>
;
<collectionCode id="ED3D2FBAFF86FFD8FB2C274C3F8DCA75" box="[1153,1236,265,291]" collectionName="CEMT" pageId="22" pageNumber="23">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF10270E3E03C96E" blockId="22.[189,1399,329,568]" pageId="22" pageNumber="23">
<materialsCitation id="3B44BD22FF86FFD8FF10270E3909CADE" ID-GBIF-Occurrence="3037418478" collectingDate="2015-03-13" collectionCode="CEMT" collectorName="Racconcia" country="Venezuela" elevation="1380" latitude="8.238334" location="Mucutuy" longLatPrecision="21" longitude="-71.28667" municipality="Arzobispo Chacon" pageId="22" pageNumber="23" specimenCount="2" specimenCount-female="2" stateProvince="Merida">
<collectingCountry id="F33BF7EFFF86FFD8FF10270E3A33CA33" box="[189,362,331,357]" name="Venezuela" pageId="22" pageNumber="23">VENEZUELA</collectingCountry>
<emphasis id="B9586B6DFF86FFD8FE29270C3AB8CA32" bold="true" box="[388,481,329,356]" pageId="22" pageNumber="23">
<collectingRegion id="49E8799DFF86FFD8FE29270C3AB8CA32" box="[388,481,329,356]" country="Venezuela" name="Merida" pageId="22" pageNumber="23">Mérida</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FF86FFD8FE54270E3962CA33" box="[505,571,331,357]" pageId="22" pageNumber="23" type="female">2 ♀♀</specimenCount>
;
<collectingMunicipality id="6BF72D05FF86FFD8FDEA270F387CCA32" box="[583,805,330,356]" pageId="22" pageNumber="23">Arzobispo Chacón</collectingMunicipality>
,
<location id="8EF3E1A4FF86FFD8FC82270E38C4CA33" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF86FFD8FC82270E38C4CA33" box="[815,925,331,357]" country="Venezuela" latitude="8.238334" longLatPrecision="21" longitude="-71.28667" municipality="Arzobispo Chacon" name="Mucutuy" pageId="22" pageNumber="23" stateProvince="Merida">Mucutuy</location>
;
<elevation id="0001504CFF86FFD8FC05270E3F66CA32" box="[936,1087,330,357]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="22" pageNumber="23" unit="m" value="1380.0">
<quantity id="4CD41A9AFF86FFD8FC05270E3F59CA33" box="[936,1024,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="22" pageNumber="23" unit="m" value="1380.0">1380 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FF86FFD8FBE1270C3F96CA33" box="[1100,1231,329,357]" degrees="8" direction="north" minutes="14" orientation="latitude" pageId="22" pageNumber="23" precision="15" seconds="18" value="8.238334">8º1418″ N</geoCoordinate>
,
<geoCoordinate id="EE18D1B8FF86FFD8FB77270C3E2BCA33" box="[1242,1394,329,357]" degrees="71" direction="west" minutes="17" orientation="longitude" pageId="22" pageNumber="23" precision="15" seconds="12" value="-71.28667">71º1712″ W</geoCoordinate>
;
<date id="FF9291BFFF86FFD8FF10272B3A3BCADE" box="[189,354,365,392]" pageId="22" pageNumber="23" value="2015-03-13">
<collectingDate id="EFD66857FF86FFD8FF10272B3A3BCADE" box="[189,354,365,392]" pageId="22" pageNumber="23" value="2015-03-13">13 Mar. 2015</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF86FFD8FEDC272B3AB6CADE" box="[369,495,366,392]" pageId="22" pageNumber="23">Racconcia</collectorName>
;
<collectionCode id="ED3D2FBAFF86FFD8FE50272B3909CADE" box="[509,592,366,392]" pageId="22" pageNumber="23">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B44BD22FF86FFD8FDDB27283BA0CAFD" ID-GBIF-Occurrence="3037418417" collectingDate="1959-07-19" collectionCode="MLP" collectorName="Bordon" country="Venezuela" location="Curupao" municipality="Guarenas" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Miranda">
<emphasis id="B9586B6DFF86FFD8FDDB272839BCCAD1" bold="true" box="[630,741,365,391]" pageId="22" pageNumber="23">
