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<document id="D5935E440AC45CBE702B60814403C41D" ID-CLB-Dataset="296270" ID-DOI="10.5281/zenodo.11205315" ID-GBIF-Dataset="58503a0e-2eb4-4e69-b274-99a538163ce2" ID-ISSN="2189-6720" ID-Zenodo-Dep="11205315" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1714415367943" checkinUser="felipe" docAuthor="SUƶU, Toshiyuki &amp; Kƚ, Seishi" docDate="2019" docId="03FD87E1A915FFA1FC0DFCADFC06BFBC" docLanguage="en" docName="BullKanagawaPrefectMus.48.21-36.pdf" docOrigin="Bulletin of the Kanagawa Prefectural Museum (Natural Science) 48" docStyle="DocumentStyle:020B98DC1285EBBD02620724ECB44EDE.1:BullKanagawaPrefectMus.2013-.journal_article.english" docStyleId="020B98DC1285EBBD02620724ECB44EDE" docStyleName="BullKanagawaPrefectMus.2013-.journal_article.english" docStyleVersion="1" docTitle="Rhinogobius Gill 1859" docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="FFC4FF99A914FFA3FFAFFFA6FFF5BC2A" masterDocTitle="Two New Lentic, Dwarf Species of Rhinogobius Gill, 1859 (Gobiidae) from Japan" masterLastPageNumber="36" masterPageNumber="21" pageNumber="22" updateTime="1715884386113" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2DA8DC7055616D01C0E6C43D959087F1">Two New Lentic, Dwarf Species of Rhinogobius Gill, 1859 (Gobiidae) from Japan</mods:title>
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<mods:namePart id="0880FA3522178AA087C8A5314D3A644E">SUƶU, Toshiyuki</mods:namePart>
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<mods:namePart id="C994CB40503310CBD7D033BB280577E5">Kƚ, Seishi</mods:namePart>
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<paragraph id="8BEB36F7A915FFA2FC0DFCADFB0ABF09" blockId="1.[930,1279,777,803]" box="[930,1279,777,803]" pageId="1" pageNumber="22">
<heading id="D0A3819BA915FFA2FC0DFCADFB0ABF09" box="[930,1279,777,803]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="22" reason="2">
Genus
<taxonomicName id="4C544D74A915FFA2FC40FCAFFB0ABF09" ID-CoL="78QB" ID-ENA="63457" authority="Gill, 1859" authorityName="Gill" authorityYear="1859" box="[1007,1279,777,803]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A915FFA2FC40FCAFFB7FBF09" bold="true" box="[1007,1162,777,803]" italics="true" pageId="1" pageNumber="22">Rhinogobius</emphasis>
<bibRefCitation id="EFC54B06A915FFA2FB3EFCADFB0ABF09" author="Gill, T. N." box="[1169,1279,779,803]" pageId="1" pageNumber="22" pagination="144 - 150" refId="ref11567" refString="Gill, T. N., 1859. Notes on a collection of Japanese fishes, made by Dr. J. Morrow. Proceedings of the Academy of Natural Sciences of Philadelphia, 11: 144 - 150." type="journal article" year="1859">Gill, 1859</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E657CA915FFA2FC83FCF2FC3BB83C" pageId="1" pageNumber="22" type="reference_group">
<paragraph id="8BEB36F7A915FFA2FC83FCF2FC35BF85" blockId="1.[812,1398,852,1284]" pageId="1" pageNumber="22">
<treatmentCitationGroup id="AB4411D9A915FFA2FC83FCF2FC35BF85" pageId="1" pageNumber="22">
<treatmentCitation id="0AF510E6A915FFA2FC83FCF2FBA3BF41" author="Gill, T. N." box="[812,1110,852,875]" page="145" pageId="1" pageNumber="22" year="1859">
<taxonomicName id="4C544D74A915FFA2FC83FCF2FBA3BF41" ID-CoL="78QB" authority="Gill, 1859: 145" authorityName="Gill" authorityPageNumber="145" authorityYear="1859" box="[812,1110,852,875]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A915FFA2FC83FCF2FC45BF41" box="[812,944,852,875]" italics="true" pageId="1" pageNumber="22">Rhinogobius</emphasis>
<bibRefCitation id="EFC54B06A915FFA2FC18FCF2FBA3BF41" author="Gill, T. N." box="[951,1110,852,875]" pageId="1" pageNumber="22" pagination="144 - 150" refId="ref11567" refString="Gill, T. N., 1859. Notes on a collection of Japanese fishes, made by Dr. J. Morrow. Proceedings of the Academy of Natural Sciences of Philadelphia, 11: 144 - 150." type="journal article" year="1859">Gill, 1859: 145</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54EF8855A915FFA2FBCBFCF2FB66BF41" box="[1124,1171,852,875]" pageId="1" pageNumber="22">type</typeStatus>
species:
<taxonomicName id="4C544D74A915FFA2FB5DFCF2FBEEBFA7" ID-CoL="4S92Y" authority="Gill, 1859" authorityName="Gill" authorityYear="1859" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="similis">
<emphasis id="B920EAE5A915FFA2FB5DFCF2FC66BFA7" italics="true" pageId="1" pageNumber="22">Rhinogobius similis</emphasis>
<bibRefCitation id="EFC54B06A915FFA2FC0FFCD0FBEEBFA7" author="Gill, T. N." box="[928,1051,886,909]" pageId="1" pageNumber="22" pagination="144 - 150" refId="ref11567" refString="Gill, T. N., 1859. Notes on a collection of Japanese fishes, made by Dr. J. Morrow. Proceedings of the Academy of Natural Sciences of Philadelphia, 11: 144 - 150." type="journal article" year="1859">Gill, 1859</bibRefCitation>
</taxonomicName>
by original designation and monotypy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEB36F7A915FFA2FC83FC1CFC3BB83C" blockId="1.[812,1398,852,1284]" pageId="1" pageNumber="22">
<emphasis id="B920EAE5A915FFA2FC83FC1CFC34BFFB" box="[812,961,954,977]" italics="true" pageId="1" pageNumber="22">Tu k u g o b i u s</emphasis>
Herre, 1927: 119 (
<typeStatus id="54EF8855A915FFA2FB67FC1CFAF4BFFB" box="[1224,1281,954,977]" pageId="1" pageNumber="22">type</typeStatus>
species:
<taxonomicName id="4C544D74A915FFA2FCEBFC7AFB19BFD9" ID-CoL="4S8YM" authority="Seale, 1910" authorityName="Seale" authorityYear="1910" box="[836,1260,988,1011]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B920EAE5A915FFA2FCEBFC7AFBADBFD9" box="[836,1112,988,1011]" italics="true" pageId="1" pageNumber="22">Rhinogobius carpenteri</emphasis>
Seale, 1910
</taxonomicName>
by original designation).
