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<document ID-DOI="10.11646/phytotaxa.595.2.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="7905867" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1683527015091" checkinUser="plazi" docAuthor="Büneker, Henrique Mallmann, Pastori, Tamara, Almeida, Pedro Schwambach De &amp; Mariath, Jorge Ernesto De Araujo" docDate="2023" docId="FB6AEA65FF93FFC022F0FA58E39AF90D" docLanguage="en" docName="phytotaxa.595.2.4.pdf" docOrigin="Phytotaxa 595 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.595.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.6:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="6" docTitle="Dyckia magnifica Büneker, Pastori, Almeida &amp; Mariath, 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="179" masterDocId="0753921DFF9BFFCA2278FFF6E238FFAA" masterDocTitle="Dyckia magnifica, a new species of Bromeliaceae (Pitcairnioideae) from Southern Brazil, and the re-establishment of D. tomentosa, of the D. selloa complex" masterLastPageNumber="185" masterPageNumber="169" pageNumber="177" updateTime="1683565175284" updateUser="tatiana" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Dyckia magnifica, a new species of Bromeliaceae (Pitcairnioideae) from Southern Brazil, and the re-establishment of D. tomentosa, of the D. selloa complex</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Büneker, Henrique Mallmann</mods:namePart>
<mods:affiliation>Laboratório de Anatomia Vegetal (LAVeg), Instituto de Biociências, Departamento de Botânica, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; Programa de Pós-graduação em Botânica, Instituto de Biociências, UFRGS, Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pastori, Tamara</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2477-0275</mods:nameIdentifier>
<mods:affiliation>Laboratório de Anatomia Vegetal (LAVeg), Instituto de Biociências, Departamento de Botânica, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; Programa de Pós-graduação em Botânica, Instituto de Biociências, UFRGS, Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; tamarapastori @ gmail. com; https: // orcid. org / 0000 - 0002 - 2477 - 0275</mods:affiliation>
<mods:nameIdentifier type="email">tamarapastori@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Almeida, Pedro Schwambach De</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-2042-3687</mods:nameIdentifier>
<mods:affiliation>Laboratório de Anatomia Vegetal (LAVeg), Instituto de Biociências, Departamento de Botânica, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; Laboratório de Palinologia, Departamento de Botânica, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, Rio de Janeiro, RJ, Brazil. &amp; Programa de Pós-graduação em Ciências Biológicas (Botânica), Museu Nacional, Universidade Federal do Rio de Janeiro (UFRJ), Quinta da Boa Vista, São Cristóvão, Rio de Janeiro, RJ, Brazil. &amp; pedro. schwambach @ ufrj. br; https: // orcid. org / 0000 - 0003 - 2042 - 3687</mods:affiliation>
<mods:nameIdentifier type="email">pedro.schwambach@ufrj.br</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mariath, Jorge Ernesto De Araujo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2776-6781</mods:nameIdentifier>
<mods:affiliation>Programa de Pós-graduação em Botânica, Instituto de Biociências, UFRGS, Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; jorge. mariath @ ufrgs. br; https: // orcid. org / 0000 - 0002 - 2776 - 6781</mods:affiliation>
<mods:nameIdentifier type="email">jorge.mariath@ufrgs.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>595</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>169</mods:start>
<mods:end>185</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/phytotaxa.595.2.4</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.595.2.4</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7905867</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D" httpUri="http://treatment.plazi.org/id/FB6AEA65FF93FFC022F0FA58E39AF90D" lastPageId="10" lastPageNumber="179" pageId="8" pageNumber="177">
<subSubSection box="[136,1086,1454,1481]" pageId="8" pageNumber="177" type="nomenclature">
<paragraph blockId="8.[136,1452,1454,1732]" box="[136,1086,1454,1481]" pageId="8" pageNumber="177">
<heading box="[136,1086,1454,1481]" fontSize="11" level="2" pageId="8" pageNumber="177" reason="4">
3.
<taxonomicName authorityName="Büneker &amp; Pastori &amp; Almeida &amp; Mariath" authorityYear="2023" box="[164,369,1454,1481]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="177" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis bold="true" box="[164,369,1454,1481]" italics="true" pageId="8" pageNumber="177">Dyckia magnifica</emphasis>
</taxonomicName>
B̧neker &amp; Mariath,
<emphasis box="[609,700,1455,1480]" italics="true" pageId="8" pageNumber="177">
<taxonomicNameLabel box="[609,696,1455,1480]" pageId="8" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
<figureCitation box="[707,866,1454,1480]" captionStart="FIGURE 1" captionStartId="4.[136,229,1811,1833]" captionTargetBox="[151,1436,187,1783]" captionTargetId="figure-24@4.[151,1436,187,1783]" captionTargetPageId="4" captionText="FIGURE 1. A–E. Dyckia magnifica (H.M. B¸neker 616 et al.) in habitat. A. Detail of part of the population in habitat on rocky banks of the Canoas river dammed by PCH Campos Novos. B. Vegetative habit. C. Habit when fertile. D. Detail of apical portion of the inflorescence. E. Immature capsules." figureDoi="http://doi.org/10.5281/zenodo.7905869" httpUri="https://zenodo.org/record/7905869/files/figure.png" pageId="8" pageNumber="177">Figures 1A–E</figureCitation>
;
<figureCitation box="[878,1009,1454,1481]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="8" pageNumber="177">2C, D, F, H</figureCitation>
, and J.
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="177" type="diagnosis">
<paragraph blockId="8.[136,1452,1454,1732]" pageId="8" pageNumber="177">
<emphasis italics="true" pageId="8" pageNumber="177">
Species morphologice proxima Dyckiae tomentosae, differt spinis triangularibus, patentibus (vs. stricto-triangularibus, aduncis), floribus secundis (vs. patentibus usque ad suberectis) cum pedicellis maioribus (
<quantity box="[897,977,1530,1551]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="8" pageNumber="177" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
vs. absentibus usque ad
<quantity box="[1224,1297,1530,1551]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="177" unit="mm" value="0.5">0.5 mm</quantity>
longis), sepalis pubescentibus (vs. tomentosis), petalis longioribus (
<quantity box="[693,783,1566,1587]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="8" pageNumber="177" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
vs.
<quantity box="[823,919,1566,1587]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.6" metricValueMin="6.0" pageId="8" pageNumber="177" unit="mm" value="6.8" valueMax="7.6" valueMin="6.0">6–7.6 mm</quantity>
longis), pubescentibus (vs. tomentosis), cum trichomis superficiei abaxialis flabeliformibus cum ramis brevis (vs. dendriticis cum ramis elongatis), et trichomis marginum digitiformibus (vs. filiformibus).
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="177" type="materials_examined">
<paragraph blockId="8.[136,1452,1454,1732]" pageId="8" pageNumber="177">
<emphasis bold="true" box="[136,194,1674,1695]" pageId="8" pageNumber="177">
<collectingRegion box="[136,190,1674,1695]" pageId="8" pageNumber="177">Type</collectingRegion>
:
</emphasis>
—
<materialsCitation box="[222,1387,1674,1696]" collectingDate="2017-02-08" collectionCode="H, M, B" country="Brazil" location="Perau do Macaco Branco" pageId="8" pageNumber="177" specimenCount="1" stateProvince="Santa Catarina">
<collectingCountry box="[222,312,1674,1695]" name="Brazil" pageId="8" pageNumber="177">BRAZIL</collectingCountry>
.