<collectingRegion id="49E8799DFF86FFD8FDDB272839BCCAD1" box="[630,741,365,391]" country="Venezuela" name="Miranda" pageId="22" pageNumber="23">Miranda</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FF86FFD8FCAE272B386BCADE" box="[771,818,364,392]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BF72D05FF86FFD8FCEE272B38ECCADE" box="[835,949,366,392]" pageId="22" pageNumber="23">Guarenas</collectingMunicipality>
,
<location id="8EF3E1A4FF86FFD8FC6E272B3F75CADE" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF86FFD8FC6E272B3F75CADE" box="[963,1068,366,392]" country="Venezuela" municipality="Guarenas" name="Curupao" pageId="22" pageNumber="23" stateProvince="Miranda">Curupao</location>
;
<date id="FF9291BFFF86FFD8FB97272B3F8ACADE" box="[1082,1235,365,392]" pageId="22" pageNumber="23" value="1959-07-19">
<collectingDate id="EFD66857FF86FFD8FB97272B3F8ACADE" box="[1082,1235,365,392]" pageId="22" pageNumber="23" value="1959-07-19">19 Jul. 1959</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF86FFD8FB4C272B3E60CADE" box="[1249,1337,366,392]" pageId="22" pageNumber="23">Bordon</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF86FFD8FF1027D43BA0CAFD" box="[189,249,401,427]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="22" pageNumber="23">MLP</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FF86FFD8FEB927D43A57CA98" ID-GBIF-Occurrence="3037418556" collectingDate="1974-05-19" collectingMethod="dung traps" collectionCode="CEMT" collectorName="S. Peck" country="Venezuela" elevation="700" location="Tachira" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Tachira">
<specimenCount id="9D2A7CF6FF86FFD8FEB927D43A1ACAFD" box="[276,323,399,427]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49E8799DFF86FFD8FEFF27D43AF6CAFD" box="[338,431,401,427]" country="Venezuela" name="Tachira" pageId="22" pageNumber="23">Táchira</collectingRegion>
,
<locationDeviation id="9A21AE5DFF86FFD8FE1127D439A0CAFD" box="[444,761,401,427]" pageId="22" pageNumber="23">4 Km SE of San Cristobal</locationDeviation>
;
<elevation id="0001504CFF86FFD8FCAB27D438CDCAFD" box="[774,916,401,427]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="23" unit="m" value="700.0">
<quantity id="4CD41A9AFF86FFD8FCAB27D4380BCAFD" box="[774,850,401,427]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="23" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
;
<collectingMethod id="526DCF68FF86FFD8FC0E27D43F7ACAFD" box="[931,1059,401,427]" pageId="22" pageNumber="23">dung traps</collectingMethod>
;
<date id="FF9291BFFF86FFD8FB9D27D43F8ACAFD" box="[1072,1235,401,427]" pageId="22" pageNumber="23" value="1974-05-19">
<collectingDate id="EFD66857FF86FFD8FB9D27D43F8ACAFD" box="[1072,1235,401,427]" pageId="22" pageNumber="23" value="1974-05-19">19 May 1974</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF86FFD8FB4C27D43E63CAFD" box="[1249,1338,401,427]" pageId="22" pageNumber="23">S. Peck</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF86FFD8FF1027F13A57CA98" box="[189,270,436,462]" collectionName="CEMT" pageId="22" pageNumber="23">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FF86FFD8FE8427F138B0CA98" ID-GBIF-Occurrence="3037418327" box="[297,1001,434,463]" collectingDate="1993-07" collectionCode="CEMT" collectorName="J. Blanco" country="Venezuela" location="Natural Paramillo" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Miranda">
<specimenCount id="9D2A7CF6FF86FFD8FE8427F13A0ECA98" box="[297,343,434,462]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
; P.