</paragraph>
</subSubSection>
<subSubSection id="C34E657CA915FFA2FC83FB87FB98B8B4" pageId="1" pageNumber="22" type="description">
<paragraph id="8BEB36F7A915FFA2FC83FB87FB98B8B4" blockId="1.[812,1398,852,1284]" pageId="1" pageNumber="22">
<taxonomicName id="4C544D74A915FFA2FC83FB87FB9FB812" authority="Liu, 1940: 215" authorityName="Liu" authorityPageNumber="215" authorityYear="1940" box="[812,1130,1057,1080]" family="Gobiidae" genus="Sinogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A915FFA2FC83FB87FC5BB812" box="[812,942,1057,1080]" italics="true" pageId="1" pageNumber="22">Sinogobius</emphasis>
Liu, 1940: 215
</taxonomicName>
[
<typeStatus id="54EF8855A915FFA2FBD3FB87FB44B812" box="[1148,1201,1057,1080]" pageId="1" pageNumber="22">type</typeStatus>
species:
<taxonomicName id="4C544D74A915FFA2FA8CFB87FA80B812" authorityName="Linnaeus" authorityYear="1758" box="[1315,1397,1057,1080]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A915FFA2FA8CFB87FA80B812" box="[1315,1397,1057,1080]" italics="true" pageId="1" pageNumber="22">Gobius</emphasis>
</taxonomicName>
(
<emphasis id="B920EAE5A915FFA2FCFFFBE5FC16B870" box="[848,995,1091,1114]" italics="true" pageId="1" pageNumber="22">S i n o g o b i u s</emphasis>
)
<emphasis id="B920EAE5A915FFA2FC50FBE5FB49B870" box="[1023,1212,1091,1114]" italics="true" pageId="1" pageNumber="22">s z e c h u a n e n s i s</emphasis>
Liu, 1940 (=
<taxonomicName id="4C544D74A915FFA2FCEBFBC3FB13B856" authority="Chen &amp; Wu, 2008" authorityName="Chen &amp; Wu" authorityYear="2008" box="[836,1254,1125,1148]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="liui">
<emphasis id="B920EAE5A915FFA2FCEBFBC3FBF0B856" box="[836,1029,1125,1148]" italics="true" pageId="1" pageNumber="22">Rhinogobius liui</emphasis>
Chen &amp; Wu, 2008
</taxonomicName>
) by original designation and monotypy].
</paragraph>
</subSubSection>
<subSubSection id="C34E657CA915FFA2FC83FB0FFAD7B92E" pageId="1" pageNumber="22" type="reference_group">
<paragraph id="8BEB36F7A915FFA2FC83FB0FFAD7B92E" blockId="1.[812,1398,852,1284]" pageId="1" pageNumber="22">
<treatmentCitationGroup id="AB4411D9A915FFA2FC83FB0FFAD7B92E" pageId="1" pageNumber="22">
<treatmentCitation id="0AF510E6A915FFA2FC83FB0FFAC4B8EA" author="Zhong, J. - S. &amp; H. - L. Wu" box="[812,1329,1193,1216]" page="149" pageId="1" pageNumber="22" year="1998">
<taxonomicName id="4C544D74A915FFA2FC83FB0FFAC4B8EA" ID-CoL="73CS" authority="Zhong &amp; Wu, 1998: 149" authorityName="Zhong &amp; Wu" authorityPageNumber="149" authorityYear="1998" box="[812,1329,1193,1216]" family="Gobiidae" genus="Pseudorhinogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A915FFA2FC83FB0FFBFCB8EA" box="[812,1033,1193,1216]" italics="true" pageId="1" pageNumber="22">Pseudorhinogobius</emphasis>
<bibRefCitation id="EFC54B06A915FFA2FBBBFB0FFAC4B8EA" author="Zhong, J. - S. &amp; H. - L. Wu" box="[1044,1329,1193,1216]" pageId="1" pageNumber="22" pagination="148 - 153" refId="ref13297" refString="Zhong, J. - S. &amp; H. - L. Wu, 1998. Pseudorhinogobius aporus, a new genus and species of gobiid fish from eastern China. Journal of Fisheries of China, 22 (2): 148 - 153. (In Chinese)." type="journal article" year="1998">Zhong &amp; Wu, 1998: 149</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54EF8855A915FFA2FAEBFB0FFA80B8EA" box="[1348,1397,1193,1216]" pageId="1" pageNumber="22">type</typeStatus>
species:
<taxonomicName id="4C544D74A915FFA2FC08FB6DFC8CB92E" ID-CoL="4P9QD" authority="Zhong &amp; Wu 1998" authorityName="Zhong &amp; Wu" authorityYear="1998" family="Gobiidae" genus="Pseudorhinogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="aporus">
<emphasis id="B920EAE5A915FFA2FC08FB6DFB23B8C8" box="[935,1238,1227,1250]" italics="true" pageId="1" pageNumber="22">Pseudorhinogobius aporus</emphasis>
<bibRefCitation id="EFC54B06A915FFA2FB4EFB6DFC8CB92E" author="Zhong, J. - S. &amp; H. - L. Wu" pageId="1" pageNumber="22" pagination="148 - 153" refId="ref13297" refString="Zhong, J. - S. &amp; H. - L. Wu, 1998. Pseudorhinogobius aporus, a new genus and species of gobiid fish from eastern China. Journal of Fisheries of China, 22 (2): 148 - 153. (In Chinese)." type="journal article" year="1998">Zhong &amp; Wu 1998</bibRefCitation>
</taxonomicName>
by original designation and monotypy).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E657CA915FFA1FCEBFA9DFDC5BC99" lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22" type="diagnosis">
<paragraph id="8BEB36F7A915FFA1FCEBFA9DFDC5BC99" blockId="1.[812,1398,1337,2102]" lastBlockId="2.[188,775,156,2086]" lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22">
<emphasis id="B920EAE5A915FFA2FCEBFA9DFC34B978" bold="true" box="[836,961,1339,1362]" pageId="1" pageNumber="22">Diagnosis.</emphasis>
