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<document id="3F1347A91E09BD7272C3BC22EF073A22" ID-DOI="10.37520/aemnp.2023.012" ID-ISSN="1804-6487" ID-Zenodo-Dep="10134165" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699897877524" checkinUser="felipe" docAuthor="Hernando, Carles" docDate="2023" docId="03FD87F98D44FFF2D2B6F8E7FD63E0B0" docLanguage="en" docName="ActaEntMusNatPra.63.1.207-212.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2023.012" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Sunius tricoliensis Hernando 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="209" masterDocId="FFC4FF818D44FFF0D16EFFB7FF9BE205" masterDocTitle="Two new micropterous species of the genus Sunius from Andalusia (Spain) (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="212" masterPageNumber="207" pageNumber="207" updateTime="1700044440017" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="BBA76509D31FBE2515CCBF2793FD4951" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F7BCF1AC300E134081379C619DCEBA5C">
<mods:title id="700D42298134192C025E569B2212DD4A">Two new micropterous species of the genus Sunius from Andalusia (Spain) (Coleoptera: Staphylinidae: Paederinae)</mods:title>
</mods:titleInfo>
<mods:name id="3A37292C34191BAEB7CC1DE4827A4611" type="personal">
<mods:role id="BBD6A43D81DD209C24D745B61FF324D5">
<mods:roleTerm id="0EF2A80AD8575242783500865F63EDB8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="21A9CF28D4CC78F331B5B71AEDFE0C33">Hernando, Carles</mods:namePart>
</mods:name>
<mods:typeOfResource id="9D686F556B4DB668757467693F59154E">text</mods:typeOfResource>
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<mods:title id="8BA0CFD27F6D206898AB526D5F03750D">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="09DC186E9B3DF61087C9737BDD24D9A5">
<mods:date id="4EA009E3285E0DCE2519CEDC0963AF8A">2023</mods:date>
<mods:detail id="C404783C8377B7E2F1B5F1141BECB4DA" type="series">
<mods:title id="BB054D146BA5EA39945FDEB4EC3E6774">Acta. Ent. Mus. Natl. Pragae</mods:title>
</mods:detail>
<mods:detail id="C6470AAF0B7E4025FC8DDBB4FA6EDA71" type="pubDate">
<mods:number id="A3F1DABABD851E12A1A848260D279648">2023-08-31</mods:number>
</mods:detail>
<mods:detail id="E96C115AF1CB64763B642A755125377B" type="volume">
<mods:number id="79F6C65AFA6698584A470CAC0B236EC8">63</mods:number>
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<mods:number id="026BB11EB74B56796FFBB31F36018540">1</mods:number>
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<mods:start id="C2DD90A3DD0ED9A4CD907E6058A9C875">207</mods:start>
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<mods:location id="71A9FE13C75FCE8A9EEEDD91FF71E051">
<mods:url id="DCA1FA5B837592E26A8D74663EC040E5">http://dx.doi.org/10.37520/aemnp.2023.012</mods:url>
</mods:location>
<mods:classification id="FCB4CA9D60836C959D7FA2E9437FC73D">journal article</mods:classification>
<mods:identifier id="EAA65716E44BF4BA66DF896D67C5831F" type="DOI">10.37520/aemnp.2023.012</mods:identifier>
<mods:identifier id="F752D639F2DE51963207CE136424B333" type="ISSN">1804-6487</mods:identifier>
<mods:identifier id="4FD9311D16BBEDAD73C32A64B2043031" type="Zenodo-Dep">10134165</mods:identifier>
<mods:identifier id="804565C1A93DDEE2FCD5F727B740A382" type="ZooBank">16CED11A-F1EA-4637-90AC-1E4E27379D24</mods:identifier>
</mods:mods>
<treatment id="03FD87F98D44FFF2D2B6F8E7FD63E0B0" LSID="urn:lsid:plazi:treatment:03FD87F98D44FFF2D2B6F8E7FD63E0B0" httpUri="http://treatment.plazi.org/id/03FD87F98D44FFF2D2B6F8E7FD63E0B0" lastPageId="2" lastPageNumber="209" pageId="0" pageNumber="207">