<location id="8EF3E1A4FF86FFD8FE2927F13902CA98" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF86FFD8FE2927F13902CA98" box="[388,603,436,462]" country="Venezuela" name="Natural Paramillo" pageId="22" pageNumber="23" stateProvince="Miranda">Natural Paramillo</location>
;
<date id="FF9291BFFF86FFD8FDCA27F1398ECA98" box="[615,727,436,462]" pageId="22" pageNumber="23" value="1993-07">
<collectingDate id="EFD66857FF86FFD8FDCA27F1398ECA98" box="[615,727,436,462]" pageId="22" pageNumber="23" value="1993-07">Jul. 1993</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF86FFD8FD4827F0380ACA98" box="[741,851,436,463]" pageId="22" pageNumber="23">J. Blanco</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF86FFD8FC3527F138B0CA98" box="[920,1001,436,462]" collectionName="CEMT" pageId="22" pageNumber="23">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FF86FFD8FBA927F13AE4CAA7" ID-GBIF-Occurrence="3037418411" collectingDate="1984-03-25" collectionCode="CEMT" collectorName="B. Joffre" country="Venezuela" location="Rio Negro" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Miranda">
<specimenCount id="9D2A7CF6FF86FFD8FBA927F13F6BCA98" box="[1028,1074,434,462]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
;
<location id="8EF3E1A4FF86FFD8FBEF27F13F9BCA99" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF86FFD8FBEF27F13F9BCA99" box="[1090,1218,436,463]" country="Venezuela" name="Rio Negro" pageId="22" pageNumber="23" stateProvince="Miranda">Rio Negro</location>
;
<date id="FF9291BFFF86FFD8FB6227F13E2BCA98" box="[1231,1394,436,463]" pageId="22" pageNumber="23" value="1984-03-25">
<collectingDate id="EFD66857FF86FFD8FB6227F13E2BCA98" box="[1231,1394,436,463]" pageId="22" pageNumber="23" value="1984-03-25">25 Mar. 1984</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF86FFD8FF10279D3A7CCAA7" box="[189,293,471,498]" pageId="22" pageNumber="23">B. Joffre</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF86FFD8FEC127923AE4CAA7" box="[364,445,471,497]" collectionName="CEMT" pageId="22" pageNumber="23">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FF86FFD8FE772792392CC943" ID-GBIF-Occurrence="3037418521" collectingDate="1989-07-01" collectionCode="BMNH" collectorName="D. Havranek" country="Venezuela" elevation="1100" location="Chorro del Indio" municipality="San Cristobal" pageId="22" pageNumber="23" specimenCount="8" specimenCount-male="1" stateProvince="Miranda">
<specimenCount id="9D2A7CF6FF86FFD8FE7727923950CAA7" box="[474,521,469,497]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
;
<locationDeviation id="9A21AE5DFF86FFD8FDB727923F09CAA7" box="[538,1104,471,498]" pageId="22" pageNumber="23">
<quantity id="4CD41A9AFF86FFD8FDB727923934CAA7" box="[538,621,471,498]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="23" unit="km" value="2.5">2.5 km</quantity>
from San Cristobal to Chorro del Indio
</locationDeviation>
;
<elevation id="0001504CFF86FFD8FBF327923FA3CAA7" box="[1118,1274,471,498]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="m" value="1100.0">
<quantity id="4CD41A9AFF86FFD8FBF327923FE1CAA4" box="[1118,1208,471,498]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
;
<specimenCount id="9D2A7CF6FF86FFD8FAA7279D3E61CAA4" box="[1290,1336,472,498]" pageId="22" pageNumber="23" type="generic">7 p.</specimenCount>
m.;
<date id="FF9291BFFF86FFD8FAC527923A77C943" pageId="22" pageNumber="23" value="1989-07-01">
<collectingDate id="EFD66857FF86FFD8FAC527923A77C943" pageId="22" pageNumber="23" value="1989-07-01">1 Jul. 1989</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF86FFD8FE9127BE3A8AC942" box="[316,467,506,533]" pageId="22" pageNumber="23">D. Havranek</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF86FFD8FDB427BE392CC943" box="[537,629,507,533]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="22" pageNumber="23">BMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B44BD22FF86FFD8FD3627BE3E0FC96E" ID-GBIF-Occurrence="3037418306" collectingDate="2009-07-20" collectionCode="CEMT" collectorName="M. Asmussen &amp; P. Colmenares &amp; H. Martinez" country="Venezuela" elevation="710" latitude="10.0" location="Aroa" longLatPrecision="21" longitude="-68.0" pageId="22" pageNumber="23" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Yaracuy">