<collectingRegion box="[321,465,1674,1696]" country="Brazil" name="Santa Catarina" pageId="8" pageNumber="177">Santa Catarina</collectingRegion>
:
<emphasis bold="true" box="[473,620,1674,1696]" pageId="8" pageNumber="177">Abdon Batista</emphasis>
,
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF93FFC22000F97CE154F935" box="[632,876,1674,1696]" country="Brazil" name="Perau do Macaco Branco" pageId="8" pageNumber="177" stateProvince="Santa Catarina">Perau do Macaco Branco</location>
, -27.609947°, -51.181137°,
<date box="[1147,1307,1674,1696]" pageId="8" pageNumber="177" value="2017-02-08">
<collectingDate box="[1147,1307,1674,1696]" pageId="8" pageNumber="177" value="2017-02-08">8 February 2017</collectingDate>
</date>
,
<emphasis box="[1316,1387,1674,1696]" italics="true" pageId="8" pageNumber="177">
<collectionCode box="[1316,1336,1674,1695]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="8" pageNumber="177" type="Herbarium">H</collectionCode>
.
<collectionCode box="[1339,1362,1674,1695]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="8" pageNumber="177" type="Herbarium">M</collectionCode>
.
<collectionCode box="[1371,1387,1674,1696]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="8" pageNumber="177" type="Herbarium">B</collectionCode>
</emphasis>
</materialsCitation>
<emphasis italics="true" pageId="8" pageNumber="177">
¸neker 616,
<collectionCode box="[237,255,1710,1731]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="177">R</collectionCode>
.
<collectionCode box="[258,276,1710,1731]" country="Denmark" name="University of Copenhagen" pageId="8" pageNumber="177" type="Herbarium">C</collectionCode>
.
<materialsCitation box="[286,418,1710,1731]" collectionCode="J, C" country="Brazil" location="Pontes" pageId="8" pageNumber="177" specimenCount="1" stateProvince="Abdon Batista">
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF93FFC22366F958E367F969" box="[286,351,1710,1731]" country="Brazil" name="Pontes" pageId="8" pageNumber="177" stateProvince="Abdon Batista">Pontes</location>
&amp;
<collectionCode box="[382,396,1710,1731]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="8" pageNumber="177" type="Herbarium">J</collectionCode>
.
<collectionCode box="[400,418,1710,1731]" country="Denmark" name="University of Copenhagen" pageId="8" pageNumber="177" type="Herbarium">C</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation box="[427,668,1710,1732]" collectionCode="HDCF" country="Brazil" location="Silva" pageId="8" pageNumber="177" specimenCount="1" stateProvince="Abdon Batista" typeStatus="holotype">
<emphasis box="[427,475,1710,1731]" italics="true" pageId="8" pageNumber="177">
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF93FFC223D3F958E3E3F969" box="[427,475,1710,1731]" country="Brazil" name="Silva" pageId="8" pageNumber="177" stateProvince="Abdon Batista">Silva</location>
</emphasis>
(
<typeStatus box="[487,576,1710,1732]" pageId="8" pageNumber="177">holotype</typeStatus>
:
<collectionCode box="[585,658,1710,1731]" pageId="8" pageNumber="177">HDCF</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="178" pageId="8" pageNumber="177" type="description">
<paragraph blockId="8.[136,1451,1778,2093]" lastBlockId="9.[136,1452,158,2093]" lastPageId="9" lastPageNumber="178" pageId="8" pageNumber="177">
<emphasis bold="true" box="[136,195,1778,1805]" italics="true" pageId="8" pageNumber="177">Herb</emphasis>
saxicolous, rhizomatous, 0.95–1.4(–2.1) m in height when in bloom;
<emphasis box="[954,1029,1779,1804]" italics="true" pageId="8" pageNumber="177">rosette</emphasis>
symmetrical or slightly asymmetrical, (25–)
<quantity box="[195,311,1814,1841]" metricMagnitude="-1" metricUnit="m" metricValue="6.4" metricValueMax="8.3" metricValueMin="4.5" pageId="8" pageNumber="177" unit="cm" value="64.0" valueMax="83.0" valueMin="45.0">45–83 cm</quantity>
in diameter.
<emphasis bold="true" box="[467,546,1815,1841]" italics="true" pageId="8" pageNumber="177">Leaves</emphasis>
<quantity box="[557,673,1814,1841]" metricMagnitude="0" metricUnit="m" metricValue="1.8796" metricValueMax="2.667" metricValueMin="1.0922" pageId="8" pageNumber="177" unit="in" value="74.0" valueMax="105.0" valueMin="43.0">43–105 in</quantity>
number, the inner leaves erect, the outer leaves patent to reflexed;
<emphasis box="[136,209,1850,1876]" italics="true" pageId="8" pageNumber="177">blades</emphasis>
(12–)23–52 ×
<quantity box="[376,503,1850,1877]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.9" metricValueMin="1.7" pageId="8" pageNumber="177" unit="cm" value="2.8" valueMax="3.9" valueMin="1.7">1.7–3.9 cm</quantity>
, narrowly triangular, rigid, adaxially green to purplish, sparsely to densely adpressed lepidote, abaxial face longitudinally nerved-striate, green to purplish, lepidote between nerves, margins spinose-serrate;
<emphasis box="[136,206,1923,1948]" italics="true" pageId="8" pageNumber="177">spines</emphasis>
4–7.6 ×
<quantity box="[310,428,1922,1949]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="5.0" metricValueMin="2.8" pageId="8" pageNumber="177" unit="mm" value="3.9" valueMax="5.0" valueMin="2.8">2.8–5 mm</quantity>
, conspicuous, triangular, patent, slightly antrorsely curved, rigid, whitish green or whitish rose at the base, whitish or yellowish in the median portion and brown at the apex, generally lepidote.
<emphasis bold="true" box="[1294,1451,1958,1985]" italics="true" pageId="8" pageNumber="177">Inflorescence</emphasis>
axillary, erect;
<emphasis box="[309,410,1994,2020]" italics="true" pageId="8" pageNumber="177">peduncle</emphasis>
<quantity box="[419,533,1994,2021]" metricMagnitude="-1" metricUnit="m" metricValue="4.8" metricValueMax="7.6" metricValueMin="2.0" pageId="8" pageNumber="177" unit="cm" value="48.0" valueMax="76.0" valueMin="20.0">20–76 cm</quantity>
long, generally slender, green to purple, glabrescent, pruinose;
<emphasis box="[1270,1451,1994,2020]" italics="true" pageId="8" pageNumber="177">peduncle bracts</emphasis>
of the median portion polystichous, stramineous, shorter than to surpassing the internodes, divergent, not hiding the peduncle;
<emphasis box="[251,595,2066,2092]" italics="true" pageId="8" pageNumber="177">fertile part of the inflorescence</emphasis>
65–108(–130) cm long, generally once-branched to slightly twice-branched;
<emphasis box="[136,306,158,184]" italics="true" pageId="9" pageNumber="178">primary bracts</emphasis>
identical to the peduncle bracts;
<emphasis box="[689,917,158,184]" italics="true" pageId="9" pageNumber="178">first order branches</emphasis>
(8–)
<quantity box="[971,1074,158,185]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" metricValueMax="12.953999999999999" metricValueMin="5.334" pageId="9" pageNumber="178" unit="in" value="36.0" valueMax="51.0" valueMin="21.0">21–51 in</quantity>
number, suberect to patent, with greenish to purple axes, glabrescent, pruinose at the base, white floccose-tomentose distally;
<emphasis box="[1193,1452,194,220]" italics="true" pageId="9" pageNumber="178">second order branches</emphasis>
(when present) usually less than 2, at the base of the first order branches;
<emphasis box="[962,1101,230,256]" italics="true" pageId="9" pageNumber="178">floral bracts</emphasis>
1–3.6 ×
<emphasis box="[1202,1235,231,256]" italics="true" pageId="9" pageNumber="178">ca.</emphasis>
<quantity box="[1242,1331,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="178" unit="mm" value="1.3">1.3 mm</quantity>
, ovate, the base green, the median portion stramineous and yellowish, the apex stramineous and brown, attenuate-acuminate, ecarinate, tomentose, shorter than the pedicels of the flowers.