<subSubSection id="C34E65648D44FFF0D2B6F8E7FA9BE562" box="[984,1280,1871,1896]" pageId="0" pageNumber="207" type="nomenclature">
<paragraph id="8BEB36EF8D44FFF0D2B6F8E7FA9BE562" blockId="0.[984,1280,1871,1896]" box="[984,1280,1871,1896]" pageId="0" pageNumber="207">
<heading id="D0A381838D44FFF0D2B6F8E7FA9BE562" box="[984,1280,1871,1896]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="207" reason="3">
<emphasis id="B920EAFD8D44FFF0D2B6F8E7FA9BE562" bold="true" box="[984,1280,1871,1896]" pageId="0" pageNumber="207">
<taxonomicName id="4C544D6C8D44FFF0D2B6F8E7FB3FE562" authority="Hernando, 2023" authorityName="Hernando" authorityYear="2023" box="[984,1188,1871,1896]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="tricoliensis" status="sp. nov.">
<emphasis id="B920EAFD8D44FFF0D2B6F8E7FB3FE562" bold="true" box="[984,1188,1871,1896]" italics="true" pageId="0" pageNumber="207">Sunius tricoliensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357868D44FFF0D5C5F8E7FA9BE562" box="[1195,1280,1872,1895]" pageId="0" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E65648D44FFF0D52EF8D8FB03E587" box="[1088,1176,1903,1922]" pageId="0" pageNumber="207" type="description">
<paragraph id="8BEB36EF8D44FFF0D52EF8D8FB03E587" blockId="0.[1088,1176,1903,1922]" box="[1088,1176,1903,1922]" pageId="0" pageNumber="207">
(
<figureCitation id="136F2A6A8D44FFF0D528F8D8FB0AE587" box="[1094,1169,1903,1922]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="0" pageNumber="207">Figs 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E65648D44FFF1D252F81EFEF3E33E" lastPageId="1" lastPageNumber="208" pageId="0" pageNumber="207" type="materials_examined">
<paragraph id="8BEB36EF8D44FFF0D252F81EFA0DE5D2" blockId="0.[828,1436,1961,2007]" pageId="0" pageNumber="207">
<emphasis id="B920EAFD8D44FFF0D252F81EFC29E5B9" bold="true" box="[828,946,1961,1980]" pageId="0" pageNumber="207">
<typeStatus id="54EF884D8D44FFF0D252F81EFCF3E5B9" box="[828,872,1961,1980]" pageId="0" pageNumber="207">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F343767F8D44FFF0D2DBF81EFC7CE5B9" box="[949,999,1961,1980]" name="Spain" pageId="0" pageNumber="207">Spain</collectingCountry>
,
<collectingRegion id="4990F80D8D44FFF0D283F81EFBDEE5B9" box="[1005,1093,1961,1980]" country="Spain" name="Andalucia" pageId="0" pageNumber="207">Andalusia</collectingRegion>
, Granada, Loja,Sierra Gorda,small sinkhole near Pico de Cabras,
<geoCoordinate id="EE6050288D44FFF0D574F873FB01E5D2" box="[1050,1178,1988,2007]" degrees="37" direction="north" minutes="06" orientation="latitude" pageId="0" pageNumber="207" precision="1" seconds="42. 64" value="37.111843">37°06′42.64″N</geoCoordinate>
<geoCoordinate id="EE6050288D44FFF0D5CEF873FA81E5D2" box="[1184,1306,1988,2007]" degrees="4" direction="west" minutes="09" orientation="longitude" pageId="0" pageNumber="207" precision="1" seconds="49. 20" value="-4.1636667">4°09′49.20″W</geoCoordinate>
,
<quantity id="4CAC9B0A8D44FFF0D44DF873FAF2E5D2" box="[1315,1385,1988,2007]" metricMagnitude="3" metricUnit="m" metricValue="1.503" pageId="0" pageNumber="207" unit="m" value="1503.0">1,503 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8BEB36EF8D45FFF1D1F9FF0AFEF3E33E" blockId="1.[151,761,184,963]" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D1F9FF0AFE8CE2D5" bold="true" box="[151,279,189,208]" pageId="1" pageNumber="208">Type material.</emphasis>
