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<document id="DE546C254BA7C486BF831870DAEBB051" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F102FFF37640FDC355A47939" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) elegantulus Cameron 1939" docType="treatment" docVersion="12" lastPageNumber="228" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="227" updateTime="1699471357828" updateUser="ExternalLinkService">
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<mods:title id="EAB10AF0B45A1218023369B5840A7EE8">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="33545E259CB5B0846A92E4BC41F9AD2C">Assing, Volker</mods:namePart>
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<mods:title id="E4D64FA900C0A4727F50EE2FE8BAD9C7">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="65E7DCD52345564EBBDB0760828E3C00">
<mods:date id="389794BA10F599A658F1FCCA4B396CCB">2017</mods:date>
<mods:detail id="4069FF09273F41E24FA39FA5357B89AB" type="pubDate">
<mods:number id="FA3FC5684C7C36FF8618B71824A69A2D">2017-12-08</mods:number>
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<mods:number id="EEEF3FEFD3DAD1354584393466919FC7">67</mods:number>
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<mods:identifier id="646B5CA7F9BBC2D15726DB658B435DA1" type="DOI">10.21248/contrib.entomol.67.2.213-246</mods:identifier>
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<treatment id="03FD87A1F102FFF37640FDC355A47939" ID-DOI="http://doi.org/10.5281/zenodo.5888792" ID-GBIF-Taxon="192500468" ID-Zenodo-Dep="5888792" LSID="urn:lsid:plazi:treatment:03FD87A1F102FFF37640FDC355A47939" httpUri="http://treatment.plazi.org/id/03FD87A1F102FFF37640FDC355A47939" lastPageId="15" lastPageNumber="228" pageId="14" pageNumber="227">
<subSubSection id="C34E653CF102FFF27640FDC351647D16" box="[869,1296,531,558]" pageId="14" pageNumber="227" type="nomenclature">
<paragraph id="8BEB36B7F102FFF27640FDC351647D16" blockId="14.[869,1296,531,591]" box="[869,1296,531,558]" pageId="14" pageNumber="227">
<heading id="D0A381DBF102FFF27640FDC351647D16" box="[869,1296,531,558]" centered="true" fontSize="12" level="2" pageId="14" pageNumber="227" reason="2">
<taxonomicName id="4C544D34F102FFF27640FDC351647D16" ID-CoL="5DDBY" authority="CAMERON, 1939" authorityName="Cameron" authorityYear="1939" box="[869,1296,531,558]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="227" phylum="Arthropoda" rank="species" species="elegantulus" subGenus="Zyras">
<emphasis id="B920EAA5F102FFF27640FDC357EF7D16" box="[869,923,531,558]" italics="true" pageId="14" pageNumber="227">Zyras</emphasis>
(
<emphasis id="B920EAA5F102FFF2768CFDC357AB7D16" box="[937,991,531,558]" italics="true" pageId="14" pageNumber="227">Zyras</emphasis>
)
<emphasis id="B920EAA5F102FFF276C8FDC350117D16" box="[1005,1125,531,558]" italics="true" pageId="14" pageNumber="227">elegantulus</emphasis>
CAMERON, 1939
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF102FFF27640FDEB50627D77" box="[869,1046,571,591]" pageId="14" pageNumber="227" type="description">
<paragraph id="8BEB36B7F102FFF27640FDEB50627D77" blockId="14.[869,1296,531,591]" box="[869,1046,571,591]" pageId="14" pageNumber="227">
(
<figureCitation id="136F2A32F102FFF2764CFDEB57CA7D77" box="[873,958,571,591]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Figs 16, 20</figureCitation>
,
<figureCitation id="136F2A32F102FFF276E0FDEB57AE7D77" box="[965,986,571,591]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="14" pageNumber="227">35</figureCitation>
,
<figureCitation id="136F2A32F102FFF276C4FDEB50657D77" box="[993,1041,571,591]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="14" pageNumber="227">53–55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF102FFF2761AFDBC51757DB8" box="[831,1281,619,641]" pageId="14" pageNumber="227" type="reference_group">