<emphasis bold="true" box="[862,955,302,329]" italics="true" pageId="9" pageNumber="178">Flowers</emphasis>
<quantity box="[966,1094,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="9" pageNumber="178" unit="cm" value="1.4" valueMax="1.6" valueMin="1.2">1.2–1.6 cm</quantity>
long, polystichous at first and becoming secund at anthesis, after anthesis becoming erect again, conspicuously pedicellate;
<emphasis box="[1173,1265,338,364]" italics="true" pageId="9" pageNumber="178">pedicels</emphasis>
clavate, twisted,
<quantity box="[136,230,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="9" pageNumber="178" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long, yellowish green, tomentose;
<emphasis box="[626,696,374,400]" italics="true" pageId="9" pageNumber="178">sepals</emphasis>
3–5 ×
<emphasis box="[777,810,375,400]" italics="true" pageId="9" pageNumber="178">ca.</emphasis>
<quantity box="[818,907,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="178" unit="mm" value="2.5">2.5 mm</quantity>
, triangular-ovate, apex obtuse to rounded, green at the base, yellow at the apex, slightly carinate, abaxial surface pubescent, covered with flabelliform trichomes, the margins with digitiform trichomes;
<emphasis box="[536,786,446,472]" italics="true" pageId="9" pageNumber="178">flabelliform trichomes</emphasis>
on the sepal abaxial surface with an uniseriate stalk of 2–3 cells, the median region expanded, multicellular, with 4–9 short branches formed by
<emphasis box="[1092,1126,483,508]" italics="true" pageId="9" pageNumber="178">ca.</emphasis>
2 cells;
<emphasis box="[1221,1451,482,508]" italics="true" pageId="9" pageNumber="178">digitiform trichomes</emphasis>
of the sepal margins with an uniseriate stalk of (1–)2(–3) cells, the median region not expanded, branches with 1–3 much-elongated cells;
<emphasis box="[391,458,554,580]" italics="true" pageId="9" pageNumber="178">petals</emphasis>
9–11 ×
<emphasis box="[551,585,555,580]" italics="true" pageId="9" pageNumber="178">ca.</emphasis>
<quantity box="[592,660,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="178" unit="mm" value="3.0">3 mm</quantity>
, not unguiculate, erect, oblong, the basal portion greenish, the median portion yellowish, the apex orange-yellow, rounded or emarginate, slightly cucullate, abaxially pubescent, covered with flabelliform trichomes, the margins with digitiform trichomes;
<emphasis box="[910,1130,626,652]" italics="true" pageId="9" pageNumber="178">dendritic trichomes</emphasis>
on the petal abaxial surface identical to the trichomes of the sepals surfaces;
<emphasis box="[683,884,662,688]" italics="true" pageId="9" pageNumber="178">filiform trichomes</emphasis>
of the petal margins identical to those of the sepal margins;
<emphasis box="[242,332,699,724]" italics="true" pageId="9" pageNumber="178">stamens</emphasis>
included during theca opening, becoming exserted;
<emphasis box="[931,1034,698,724]" italics="true" pageId="9" pageNumber="178">filaments</emphasis>
<quantity box="[1043,1151,698,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="9" pageNumber="178" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
long, free above the short hypanthium, the antesepalous ones adnate to the sepals for
<emphasis box="[804,838,735,760]" italics="true" pageId="9" pageNumber="178">ca.</emphasis>
<quantity box="[845,934,734,761]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="178" unit="mm" value="0.4">0.4 mm</quantity>
, the antepetalous ones adnate to the petals for
<emphasis box="[136,170,771,796]" italics="true" pageId="9" pageNumber="178">ca.</emphasis>
<quantity box="[177,245,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="178" unit="mm" value="1.0">1 mm</quantity>
;
<emphasis box="[257,313,770,796]" italics="true" pageId="9" pageNumber="178">pistil</emphasis>
<quantity box="[320,428,770,797]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="9" pageNumber="178" unit="mm" value="10.5" valueMax="12.0" valueMin="9.0">9–12 mm</quantity>
long;
<emphasis box="[499,563,771,796]" italics="true" pageId="9" pageNumber="178">ovary</emphasis>
2 ×
<quantity box="[615,732,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="9" pageNumber="178" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
;
<emphasis box="[744,795,770,796]" italics="true" pageId="9" pageNumber="178">style</emphasis>
<quantity box="[802,896,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="9" pageNumber="178" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long;
<emphasis box="[968,1043,771,796]" italics="true" pageId="9" pageNumber="178">stigma</emphasis>
exserted at anthesis, stigmatic lobes conduplicate, slightly spirally twisted at pre-anthesis, becoming suberect afterward.
<emphasis bold="true" box="[1070,1174,806,833]" italics="true" pageId="9" pageNumber="178">Capsules</emphasis>
ovoid, dark brown when mature.
<emphasis bold="true" box="[228,293,842,869]" italics="true" pageId="9" pageNumber="178">Seeds</emphasis>
oblanceoloid.