<materialsCitation id="3B3C3CB28D45FFF1D074FF0AFE88E325" collectingDate="2022-04-18" collectionCode="MCNB" collectorName="C. Hernando" country="Spain" county="Pico de Cabras" elevation="1503" latitude="37.111843" location="Granada" longLatPrecision="1" longitude="-4.1636667" municipality="Sa Gorda" pageId="1" pageNumber="208" specimenCount="2" typeStatus="holotype">
<typeStatus id="54EF884D8D45FFF1D074FF0AFEE8E2D5" box="[282,371,189,208]" pageId="1" pageNumber="208" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED45AE2A8D45FFF1D0FBFF0AFE4CE2D5" box="[405,471,189,208]" country="Spain" httpUri="http://grbio.org/cool/fs2t-231f" name="Museu de Ciències Naturals de Barcelona" pageId="1" pageNumber="208" type="Museum">MCNB</collectionCode>
), “
<collectingCountry id="F343767F8D45FFF1D085FF0AFDB1E2D5" box="[491,554,189,208]" name="Spain" pageId="1" pageNumber="208">SPAIN</collectingCountry>
,
<location id="8E8B60348D45FFF1D35EFF0AFDE2E2D5" LSID="urn:lsid:plazi:treatment:03FD87F98D44FFF2D2B6F8E7FD63E0B0:8E8B60348D45FFF1D35EFF0AFDE2E2D5" box="[560,633,189,208]" country="Spain" county="Pico de Cabras" latitude="37.111843" longLatPrecision="1" longitude="-4.1636667" municipality="Sa Gorda" name="Granada" pageId="1" pageNumber="208">Granada</location>
,
<location id="8E8B60348D45FFF1D3EEFF0AFD33E2D5" LSID="urn:lsid:plazi:treatment:03FD87F98D44FFF2D2B6F8E7FD63E0B0:8E8B60348D45FFF1D3EEFF0AFD33E2D5" box="[640,680,189,208]" country="Spain" county="Pico de Cabras" latitude="37.111843" longLatPrecision="1" longitude="-4.1636667" municipality="Sa Gorda" name="Loja" pageId="1" pageNumber="208">Loja</location>
,
<collectingMunicipality id="6B8FAC958D45FFF1D3C1FF0AFD63E2D5" box="[687,760,188,208]" pageId="1" pageNumber="208">
<superScript id="7C219BA78D45FFF1D3C1FF0AFD5BE2C2" attach="left" box="[687,704,188,208]" fontSize="5" pageId="1" pageNumber="208">Sa</superScript>
Gorda
</collectingMunicipality>
/ circa
<collectingCounty id="628A4E638D45FFF1D1BFFF6FFEC9E2EE" box="[209,338,216,235]" pageId="1" pageNumber="208">Pico de Cabras</collectingCounty>
,
<quantity id="4CAC9B0A8D45FFF1D034FF6FFE00E2EE" box="[346,411,216,235]" metricMagnitude="3" metricUnit="m" metricValue="1.503" pageId="1" pageNumber="208" unit="m" value="1503.0">
<elevation id="0079D1DC8D45FFF1D034FF6FFE00E2EE" box="[346,411,216,235]" metricMagnitude="3" metricUnit="m" metricValue="1.503" pageId="1" pageNumber="208" unit="m" value="1503.0">1503 m</elevation>
</quantity>
/
<geoCoordinate id="EE6050288D45FFF1D0C5FF6FFDABE2EE" box="[427,560,216,235]" degrees="37" direction="north" minutes="06" orientation="latitude" pageId="1" pageNumber="208" precision="1" seconds="42.64" value="37.111843">37º06’42.64’’N</geoCoordinate>
<geoCoordinate id="EE6050288D45FFF1D35AFF6FFD29E2EE" box="[564,690,216,235]" degrees="4" direction="west" minutes="09" orientation="longitude" pageId="1" pageNumber="208" precision="1" seconds="49.20" value="-4.1636667">4º09’49.20’’W</geoCoordinate>
/ Under stones buried deep in the soil /
<date id="FFEA102F8D45FFF1D0CDFF44FD98E303" box="[419,515,243,262]" pageId="1" pageNumber="208" value="2022-04-18">
<collectingDate id="EFAEE9C78D45FFF1D0CDFF44FD98E303" box="[419,515,243,262]" pageId="1" pageNumber="208" value="2022-04-18">18.IV.2022</collectingDate>
</date>
,
<collectorName id="26A153398D45FFF1D362FF44FDE2E303" box="[524,633,243,262]" pageId="1" pageNumber="208">C. Hernando</collectorName>
leg.”, plus red
<typeStatus id="54EF884D8D45FFF1D1F9FEBAFF79E325" box="[151,226,269,288]" pageId="1" pageNumber="208" type="holotype">holotype</typeStatus>
label
</materialsCitation>
.