<emphasis id="B9586B6DFF86FFD8FD3627BE385AC943" bold="true" box="[667,771,507,533]" pageId="22" pageNumber="23">
<collectingRegion id="49E8799DFF86FFD8FD3627BE385AC943" box="[667,771,507,533]" country="Venezuela" name="Yaracuy" pageId="22" pageNumber="23">Yaracuy</collectingRegion>
</emphasis>
<specimenCount id="9D2A7CF6FF86FFD8FCB327BE383AC942" box="[798,867,504,533]" pageId="22" pageNumber="23" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D2A7CF6FF86FFD8FCDE27BE38F8C943" box="[883,929,507,533]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49E8799DFF86FFD8FC1F27BF3F53C942" box="[946,1034,506,532]" country="Venezuela" name="Bolivar" pageId="22" pageNumber="23">Bolivar</collectingRegion>
<location id="8EF3E1A4FF86FFD8FBBC27BE3F16C943" LSID="urn:lsid:plazi:treatment:03850669FF85FFD6FE6425543A2AC9B3:8EF3E1A4FF86FFD8FBBC27BE3F16C943" box="[1041,1103,507,533]" country="Venezuela" latitude="10.0" longLatPrecision="21" longitude="-68.0" name="Aroa" pageId="22" pageNumber="23" stateProvince="Yaracuy">Aroa</location>
;
<elevation id="0001504CFF86FFD8FBF027BE3FB2C942" box="[1117,1259,506,533]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="22" pageNumber="23" unit="m" value="710.0">
<quantity id="4CD41A9AFF86FFD8FBF027BE3FF0C943" box="[1117,1193,507,533]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="22" pageNumber="23" unit="m" value="710.0">710 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FF86FFD8FB5727BC3E2BC943" box="[1274,1394,505,533]" degrees="10" direction="north" minutes="0" orientation="latitude" pageId="22" pageNumber="23" precision="15" seconds="0" value="10.0">10º00″ N</geoCoordinate>
,
<geoCoordinate id="EE18D1B8FF86FFD8FF1024593A61C96E" box="[189,312,540,568]" degrees="68" direction="west" minutes="0" orientation="longitude" pageId="22" pageNumber="23" precision="15" seconds="0" value="-68.0">68º00″ W</geoCoordinate>
; heces humanas;
<date id="FF9291BFFF86FFD8FDA8245B39C1C96E" box="[517,664,542,568]" pageId="22" pageNumber="23" value="2009-07-20">
<collectingDate id="EFD66857FF86FFD8FDA8245B39C1C96E" box="[517,664,542,568]" pageId="22" pageNumber="23" value="2009-07-20">20 Jul. 2009</collectingDate>
</date>
;
<collectorName id="26D9D2A9FF86FFD8FD09245B381EC96E" box="[676,839,542,568]" pageId="22" pageNumber="23">M. Asmüssen</collectorName>
,
<collectorName id="26D9D2A9FF86FFD8FCFE245B38A3C96E" box="[851,1018,542,568]" pageId="22" pageNumber="23">P. Colmenares</collectorName>
and
<collectorName id="26D9D2A9FF86FFD8FB9E245B3F98C96E" box="[1075,1217,542,568]" pageId="22" pageNumber="23">H. Martínez</collectorName>
leg.;
<collectionCode id="ED3D2FBAFF86FFD8FAAE245B3E0FC96E" box="[1283,1366,542,568]" collectionName="CEMT" pageId="22" pageNumber="23">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FF86FFD6FF10241A38A0C958" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="8B93B77FFF86FFD8FF10241A3A32C92F" blockId="22.[189,363,607,633]" box="[189,363,607,633]" pageId="22" pageNumber="23">
<heading id="D0DB0013FF86FFD8FF10241A3A32C92F" bold="true" box="[189,363,607,633]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B9586B6DFF86FFD8FF10241A3A32C92F" bold="true" box="[189,363,607,633]" pageId="22" pageNumber="23">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF4724CB3A73C9FE" blockId="22.[189,590,654,715]" box="[234,298,654,680]" pageId="22" pageNumber="23">
<emphasis id="B9586B6DFF86FFD8FF4724CB3A73C9FE" bold="true" box="[234,298,654,680]" pageId="22" pageNumber="23">Male</emphasis>
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF1024F43917C99D" blockId="22.[189,590,654,715]" box="[189,590,689,715]" pageId="22" pageNumber="23">
MEASUREMENTS. Length:
<quantity id="4CD41A9AFF86FFD8FE7324F43910C99D" box="[478,585,689,715]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="22" pageNumber="23" unit="mm" value="28.7">28.7 mm</quantity>
.
