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<document id="809C9895D22F5798AD15A026BFD6F3B1" ID-CLB-Dataset="31353" ID-DOI="10.11646/zootaxa.4353.2.1" ID-GBIF-Dataset="d93b2bd2-75e1-467c-89d3-e2faee3b10a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1065124" ID-ZooBank="D2902BF9-0213-40F3-91A4-EC4D4FDB3B27" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511423577451" checkinUser="plazi" docAuthor="Serrano, Artur R. M., Capela, Rúben A. &amp; Santos, Carmen Van-Dúnem Neto" docDate="2017" docId="03EFF252F3087F6F14BFA6D5FE3CF9BD" docLanguage="en" docName="zootaxa.4353.2.1.pdf" docOrigin="Zootaxa 4353 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systolocranius tibialis Lecordier 1972" docType="treatment" docVersion="8" lastPageNumber="213" masterDocId="FFD68A2AF3047F631428A273FFF3FFBF" masterDocTitle="Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="213" updateTime="1698503882738" updateUser="ExternalLinkService">
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<mods:title id="ADCA7FFD4FB96F388CC8BC2F93188636">Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="0AAE720AD483472F2A360CF1E7B67B5F">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="6E8EF463FA78D0A8F8E87674D64CE97D">Capela, Rúben A.</mods:namePart>
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<mods:namePart id="13521A542365E24CCC59B5D660AC9BEC">Santos, Carmen Van-Dúnem Neto</mods:namePart>
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<treatment id="03EFF252F3087F6F14BFA6D5FE3CF9BD" ID-DOI="http://doi.org/10.5281/zenodo.6005630" ID-GBIF-Taxon="137541492" ID-Zenodo-Dep="6005630" LSID="urn:lsid:plazi:treatment:03EFF252F3087F6F14BFA6D5FE3CF9BD" httpUri="http://treatment.plazi.org/id/03EFF252F3087F6F14BFA6D5FE3CF9BD" lastPageNumber="213" pageId="12" pageNumber="213">
<subSubSection id="C35C10CFF3087F6F14BFA6D5FD98FB7F" box="[151,619,1190,1216]" pageId="12" pageNumber="213" type="nomenclature">
<paragraph id="8BF94344F3087F6F14BFA6D5FD98FB7F" blockId="12.[151,619,1190,1216]" box="[151,619,1190,1216]" pageId="12" pageNumber="213">
<heading id="D0B1F428F3087F6F14BFA6D5FD98FB7F" bold="true" box="[151,619,1190,1216]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<taxonomicName id="4C4638C7F3087F6F14BFA6D5FD98FB7F" ID-CoL="54528" authority="Lecordier, 1972" authorityName="Lecordier" authorityYear="1972" box="[151,619,1190,1216]" class="Insecta" family="Carabidae" genus="Systolocranius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B9329F56F3087F6F14BFA6D5FD98FB7F" bold="true" box="[151,619,1190,1216]" pageId="12" pageNumber="213">
<emphasis id="B9329F56F3087F6F14BFA6D5FE52FB7F" bold="true" box="[151,417,1190,1216]" italics="true" pageId="12" pageNumber="213">Systolocranius tibialis</emphasis>
<bibRefCitation id="EFD73EB5F3087F6F1580A6D5FD98FB7F" author="Lecordier" box="[424,619,1190,1216]" pageId="12" pageNumber="213" refString="Lecordier, C. (1972) Note sur les Oodinae (Coleoptera Carabidae) d`Afrique Noire. Revue Zoologique et Botanique d`Afrique, 86 (1 - 2), 1 - 16." type="journal article" year="1972">Lecordier, 1972</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C10CFF3087F6F14BFA69EFCA2FAB9" box="[151,849,1261,1286]" pageId="12" pageNumber="213" type="distribution">