<taxonomicName id="4C544D74A915FFA2FC77FA9FFB83B979" authorityName="Gill" authorityYear="1859" box="[984,1142,1337,1363]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A915FFA2FC77FA9FFB83B979" box="[984,1142,1337,1363]" italics="true" pageId="1" pageNumber="22">Rhinogobius</emphasis>
</taxonomicName>
belongs to the gobiid subfamily
<taxonomicName id="4C544D74A915FFA2FC36FAC4FBD2B953" box="[921,1063,1378,1401]" pageId="1" pageNumber="22" rank="subFamily" subFamily="Gobionellinae">Gobionellinae</taxonomicName>
(sensu
<bibRefCitation id="EFC54B06A915FFA2FBD9FAC4FB0BB953" author="Pezold, F. L." box="[1142,1278,1378,1401]" pageId="1" pageNumber="22" pagination="634 - 643" refId="ref12062" refString="Pezold, F. L., 1993. Evidence for a monophyletic Gobiinae. Copeia, 1993: 634 - 643." type="journal article" year="1993">Pezold, 1993</bibRefCitation>
,
<bibRefCitation id="EFC54B06A915FFA2FAA6FAC4FACBB953" author="Pezold, F. L." box="[1289,1342,1378,1401]" pageId="1" pageNumber="22" pagination="87 - 97" refId="ref12085" refString="Pezold, F. L., 2011. Systematics of Gobionellidae, Chapter 1.6. In Patzner, R. A., J. L. van Tassel, M. Kovacic &amp; B. G. Kapoor (eds.), The biology of gobies, pp. 87 - 97. CRC Press, Taylor and Francis Group, Science Publishers, Enfield, New Hampshire." type="book chapter" year="2011">2011</bibRefCitation>
), and is distinguished from the other gobionelline genera by the following combination of characters (
<bibRefCitation id="EFC54B06A915FFA2FB0AFA16FA86B9ED" author="Chen, I-S. &amp; K. - T. Shao" box="[1189,1395,1456,1479]" pageId="1" pageNumber="22" pagination="200 - 214" refId="ref11402" refString="Chen, I-S. &amp; K. - T. Shao, 1996. A taxonomic review of the gobiid fish genus Rhinogobius Gill, 1859, from Taiwan, with description of three new species. Zoological Studies, 35: 200 - 214." type="journal article" year="1996">Chen &amp; Shao, 1996</bibRefCitation>
;
<bibRefCitation id="EFC54B06A915FFA2FC83FA71FC11B9C4" author="Yang, J. - Q. &amp; H. - L. Wu &amp; I. - S. Chen" box="[812,996,1493,1519]" pageId="1" pageNumber="22" pagination="379 - 385" refId="ref13244" refString="Yang, J. - Q., H. - L. Wu &amp; I. - S. Chen, 2008. A new species of Rhinogobius (Teleostei: Gobiidae) from the Feiyunjiang Basin in Zhejiang Province, China. Ichthyological Research, 55: 379 - 385." type="journal article" year="2008">
Yang
<emphasis id="B920EAE5A915FFA2FCC8FA73FC55B9C5" box="[871,928,1493,1519]" italics="true" pageId="1" pageNumber="22">et al.</emphasis>
, 2008
</bibRefCitation>
;
<bibRefCitation id="EFC54B06A915FFA2FC40FA71FB42B9C4" author="Suzuki, T. &amp; K. Shibukawa &amp; H. Senou &amp; I-S. Chen" box="[1007,1207,1493,1519]" pageId="1" pageNumber="22" refId="ref12699" refString="Suzuki, T., K. Shibukawa, H. Senou &amp; I-S. Chen, 2015. Redescription of Rhinogobius similis Gill 1859 (Gobiidae: Gobionellinae), the type species of the genus Rhinogobius Gill 1859, with designation of the neotype. Ichthyological Research, DOI 10.1007 / s 10228 - 015 - 0494 - 3." type="book" year="2015">
Suzuki
<emphasis id="B920EAE5A915FFA2FB94FA73FB81B9C5" box="[1083,1140,1493,1519]" italics="true" pageId="1" pageNumber="22">et al.</emphasis>
, 2015
</bibRefCitation>
, 2017): first dorsal fin with 5–7 spines; second dorsal fin with a single spine and 6–11 segmented rays; anal fin with a single spine and 5–11 segmented rays; pectoral fin with 14–23 segmented rays; pelvic fin with a single spine and five segmented rays; 25–44 longitudinal scales; 7–16 transverse scales; P-V 3/2211 0/9 or, in a few exceptions, its derived pattern; 10– 11+15–18=25–29 vertebrae; snout, cheek and operculum naked; body largely with ctenoid scales; cheek with a longitudinal pattern of sensory papillae (sensu
<bibRefCitation id="EFC54B06A915FFA2FA82F893FC90BB59" author="Hoese, D. F." pageId="1" pageNumber="22" pagination="223 - 230" refId="ref11608" refString="Hoese, D. F., 1983. Sensory papilla patterns of the cheek lateralis system in the gobiid fishes Acentrogobius and Glossogobius, and their significance for the classification of gobioid fishes. Records of the Australian Museum, 35: 223 - 230." type="journal article" year="1983">Hoese, 1983</bibRefCitation>
), except for a single species,
<taxonomicName id="4C544D74A915FFA2FB02F8FCFAE8BB5E" authorityName="Gill" authorityYear="1859" box="[1197,1309,1882,1908]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="similis">
<emphasis id="B920EAE5A915FFA2FB02F8FCFAE8BB5E" box="[1197,1309,1882,1908]" italics="true" pageId="1" pageNumber="22">R. similis</emphasis>
</taxonomicName>
, having several short transverse rows of sensory papillae below the eye; gill opening moderate in size, its anteroventral end extending to a vertical through posterior margin of preopercle; pelvic fins fused medially into a circular/ ovoid disc via frenum (between spines) and connecting membrane (between innermost rays).