<paragraph id="8BEB36B7F102FFF2761AFDBC51757DB8" blockId="14.[831,1281,619,641]" box="[831,1281,619,641]" pageId="14" pageNumber="227">
<treatmentCitationGroup id="AB441199F102FFF2761AFDBC51757DB8" box="[831,1281,619,641]" pageId="14" pageNumber="227">
<treatmentCitation id="0AF510A6F102FFF2761AFDBC508A7DB8" author="CAMERON, M." box="[831,1278,619,641]" page="20" pageId="14" pageNumber="227" year="1939">
<taxonomicName id="4C544D34F102FFF2761AFDBC508A7DB8" ID-CoL="5DDBY" authority="CAMERON, 1939 b: 20" authorityName="Cameron" authorityPageNumber="20" authorityYear="1939" box="[831,1278,619,641]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="227" phylum="Arthropoda" rank="species" species="elegantulus" subGenus="Zyras">
<emphasis id="B920EAA5F102FFF2761AFDBC57077DB9" box="[831,883,620,641]" italics="true" pageId="14" pageNumber="227">Zyras</emphasis>
(
<emphasis id="B920EAA5F102FFF276A4FDBC57C17DB9" box="[897,949,620,641]" italics="true" pageId="14" pageNumber="227">Zyras</emphasis>
)
<emphasis id="B920EAA5F102FFF276E6FDBB50587DB8" box="[963,1068,619,640]" italics="true" pageId="14" pageNumber="227">elegantulus</emphasis>
<bibRefCitation id="EFC54B46F102FFF27114FDBB508A7DB8" author="CAMERON, M." box="[1073,1278,619,641]" pageId="14" pageNumber="227" pagination="1 - 29" refId="ref26173" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON, 1939b: 20</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF102FFF2761AFD7950687CF9" pageId="14" pageNumber="227" type="materials_examined">
<paragraph id="8BEB36B7F102FFF2761AFD7950687CF9" blockId="14.[831,1437,681,961]" pageId="14" pageNumber="227">
<emphasis id="B920EAA5F102FFF2761AFD79503F7DF9" bold="true" box="[831,1099,681,705]" pageId="14" pageNumber="227">Type material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF102FFF2717CFD7A51E97C79" ID-GBIF-Occurrence="3455148305" pageId="14" pageNumber="227" typeStatus="lectotype">
<typeStatus id="54EF8815F102FFF2717CFD7A50B67DFA" box="[1113,1218,682,706]" pageId="14" pageNumber="227" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D52FD3EF102FFF271ECFD7A50AA7DF9" box="[1225,1246,682,705]" pageId="14" pageNumber="227">♂</specimenCount>
, present designation: “
<collectorName id="26A15361F102FFF276A4FD19506F7DD9" box="[897,1051,713,737]" pageId="14" pageNumber="227">F. C. Drescher</collectorName>
, G. Tangkoeban Prahoe, 4000–5000
<collectingMunicipality id="6B8FACCDF102FFF2761AFD3A57077C39" box="[831,883,746,769]" pageId="14" pageNumber="227">Voet</collectingMunicipality>
,
<location id="8E8B606CF102FFF2765AFD3A57AB7C3A" LSID="urn:lsid:plazi:treatment:03FD87A1F102FFF37640FDC355A47939:8E8B606CF102FFF2765AFD3A57AB7C3A" box="[895,991,746,770]" municipality="Voet" name="Preanger" pageId="14" pageNumber="227" stateProvince="Java">Preanger</location>
,
<collectingRegion id="4990F855F102FFF276CFFD3A506C7C39" box="[1002,1048,746,769]" country="Indonesia" name="Jawa Timur" pageId="14" pageNumber="227">Java</collectingRegion>
,
<date id="FFEA1077F102FFF27106FD3950D87C39" box="[1059,1196,745,769]" pageId="14" pageNumber="227" value="1933-01-12" valueMax="1933-01-18" valueMin="1933-01-12">
<collectingDate id="EFAEE99FF102FFF27106FD3950D87C39" box="[1059,1196,745,769]" pageId="14" pageNumber="227" value="1933-01-12" valueMax="1933-01-18" valueMin="1933-01-12">12–18.I.1933</collectingDate>
</date>
/ Syntype / M. Cameron. Bequest. B.M. 1955-147 / Lectotypus ♂
<taxonomicName id="4C544D34F102FFF27047FCDA50517C79" authority="Cameron" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="227" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B920EAA5F102FFF27047FCDA57C77C79" italics="true" pageId="14" pageNumber="227">Zyras elegantulus</emphasis>
Cameron
</taxonomicName>
, desig.
<determinerName id="7E37DA73F102FFF27158FCFA50AB7C79" box="[1149,1247,809,833]" pageId="14" pageNumber="227">
V.