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF1024B63A7FC8EA" blockId="22.[187,1399,754,956]" pageId="22" pageNumber="23">
HEAD. Surface of anterior third rugose not reaching base of cephalic carina. Clypeus central emargination separating two rounded and dorsally angled clypeal teeth (
<figureCitation id="1317ABFAFF86FFD8FC2825503F49C866" box="[901,1040,789,816]" captionStart="Fig" captionStartId="23.[189,232,1927,1953]" captionTargetBox="[210,1378,272,1894]" captionTargetId="figure-59@23.[204,1383,265,1899]" captionTargetPageId="23" captionText="Fig. 7. Dichotomius deyrollei (Harold, 1869). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the head cephalic carina. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Metasternum dense pilosity." figureDoi="http://doi.org/10.5281/zenodo.4542121" httpUri="https://zenodo.org/record/4542121/files/figure.png" pageId="22" pageNumber="23">Fig. 7A, D</figureCitation>
). Cephalic carina higher than wide, antero-posteriorly flattened; anterior surface with well-marked punctures. Median tubercle approximately twice as high as lateral and gently inclined anteriorly, expanded and emarginate apically. Lateral tubercles, in frontal view, with two acute apices (
<figureCitation id="1317ABFAFF86FFD8FCFA253A38EFC8CC" box="[855,950,895,922]" captionStart="Fig" captionStartId="23.[189,232,1927,1953]" captionTargetBox="[210,1378,272,1894]" captionTargetId="figure-59@23.[204,1383,265,1899]" captionTargetPageId="23" captionText="Fig. 7. Dichotomius deyrollei (Harold, 1869). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the head cephalic carina. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Metasternum dense pilosity." figureDoi="http://doi.org/10.5281/zenodo.4542121" httpUri="https://zenodo.org/record/4542121/files/figure.png" pageId="22" pageNumber="23">Fig. 7D</figureCitation>
). Ventral clypeal process truncate (see
<figureCitation id="1317ABFAFF86FFD8FF1025E73A41C8EA" box="[189,280,930,956]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. AD. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. EF. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). GH. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). IJ. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="22" pageNumber="23">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF1025A139E4CF87" blockId="22.[188,1400,996,1234]" pageId="22" pageNumber="23">
PRONOTUM. Wider than long. Pronotal disc shiny and smooth, covered by fine punctures. Antero-medial edge with strong convex declivity, which in the centre could accommodate the large frontal carina (
<figureCitation id="1317ABFAFF86FFD8FF6B226F3A7FCF12" box="[198,294,1066,1092]" captionStart="Fig" captionStartId="23.[189,232,1927,1953]" captionTargetBox="[210,1378,272,1894]" captionTargetId="figure-59@23.[204,1383,265,1899]" captionTargetPageId="23" captionText="Fig. 7. Dichotomius deyrollei (Harold, 1869). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the head cephalic carina. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Metasternum dense pilosity." figureDoi="http://doi.org/10.5281/zenodo.4542121" httpUri="https://zenodo.org/record/4542121/files/figure.png" pageId="22" pageNumber="23">Fig. 7A</figureCitation>
). Row of interrupted ocellate punctures arranged along anterior and posterior margins, fewer and scattered on lateral margins. Hypomeron anterior edge, presenting rounded ocellate punctures with long setae, punctures separated by one time the width of a puncture. Hypomeron medial disc smooth (see
<figureCitation id="1317ABFAFF86FFD8FEE222D13AF1CFF8" box="[335,424,1172,1198]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. AD. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. EF. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). GH. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). IJ. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="22" pageNumber="23">Fig. 2E</figureCitation>
), without punctures. External margin bearing single row of organized long setae, all approximately of same length (see