<typeStatus id="54EF884D8D45FFF1D074FEBAFEE8E325" box="[282,371,269,288]" pageId="1" pageNumber="208" type="paratype">PARATYPES</typeStatus>
: 4 JJ
<specimenCount id="9D52FD668D45FFF1D0DCFEBAFE7BE325" box="[434,480,263,288]" count="5" pageId="1" pageNumber="208" type="female">5 ♀♀</specimenCount>
(
<collectionCode id="ED45AE2A8D45FFF1D085FEBAFDB2E325" box="[491,553,269,288]" pageId="1" pageNumber="208">CHCB</collectionCode>
), same data as
<typeStatus id="54EF884D8D45FFF1D3C6FEBAFD6EE325" box="[680,757,269,288]" pageId="1" pageNumber="208" type="holotype">holotype</typeStatus>
, plus red
<typeStatus id="54EF884D8D45FFF1D18FFE9FFEB1E33E" box="[225,298,296,315]" pageId="1" pageNumber="208" type="paratype">paratype</typeStatus>
labels.
</paragraph>
</subSubSection>
<subSubSection id="C34E65648D45FFF1D1F9FEFBFB92E010" pageId="1" pageNumber="208" type="description">
<paragraph id="8BEB36EF8D45FFF1D1F9FEFBFE84E386" blockId="1.[151,761,184,963]" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D1F9FEFBFEF7E366" bold="true" box="[151,364,331,355]" pageId="1" pageNumber="208">
Description.
<emphasis id="B920EAFD8D45FFF1D043FEFCFEF7E366" box="[301,364,331,355]" italics="true" pageId="1" pageNumber="208">Male.</emphasis>
</emphasis>
Habitus as in
<figureCitation id="136F2A6A8D45FFF1D378FEFBFDC3E366" box="[534,600,332,355]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208">Fig. 1</figureCitation>
. Body length:
<quantity id="4CAC9B0A8D45FFF1D1F9FEDBFE80E386" box="[151,283,364,387]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="1" pageNumber="208" unit="mm" value="3.3" valueMax="3.5" valueMin="3.1">3.1–3.5 mm</quantity>
.
</paragraph>
<paragraph id="8BEB36EF8D45FFF1D1DBFE3BFD63E3C6" blockId="1.[151,761,184,963]" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D1DBFE3BFEA9E3A6" box="[181,306,396,419]" italics="true" pageId="1" pageNumber="208">Colouration</emphasis>
. Forebody uniformly reddish brown (
<figureCitation id="136F2A6A8D45FFF1D3C3FE3BFD71E3A6" box="[685,746,396,419]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208">Fig. 2</figureCitation>
); abdomen dark brown; legs and antennae yellowish brown.
</paragraph>
<paragraph id="8BEB36EF8D45FFF1D1DBFE7BFEE1E066" blockId="1.[151,761,184,963]" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D1DBFE7BFF74E3E6" box="[181,239,460,483]" italics="true" pageId="1" pageNumber="208">Head</emphasis>
slightly longer than wide (0.41/
<quantity id="4CAC9B0A8D45FFF1D33DFE7BFD21E3E6" box="[595,698,460,483]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="208" unit="mm" value="0.39">0.39 mm</quantity>
), distinctly dilated posteriorly; with coarse, sparse punctation; lacking microsculpture; length of eyes less than half of length of postocular region in dorsal view (
<figureCitation id="136F2A6A8D45FFF1D334FD9BFD01E046" box="[602,666,556,579]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208">Fig. 2</figureCitation>
). Length of antennae:
<quantity id="4CAC9B0A8D45FFF1D04FFDFBFEEDE066" box="[289,374,588,611]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="208" unit="mm" value="0.8">0.8 mm</quantity>
.