<bibRefCitation id="EFC54B46F102FFF271BDFCF950AB7C79" author="ASSING, V." box="[1176,1247,809,833]" pageId="14" pageNumber="227" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
</determinerName>
2017” (
<collectionCode id="ED45AE72F102FFF27019FCF951E67C79" box="[1340,1426,809,833]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="14" pageNumber="227">BMNH</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B3C3CEAF102FFF2761AFC99506D7CF9" ID-GBIF-Occurrence="3450187342" pageId="14" pageNumber="227" typeStatus="paralectotype">
<typeStatus id="54EF8815F102FFF2761AFC9957A47C59" box="[831,976,841,865]" pageId="14" pageNumber="227" type="paralectotype">Paralectotype</typeStatus>
♀: “
<collectorName id="26A15361F102FFF27124FC9950ED7C59" box="[1025,1177,841,865]" pageId="14" pageNumber="227">F. C. Drescher</collectorName>
,
<collectorName id="26A15361F102FFF27186FC9951EC7C59" box="[1187,1432,841,865]" pageId="14" pageNumber="227">G. Tangkoeban Prahoe</collectorName>
, 4000–5000
<collectingMunicipality id="6B8FACCDF102FFF2769FFCBA579A7CB9" box="[954,1006,874,897]" pageId="14" pageNumber="227">Voet</collectingMunicipality>
,
<location id="8E8B606CF102FFF276DDFCBA502C7CBA" LSID="urn:lsid:plazi:treatment:03FD87A1F102FFF37640FDC355A47939:8E8B606CF102FFF276DDFCBA502C7CBA" box="[1016,1112,874,898]" municipality="Voet" name="Preanger" pageId="14" pageNumber="227" stateProvince="Java">Preanger</location>
,
<collectingRegion id="4990F855F102FFF27144FCBA50FB7CB9" box="[1121,1167,874,897]" country="Indonesia" name="Jawa Timur" pageId="14" pageNumber="227">Java</collectingRegion>
,
<date id="FFEA1077F102FFF271BCFCB950837CB9" box="[1177,1271,873,897]" pageId="14" pageNumber="227" value="1934-07">
<collectingDate id="EFAEE99FF102FFF271BCFCB950837CB9" box="[1177,1271,873,897]" pageId="14" pageNumber="227" value="1934-07">VII.1934</collectingDate>
</date>
/
<taxonomicName id="4C544D34F102FFF27029FCBA57A67C99" authority="Cam. Cotype" authorityName="Cam. Cotype" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="227" phylum="Arthropoda" rank="species" species="elegantulus">
<collectorName id="26A15361F102FFF27029FCBA57A67C99" pageId="14" pageNumber="227">
<emphasis id="B920EAA5F102FFF27029FCBA51E87CB9" box="[1292,1436,873,897]" italics="true" pageId="14" pageNumber="227">Z. elegantulus</emphasis>
Cam. Cotype
</collectorName>
</taxonomicName>
/
<typeStatus id="54EF8815F102FFF276D6FC59503C7C99" box="[1011,1096,905,929]" pageId="14" pageNumber="227" type="syntype">Syntype</typeStatus>
/
<collectorName id="26A15361F102FFF2714CFC5A512A7C99" box="[1129,1374,905,929]" pageId="14" pageNumber="227">M. Cameron. Bequest.</collectorName>
B.M. 1955-147” (
<collectionCode id="ED45AE72F102FFF2769EFC7950657CF9" box="[955,1041,937,961]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="14" pageNumber="227">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF102FFF2761AFC39500E7BB9" pageId="14" pageNumber="227" type="discussion">
<paragraph id="8BEB36B7F102FFF2761AFC39500E7BB9" blockId="14.[831,1437,1001,1153]" pageId="14" pageNumber="227">
<emphasis id="B920EAA5F102FFF2761AFC3957DA7B39" bold="true" box="[831,942,1001,1025]" pageId="14" pageNumber="227">Comment</emphasis>
: The original description is based on an unspecified number of
<typeStatus id="54EF8815F102FFF2716DFBDA50D07B1A" box="[1096,1188,1034,1058]" pageId="14" pageNumber="227" type="syntype">syntypes</typeStatus>
from “G. Tangkoeban Prahoe” (
<bibRefCitation id="EFC54B46F102FFF27680FBF950147B79" author="CAMERON, M." box="[933,1120,1065,1089]" pageId="14" pageNumber="227" pagination="1 - 29" refId="ref26173" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON 1939b</bibRefCitation>
). Two
<typeStatus id="54EF8815F102FFF2718DFBFA51727B7A" box="[1192,1286,1066,1090]" pageId="14" pageNumber="227" type="syntype">syntypes</typeStatus>
, a male and a female, were located in the Cameron collection. The male is designated as the
<typeStatus id="54EF8815F102FFF27137FBB950027BB9" box="[1042,1142,1129,1153]" pageId="14" pageNumber="227" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF102FFF3761AFB7955537A99" lastPageId="15" lastPageNumber="228" pageId="14" pageNumber="227" type="description">