</paragraph>
<paragraph blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="178">
<emphasis bold="true" box="[189,361,878,904]" pageId="9" pageNumber="178">Leaf anatomy:</emphasis>
—The analysis of the leaf anatomy of
<taxonomicName authorityName="Büneker &amp; Pastori &amp; Almeida &amp; Mariath" authorityYear="2023" box="[766,908,878,904]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="9" pageNumber="178" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis box="[766,908,878,904]" italics="true" pageId="9" pageNumber="178">D.magnifica</emphasis>
</taxonomicName>
enabled the identification of the following tissues: epidermis, water-storage parenchyma, chlorenchyma, vascular bundles, spongy parenchyma, ground parenchyma and mechanical hypodermis (
<figureCitation box="[424,611,950,976]" captionStart="FIGURE 3" captionStartId="7.[136,229,1679,1701]" captionTargetBox="[151,1436,497,1652]" captionTargetId="figure-179@7.[151,1436,497,1654]" captionTargetPageId="7" captionText="FIGURE 3. Leaf anatomy of Dyckia tomentosa (A, C, E, H, J and K) and Dyckia magnifica (B, D, F, G and I). A–B. Transverse section of blade, without coloration, indicating the adaxial surface (AD) and the abaxial surface (AB). C–D. Transverse section of blade showing the identified tissues: hypodermis (hy), water-storage parenchyma (wp), chlorenchyma (c), vascular bundle (vb), ground parenchyma (p), spongy parenchyma (sp). E–J. Transverse section of blade of material fixed and stained with toluidine blue O. E. Adaxial surface of blade showing the epidermis, the silica bodies (sb) of the epidermis, and the hypodermis (hy). F. Abaxial surface of the blade showing the stomatal complex (st). G. Transverse section of the blade showing the leaf margin and the lignified hypodermis (hy) on both surfaces. H. Tissues identified in the central section of the transverse section, water-storage parenchyma (wp), chlorenchyma (c), ground parenchyma (p) and peltate trichomes (pt). I. Vascular bundle showing the presence of sclerenchyma fibers cap (fb) and the position of phloem (ph) and xylem (x). J. Water-storage parenchyma (wp) and ground parenchyma (p). K. Transverse section of fresh material stained with astra-blue - safranin, showing the sclerenchyma fiber caps (fb) associated with vascular bundles, spongy parenchyma (sp) and ground parenchyma (p) and mechanical hypodermis (hy)." figureDoi="http://doi.org/10.5281/zenodo.7905873" httpUri="https://zenodo.org/record/7905873/files/figure.png" pageId="9" pageNumber="178">Fig. 3B and 3D</figureCitation>
). The epidermis consists of cells that contain silica bodies and present a predominant thickening in the cell walls. The stomatal complexes are located on the abaxial surface of the leaves upward the level of the epidermal cells (
<figureCitation box="[584,669,1022,1048]" captionStart="FIGURE 3" captionStartId="7.[136,229,1679,1701]" captionTargetBox="[151,1436,497,1652]" captionTargetId="figure-179@7.[151,1436,497,1654]" captionTargetPageId="7" captionText="FIGURE 3. Leaf anatomy of Dyckia tomentosa (A, C, E, H, J and K) and Dyckia magnifica (B, D, F, G and I). A–B. Transverse section of blade, without coloration, indicating the adaxial surface (AD) and the abaxial surface (AB). C–D. Transverse section of blade showing the identified tissues: hypodermis (hy), water-storage parenchyma (wp), chlorenchyma (c), vascular bundle (vb), ground parenchyma (p), spongy parenchyma (sp). E–J. Transverse section of blade of material fixed and stained with toluidine blue O. E. Adaxial surface of blade showing the epidermis, the silica bodies (sb) of the epidermis, and the hypodermis (hy). F. Abaxial surface of the blade showing the stomatal complex (st). G. Transverse section of the blade showing the leaf margin and the lignified hypodermis (hy) on both surfaces. H. Tissues identified in the central section of the transverse section, water-storage parenchyma (wp), chlorenchyma (c), ground parenchyma (p) and peltate trichomes (pt). I. Vascular bundle showing the presence of sclerenchyma fibers cap (fb) and the position of phloem (ph) and xylem (x). J. Water-storage parenchyma (wp) and ground parenchyma (p). K. Transverse section of fresh material stained with astra-blue - safranin, showing the sclerenchyma fiber caps (fb) associated with vascular bundles, spongy parenchyma (sp) and ground parenchyma (p) and mechanical hypodermis (hy)." figureDoi="http://doi.org/10.5281/zenodo.7905873" httpUri="https://zenodo.org/record/7905873/files/figure.png" pageId="9" pageNumber="178">Fig. 3F</figureCitation>
). The sub-stomatal chambers are associated with spongy parenchyma, which provides a larger dimension to the area of these chambers and is located between the vascular bundles. The mesophyll presents, on the abaxial and adaxial surfaces, a mechanical hypodermis rich in lignins, consisting of three or four layers of cells (
<figureCitation box="[389,479,1130,1156]" captionStart="FIGURE 3" captionStartId="7.[136,229,1679,1701]" captionTargetBox="[151,1436,497,1652]" captionTargetId="figure-179@7.[151,1436,497,1654]" captionTargetPageId="7" captionText="FIGURE 3. Leaf anatomy of Dyckia tomentosa (A, C, E, H, J and K) and Dyckia magnifica (B, D, F, G and I). A–B. Transverse section of blade, without coloration, indicating the adaxial surface (AD) and the abaxial surface (AB). C–D. Transverse section of blade showing the identified tissues: hypodermis (hy), water-storage parenchyma (wp), chlorenchyma (c), vascular bundle (vb), ground parenchyma (p), spongy parenchyma (sp). E–J. Transverse section of blade of material fixed and stained with toluidine blue O. E. Adaxial surface of blade showing the epidermis, the silica bodies (sb) of the epidermis, and the hypodermis (hy). F. Abaxial surface of the blade showing the stomatal complex (st). G. Transverse section of the blade showing the leaf margin and the lignified hypodermis (hy) on both surfaces. H. Tissues identified in the central section of the transverse section, water-storage parenchyma (wp), chlorenchyma (c), ground parenchyma (p) and peltate trichomes (pt). I. Vascular bundle showing the presence of sclerenchyma fibers cap (fb) and the position of phloem (ph) and xylem (x). J. Water-storage parenchyma (wp) and ground parenchyma (p). K. Transverse section of fresh material stained with astra-blue - safranin, showing the sclerenchyma fiber caps (fb) associated with vascular bundles, spongy parenchyma (sp) and ground parenchyma (p) and mechanical hypodermis (hy)." figureDoi="http://doi.org/10.5281/zenodo.7905873" httpUri="https://zenodo.org/record/7905873/files/figure.png" pageId="9" pageNumber="178">Fig. 3G</figureCitation>
). On the adaxial surface it is possible to identify water-storage parenchyma comprising three to four layers of isodiametric cells and then five to six layers of anticlinally elongated cells, with
<emphasis box="[1321,1355,1167,1192]" italics="true" pageId="9" pageNumber="178">ca.</emphasis>
<quantity box="[1364,1452,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="178" unit="mm" value="1.2">1.2 mm</quantity>
height. This parenchyma extends to the region close to the vascular bundles, where it is interspersed with projections of chlorenchyma, 50–100 μm height. The bundle sheath is continuous around the vascular bundle. The collateral vascular bundles are distributed linearly throughout the leaf section and have lignified sclerenchymatic fibers at the poles of the xylem and of the phloem (