<figureCitation id="1317ABFAFF86FFD8FDFA22F239E9CF87" box="[599,688,1207,1233]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. AD. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. EF. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). GH. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). IJ. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="22" pageNumber="23">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF1022BC3F46CEB0" blockId="22.[189,1401,1272,1510]" pageId="22" pageNumber="23">
ABDOMEN. Mesosternum and metepisternum fully covered by dense ocellate-setigerous punctures. Metasternum setigerous punctures restricted mainly to lateral and anterior areas (
<figureCitation id="1317ABFAFF86FFD8FB2823593FBCCE60" box="[1157,1253,1308,1334]" captionStart="Fig" captionStartId="23.[189,232,1927,1953]" captionTargetBox="[210,1378,272,1894]" captionTargetId="figure-59@23.[204,1383,265,1899]" captionTargetPageId="23" captionText="Fig. 7. Dichotomius deyrollei (Harold, 1869). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the head cephalic carina. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Metasternum dense pilosity." figureDoi="http://doi.org/10.5281/zenodo.4542121" httpUri="https://zenodo.org/record/4542121/files/figure.png" pageId="22" pageNumber="23">Fig. 7H</figureCitation>
). Pilosity of anterior lobe of metasternum connecting with lateral pilosity through margin of mesocoxa (
<figureCitation id="1317ABFAFF86FFD8FAA7237A3E32CE0F" box="[1290,1387,1343,1369]" captionStart="Fig" captionStartId="23.[189,232,1927,1953]" captionTargetBox="[210,1378,272,1894]" captionTargetId="figure-59@23.[204,1383,265,1899]" captionTargetPageId="23" captionText="Fig. 7. Dichotomius deyrollei (Harold, 1869). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the head cephalic carina. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Metasternum dense pilosity." figureDoi="http://doi.org/10.5281/zenodo.4542121" httpUri="https://zenodo.org/record/4542121/files/figure.png" pageId="22" pageNumber="23">Fig. 7H</figureCitation>
). Lateral surface of metasternum covered by ocellate setigerous punctures separated by twice width of a puncture, setae long and dense, covering lateral surface of metasternum and metepisternal suture. Ocellate setigerous punctures along antero-lateral margins, denser at lateral areas. Pygidium surface covered by fine punctures, basal margin with scattered ocellate punctures.
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF10204B384ACD1C" blockId="22.[186,1398,1549,1611]" pageId="22" pageNumber="23">
ELYTRA. Interstriae microsculpture shagreened (see
<figureCitation id="1317ABFAFF86FFD8FCA720483802CD71" box="[778,859,1549,1575]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. AD. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. EF. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). GH. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). IJ. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="22" pageNumber="23">Fig. 2J</figureCitation>
). Striae deeply impressed, crenulate, punctures separated by diameter of a puncture on elytral disc.
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF10203739DACDE6" blockId="22.[189,1398,1650,1712]" pageId="22" pageNumber="23">LEGS. Metacoxa posterior edge bearing single row of disorganized setigerous punctures, separated by half or less the diameter of a puncture.</paragraph>
<paragraph id="8B93B77FFF86FFD8FF102092386DCDA7" blockId="22.[189,820,1750,1777]" box="[189,820,1750,1777]" pageId="22" pageNumber="23">
PARAMERES AND LAMELLA COPULATRIX. As
<figureCitation id="1317ABFAFF86FFD8FD3B209339A9CDA7" box="[662,752,1750,1777]" captionStart="Fig" captionStartId="23.[189,232,1927,1953]" captionTargetBox="[210,1378,272,1894]" captionTargetId="figure-59@23.[204,1383,265,1899]" captionTargetPageId="23" captionText="Fig. 7. Dichotomius deyrollei (Harold, 1869). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the head cephalic carina. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Metasternum dense pilosity." figureDoi="http://doi.org/10.5281/zenodo.4542121" httpUri="https://zenodo.org/record/4542121/files/figure.png" pageId="22" pageNumber="23">Fig. 7C</figureCitation>
, EF.