</paragraph>
</subSubSection>
<subSubSection id="C34E657CA916FFA1FF7AFF65FC06BFBC" pageId="2" pageNumber="23" type="discussion">
<paragraph id="8BEB36F7A916FFA1FF7AFF65FEADBF08" blockId="2.[188,775,156,2086]" pageId="2" pageNumber="23">
<emphasis id="B920EAE5A916FFA1FF7AFF65FEC8BCF0" bold="true" box="[213,317,195,218]" pageId="2" pageNumber="23">Remarks.</emphasis>
<taxonomicName id="4C544D74A916FFA1FEE8FF67FE27BCF1" authorityName="Gill" authorityYear="1859" box="[327,466,193,219]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FEE8FF67FE27BCF1" box="[327,466,193,219]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
is currently known as the most species-rich freshwater gobiid genus, comprising 83 valid species (Suzuki
<emphasis id="B920EAE5A916FFA1FECFFEA9FE6DBD03" box="[352,408,271,297]" italics="true" pageId="2" pageNumber="23">et al.</emphasis>
, 2017;
<bibRefCitation id="EFC54B06A916FFA1FE4AFEB7FCF6BD02" author="Takahashi, S. &amp; T. Okazaki" box="[485,771,273,296]" pageId="2" pageNumber="23" refId="ref12811" refString="Takahashi, S. &amp; T. Okazaki, 2017. Rhinogobius biwaensis, a new gobiid fish of the &quot; yoshinobori &quot; species complex, Rhinogobius spp., endemic to Lake Biwa, Japan. Ichthyological Research, DOI 10.1007 / s 10228 - 017 - 0577 - 4." type="book" year="2017">Takahashi &amp; Okazaki, 2017</bibRefCitation>
;
<bibRefCitation id="EFC54B06A916FFA1FF12FE91FE9DBD64" author="Endruweit, M." box="[189,360,311,334]" pageId="2" pageNumber="23" pagination="284 - 310" refId="ref11488" refString="Endruweit, M., 2018. Description of four new species of freshwater gobies from the Black River drainage in China and Vietnam (Teleostei: Gobiidae). Zootaxa, 4486 (3): 284 - 310." type="journal article" year="2018">Endruweit, 2018</bibRefCitation>
;
<bibRefCitation id="EFC54B06A916FFA1FEDCFE91FDFEBD64" author="Li, F. &amp; S. Li &amp; J. - K. Chen" box="[371,523,310,336]" pageId="2" pageNumber="23" pagination="1 - 10" refId="ref11820" refString="Li, F., S. Li &amp; J. - K. Chen, 2018. Rhinogobius immaculatus, a new species of freshwater goby (Teleostei: Gobiidae) from the Qiantang River, China. Zoological Research, 39 (6): 1 - 10." type="journal article" year="2018">
Li
<emphasis id="B920EAE5A916FFA1FE20FE90FE3DBD7A" box="[399,456,310,336]" italics="true" pageId="2" pageNumber="23">et al.</emphasis>
, 2018
</bibRefCitation>
;
<bibRefCitation id="EFC54B06A916FFA1FDBAFE91FD49BD64" author="Wu, Q. &amp; X. Deng &amp; Y. Wang &amp; Y. Liu" box="[533,700,310,336]" pageId="2" pageNumber="23" pagination="118 - 129" refId="ref13074" refString="Wu, Q., X. Deng, Y. Wang &amp; Y. Liu, 2018. Rhinogobius maculagenys, A new species of freshwater goby (Teleostei: Gobiidae) from Hunan, China. Zootaxa, 4476 (1): 118 - 129." type="journal article" year="2018">
Wu
<emphasis id="B920EAE5A916FFA1FDEFFE90FD8CBD7A" box="[576,633,310,336]" italics="true" pageId="2" pageNumber="23">et al.</emphasis>
, 2018
</bibRefCitation>
;
<bibRefCitation id="EFC54B06A916FFA1FD69FE91FEE8BD5F" author="Xia J. - H. &amp; H. - L. Wu &amp; C. - H. Li &amp; Y. - Q. Wu &amp; S. - H. Liu" pageId="2" pageNumber="23" pagination="553 - 562" refId="ref13123" refString="Xia J. - H., H. - L. Wu, C. - H. Li, Y. - Q. Wu &amp; S. - H. Liu, 2018. A new species of Rhinogobius (Pisces: Gobiidae), with analyses of its DNA barcode. Zootaxa, 4407 (4): 553 - 562." type="journal article" year="2018">
Xia
<emphasis id="B920EAE5A916FFA1FD5EFE90FF2CBD5D" italics="true" pageId="2" pageNumber="23">et al.</emphasis>