<paragraph id="8BF94344F3087F6F14BFA69EFCA2FAB9" blockId="12.[151,1436,1261,1538]" box="[151,849,1261,1286]" pageId="12" pageNumber="213">
<emphasis id="B9329F56F3087F6F14BFA69EFDC8FAB9" bold="true" box="[151,571,1261,1286]" pageId="12" pageNumber="213">
Distribution in
<collectingCountry id="F35103D4F3087F6F1566A69EFE50FAB9" box="[334,419,1261,1286]" name="Angola" pageId="12" pageNumber="213">Angola</collectingCountry>
(Provinces):
</emphasis>
1)
<collectingRegion id="49828DA6F3087F6F1648A69EFD1CFABA" box="[608,751,1261,1286]" country="Angola" name="Lunda Norte" pageId="12" pageNumber="213">
Lunda
<collectingRegion id="49828DA6F3087F6F1687A69DFD1CFABA" box="[687,751,1262,1285]" country="Sudan" name="Ash Shimaliyya" pageId="12" pageNumber="213">Norte</collectingRegion>
</collectingRegion>
; 2)
<collectingRegion id="49828DA6F3087F6F170BA69EFCBDFAB9" box="[803,846,1261,1286]" country="Angola" name="Bie" pageId="12" pageNumber="213">Bié</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C10CFF3087F6F14EFA762FB7EFACD" pageId="12" pageNumber="213" type="materials_examined">
<paragraph id="8BF94344F3087F6F14EFA762FB7EFACD" blockId="12.[151,1436,1261,1538]" pageId="12" pageNumber="213">
<materialsCitation id="3B2E4919F3087F6F14EFA762FCCBFAF1" ID-GBIF-Occurrence="1701951522" collectingDate="2014-11-03" collectionCode="ASC" collectorName="Serrano &amp; R. Capela" elevation="1611" latitude="-13.74625" location="BIE" longLatPrecision="1" longitude="17.18396" pageId="12" pageNumber="213" specimenCount="3" specimenCount-female="3" stateProvince="BIE">
<emphasis id="B9329F56F3087F6F14EFA762FE42FA95" bold="true" box="[199,433,1297,1322]" pageId="12" pageNumber="213">Material examined.</emphasis>
Satchijamba-Somakwanza (
<geoCoordinate id="EE722583F3087F6F16DBA761FC4AFA95" box="[755,953,1297,1322]" direction="south" orientation="latitude" pageId="12" pageNumber="213" precision="1" value="-13.74625">13° 44´46.50´´ S</geoCoordinate>
,
<geoCoordinate id="EE722583F3087F6F17EEA761FB73FA96" box="[966,1152,1297,1322]" direction="east" orientation="longitude" pageId="12" pageNumber="213" precision="1" value="17.18396">17° 11´2.26´´ E</geoCoordinate>
,
<quantity id="4CBEEEA1F3087F6F10A4A762FB12FA96" box="[1164,1249,1297,1322]" metricMagnitude="3" metricUnit="m" metricValue="1.611" pageId="12" pageNumber="213" unit="m" value="1611.0">
<elevation id="006BA477F3087F6F10A4A762FB12FA96" box="[1164,1249,1297,1322]" metricMagnitude="3" metricUnit="m" metricValue="1.611" pageId="12" pageNumber="213" unit="m" value="1611.0">1611 m</elevation>
</quantity>
alt., 303) (
<collectingRegion id="49828DA6F3087F6F1176A762FA63FA95" box="[1374,1424,1297,1322]" country="Angola" name="Bie" pageId="12" pageNumber="213">BIÉ</collectingRegion>
), 2.
<date id="FFF86584F3087F6F1481A745FED6FAF1" box="[169,293,1334,1358]" pageId="12" pageNumber="213" value="2014-11-03">
<collectingDate id="EFBC9C6CF3087F6F1481A745FED6FAF1" box="[169,293,1334,1358]" pageId="12" pageNumber="213" value="2014-11-03">XI.2014, 3</collectingDate>
</date>
♂,
<specimenCount id="9D4088CDF3087F6F1560A745FE9FFAF1" box="[328,364,1334,1358]" pageId="12" pageNumber="213" type="female">3♀</specimenCount>
, DO,
<collectingRegion id="49828DA6F3087F6F1599A745FE3BFAF1" box="[433,456,1334,1358]" country="Nigeria" name="Rivers" pageId="12" pageNumber="213">A</collectingRegion>
.