<figureCitation box="[571,649,1310,1336]" captionStart="FIGURE 3" captionStartId="7.[136,229,1679,1701]" captionTargetBox="[151,1436,497,1652]" captionTargetId="figure-179@7.[151,1436,497,1654]" captionTargetPageId="7" captionText="FIGURE 3. Leaf anatomy of Dyckia tomentosa (A, C, E, H, J and K) and Dyckia magnifica (B, D, F, G and I). A–B. Transverse section of blade, without coloration, indicating the adaxial surface (AD) and the abaxial surface (AB). C–D. Transverse section of blade showing the identified tissues: hypodermis (hy), water-storage parenchyma (wp), chlorenchyma (c), vascular bundle (vb), ground parenchyma (p), spongy parenchyma (sp). E–J. Transverse section of blade of material fixed and stained with toluidine blue O. E. Adaxial surface of blade showing the epidermis, the silica bodies (sb) of the epidermis, and the hypodermis (hy). F. Abaxial surface of the blade showing the stomatal complex (st). G. Transverse section of the blade showing the leaf margin and the lignified hypodermis (hy) on both surfaces. H. Tissues identified in the central section of the transverse section, water-storage parenchyma (wp), chlorenchyma (c), ground parenchyma (p) and peltate trichomes (pt). I. Vascular bundle showing the presence of sclerenchyma fibers cap (fb) and the position of phloem (ph) and xylem (x). J. Water-storage parenchyma (wp) and ground parenchyma (p). K. Transverse section of fresh material stained with astra-blue - safranin, showing the sclerenchyma fiber caps (fb) associated with vascular bundles, spongy parenchyma (sp) and ground parenchyma (p) and mechanical hypodermis (hy)." figureDoi="http://doi.org/10.5281/zenodo.7905873" httpUri="https://zenodo.org/record/7905873/files/figure.png" pageId="9" pageNumber="178">Fig. 3I</figureCitation>
). The diameter of the vascular bundles varies along the cross section of the leaf. The larger caliber bundles have fiber caps of the same thickness, both at the xylem pole and at the phloem pole (
<figureCitation box="[144,223,1382,1408]" captionStart="FIGURE 3" captionStartId="7.[136,229,1679,1701]" captionTargetBox="[151,1436,497,1652]" captionTargetId="figure-179@7.[151,1436,497,1654]" captionTargetPageId="7" captionText="FIGURE 3. Leaf anatomy of Dyckia tomentosa (A, C, E, H, J and K) and Dyckia magnifica (B, D, F, G and I). A–B. Transverse section of blade, without coloration, indicating the adaxial surface (AD) and the abaxial surface (AB). C–D. Transverse section of blade showing the identified tissues: hypodermis (hy), water-storage parenchyma (wp), chlorenchyma (c), vascular bundle (vb), ground parenchyma (p), spongy parenchyma (sp). E–J. Transverse section of blade of material fixed and stained with toluidine blue O. E. Adaxial surface of blade showing the epidermis, the silica bodies (sb) of the epidermis, and the hypodermis (hy). F. Abaxial surface of the blade showing the stomatal complex (st). G. Transverse section of the blade showing the leaf margin and the lignified hypodermis (hy) on both surfaces. H. Tissues identified in the central section of the transverse section, water-storage parenchyma (wp), chlorenchyma (c), ground parenchyma (p) and peltate trichomes (pt). I. Vascular bundle showing the presence of sclerenchyma fibers cap (fb) and the position of phloem (ph) and xylem (x). J. Water-storage parenchyma (wp) and ground parenchyma (p). K. Transverse section of fresh material stained with astra-blue - safranin, showing the sclerenchyma fiber caps (fb) associated with vascular bundles, spongy parenchyma (sp) and ground parenchyma (p) and mechanical hypodermis (hy)." figureDoi="http://doi.org/10.5281/zenodo.7905873" httpUri="https://zenodo.org/record/7905873/files/figure.png" pageId="9" pageNumber="178">Fig. 3I</figureCitation>
). The smaller caliber bundles have thicker sclerenchyma fibers at the phloem pole. On the abaxial surface, it is possible to identify spongy parenchyma interspersed with the ground parenchyma. Similarly to
<taxonomicName authorityName="Mez" authorityYear="1896" box="[1251,1402,1419,1444]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="9" pageNumber="178" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis box="[1251,1402,1419,1444]" italics="true" pageId="9" pageNumber="178">D. tomentosa</emphasis>
</taxonomicName>
, the spongy parenchyma in
<taxonomicName authorityName="Büneker &amp; Pastori &amp; Almeida &amp; Mariath" authorityYear="2023" box="[394,539,1454,1480]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="9" pageNumber="178" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis box="[394,539,1454,1480]" italics="true" pageId="9" pageNumber="178">D. magnifica</emphasis>
</taxonomicName>
begins in the region between the vascular bundles and extends to the region of the epidermis, where the stomata and trichomes are located. In
<taxonomicName authorityName="Büneker &amp; Pastori &amp; Almeida &amp; Mariath" authorityYear="2023" box="[811,958,1490,1516]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="9" pageNumber="178" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis box="[811,958,1490,1516]" italics="true" pageId="9" pageNumber="178">D. magnifica</emphasis>
</taxonomicName>
the spongy parenchyma measures 600–800 μm height × 70–120 μm width. The ground parenchyma in this species measures 600–700 μm height × 150–200 μm width, begins in the region of the vascular bundles and extends to the region of the mechanical hypodermis.
</paragraph>
<paragraph blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="178">
<emphasis bold="true" box="[189,428,1598,1624]" pageId="9" pageNumber="178">Pollen morphology:</emphasis>
—The pollen grains of
<taxonomicName authorityName="Büneker &amp; Pastori &amp; Almeida &amp; Mariath" authorityYear="2023" box="[699,848,1598,1624]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="9" pageNumber="178" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis box="[699,848,1598,1624]" italics="true" pageId="9" pageNumber="178">D. magnifica</emphasis>
</taxonomicName>
are oblate, heteropolar, medium in size (25–50 µm) (
<bibRefCitation author="Hesse, M. &amp; Halbritter, H. &amp; Zetter, R. &amp; Weber, M. &amp; Buchner, R. &amp; Frosch-Radivo, A. &amp; Ulrich, S." box="[144,345,1634,1660]" pageId="9" pageNumber="178" refId="ref12957" refString="Hesse, M., Halbritter, H., Zetter, R., Weber, M., Buchner, R., Frosch-Radivo, A. &amp; Ulrich, S. (2009) Pollen Terminology. An illustrated handbook. Springer, Wien, 264 pp." type="book" year="2009">
Hesse
<emphasis box="[220,277,1634,1660]" italics="true" pageId="9" pageNumber="178">et al.</emphasis>
2009
</bibRefCitation>
), and sulcate of the simple sulcus
<typeStatus box="[733,781,1635,1661]" pageId="9" pageNumber="178">type</typeStatus>
(
<bibRefCitation author="Halbritter, H. &amp; Hesse, M." box="[797,1080,1634,1661]" pageId="9" pageNumber="178" pagination="87 - 99" refId="ref12793" refString="Halbritter, H. &amp; Hesse, M. (1993) Sulcus morphology in some monocot families. Grana 32: 87 - 99. https: // doi. org / 10.1080 / 00173139309429459" type="journal article" year="1993">Halbritter &amp; Hesse 1993</bibRefCitation>
). The ornamentation is reticulate to micro-reticulate and heterobrochate (
<bibRefCitation author="Punt, W. &amp; Hoen, P. P. &amp; Blackmore, S. &amp; Nilsson, S. &amp; Thomas, A. Le." box="[585,771,1670,1696]" pageId="9" pageNumber="178" pagination="1 - 81" refId="ref14263" refString="Punt, W., Hoen, P. P., Blackmore, S., Nilsson, S. &amp; Thomas, A. Le. (2007) Glossary of pollen and spore terminology. Review of Paleobotany and Palynology 143: 1 - 81. https: // doi. org / 10.1016 / j. revpalbo. 2006.06.008" type="journal article" year="2007">
Punt
<emphasis box="[647,705,1670,1696]" italics="true" pageId="9" pageNumber="178">et al.</emphasis>
2007
</bibRefCitation>
) (
<figureCitation box="[795,888,1670,1696]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="9" pageNumber="178">Fig. 2O</figureCitation>
). Eventual discontinuities in the reticulum can be observed, specifically in the proximal polar region (
<figureCitation box="[717,802,1706,1732]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="9" pageNumber="178">Fig. 2P</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="178" type="etymology">
<paragraph blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="178">