, 2018
</bibRefCitation>
; present study). As indicated by
<bibRefCitation id="EFC54B06A916FFA1FDD6FEF8FF0ABDB6" author="Chen, I-S. &amp; K. - T. Shao" pageId="2" pageNumber="23" pagination="200 - 214" refId="ref11402" refString="Chen, I-S. &amp; K. - T. Shao, 1996. A taxonomic review of the gobiid fish genus Rhinogobius Gill, 1859, from Taiwan, with description of three new species. Zoological Studies, 35: 200 - 214." type="journal article" year="1996">Chen &amp; Shao (1996)</bibRefCitation>
and
<bibRefCitation id="EFC54B06A916FFA1FE9DFE23FDF6BDB6" author="Suzuki, T. &amp; K. Shibukawa &amp; H. Senou &amp; I-S. Chen" box="[306,515,388,414]" pageId="2" pageNumber="23" refId="ref12699" refString="Suzuki, T., K. Shibukawa, H. Senou &amp; I-S. Chen, 2015. Redescription of Rhinogobius similis Gill 1859 (Gobiidae: Gobionellinae), the type species of the genus Rhinogobius Gill 1859, with designation of the neotype. Ichthyological Research, DOI 10.1007 / s 10228 - 015 - 0494 - 3." type="book" year="2015">
Suzuki
<emphasis id="B920EAE5A916FFA1FED1FE22FE42BDB4" box="[382,439,388,414]" italics="true" pageId="2" pageNumber="23">et al.</emphasis>
(2015)
</bibRefCitation>
, the genus is divided into two distinct groups; one comprises only a single species
<taxonomicName id="4C544D74A916FFA1FF13FE77FEDDBDC1" authorityName="Gill" authorityYear="1859" box="[188,296,465,491]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="similis">
<emphasis id="B920EAE5A916FFA1FF13FE77FEDDBDC1" box="[188,296,465,491]" italics="true" pageId="2" pageNumber="23">R. similis</emphasis>
</taxonomicName>
, whereas the other includes all the remaining species.
<taxonomicName id="4C544D74A916FFA1FEB3FE5EFDF8BE38" authorityName="Gill" authorityYear="1859" box="[284,525,504,530]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="similis">
<emphasis id="B920EAE5A916FFA1FEB3FE5EFDF8BE38" box="[284,525,504,530]" italics="true" pageId="2" pageNumber="23">Rhinogobius similis</emphasis>
</taxonomicName>
differs from the other congeners by having large ctenoid scales on the nape (vs. nape naked or with cycloid scales in the others) and several short transverse rows of sensory papillae on the cheek (vs. no distinct transverse rows of sensory papillae on cheek). We here assign all species of the genus but
<taxonomicName id="4C544D74A916FFA1FD2EFD1DFD1FBEFF" authorityName="Gill" authorityYear="1859" box="[641,746,699,725]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="similis">
<emphasis id="B920EAE5A916FFA1FD2EFD1DFD1FBEFF" box="[641,746,699,725]" italics="true" pageId="2" pageNumber="23">R. similis</emphasis>
</taxonomicName>
to the “
<taxonomicName id="4C544D74A916FFA1FF40FD44FE18BED6" authorityName="Temminck &amp; Schlegel" authorityYear="1845" box="[239,493,738,764]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="brunneus">
<emphasis id="B920EAE5A916FFA1FF40FD44FE18BED6" box="[239,493,738,764]" italics="true" pageId="2" pageNumber="23">Rhinogobius brunneus</emphasis>
</taxonomicName>
complex”, following
<bibRefCitation id="EFC54B06A916FFA1FD7FFD42FEA0BF08" author="Chen, I-S. &amp; K. - T. Shao" pageId="2" pageNumber="23" pagination="200 - 214" refId="ref11402" refString="Chen, I-S. &amp; K. - T. Shao, 1996. A taxonomic review of the gobiid fish genus Rhinogobius Gill, 1859, from Taiwan, with description of three new species. Zoological Studies, 35: 200 - 214." type="journal article" year="1996">Chen &amp; Shao (1996)</bibRefCitation>
.