</paragraph>
<paragraph id="8BEB36EF8D45FFF1D1DBFDDBFE69E106" blockId="1.[151,761,184,963]" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D1DBFDDBFE84E086" box="[181,287,620,643]" italics="true" pageId="1" pageNumber="208">Pronotum</emphasis>
as long as head and slightly narrower than head (
<quantity id="4CAC9B0A8D45FFF1D1B7FD3BFEA4E0A6" box="[217,319,652,675]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="1" pageNumber="208" unit="mm" value="0.36">0.36 mm</quantity>
), slightly oblong, evenly narrowing posteriad, with posterior margin narrower than anterior one; punctation less impressed than that of head, but denser (
<figureCitation id="136F2A6A8D45FFF1D1F1FD5BFF7AE106" box="[159,225,748,771]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208">Fig. 2</figureCitation>
); lacking microsculpture.
</paragraph>
<paragraph id="8BEB36EF8D45FFF1D1DBFCBBFD30E186" blockId="1.[151,761,184,963]" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D1DBFCBBFF63E126" box="[181,248,780,803]" italics="true" pageId="1" pageNumber="208">Elytra</emphasis>
as wide (
<quantity id="4CAC9B0A8D45FFF1D00BFCBBFE51E126" box="[357,458,780,803]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="208" unit="mm" value="0.35">0.35 mm</quantity>
) and distinctly shorter than pronotum (
<quantity id="4CAC9B0A8D45FFF1D07DFC9BFEE5E146" box="[275,382,812,835]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="208" unit="mm" value="0.37">0.37 mm</quantity>
); punctation somewhat finer than pronotum, dense and weakly defined, surface between punctures somewhat rugose. Hind wings reduced.
</paragraph>
<paragraph id="8BEB36EF8D45FFF1D1DBFC3BFA1CE330" blockId="1.[151,761,184,963]" lastBlockId="1.[828,1437,190,949]" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D1DBFC3BFE82E1A6" box="[181,281,908,931]" italics="true" pageId="1" pageNumber="208">Abdomen</emphasis>
about 1.1 times as wide as elytra, widest at segment VI; punctation fine and moderately dense; surface with shallow microsculpture; posterior margin of tergite VII without palisade fringe. Sternite VII with weakly concave posterior margin, pubescence unmodified; sternite VIII with broad U-shaped incision at its posterior margin (
<figureCitation id="136F2A6A8D45FFF1D454FEA9FAE1E330" box="[1338,1402,286,309]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BEB36EF8D45FFF1D234FE89FB70E3D0" blockId="1.[828,1437,190,949]" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D234FE89FC58E350" box="[858,963,318,341]" italics="true" pageId="1" pageNumber="208">Aedeagus</emphasis>
(
<figureCitation id="136F2A6A8D45FFF1D2BBFE89FBA2E350" box="[981,1081,318,341]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208">Figs 4–5</figureCitation>
) with ventral process distinctly longer than basal part of aedeagus; endophallus with small series of stout sclerotized spines on basal lobe (
<figureCitation id="136F2A6A8D45FFF1D456FEC9FAE3E390" box="[1336,1400,382,405]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208">Fig. 4</figureCitation>
) in lateral view, with two internal preapical structures with falciform apices (
<figureCitation id="136F2A6A8D45FFF1D299FE09FBACE3D0" box="[1015,1079,446,469]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208">Fig. 5</figureCitation>
) in ventral view.
</paragraph>
<paragraph id="8BEB36EF8D45FFF1D234FE6AFB92E010" blockId="1.[828,1437,190,949]" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D234FE6AFC28E3F0" bold="true" box="[858,947,477,501]" italics="true" pageId="1" pageNumber="208">Female.</emphasis>
General appearance as male, except for unmodified sternite VIII.