<emphasis bold="true" box="[189,328,1742,1768]" pageId="9" pageNumber="178">Etymology:</emphasis>
—The specific epithet “
<taxonomicName authorityName="Büneker &amp; Pastori &amp; Almeida &amp; Mariath" authorityYear="2023" box="[599,711,1742,1768]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="9" pageNumber="178" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis box="[599,711,1742,1768]" italics="true" pageId="9" pageNumber="178">magnifica</emphasis>
</taxonomicName>
”, reflects the beauty of the new species. This epithet was created and disseminated by the
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1943" box="[410,489,1778,1804]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="9" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis box="[410,489,1778,1804]" italics="true" pageId="9" pageNumber="178">Dyckia</emphasis>
</taxonomicName>
cultivation enthusiast Constantino Gastaldi, who distributed specimens of it to several collectors and botanical collections under this name.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="178" type="distribution">
<paragraph blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="178">
<emphasis bold="true" box="[189,472,1850,1876]" pageId="9" pageNumber="178">Distribution &amp; habitat:</emphasis>
—The species occurs as a saxicolous heliophyte on rocky outcrops or escarpments of basalt located near rivers. Five populations of the species are known in southern
<collectingCountry box="[1041,1109,1886,1912]" name="Brazil" pageId="9" pageNumber="178">Brazil</collectingCountry>
in the states of
<collectingRegion country="Brazil" name="Rio Grande do Sul" pageId="9" pageNumber="178">Rio Grande do Sul</collectingRegion>
and
<collectingRegion box="[225,388,1922,1948]" country="Brazil" name="Santa Catarina" pageId="9" pageNumber="178">Santa Catarina</collectingRegion>
(
<figureCitation box="[402,471,1922,1948]" captionStart="FIGURE 4" captionStartId="10.[136,229,1356,1378]" captionTargetBox="[309,1278,651,1322]" captionTargetId="figure-248@10.[305,1282,641,1332]" captionTargetPageId="10" captionText="FIGURE 4. Distribution map of Dyckia tomentosa (blue dots) and Dyckia magnifica (red dots) in southern Brazil." figureDoi="http://doi.org/10.5281/zenodo.7905875" httpUri="https://zenodo.org/record/7905875/files/figure.png" pageId="9" pageNumber="178">Fig. 4</figureCitation>
). In
<collectingRegion box="[519,741,1922,1949]" country="Brazil" name="Santa Catarina" pageId="9" pageNumber="178">Santa Catarina state</collectingRegion>
, three of them were found on rocky escarpments on the banks of the Canoas river, in Perau do Macaco Branco (
<figureCitation box="[648,766,1958,1984]" captionStart="FIGURE 1" captionStartId="4.[136,229,1811,1833]" captionTargetBox="[151,1436,187,1783]" captionTargetId="figure-24@4.[151,1436,187,1783]" captionTargetPageId="4" captionText="FIGURE 1. A–E. Dyckia magnifica (H.M. B¸neker 616 et al.) in habitat. A. Detail of part of the population in habitat on rocky banks of the Canoas river dammed by PCH Campos Novos. B. Vegetative habit. C. Habit when fertile. D. Detail of apical portion of the inflorescence. E. Immature capsules." figureDoi="http://doi.org/10.5281/zenodo.7905869" httpUri="https://zenodo.org/record/7905869/files/figure.png" pageId="9" pageNumber="178">Fig. 1A–E</figureCitation>
) and in Perau da Cabrita in the municipality of Abdon Batista, one on a rocky escarpment that forms a waterfall in the municipality of Anita Garibaldi; and another one on rocks along the margins of the Jacutinga river in the municipality of Concórdia. In
<collectingRegion box="[946,1218,2030,2057]" country="Brazil" name="Rio Grande do Sul" pageId="9" pageNumber="178">Rio Grande do sul state</collectingRegion>
, one population was located rocky escarpment in the municipality of Machadinho.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="179" type="discussion">
<paragraph blockId="10.[136,1452,157,616]" pageId="10" pageNumber="179">
<emphasis bold="true" box="[189,357,157,183]" pageId="10" pageNumber="179">Observations:</emphasis>
—
<taxonomicName authorityName="Büneker &amp; Pastori &amp; Almeida &amp; Mariath" authorityYear="2023" box="[385,589,158,184]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="10" pageNumber="179" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis box="[385,589,158,184]" italics="true" pageId="10" pageNumber="179">Dyckia magnifica</emphasis>
</taxonomicName>
is morphologically related to
<taxonomicName authorityName="Mez" authorityYear="1896" box="[950,1104,159,184]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="10" pageNumber="179" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis box="[950,1104,159,184]" italics="true" pageId="10" pageNumber="179">D. tomentosa</emphasis>
</taxonomicName>
, but differs by having patent triangular spines (
<figureCitation box="[342,465,194,220]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2C–D</figureCitation>
) (
<emphasis box="[491,522,195,220]" italics="true" pageId="10" pageNumber="179">vs.</emphasis>
aduncate narrow-triangular (
<figureCitation box="[854,979,194,220]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2A–B</figureCitation>
)); secund flowers (
<figureCitation box="[1200,1286,194,220]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2F</figureCitation>
) (
<emphasis box="[1312,1343,195,220]" italics="true" pageId="10" pageNumber="179">vs.</emphasis>
patent to suberect (
<figureCitation box="[244,327,230,256]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2E</figureCitation>
)) with larger pedicels (
<quantity box="[591,684,230,256]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="10" pageNumber="179" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
(
<figureCitation box="[699,785,230,256]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2H</figureCitation>
)
<emphasis box="[800,830,231,256]" italics="true" pageId="10" pageNumber="179">vs.</emphasis>
lacking or up to
<quantity box="[1017,1102,230,257]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="179" unit="mm" value="0.5">0.5 mm</quantity>
in length (
<figureCitation box="[1221,1309,230,256]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2G</figureCitation>
)); pubescent sepals (
<figureCitation box="[219,306,266,292]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2H</figureCitation>
) (
<emphasis box="[330,361,267,292]" italics="true" pageId="10" pageNumber="179">vs.</emphasis>
tomentose (
<figureCitation box="[497,585,266,292]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2G</figureCitation>
)); longer petals (9–11 (
<figureCitation box="[844,931,266,292]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2H</figureCitation>
)
<emphasis box="[946,976,267,292]" italics="true" pageId="10" pageNumber="179">vs.</emphasis>
<quantity box="[982,1096,266,292]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.6" metricValueMin="6.0" pageId="10" pageNumber="179" unit="mm" value="6.8" valueMax="7.6" valueMin="6.0">6–7.6 mm</quantity>
in length (
<figureCitation box="[1214,1303,266,292]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2G</figureCitation>
)), pubescent, flabelliform trichomes on the abaxial surface with short branches (
<figureCitation box="[870,957,302,328]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2H</figureCitation>
) (
<emphasis box="[980,1011,303,328]" italics="true" pageId="10" pageNumber="179">vs.</emphasis>
dendritic with elongated branches (
<figureCitation captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2G</figureCitation>
)), digitiform trichomes on the margins (
<figureCitation box="[622,709,338,364]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2H</figureCitation>
) (
<emphasis box="[734,757,339,364]" italics="true" pageId="10" pageNumber="179">vs</emphasis>
. filiform (
<figureCitation box="[874,963,338,364]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2G</figureCitation>
)).