</paragraph>
<paragraph id="8BEB36F7A916FFA1FF7AFC94FC06BFBC" blockId="2.[188,775,156,2086]" lastBlockId="2.[850,1437,154,920]" pageId="2" pageNumber="23">
<bibRefCitation id="EFC54B06A916FFA1FF7AFC94FE72BF63" author="Endruweit, M." box="[213,391,818,841]" pageId="2" pageNumber="23" pagination="549 - 560" refId="ref11448" refString="Endruweit, M., 2017. Neotype designation for Rhinogobius carpenteri Seale, 1910, and its placement in Tukugobius Herre 1927 (Teleostei: Gobiidae). Zootaxa, 4277 (4): 549 - 560." type="journal article" year="2017">Endruweit (2017)</bibRefCitation>
resurrected
<taxonomicName id="4C544D74A916FFA1FDAEFC96FCF6BF63" authority="Herre, 1927" authorityName="Herre" authorityYear="1927" box="[513,771,816,842]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FDAEFC96FD75BF60" box="[513,640,816,842]" italics="true" pageId="2" pageNumber="23">Tukugobius</emphasis>
Herre, 1927
</taxonomicName>
, previously regarded as a junior synonym of
<taxonomicName id="4C544D74A916FFA1FDDBFCF1FD0ABF5B" authorityName="Gill" authorityYear="1859" box="[628,767,855,881]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FDDBFCF1FD0ABF5B" box="[628,767,855,881]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
, as valid; assigning three described species known from the
<collectingCountry id="F3437667A916FFA1FF4BFC00FEAEBF97" box="[228,347,934,957]" name="Philippines" pageId="2" pageNumber="23">Philippines</collectingCountry>
,
<taxonomicName id="4C544D74A916FFA1FEC9FC03FCF7BF97" authority="Herre, 1927" authorityName="Herre" authorityYear="1927" box="[358,770,933,959]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="bucculentus">
<emphasis id="B920EAE5A916FFA1FEC9FC03FD8EBF95" box="[358,635,933,959]" italics="true" pageId="2" pageNumber="23">Tukugobius bucculentus</emphasis>
Herre, 1927
</taxonomicName>
,
<taxonomicName id="4C544D74A916FFA1FF13FC6AFDA3BFCE" authority="Seale, 1910" authorityName="Seale" authorityYear="1910" box="[188,598,972,998]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B920EAE5A916FFA1FF13FC6AFE25BFCC" box="[188,464,972,998]" italics="true" pageId="2" pageNumber="23">Rhinogobius carpenteri</emphasis>
Seale, 1910
</taxonomicName>
(
<typeStatus id="54EF8855A916FFA1FDCAFC6BFD61BFCE" box="[613,660,973,996]" pageId="2" pageNumber="23">type</typeStatus>
species of
<taxonomicName id="4C544D74A916FFA1FF13FC54FECEB826" authorityName="Herre" authorityYear="1927" box="[188,315,1010,1036]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FF13FC54FECEB826" box="[188,315,1010,1036]" italics="true" pageId="2" pageNumber="23">Tukugobius</emphasis>
</taxonomicName>
), and
<taxonomicName id="4C544D74A916FFA1FED3FC54FCF6B821" authority="Herre, 1927" authorityName="Herre" authorityYear="1927" box="[380,771,1010,1036]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="philippinus">
<emphasis id="B920EAE5A916FFA1FED3FC54FD8AB826" box="[380,639,1010,1036]" italics="true" pageId="2" pageNumber="23">Tukugobius philippinus</emphasis>
Herre, 1927
</taxonomicName>
, to
<taxonomicName id="4C544D74A916FFA1FF7AFBBFFEA1B819" authorityName="Herre" authorityYear="1927" box="[213,340,1049,1075]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FF7AFBBFFEA1B819" box="[213,340,1049,1075]" italics="true" pageId="2" pageNumber="23">Tukugobius</emphasis>
</taxonomicName>
, with all others to
<taxonomicName id="4C544D74A916FFA1FDA2FBBFFD6DB819" authorityName="Gill" authorityYear="1859" box="[525,664,1049,1075]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FDA2FBBFFD6DB819" box="[525,664,1049,1075]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
. However, on the basis of the following reasons, we do not concur with his conclusion. First of all, it is too early to resurrect
<taxonomicName id="4C544D74A916FFA1FF12FB28FECAB882" authorityName="Herre" authorityYear="1927" box="[189,319,1166,1192]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FF12FB28FECAB882" box="[189,319,1166,1192]" italics="true" pageId="2" pageNumber="23">Tukugobius</emphasis>
</taxonomicName>
as valid, based on these three species. Like
<taxonomicName id="4C544D74A916FFA1FF13FB13FEADB8E5" authorityName="Seale" authorityYear="1910" box="[188,344,1205,1231]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B920EAE5A916FFA1FF13FB13FEADB8E5" box="[188,344,1205,1231]" italics="true" pageId="2" pageNumber="23">R. carpenteri</emphasis>
</taxonomicName>
, the
<typeStatus id="54EF8855A916FFA1FE3EFB11FE4AB8E4" box="[401,447,1207,1230]" pageId="2" pageNumber="23">type</typeStatus>
specimens of
<taxonomicName id="4C544D74A916FFA1FDF2FB13FCF3B8E5" authorityName="Herre" authorityYear="1927" box="[605,774,1205,1231]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="bucculentus">
<emphasis id="B920EAE5A916FFA1FDF2FB13FCF3B8E5" box="[605,774,1205,1231]" italics="true" pageId="2" pageNumber="23">T. bucculentus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D74A916FFA1FF44FB7AFE78B8DC" authorityName="Herre" authorityYear="1927" box="[235,397,1244,1270]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="philippinus">
<emphasis id="B920EAE5A916FFA1FF44FB7AFE78B8DC" box="[235,397,1244,1270]" italics="true" pageId="2" pageNumber="23">T. philippinus</emphasis>
</taxonomicName>
were destroyed during the Second World War (
<bibRefCitation id="EFC54B06A916FFA1FEEFFAA3FE09B936" author="Eschmeyer, W. N." box="[320,508,1285,1308]" pageId="2" pageNumber="23" refId="ref11529" refString="Eschmeyer, W. N., 1998. Catalog of Fishes. Center for Biodiversity Research and Information, Special Publication 1. 3 volumes. 2905 pp. California Academy of Sciences, San Francisco." type="book" year="1998">Eschmeyer, 1998</bibRefCitation>
;