</paragraph>
</subSubSection>
<subSubSection id="C34E65648D45FFF2D252FDA9FEE3E390" lastPageId="2" lastPageNumber="209" pageId="1" pageNumber="208" type="diagnosis">
<paragraph id="8BEB36EF8D45FFF2D252FDA9FEE3E390" blockId="1.[828,1437,190,949]" lastBlockId="2.[151,760,190,693]" lastPageId="2" lastPageNumber="209" pageId="1" pageNumber="208">
<emphasis id="B920EAFD8D45FFF1D252FDA9FBADE030" bold="true" box="[828,1078,542,565]" pageId="1" pageNumber="208">Differential diagnosis.</emphasis>
Using the key of
<bibRefCitation id="EFC54B1E8D45FFF1D594FDA9FA03E030" author="ASSING V." box="[1274,1432,542,565]" pageId="1" pageNumber="208" pagination="151 - 193" refId="ref3340" refString="ASSING V. 2011: The Sunius species of the Palaeartic region (Coleoptera: Staphylinidae: Paederinae). Linzer Biologische Beitrage 43: 151 - 193." type="journal article" year="2011">ASSINǤ (2011)</bibRefCitation>
, this species would key out at couplet 85 together with the Betic-Rifean species with spines on the endophallus; however, it is easily separated from all of them by a very small and poorly developed group of spines (
<figureCitation id="136F2A6A8D45FFF1D465FD29FAD1E0B0" box="[1291,1354,670,693]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208">Fig. 4</figureCitation>
). Based on the general shape of the aedeagus,
<taxonomicName id="4C544D6C8D45FFF1D58FFD09FAEEE0D0" authority="Hernando, 2023" authorityName="Hernando" authorityYear="2023" box="[1249,1397,702,725]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="species" species="tricoliensis" status="sp. nov.">
<emphasis id="B920EAFD8D45FFF1D58FFD09FAEEE0D0" box="[1249,1397,702,725]" italics="true" pageId="1" pageNumber="208">S. tricoliensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357868D45FFF1D410FD09FCF7E0F0" pageId="1" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
, is quite similar to
<taxonomicName id="4C544D6C8D45FFF1D55BFD69FAEBE0F0" authority="(Coiffait, 1980)" baseAuthorityName="Coiffait" baseAuthorityYear="1980" box="[1077,1392,734,757]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="species" species="nevadensis">
<emphasis id="B920EAFD8D45FFF1D55BFD69FB5FE0F0" box="[1077,1220,734,757]" italics="true" pageId="1" pageNumber="208">S. nevadensis</emphasis>
(Coiffait, 1980)
</taxonomicName>
and
<taxonomicName id="4C544D6C8D45FFF1D252FD49FBD0E110" authority="Assing, 2008" authorityName="Assing" authorityYear="2008" box="[828,1099,766,789]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="species" species="tronqueti">
<emphasis id="B920EAFD8D45FFF1D252FD49FC2CE110" box="[828,951,766,789]" italics="true" pageId="1" pageNumber="208">S. tronqueti</emphasis>
Assing, 2008
</taxonomicName>
, both from Sierra Nevada, from which it is distinguished by the different shape of the aedeagus and the structures of the endophallus. For illustrations of
<taxonomicName id="4C544D6C8D45FFF1D234FCE9FC77E170" baseAuthorityName="Coiffait" baseAuthorityYear="1980" box="[858,1004,862,885]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="species" species="nevadensis">
<emphasis id="B920EAFD8D45FFF1D234FCE9FCFCE170" box="[858,871,862,885]" italics="true" pageId="1" pageNumber="208">S</emphasis>
.
<emphasis id="B920EAFD8D45FFF1D219FCE9FC77E170" box="[887,1004,862,885]" italics="true" pageId="1" pageNumber="208">nevadensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D6C8D45FFF1D54DFCE9FB3BE170" authorityName="Assing" authorityYear="2008" box="[1059,1184,862,885]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="species" species="tronqueti">
<emphasis id="B920EAFD8D45FFF1D54DFCE9FBABE170" box="[1059,1072,862,885]" italics="true" pageId="1" pageNumber="208">S</emphasis>
.