<collectorName id="26B32692F3087F6F15FCA745FDD8FAF1" box="[468,555,1334,1358]" pageId="12" pageNumber="213">Serrano</collectorName>
&amp;
<collectorName id="26B32692F3087F6F1667A745FD4DFAF1" box="[591,702,1333,1358]" pageId="12" pageNumber="213">R. Capela</collectorName>
leg.,
<collectionCode id="ED57DB81F3087F6F16D6A745FCCBFAF1" box="[766,824,1334,1358]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15857" name="Northern Arizona University" pageId="12" pageNumber="213">ASC</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B2E4919F3087F6F176BA746FB7AFACD" ID-GBIF-Occurrence="1701951560" collectingDate="2015-11-01" collectionCode="ASC" collectorName="Serrano &amp; R. Capela" elevation="1511" latitude="-13.828055" location="BIE" longLatPrecision="21" longitude="17.32389" pageId="12" pageNumber="213" specimenCount="1" stateProvince="BIE">
Mumbué (Cuanza riverhead) (
<geoCoordinate id="EE722583F3087F6F10B0A745FAC8FAF1" box="[1176,1339,1334,1358]" direction="south" orientation="latitude" pageId="12" pageNumber="213" precision="15" value="-13.828055">13° 49´41´´ S</geoCoordinate>
,
<geoCoordinate id="EE722583F3087F6F1160A745FF2CFACE" direction="east" orientation="longitude" pageId="12" pageNumber="213" precision="15" value="17.32389">17° 19´26´´ E</geoCoordinate>
,
<quantity id="4CBEEEA1F3087F6F14C3A72AFEB3FACE" box="[235,320,1369,1394]" metricMagnitude="3" metricUnit="m" metricValue="1.511" pageId="12" pageNumber="213" unit="m" value="1511.0">
<elevation id="006BA477F3087F6F14C3A72AFEB3FACE" box="[235,320,1369,1394]" metricMagnitude="3" metricUnit="m" metricValue="1.511" pageId="12" pageNumber="213" unit="m" value="1511.0">1511 m</elevation>
</quantity>
alt., 303) (
<collectingRegion id="49828DA6F3087F6F1592A72AFE1FFACD" box="[442,492,1369,1394]" country="Angola" name="Bie" pageId="12" pageNumber="213">BIÉ</collectingRegion>
), 817.
<date id="FFF86584F3087F6F1613A72AFD47FACD" box="[571,692,1369,1394]" pageId="12" pageNumber="213" value="2015-11-01">
<collectingDate id="EFBC9C6CF3087F6F1613A72AFD47FACD" box="[571,692,1369,1394]" pageId="12" pageNumber="213" value="2015-11-01">XI.2015, 1</collectingDate>
</date>
♂, PF,
<collectingRegion id="49828DA6F3087F6F172DA729FCEFFACD" box="[773,796,1370,1394]" country="Nigeria" name="Rivers" pageId="12" pageNumber="213">A</collectingRegion>
.