<bibRefCitation id="EFC54B06A916FFA1FDA6FAA3FD51B936" author="Kottelat, M." box="[521,676,1285,1308]" pageId="2" pageNumber="23" pagination="1 - 663" refId="ref11770" refString="Kottelat, M., 2013. The fishes of the inland waters of southeast Asia: a catalogue and core bibiography of the fishes known to occur in freshwaters, mangroves and estuaries. Raffles Bulletin of Zoology, Supplement, (27): 1 - 663." type="journal article" year="2013">Kottelat, 2013</bibRefCitation>
), and the taxonomic status of these species is open to debate. For these two species,
<bibRefCitation id="EFC54B06A916FFA1FE27FAF5FDB1B940" author="Endruweit, M." box="[392,580,1363,1386]" pageId="2" pageNumber="23" pagination="549 - 560" refId="ref11448" refString="Endruweit, M., 2017. Neotype designation for Rhinogobius carpenteri Seale, 1910, and its placement in Tukugobius Herre 1927 (Teleostei: Gobiidae). Zootaxa, 4277 (4): 549 - 560." type="journal article" year="2017">Endruweit (2017)</bibRefCitation>
merely examined the non-type specimens from the
<collectingCountry id="F3437667A916FFA1FD9EFADCFD5EB9BB" box="[561,683,1402,1425]" name="Philippines" pageId="2" pageNumber="23">Philippines</collectingCountry>
and did not state the reasons why these were congeneric with
<taxonomicName id="4C544D74A916FFA1FD42FA39FEC2B9CA" authorityName="Seale" authorityYear="1910" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B920EAE5A916FFA1FD42FA39FEC2B9CA" italics="true" pageId="2" pageNumber="23">R. carpenteri</emphasis>
</taxonomicName>
, only stating, “
<taxonomicName id="4C544D74A916FFA1FE4EFA60FD19B9CA" authorityName="Seale" authorityYear="1910" box="[481,748,1478,1504]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B920EAE5A916FFA1FE4EFA60FD19B9CA" box="[481,748,1478,1504]" italics="true" pageId="2" pageNumber="23">Tukugobius carpenteri</emphasis>
</taxonomicName>
is readily distinguished from all species currently allocated in
<taxonomicName id="4C544D74A916FFA1FF13F9B2FEBDBA04" authorityName="Gill" authorityYear="1859" box="[188,328,1556,1582]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FF13F9B2FEBDBA04" box="[188,328,1556,1582]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
by seven pterygiophores supporting the first dorsal fin (vs.
<quantity id="4CAC9B12A916FFA1FEE4F99AFE87BA79" box="[331,370,1596,1619]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="23" unit="in" value="6.0">6 in</quantity>
all species from continental Asia,
<collectingCountry id="F3437667A916FFA1FD68F99AFCF6BA79" box="[711,771,1596,1619]" name="Japan" pageId="2" pageNumber="23">Japan</collectingCountry>
,
<collectingCountry id="F3437667A916FFA1FF12F9C5FEF9BA50" box="[189,268,1635,1658]" name="Taiwan" pageId="2" pageNumber="23">Taiwan</collectingCountry>
, and Hainan), the first pterygiophore of second dorsal fin inserted in interneural space 9 (vs. 8)”, These characters need further investigation based on broader sampling. For example, some specimens of
<taxonomicName id="4C544D74A916FFA1FDD5F970FF2EBB3C" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B920EAE5A916FFA1FDD5F970FCF3BADA" box="[634,774,1750,1776]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
sp.
</taxonomicName>
BF (one of the new species described below) and
<taxonomicName id="4C544D74A916FFA1FF12F882FE84BB17" box="[189,369,1828,1854]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B920EAE5A916FFA1FF12F882FEB9BB14" box="[189,332,1828,1854]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
sp.
</taxonomicName>
<collectionCode id="ED45AE32A916FFA1FED6F880FE69BB17" box="[377,412,1830,1853]" country="0" name="Buffalo Bill Museum" pageId="2" pageNumber="23">BB</collectionCode>
have seven pterygiophores for the first dorsal fin, and
<taxonomicName id="4C544D74A916FFA1FE21F8EDFDEABB4F" baseAuthorityName="Mizuno" baseAuthorityYear="1960" box="[398,543,1867,1893]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="fluminues">
<emphasis id="B920EAE5A916FFA1FE21F8EDFDEABB4F" box="[398,543,1867,1893]" italics="true" pageId="2" pageNumber="23">R. fluminues</emphasis>
</taxonomicName>
, and some specimens of
<taxonomicName id="4C544D74A916FFA1FF75F8D4FE64BBA1" box="[218,401,1906,1932]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B920EAE5A916FFA1FF75F8D4FE9FBBA6" box="[218,362,1906,1932]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
sp.
</taxonomicName>
YB have the first pterygiophore of second dorsal fin inserting behind neural spine of ninth vertebra (TS, personal investigation). These inter- or intra-specific variations suggest that the characters shown by
<bibRefCitation id="EFC54B06A916FFA1FF4EF7A9FE56B40C" author="Endruweit, M." box="[225,419,2063,2086]" pageId="2" pageNumber="23" pagination="549 - 560" refId="ref11448" refString="Endruweit, M., 2017. Neotype designation for Rhinogobius carpenteri Seale, 1910, and its placement in Tukugobius Herre 1927 (Teleostei: Gobiidae). Zootaxa, 4277 (4): 549 - 560." type="journal article" year="2017">Endruweit (2017)</bibRefCitation>
are not sufficient to distinguish
<taxonomicName id="4C544D74A916FFA1FCFDFF3CFC21BC9E" authorityName="Herre" authorityYear="1927" box="[850,980,154,180]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FCFDFF3CFC21BC9E" box="[850,980,154,180]" italics="true" pageId="2" pageNumber="23">Tukugobius</emphasis>
</taxonomicName>
from
<taxonomicName id="4C544D74A916FFA1FBBAFF3AFB63BC99" authorityName="Gill" authorityYear="1859" box="[1045,1174,156,179]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FBBAFF3AFB63BC99" box="[1045,1174,156,179]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
. Furthermore,