<emphasis id="B920EAFD8D45FFF1D551FCE9FB3BE170" box="[1087,1184,862,885]" italics="true" pageId="1" pageNumber="208">tronqueti</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFC54B1E8D45FFF1D5BFFCE9FAE8E170" author="ASSING V." box="[1233,1395,862,885]" pageId="1" pageNumber="208" pagination="5 - 135" refId="ref3266" refString="ASSING V. 2008 a: A revision of the Sunius species of the Western Palaeartic region and Middle Asia (Coleoptera: Staphylinidae: Paederinae). Linzer Biologische Beitrage 40: 5 - 135." type="journal article" year="2008">ASSINǤ (2008a</bibRefCitation>
, b). Although apparently not closely related, it differs from the sympatric species described below, which lives at lower altitudes, in having smaller eyes, pronotum with the posterior margin clearly narrower than the anterior one, and in a very different shape of the aedeagus (
<figureCitation id="136F2A6A8D46FFF2D324FF49FD32E310" box="[586,681,254,277]" captionStart="Figs 1–5" captionStartId="1.[153,190,2059,2078]" captionTargetBox="[184,1436,996,2031]" captionTargetId="figure-450@1.[151,1436,996,2031]" captionTargetPageId="1" captionText="Figs 1–5.Sunius tricoliensis sp. nov.J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134167" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="2" pageNumber="209">Figs 4–5</figureCitation>
,
<figureCitation id="136F2A6A8D46FFF2D3DDFF49FD77E310" box="[691,748,254,277]" captionStart="Figs 6 –10" captionStartId="2.[151,187,2059,2078]" captionTargetBox="[151,1436,1022,2030]" captionTargetId="figure-427@2.[151,1436,1021,2030]" captionTargetPageId="2" captionText="Figs 6 –10.Sunius lauxaensis sp. nov.,J. 6 – habitus in dorsal view; 7 – forebody;8 –sternite VIII; 9 – aedeagus in lateral view; 10 – aedeagus in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10134169" httpUri="https://zenodo.org/record/10134169/files/figure.png" pageId="2" pageNumber="209">9–10</figureCitation>
).
<emphasis id="B920EAFD8D46FFF2D1F9FEA9FE83E330" bold="true" box="[151,280,286,309]" pageId="2" pageNumber="209">Etymology.</emphasis>
In reference to Tricolia, the name given to the city of Loja in Roman times due to its settlement on three hills, in whose municipal district the
<typeStatus id="54EF884D8D46FFF2D372FEE9FDD1E370" box="[540,586,350,373]" pageId="2" pageNumber="209">type</typeStatus>
locality lies; the name is an adjective.
</paragraph>
</subSubSection>
<caption id="DF2B66678D45FFF1D1F7F7BCFA05EA1B" ID-DOI="http://doi.org/10.5281/zenodo.10134167" ID-Zenodo-Dep="10134167" box="[153,1438,2053,2078]" httpUri="https://zenodo.org/record/10134167/files/figure.png" pageId="1" pageNumber="208" startId="1.[153,190,2059,2078]" targetBox="[184,1436,996,2031]" targetPageId="1" targetType="figure">
<paragraph id="8BEB36EF8D45FFF1D1F7F7BCFA05EA1B" blockId="1.[153,1438,2053,2078]" box="[153,1438,2053,2078]" pageId="1" pageNumber="208">
Figs 1–5.
<taxonomicName id="4C544D6C8D45FFF1D187F7BCFE18EA1B" authority="Hernando, 2023" authorityName="Hernando" authorityYear="2023" box="[233,387,2059,2078]" class="Insecta" family="Staphylinidae" genus="Sunius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="208" phylum="Arthropoda" rank="species" species="tricoliensis" status="sp. nov.">
<emphasis id="B920EAFD8D45FFF1D187F7BCFE18EA1B" box="[233,387,2059,2078]" italics="true" pageId="1" pageNumber="208">Sunius tricoliensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357868D45FFF1D0E9F7BCFE52EA1B" box="[391,457,2059,2078]" pageId="1" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
J. 1– habitus in dorsal view; 2 – forebody; 3 – sternite VIII; 4 – aedeagus in lateral view; 5 – aedeagus in ventral view.