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF10215D3AB7CC64" blockId="22.[189,494,1816,1842]" box="[189,494,1816,1842]" pageId="22" pageNumber="23">
<heading id="D0DB0013FF86FFD8FF10215D3AB7CC64" bold="true" box="[189,494,1816,1842]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B9586B6DFF86FFD8FF10215D3AB7CC64" bold="true" box="[189,494,1816,1842]" pageId="22" pageNumber="23">Morphological variation</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF4721023A9DCC37" blockId="22.[189,722,1862,1924]" box="[234,452,1862,1889]" pageId="22" pageNumber="23">
<emphasis id="B9586B6DFF86FFD8FF4721023A08CC37" bold="true" box="[234,337,1863,1889]" pageId="22" pageNumber="23">Females</emphasis>
(
<figureCitation id="1317ABFAFF86FFD8FECC21033AE2CC37" box="[353,443,1862,1889]" captionStart="Fig" captionStartId="23.[189,232,1927,1953]" captionTargetBox="[210,1378,272,1894]" captionTargetId="figure-59@23.[204,1383,265,1899]" captionTargetPageId="23" captionText="Fig. 7. Dichotomius deyrollei (Harold, 1869). AB. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the head cephalic carina. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Metasternum dense pilosity." figureDoi="http://doi.org/10.5281/zenodo.4542121" httpUri="https://zenodo.org/record/4542121/files/figure.png" pageId="22" pageNumber="23">Fig. 7B</figureCitation>
)
</paragraph>
<paragraph id="8B93B77FFF86FFD8FF10212F398BCCD2" blockId="22.[189,722,1862,1924]" box="[189,722,1898,1924]" pageId="22" pageNumber="23">As males except for the following characters:</paragraph>
<paragraph id="8B93B77FFF86FFD8FF1021E93A76CCBE" blockId="22.[189,1399,1963,2024]" pageId="22" pageNumber="23">HEAD. Surface transversely rugose, denser on anterior portion. Cephalic carina with pair of high medial tubercles.</paragraph>
<caption id="DF53E7F7FF87FFD9FF1021C239EBCCBE" ID-DOI="http://doi.org/10.5281/zenodo.4542121" ID-Zenodo-Dep="4542121" httpUri="https://zenodo.org/record/4542121/files/figure.png" pageId="23" pageNumber="24" startId="23.[189,232,1927,1953]" targetBox="[210,1378,272,1894]" targetPageId="23">
<paragraph id="8B93B77FFF87FFD9FF1021C239EBCCBE" blockId="23.[189,1399,1926,2025]" pageId="23" pageNumber="24">
<emphasis id="B9586B6DFF87FFD9FF1021C23A52CCF4" bold="true" box="[189,267,1927,1954]" pageId="23" pageNumber="24">Fig. 7.</emphasis>
<taxonomicName id="4C2CCCFCFF87FFD9FEB921CD398DCCF5" authority="(Harold, 1869)" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[276,724,1928,1955]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="B9586B6DFF87FFD9FEB921CD3940CCF4" box="[276,537,1928,1954]" italics="true" pageId="23" pageNumber="24">Dichotomius deyrollei</emphasis>
(
<bibRefCitation id="EFBDCA8EFF87FFD9FD8421CD3993CCF5" author="Harold E." box="[553,714,1928,1955]" pageId="23" pageNumber="24" pagination="124 - 144" refId="ref29859" refString="Harold E. 1869. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 124 - 144." type="journal article" year="1869">Harold, 1869</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B9586B6DFF87FFD9FD7221C2384FCCF7" bold="true" box="[735,790,1927,1953]" pageId="23" pageNumber="24">AB</emphasis>
. Dorsal habitus.
<emphasis id="B9586B6DFF87FFD9FC4D21C238ACCCF7" bold="true" box="[992,1013,1927,1953]" pageId="23" pageNumber="24">A</emphasis>
. ♂.