<bibRefCitation id="EFC54B06A916FFA1FA9EFF3AFC6DBCF0" author="Endruweit, M." pageId="2" pageNumber="23" pagination="549 - 560" refId="ref11448" refString="Endruweit, M., 2017. Neotype designation for Rhinogobius carpenteri Seale, 1910, and its placement in Tukugobius Herre 1927 (Teleostei: Gobiidae). Zootaxa, 4277 (4): 549 - 560." type="journal article" year="2017">Endruweit (2017)</bibRefCitation>
indicated that some differentiations between
<taxonomicName id="4C544D74A916FFA1FA2CFF67FC30BD28" authorityName="Seale" authorityYear="1910" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="carpenteri">
<emphasis id="B920EAE5A916FFA1FA2CFF67FC30BD28" italics="true" pageId="2" pageNumber="23">R. carpenteri</emphasis>
</taxonomicName>
(
<typeStatus id="54EF8855A916FFA1FC7BFF4CFBF4BD2B" box="[980,1025,234,257]" pageId="2" pageNumber="23">type</typeStatus>
species of
<taxonomicName id="4C544D74A916FFA1FBDAFF4EFB03BD28" authorityName="Herre" authorityYear="1927" box="[1141,1270,232,258]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FBDAFF4EFB03BD28" box="[1141,1270,232,258]" italics="true" pageId="2" pageNumber="23">Tukugobius</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C544D74A916FFA1FA9DFF4EFA6EBD28" authorityName="Gill" authorityYear="1859" box="[1330,1435,232,258]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="similis">
<emphasis id="B920EAE5A916FFA1FA9DFF4EFA6EBD28" box="[1330,1435,232,258]" italics="true" pageId="2" pageNumber="23">R. similis</emphasis>
</taxonomicName>
(
<typeStatus id="54EF8855A916FFA1FCF4FEB7FC7CBD02" box="[859,905,273,296]" pageId="2" pageNumber="23">type</typeStatus>
species of
<taxonomicName id="4C544D74A916FFA1FC53FEA9FB7FBD03" authorityName="Gill" authorityYear="1859" box="[1020,1162,271,297]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FC53FEA9FB7FBD03" box="[1020,1162,271,297]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
), such as configuration of sensory-papillae rows on the cheek (i.e., transverse rows absent vs. present) and squamation of pectoral-fin base (i.e., naked vs. scaly), “may possess intrinsic value at generic level.” Nevertheless, in the gobioid fishes, more than a few examples are known to have these characters as intrageneric variations (see, e.g.,
<bibRefCitation id="EFC54B06A916FFA1FB10FE5CFA64BE3B" author="Akihito &amp; K. Sakamoto &amp; Y. Ikeda &amp; M. Aizawa" box="[1215,1425,504,530]" pageId="2" pageNumber="23" pagination="2109 - 2211" refId="ref11293" refString="Akihito, K. Sakamoto, Y. Ikeda &amp; M. Aizawa, 2013. Gobioidei. In Nakabo, T. (ed.), Fishes of Japan with pictorial keys to the species, 3 rd edition. pp. 1347 - 1608, 2109 - 2211. Tokai University Press, Kanagawa. (In Japanese)." type="journal article" year="2013">
Akihito
<emphasis id="B920EAE5A916FFA1FABCFE5EFAB8BE38" box="[1299,1357,504,530]" italics="true" pageId="2" pageNumber="23">et al.</emphasis>
, 2013
</bibRefCitation>
). And, as noted above, the presence of the transverse rows of sensory papillae does not only distinguish
<taxonomicName id="4C544D74A916FFA1FCFCFDCBFC36BEAD" authorityName="Gill" authorityYear="1859" box="[851,963,621,647]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="similis">
<emphasis id="B920EAE5A916FFA1FCFCFDCBFC36BEAD" box="[851,963,621,647]" italics="true" pageId="2" pageNumber="23">R. similis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C544D74A916FFA1FBA7FDCBFB65BEAD" authorityName="Herre" authorityYear="1927" box="[1032,1168,621,647]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FBA7FDCBFB65BEAD" box="[1032,1168,621,647]" italics="true" pageId="2" pageNumber="23">Tukugobius</emphasis>
</taxonomicName>
, but also from the other species of
<taxonomicName id="4C544D74A916FFA1FC6AFD32FBA2BE84" authorityName="Gill" authorityYear="1859" box="[965,1111,660,686]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FC6AFD32FBA2BE84" box="[965,1111,660,686]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
; if following his scenario, we should split further his
<taxonomicName id="4C544D74A916FFA1FBEBFD1DFB26BEFF" authorityName="Gill" authorityYear="1859" box="[1092,1235,699,725]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FBEBFD1DFB26BEFF" box="[1092,1235,699,725]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
into two genera. It appears to be merely artificial, less-necessary splitting. We, therefore, do not accept his idea on the limits of
<taxonomicName id="4C544D74A916FFA1FCFDFC96FC2BBF60" authorityName="Gill" authorityYear="1859" box="[850,990,816,842]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FCFDFC96FC2BBF60" box="[850,990,816,842]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C544D74A916FFA1FC4AFC96FB90BF60" authorityName="Herre" authorityYear="1927" box="[997,1125,816,842]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FC4AFC96FB90BF60" box="[997,1125,816,842]" italics="true" pageId="2" pageNumber="23">Tukugobius</emphasis>
</taxonomicName>
, and, in this paper, regard the latter (
<taxonomicName id="4C544D74A916FFA1FC37FCF1FBE9BF5B" authorityName="Herre" authorityYear="1927" box="[920,1052,855,881]" family="Gobiidae" genus="Tukugobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FC37FCF1FBE9BF5B" box="[920,1052,855,881]" italics="true" pageId="2" pageNumber="23">Tukugobius</emphasis>
</taxonomicName>
) as a junior synonym of the former (
<taxonomicName id="4C544D74A916FFA1FCF5FCD8FC10BFB2" authorityName="Gill" authorityYear="1859" box="[858,997,894,920]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B920EAE5A916FFA1FCF5FCD8FC10BFB2" box="[858,997,894,920]" italics="true" pageId="2" pageNumber="23">Rhinogobius</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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