</paragraph>
</caption>
<subSubSection id="C34E65648D46FFF2D1F9FE29FD63E0B0" pageId="2" pageNumber="209" type="distribution">
<paragraph id="8BEB36EF8D46FFF2D1F9FE29FD63E0B0" blockId="2.[151,760,190,693]" pageId="2" pageNumber="209">
<emphasis id="B920EAFD8D46FFF2D1F9FE29FE7EE3B0" bold="true" box="[151,485,414,437]" pageId="2" pageNumber="209">Distribution and bionomics.</emphasis>
The known distribution is limited to the high plateaus near Pico de Cabras, the second-highest altitude in the Sierra Gorda de Loja.
<typeStatus id="54EF884D8D46FFF2D3ADFE69FD63E3F0" box="[707,760,478,501]" pageId="2" pageNumber="209">Type</typeStatus>
specimens were collected under stones buried deep in the soil in a doline with herbaceous vegetation and tufts of
<taxonomicName id="4C544D6C8D46FFF2D1F9FD89FEA7E050" box="[151,316,574,597]" class="Liliopsida" family="Asphodelaceae" genus="Asphodelus" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="209" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B920EAFD8D46FFF2D1F9FD89FE89E050" box="[151,274,574,597]" italics="true" pageId="2" pageNumber="209">Asphodelus</emphasis>
sp.
</taxonomicName>
, at an altitude above
<quantity id="4CAC9B0A8D46FFF2D340FD89FD13E050" box="[558,648,574,597]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="209" unit="m" value="1500.0">1,500 m</quantity>
(
<figureCitation id="136F2A6A8D46FFF2D3F4FD89FD71E050" box="[666,746,574,597]" captionStart="Figs 11–12" captionStartId="3.[151,188,2079,2098]" captionTargetBox="[166,1421,178,2067]" captionTargetPageId="3" captionText="Figs 11–12. Habitat of Sunius in Sierra Gorda de Loja. 11 – type locality of Sunius tricoliensis sp. nov.; 12 – type locality of Sunius lauxaensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10134171" httpUri="https://zenodo.org/record/10134171/files/figure.png" pageId="2" pageNumber="209">Fig. 11</figureCitation>
), together with
<taxonomicName id="4C544D6C8D46FFF2D049FDE9FF54E090" authority="Pace, 1983" authorityName="Pace" authorityYear="1983" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="species" species="baetica" subGenus="Trachyglutosipalia">
<emphasis id="B920EAFD8D46FFF2D049FDE9FE1CE070" box="[295,391,606,629]" italics="true" pageId="2" pageNumber="209">Geostiba</emphasis>
(
<emphasis id="B920EAFD8D46FFF2D0FBFDE9FDC5E070" box="[405,606,606,629]" italics="true" pageId="2" pageNumber="209">Trachyglutosipalia</emphasis>
)
<emphasis id="B920EAFD8D46FFF2D302FDE9FD21E070" box="[620,698,606,629]" italics="true" pageId="2" pageNumber="209">baetica</emphasis>
Pace, 1983
</taxonomicName>
,
<taxonomicName id="4C544D6C8D46FFF2D1B6FDC9FE1BE090" box="[216,384,638,661]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B920EAFD8D46FFF2D1B6FDC9FEC7E090" box="[216,348,638,661]" italics="true" pageId="2" pageNumber="209">Scydmaenus</emphasis>
sp.
</taxonomicName>
(both
<taxonomicName id="4C544D6C8D46FFF2D0ACFDC9FDC2E090" box="[450,601,638,661]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
), and the endogean ant
<taxonomicName id="4C544D6C8D46FFF2D19AFD29FD6FE0B0" authority="(Baroni Urbani, 1978)" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[244,756,670,693]" class="Insecta" family="Formicidae" genus="Stigmatomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="species" species="gaetulicum">
<emphasis id="B920EAFD8D46FFF2D19AFD29FD99E0B0" box="[244,514,670,693]" italics="true" pageId="2" pageNumber="209">Stigmatomma gaetulicum</emphasis>
(Baroni Urbani, 1978)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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