<collectorName id="26B32692F3087F6F170FA729FC8DFACD" box="[807,894,1370,1394]" pageId="12" pageNumber="213">Serrano</collectorName>
&amp;
<collectorName id="26B32692F3087F6F178AA729FBE2FACD" box="[930,1041,1369,1394]" pageId="12" pageNumber="213">R. Capela</collectorName>
leg.,
<collectionCode id="ED57DB81F3087F6F1067A729FB7AFACD" box="[1103,1161,1370,1394]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15857" name="Northern Arizona University" pageId="12" pageNumber="213">ASC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C10CFF3087F6F14EFA70EFE3CF9BD" pageId="12" pageNumber="213" type="discussion">
<paragraph id="8BF94344F3087F6F14EFA70EFE3CF9BD" blockId="12.[151,1436,1261,1538]" pageId="12" pageNumber="213">
<emphasis id="B9329F56F3087F6F14EFA70EFEC8FA29" bold="true" box="[199,315,1405,1430]" pageId="12" pageNumber="213">Remarks.</emphasis>
<collectingRegion id="49828DA6F3087F6F156AA70DFEA5FA29" box="[322,342,1406,1430]" country="Nigeria" name="Rivers" pageId="12" pageNumber="213">A</collectingRegion>
species spread throughout
<collectingCountry id="F35103D4F3087F6F16A0A70EFD28FA29" box="[648,731,1405,1430]" name="Angola" pageId="12" pageNumber="213">Angola</collectingCountry>
and D. R. of the
<collectingCountry id="F35103D4F3087F6F17B6A70DFC1AFA29" box="[926,1001,1406,1430]" name="Democratic Republic of the Congo" pageId="12" pageNumber="213">
<collectingRegion id="49828DA6F3087F6F17B6A70DFC1AFA29" box="[926,1001,1406,1430]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="12" pageNumber="213">Congo</collectingRegion>
</collectingCountry>
(
<collectingRegion id="49828DA6F3087F6F17D0A70DFBAAFA29" box="[1016,1113,1406,1430]" country="Democratic Republic of the Congo" name="Katanga" pageId="12" pageNumber="213">Katanga</collectingRegion>
) (
<bibRefCitation id="EFD73EB5F3087F6F1047A70EFAD2FA29" author="Lecordier" box="[1135,1313,1405,1430]" pageId="12" pageNumber="213" refString="Lecordier, C. (1972) Note sur les Oodinae (Coleoptera Carabidae) d`Afrique Noire. Revue Zoologique et Botanique d`Afrique, 86 (1 - 2), 1 - 16." type="journal article" year="1972">Lecordier 1972</bibRefCitation>
). Adults in the first locality were found in activity on sandy whitish soil within a secondary open forest during night together with other ground beetles [e.g.
<taxonomicName id="4C4638C7F3087F6F1629A7B5FD1BFA61" box="[513,744,1477,1502]" class="Insecta" family="Carabidae" genus="Systolocranius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="subSpecies" species="tenebrosus" subSpecies="fatalis">
<emphasis id="B9329F56F3087F6F1629A7B5FD1BFA61" box="[513,744,1477,1502]" italics="true" pageId="12" pageNumber="213">S. tenebrosus fatalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4638C7F3087F6F16D1A7B5FBADFA61" authorityName="Boheman" authorityYear="1848" box="[761,1118,1477,1502]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="bipustulatus" subGenus="Pachydinodes">
<emphasis id="B9329F56F3087F6F16D1A7B5FCE0FA62" box="[761,787,1478,1501]" italics="true" pageId="12" pageNumber="213">C.</emphasis>
(
<emphasis id="B9329F56F3087F6F170DA7B6FC31FA61" box="[805,962,1477,1502]" italics="true" pageId="12" pageNumber="213">Pachydinodes</emphasis>
)
<emphasis id="B9329F56F3087F6F17FDA7B6FBADFA61" box="[981,1118,1477,1502]" italics="true" pageId="12" pageNumber="213">bipustulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4638C7F3087F6F1047A7B5FAFFFA61" authorityName="Guerin-Meneville" authorityYear="1845" box="[1135,1292,1477,1502]" class="Insecta" family="Carabidae" genus="Termophilum" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="massilicata">
<emphasis id="B9329F56F3087F6F1047A7B5FAFFFA61" box="[1135,1292,1477,1502]" italics="true" pageId="12" pageNumber="213">T. massilicata</emphasis>
</taxonomicName>
] It is a new record for the
<collectingRegion id="49828DA6F3087F6F151EA79AFE38F9BD" box="[310,459,1513,1538]" country="Angola" name="Bie" pageId="12" pageNumber="213">Bié Province</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>