<paragraph id="8BEB36B7F102FFF2761AFB7950B77939" blockId="14.[831,1437,1193,2081]" pageId="14" pageNumber="227">
<emphasis id="B920EAA5F102FFF2761AFB7957AC7BF9" bold="true" box="[831,984,1193,1217]" pageId="14" pageNumber="227">Redescription</emphasis>
: Body length
<quantity id="4CAC9B52F102FFF271B3FB79516C7BF9" box="[1174,1304,1193,1217]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.7" metricValueMin="5.3" pageId="14" pageNumber="227" unit="mm" value="5.5" valueMax="5.7" valueMin="5.3">5.3–5.7 mm</quantity>
; length of forebody
<quantity id="4CAC9B52F102FFF27696FB1950737BD9" box="[947,1031,1225,1249]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="227" unit="mm" value="2.5">2.5 mm</quantity>
. Coloration (
<figureCitation id="136F2A32F102FFF27188FB1951567BD9" box="[1197,1314,1225,1249]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Figs 16, 20</figureCitation>
,
<figureCitation id="136F2A32F102FFF27009FB19513D7BD9" box="[1324,1353,1225,1249]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="14" pageNumber="227">35</figureCitation>
): head pale-brown to reddish-brown; pronotum pale-reddish; elytra yellowish to yellowish-red, with the postero-lateral portions more or less extensively infuscate (very extensively so in the
<typeStatus id="54EF8815F102FFF276C2FA99500F7A59" box="[999,1147,1353,1377]" pageId="14" pageNumber="227" type="paralectotype">paralectotype</typeStatus>
); abdomen pale-reddish to dark-reddish, with the posterior halves of tergites V–VII darker; legs pale-yellowish; antennae dark-brown with antennomeres I–II pale-reddish to reddish-brown and X–XI yellowish-brown to brown; maxillary palpi reddish with the apical palpomere yellowish.
</paragraph>
<paragraph id="8BEB36B7F102FFF2761AF9D950807839" blockId="14.[831,1437,1193,2081]" pageId="14" pageNumber="227">
Head (
<figureCitation id="136F2A32F102FFF276A8F9D957A37919" box="[909,983,1545,1569]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Fig. 20</figureCitation>
) moderately transverse, median portion extensively impunctate; punctures in lateral portions sparse and coarse. Eyes longer than postocular region in dorsal view. Antenna (
<figureCitation id="136F2A32F102FFF27174F9B950EF79B9" box="[1105,1179,1641,1665]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Fig. 16</figureCitation>
) approximately
<quantity id="4CAC9B52F102FFF2706EF9B951E979B9" box="[1355,1437,1641,1665]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="227" unit="mm" value="1.5">1.5 mm</quantity>
long; antennomeres IV–X transverse, of gradually increasing width and increasingly transverse, X approximately twice as broad as long, and XI approximately as long as the combined length of IX and X.
</paragraph>
<paragraph id="8BEB36B7F102FFF2761AF8DA57CB78D9" blockId="14.[831,1437,1193,2081]" pageId="14" pageNumber="227">
Pronotum (
<figureCitation id="136F2A32F102FFF276E0F8D9507B7819" box="[965,1039,1801,1825]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Fig. 20</figureCitation>
) large in relation to head, 1.15– 1.18 times as broad as long and 1.36–1.43 times as broad as head, broadest near anterior angles, moderately convex in cross-section; punctation rather coarse, sparse, and very irregularly distributed, with extensive impunctate areas on either side of midline; midline broadly impunctate.
</paragraph>
<paragraph id="8BEB36B7F102FFF3761AF83955D87E59" blockId="14.[831,1437,1193,2081]" lastBlockId="15.[151,756,233,1249]" lastPageId="15" lastPageNumber="228" pageId="14" pageNumber="227">
Elytra (
<figureCitation id="136F2A32F102FFF276BEF83957927739" box="[923,998,2025,2049]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Fig. 20</figureCitation>
) 0.75–0.79 times as long as pronotum; punctation variable, moderately coarse to coarse, moderately sparse to moderately dense, defined, sparser posteriorly than anteriorly. Hind wings present. Metatarsomere I slender, slightly to distinctly shorter than the combined length of II–IV.