</paragraph>
<paragraph blockId="10.[136,1452,157,616]" pageId="10" pageNumber="179">
Differences in rosette size and leaf color can be observed among populations of the species. Specimens from Perau do Macaco Branco and Perau da Cabrita, in Abdon Batista,
<collectingRegion box="[906,1133,410,436]" country="Brazil" name="Santa Catarina" pageId="10" pageNumber="179">Santa Catarina state</collectingRegion>
possess significantly larger rosettes than those in other known populations, and the leaves blades are dark green (
<figureCitation box="[1075,1195,446,472]" captionStart="FIGURE 1" captionStartId="4.[136,229,1811,1833]" captionTargetBox="[151,1436,187,1783]" captionTargetId="figure-24@4.[151,1436,187,1783]" captionTargetPageId="4" captionText="FIGURE 1. A–E. Dyckia magnifica (H.M. B¸neker 616 et al.) in habitat. A. Detail of part of the population in habitat on rocky banks of the Canoas river dammed by PCH Campos Novos. B. Vegetative habit. C. Habit when fertile. D. Detail of apical portion of the inflorescence. E. Immature capsules." figureDoi="http://doi.org/10.5281/zenodo.7905869" httpUri="https://zenodo.org/record/7905869/files/figure.png" pageId="10" pageNumber="179">Fig. 1A–C</figureCitation>
) or rarely pale pinkish. The known population in the municipality of Anita Garibaldi has individuals with smaller rosettes (up to
<emphasis box="[1339,1372,483,508]" italics="true" pageId="10" pageNumber="179">ca.</emphasis>
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in diameter) and leaves with more flexible and pinkish-purple blades (
<figureCitation box="[928,1017,518,544]" captionStart="FIGURE 2" captionStartId="5.[136,229,1810,1832]" captionTargetBox="[151,1436,190,1786]" captionTargetId="figure-19@5.[151,1436,190,1786]" captionTargetPageId="5" captionText="FIGURE 2. A–L. Main diagnostic characteristics differentiating Dyckia tomentosa (A, B, E, G, I, K, M and N) and Dyckia magnifica (C, D, F, H, J, L, O and P). H.M. B¸neker 607 A, B, G and I; H.M. B¸neker 251 et al. E.; H.M. B¸neker 721 &amp; L. Witeck C, D, F, H, K and L. A and C. Detail of adaxial surface of a leaf spine. B and D. Detail of abaxial surface of a leaf spine. E–F. Detail of an inflorescence branch with flowers at various stages of development. G–H. Detailed lateral view of a flower at anthesis. I and J. Detail of trichomes on the surface and margins of petals. K and L. Lateral view of stigma at anthesis. M–P. Pollen under SEM. M. Detail of sulcus where the margin is observed. N. General aspect where general discontinuities in the tectum of the reticulum can be seen. O. Distal polar view where sulcus and phenomenon of harmomegathy are observed. P. Proximal polar view where the region of the tectum of the reticulum with minimal discontinuities can be seen." figureDoi="http://doi.org/10.5281/zenodo.7905871" httpUri="https://zenodo.org/record/7905871/files/figure.png" pageId="10" pageNumber="179">Fig. 2C</figureCitation>
). Specimens from the “barro amarelo” population, in Machadinho,
<collectingRegion box="[458,737,554,580]" country="Brazil" name="Rio Grande do Sul" pageId="10" pageNumber="179">Rio Grande do Sul state</collectingRegion>
, have rosettes intermediate in size between Abdon Batista and Anita Garibaldi populations, and light green bright leaves.
</paragraph>
</subSubSection>
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<emphasis bold="true" box="[136,254,1356,1378]" pageId="10" pageNumber="179">FIGURE 4.</emphasis>
Distribution map of
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<emphasis box="[455,628,1357,1378]" italics="true" pageId="10" pageNumber="179">Dyckia tomentosa</emphasis>
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(blue dots) and
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<emphasis box="[783,953,1357,1378]" italics="true" pageId="10" pageNumber="179">Dyckia magnifica</emphasis>
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(red dots) in southern Brazil.
</paragraph>
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<typeStatus box="[189,573,1425,1451]" isOtherTypeStatus="true" pageId="10" pageNumber="179">Additional specimens examined</typeStatus>
(
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):
</emphasis>
—
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.
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:
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,
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,
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<collectingDate box="[234,427,1461,1487]" pageId="10" pageNumber="179" value="2017-02-08">8 February 2017</collectingDate>
</date>
,
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<emphasis box="[441,465,1462,1487]" italics="true" pageId="10" pageNumber="179">H</emphasis>
</collectionCode>
</materialsCitation>
<emphasis box="[464,871,1461,1487]" italics="true" pageId="10" pageNumber="179">
.
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.
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<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF91FFC02384FA43E020FA65" box="[508,536,1461,1487]" country="Brazil" name="Bu" pageId="10" pageNumber="179" stateProvince="Abdon Batista">Bu</location>
̈neker 616,
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.
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.
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&amp;
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</materialsCitation>
.
<materialsCitation box="[846,867,1461,1487]" collectionCode="C" country="Brazil" isTailEnumeration="true" location="Silva" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Abdon Batista">
<collectionCode box="[846,867,1461,1487]" country="Denmark" name="University of Copenhagen" pageId="10" pageNumber="179" type="Herbarium">C</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation collectingDate="2018-02-08" collectionCode="HDCF, H" country="Brazil" location="Perau da Cabrita" municipality="Pontes" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Abdon Batista">
<emphasis box="[881,957,1462,1487]" italics="true" pageId="10" pageNumber="179">
<collectingMunicipality box="[881,957,1462,1487]" pageId="10" pageNumber="179">Pontes</collectingMunicipality>
</emphasis>
(
<collectionCode box="[974,1055,1461,1487]" pageId="10" pageNumber="179">HDCF</collectionCode>
);
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF91FFC0264DFA43E6C0FA65" box="[1077,1272,1461,1487]" country="Brazil" municipality="Pontes" name="Perau da Cabrita" pageId="10" pageNumber="179" stateProvince="Abdon Batista">Perau da Cabrita</location>
, -27.613359°, - 51.181929°,
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<collectingDate box="[279,466,1497,1523]" pageId="10" pageNumber="179" value="2018-02-08">8 February 2018</collectingDate>
</date>
,
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</collectionCode>
</materialsCitation>
<emphasis box="[500,1045,1497,1523]" italics="true" pageId="10" pageNumber="179">
.
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<collectionCode box="[503,530,1498,1523]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="10" pageNumber="179" type="Herbarium">M</collectionCode>
</materialsCitation>
. B¸neker 718, J.
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</materialsCitation>
.
<materialsCitation box="[737,881,1497,1523]" collectionCode="L" country="Brazil" location="Nardin" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Abdon Batista">
D.
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,
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</materialsCitation>
.
<materialsCitation box="[891,1016,1497,1523]" collectionCode="J" country="Brazil" location="Witeck" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Abdon Batista">
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF91FFC02103FA2FE1FEFA59" box="[891,966,1497,1523]" country="Brazil" name="Witeck" pageId="10" pageNumber="179" stateProvince="Abdon Batista">Witeck</location>
&amp;
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</materialsCitation>
.