<emphasis id="B9586B6DFF87FFD9FB8421CD3F64CCF4" bold="true" box="[1065,1085,1928,1954]" pageId="23" pageNumber="24">B</emphasis>
. ♀.
<emphasis id="B9586B6DFF87FFD9FBDF21C23FDECCF7" bold="true" box="[1138,1159,1927,1953]" pageId="23" pageNumber="24">C</emphasis>
. Lateral view of the aedeagus.
<emphasis id="B9586B6DFF87FFD9FE9A21EE3A15CC93" bold="true" box="[311,332,1963,1989]" pageId="23" pageNumber="24">D</emphasis>
. Anterior view of the head cephalic carina.
<emphasis id="B9586B6DFF87FFD9FCE021EE3838CC93" bold="true" box="[845,865,1963,1989]" pageId="23" pageNumber="24">E</emphasis>
. Ventral view of the aedeagus.
<emphasis id="B9586B6DFF87FFD9FB7D21EE3E5ECC93" bold="true" box="[1232,1287,1963,1989]" pageId="23" pageNumber="24">FG</emphasis>
. Lamella copulatrix.
<emphasis id="B9586B6DFF87FFD9FEEE218B3A03CCBE" bold="true" box="[323,346,1998,2024]" pageId="23" pageNumber="24">H</emphasis>
. Metasternum dense pilosity.
</paragraph>
</caption>
<paragraph id="8B93B77FFF88FFD6FF10274C3888CADB" blockId="24.[189,1399,265,397]" pageId="24" pageNumber="25">PRONOTUM. With weak medial knob, observable in lateral view. Uninterrupted ocellate punctures throughout lateral margins. Hypomeron punctures ocellate and setigerous, equally spaced; setae longer and denser than in males. Metasternum almost completely covered by ocellate setigerous punctures, metasternal lobe smooth with fine and deeply impressed punctures.</paragraph>
<paragraph id="8B93B77FFF88FFD6FF1027F139BBCA98" blockId="24.[189,738,435,462]" box="[189,738,435,462]" pageId="24" pageNumber="25">ELYTRA. Smooth, with scattered fine punctures.</paragraph>
<paragraph id="8B93B77FFF88FFD6FF1027B038A0C958" blockId="24.[189,1017,500,527]" box="[189,1017,500,527]" pageId="24" pageNumber="25">ABDOMEN. Sternites dense ocellate punctures covering anterior margin.</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FF88FFD6FF1024703A2AC9B3" pageId="24" pageNumber="25" type="discussion">
<paragraph id="8B93B77FFF88FFD6FF1024703A76C919" blockId="24.[189,303,565,591]" box="[189,303,565,591]" pageId="24" pageNumber="25">
<heading id="D0DB0013FF88FFD6FF1024703A76C919" bold="true" box="[189,303,565,591]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B9586B6DFF88FFD6FF1024703A76C919" bold="true" box="[189,303,565,591]" pageId="24" pageNumber="25">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFF88FFD6FF1024263A2AC9B3" blockId="24.[189,1399,611,742]" pageId="24" pageNumber="25">
This species occurs in
<collectingCountry id="F33BF7EFFF88FFD6FE7E2426391EC92B" box="[467,583,611,637]" name="Colombia" pageId="24" pageNumber="25">Colombia</collectingCountry>
and
<collectingCountry id="F33BF7EFFF88FFD6FD2B2426385AC92B" box="[646,771,611,637]" name="Venezuela" pageId="24" pageNumber="25">Venezuela</collectingCountry>
. The distribution of the species closely follows that of the Andes Mountains. The original description notes that the
<typeStatus id="549709DDFF88FFD6FC6A24C338A3C9F6" box="[967,1018,646,672]" pageId="24" pageNumber="25">type</typeStatus>
specimens were from “
<collectingCountry id="F33BF7EFFF88FFD6FAB724C33E3EC9F6" box="[1306,1383,646,672]" name="Brazil" pageId="24" pageNumber="25">Brésil</collectingCountry>
”, without any other specification of the locality. However, as far as we know this Andean species does not occur in
<collectingCountry id="F33BF7EFFF88FFD6FE89248E3A36C9B3" box="[292,367,715,741]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>