</paragraph>
<paragraph id="8BEB36B7F103FFF375B2FEB9563E7B39" blockId="15.[151,756,233,1249]" pageId="15" pageNumber="228">
Abdomen (
<figureCitation id="136F2A32F103FFF3743FFEB955127EB9" box="[282,358,361,385]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="15" pageNumber="228">Fig. 35</figureCitation>
) narrower than elytra, with deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of weakly defined non-setiferous punctures; tergite III with two lateral punctures on either side and with approximately 15 setiferous punctures at posterior margin; tergite IV–V with a median pair of punctures, with 3–5 lateral setiferous punctures on either side, and with approximately 20 setiferous punctures at posterior margins; tergite VI with a narrow transverse row of coarse non-setiferous punctures anteriorly, with a median pair of punctures, with a lateral transverse series of three setiferous punctures on either side, and with approximately 20 setiferous punctures at posterior margin; tergite VII with a narrow transverse band of non-setiferous punctures anteriorly and with two transverse series of setiferous punctures in posterior portion, posterior margin with palisade fringe; tergite VIII with few setiferous punctures bearing long black setae only near posterior margin, posterior margin convex, in the middle truncate or indistinctly concave.
</paragraph>
<paragraph id="8BEB36B7F103FFF375B2FBDA55397B99" blockId="15.[151,756,233,1249]" pageId="15" pageNumber="228">
♂: posterior margin of sternite VIII convex; median lobe of aedeagus (
<figureCitation id="136F2A32F103FFF37439FBF955E77B79" box="[284,403,1065,1089]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="15" pageNumber="228">Figs 53–54</figureCitation>
)
<quantity id="4CAC9B52F103FFF374BBFBF955887B79" box="[414,508,1065,1089]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="15" pageNumber="228" unit="mm" value="0.57">0.57 mm</quantity>
long, of compact shape, ventral process short and broad; paramere (
<figureCitation id="136F2A32F103FFF37745FB9956DE7B59" box="[608,682,1097,1121]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="15" pageNumber="228">Fig. 55</figureCitation>
) much longer than median lobe,
<quantity id="4CAC9B52F103FFF37490FBB956607BB9" box="[437,532,1129,1153]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="15" pageNumber="228" unit="mm" value="0.84">0.84 mm</quantity>
long, and with very short apical lobe.
</paragraph>
<paragraph id="8BEB36B7F103FFF375B2FB7A549D7BD9" blockId="15.[151,756,233,1249]" pageId="15" pageNumber="228">♀: posterior margin of sternite VIII concave in the middle.</paragraph>
<paragraph id="8BEB36B7F103FFF375B2FAD955537A99" blockId="15.[151,756,1289,1441]" pageId="15" pageNumber="228">
<emphasis id="B920EAA5F103FFF375B2FAD955187A19" bold="true" box="[151,364,1289,1313]" pageId="15" pageNumber="228">Comparative notes</emphasis>
: This species is characterized by a rather large pronotum (in relation to head), its coloration, a pronotum with very irregular punctation, the punctation pattern of the abdomen, and by the morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF103FFF375B2FA1955A47939" pageId="15" pageNumber="228" type="distribution">
<paragraph id="8BEB36B7F103FFF375B2FA1955A47939" blockId="15.[151,756,1481,1538]" pageId="15" pageNumber="228">
<emphasis id="B920EAA5F103FFF375B2FA1955547AD9" bold="true" box="[151,288,1481,1505]" pageId="15" pageNumber="228">Distribution</emphasis>
:
<taxonomicName id="4C544D34F103FFF37415FA1A559C7AD9" box="[304,488,1481,1505]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B920EAA5F103FFF37415FA1A559C7AD9" box="[304,488,1481,1505]" italics="true" pageId="15" pageNumber="228">Zyras elegantulus</emphasis>
</taxonomicName>
has been recorded only from the
<typeStatus id="54EF8815F103FFF375DFFA3A555C793A" box="[250,296,1514,1538]" pageId="15" pageNumber="228">type</typeStatus>
locality in
<collectingRegion id="4990F855F103FFF374BBFA3A55B87939" box="[414,460,1514,1537]" country="Indonesia" name="Jawa Timur" pageId="15" pageNumber="228">Java</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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