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<collectionCode box="[1020,1041,1497,1523]" country="Denmark" name="University of Copenhagen" pageId="10" pageNumber="179" type="Herbarium">C</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation box="[1051,1244,1497,1523]" collectionCode="HDCF" country="Brazil" location="Silva" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Abdon Batista">
<emphasis box="[1051,1141,1497,1523]" italics="true" pageId="10" pageNumber="179">
da
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF91FFC02645FA2FE64DFA59" box="[1085,1141,1497,1523]" country="Brazil" name="Silva" pageId="10" pageNumber="179" stateProvince="Abdon Batista">Silva</location>
</emphasis>
(
<collectionCode box="[1155,1236,1497,1523]" pageId="10" pageNumber="179">HDCF</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2018-02-09" collectionCode="H" country="Brazil" location="Planeta Bromelia" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Anita Garibaldi">
<emphasis bold="true" box="[1255,1445,1497,1523]" pageId="10" pageNumber="179">
<collectingRegion box="[1255,1445,1497,1523]" pageId="10" pageNumber="179">Anita Garibaldi</collectingRegion>
</emphasis>
, -27.632003°, -51.163333°,
<date box="[446,637,1533,1559]" pageId="10" pageNumber="179" value="2018-02-09">
<collectingDate box="[446,637,1533,1559]" pageId="10" pageNumber="179" value="2018-02-09">9 February 2018</collectingDate>
</date>
, floresceu na coleç„o de
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF91FFC021E3FA0BE666F9BD" box="[923,1118,1533,1559]" country="Brazil" name="Planeta Bromelia" pageId="10" pageNumber="179" stateProvince="Anita Garibaldi">Planeta Bromélia</location>
em dezembro de 2019,
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<emphasis box="[1394,1418,1534,1559]" italics="true" pageId="10" pageNumber="179">H</emphasis>
</collectionCode>
</materialsCitation>
<emphasis italics="true" pageId="10" pageNumber="179">
.
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<collectionCode box="[1421,1448,1534,1559]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="10" pageNumber="179" type="Herbarium">M</collectionCode>
</materialsCitation>
. B¸neker 721 &amp; L.
</emphasis>
<materialsCitation box="[350,503,1569,1595]" collectionCode="ICN" country="Brazil" location="Witeck" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Anita Garibaldi">
<emphasis box="[350,425,1569,1595]" italics="true" pageId="10" pageNumber="179">
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF91FFC02326F9D7E391F991" box="[350,425,1569,1595]" country="Brazil" name="Witeck" pageId="10" pageNumber="179" stateProvince="Anita Garibaldi">Witeck</location>
</emphasis>
(
<collectionCode box="[442,496,1569,1595]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="10" pageNumber="179">ICN</collectionCode>
)
</materialsCitation>
;
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<locationName box="[517,643,1568,1595]" pageId="10" pageNumber="179">
<collectingMunicipality box="[517,643,1568,1595]" pageId="10" pageNumber="179">Concórdia</collectingMunicipality>
</locationName>
</emphasis>
,
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,
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<collectingDate box="[829,984,1569,1595]" pageId="10" pageNumber="179" value="2000-05-12">12 May 2000</collectingDate>
</date>
,
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<emphasis box="[997,1016,1570,1595]" italics="true" pageId="10" pageNumber="179">L</emphasis>
</collectionCode>
</materialsCitation>
<emphasis box="[1015,1044,1570,1595]" italics="true" pageId="10" pageNumber="179">
.
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<collectionCode box="[1019,1040,1570,1595]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="10" pageNumber="179" type="Herbarium">E</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation box="[1052,1308,1569,1595]" collectionCode="HBR" country="Brazil" location="Markiewicz" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Concordia">
<emphasis box="[1052,1219,1569,1595]" italics="true" pageId="10" pageNumber="179">
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF91FFC02664F9D7E6A7F991" box="[1052,1183,1569,1595]" country="Brazil" name="Markiewicz" pageId="10" pageNumber="179" stateProvince="Concordia">Markiewicz</location>
41
</emphasis>
(
<collectionCode box="[1236,1299,1569,1595]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14907" lsid="urn:lsid:biocol.org:col:14907" name="Universidade Federal de Santa Catarina" pageId="10" pageNumber="179" type="Herbarium">HBR</collectionCode>
)
</materialsCitation>
.
<materialsCitation collectingDate="2002-04-11" collectingDateMax="2020-02-20" collectingDateMin="2002-04-11" collectionCode="A, C, S, ICN" collectorName="D. Nilson" country="Brazil" location="Jardim Botanico de Porto Alegre" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Rio Grande do Sul">
<collectingRegion country="Brazil" name="Rio Grande do Sul" pageId="10" pageNumber="179">Rio Grande do Sul</collectingRegion>
:
<emphasis bold="true" box="[223,374,1605,1631]" pageId="10" pageNumber="179">Machadinho</emphasis>
,
<date box="[387,544,1605,1631]" pageId="10" pageNumber="179" value="2002-04-11">
<collectingDate box="[387,544,1605,1631]" pageId="10" pageNumber="179" value="2002-04-11">11 April 2002</collectingDate>
</date>
, floresceu na coleç„o do
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF91FFC02140F9B3E69EF9F5" box="[824,1190,1605,1631]" country="Brazil" name="Jardim Botanico de Porto Alegre" pageId="10" pageNumber="179" stateProvince="Rio Grande do Sul">Jardim Botânico de Porto Alegre</location>
em dezembro de 2019,
<emphasis box="[136,456,1641,1667]" italics="true" pageId="10" pageNumber="179">
<collectionCode box="[136,156,1641,1667]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="10" pageNumber="179" type="Herbarium">A</collectionCode>
.
<collectorName box="[160,268,1641,1667]" pageId="10" pageNumber="179">D. Nilson</collectorName>
&amp;
<collectionCode box="[305,327,1641,1667]" country="Denmark" name="University of Copenhagen" pageId="10" pageNumber="179" type="Herbarium">C</collectionCode>
.
<collectionCode box="[331,348,1641,1667]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="10" pageNumber="179" type="Herbarium">S</collectionCode>
. Fior s.n.
</emphasis>
(
<collectionCode box="[472,526,1641,1667]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="10" pageNumber="179">ICN</collectionCode>
); Barro Amarelo, floresceu na coleç„o de Planeta Bromélia em
<date box="[1244,1447,1641,1667]" pageId="10" pageNumber="179" value="2020-02-20">
<collectingDate box="[1244,1447,1641,1667]" pageId="10" pageNumber="179" value="2020-02-20">20 February 2020</collectingDate>
</date>
,
<collectionCode box="[136,159,1677,1703]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="10" pageNumber="179" type="Herbarium">A</collectionCode>
</materialsCitation>
.
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<collectionCode box="[163,185,1677,1703]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="179">R</collectionCode>
</materialsCitation>
.
<materialsCitation box="[196,414,1677,1703]" collectionCode="ICN" country="Brazil" location="Maixner" pageId="10" pageNumber="179" specimenCount="1" stateProvince="Machadinho">
<emphasis box="[196,337,1678,1703]" italics="true" pageId="10" pageNumber="179">
<location LSID="urn:lsid:plazi:treatment:FB6AEA65FF93FFC022F0FA58E39AF90D:761C0DA8FF91FFC022BCF978E31BF90D" box="[196,291,1678,1703]" country="Brazil" name="Maixner" pageId="10" pageNumber="179" stateProvince="Machadinho">Maixner</location>
s.n.
</emphasis>
(
<collectionCode box="[352,406,1677,1703]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="10" pageNumber="179">ICN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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