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<document id="2084D641FC28149A38A720F650F3C060" ID-DOI="10.11646/zootaxa.5361.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10150449" ID-ZooBank="B960FE5C-095B-455B-9639-52ACE8CEFD22" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700246390697" checkinUser="plazi" docAuthor="Zhao, Ming-Zhi &amp; Ahrens, Dirk" docDate="2023" docId="03EF6024B450CE4810FD33C1FED6FB21" docLanguage="en" docName="zootaxa.5361.2.3.pdf" docOrigin="Zootaxa 5361 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5361.2.3/52178" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maladera zhanchaoi Zhao &amp; Ahrens 2023, new species" docType="treatment" docVersion="2" lastPageNumber="210" masterDocId="FFD6185CB44BCE55106A3037FF9CFFC8" masterDocTitle="New species of the tribe Sericini Kirby, 1837 from China, with further updates on their taxonomy and distribution (Coleoptera: Scarabaeidae: Sericinae)" masterLastPageNumber="220" masterPageNumber="181" pageNumber="208" updateTime="1700246908677" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E0D00DA058289B847E59CCF1D2FFB85E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2DDE6E6F91BC15B0968F9DB760CB357D">
<mods:title id="0588F57FF6F05532741D492267AB54BC">New species of the tribe Sericini Kirby, 1837 from China, with further updates on their taxonomy and distribution (Coleoptera: Scarabaeidae: Sericinae)</mods:title>
</mods:titleInfo>
<mods:name id="9EE7B697F9FE550F5E60B912EC26336F" type="personal">
<mods:role id="B5E0D6521ED6FEC0C617ADD81FBD29C2">
<mods:roleTerm id="9AE70ACCFAD27327EBBCD4FA49591ACA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7FB823D9541E0C80467A658DC126ACD4">Zhao, Ming-Zhi</mods:namePart>
<mods:nameIdentifier id="08DD343A23531DA12D9F9B6D199046BC" type="ORCID">0000-0002-3158-6177</mods:nameIdentifier>
<mods:affiliation id="A1F7CEC73A2E0DCDE0AE97BB5351BE55">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou 510642, P. R. China zhaomzhai @ 126. com; https: // orcid. org / 0000 - 0002 - 3158 - 6177</mods:affiliation>
</mods:name>
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<mods:roleTerm id="9D007A603A4A0BCEA48CA2DC6EBDD5FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C847C19570D20904A511278F96563F0B">Ahrens, Dirk</mods:namePart>
<mods:affiliation id="E5F08F54959EA072F5824C2D47D5B593">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6DA7E9226BF5B9E0C2C090FD26F4134F">text</mods:typeOfResource>
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<mods:title id="A87CD29EE8900BD107485377975F81E6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="41AFC74328828CC52B4DC2D9D6886B9F">
<mods:date id="8769AF6C4093261C9A7F2F7D17A6D4A8">2023</mods:date>
<mods:detail id="1FF20C6E7C7A1DE41C88CCDAD049333A" type="pubDate">
<mods:number id="018F93E28F71174F6C5BB3C5888D0988">2023-11-01</mods:number>
</mods:detail>
<mods:detail id="287BBB1515F41DFBC2EE51655E644F29" type="volume">
<mods:number id="1FC3272D84E96B17F498929BD321C635">5361</mods:number>
</mods:detail>
<mods:detail id="7F0848D28CEE77C72128665D453B5C00" type="issue">
<mods:number id="FD69AB8540F5337B192746B5046DECE0">2</mods:number>
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<mods:url id="249DCF32B654E2EEFEBC18EF07A90D25">https://www.mapress.com/zt/article/download/zootaxa.5361.2.3/52178</mods:url>
</mods:location>
<mods:classification id="F066BF79667BE614DA496BB286B3B01B">journal article</mods:classification>
<mods:identifier id="65E657A9F7FDA346A24FB952DB37F790" type="DOI">10.11646/zootaxa.5361.2.3</mods:identifier>
<mods:identifier id="9779C6AD98241CDBB9A79CA4A559B7B9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B9ABEFC1A6FA818F34D06A3FFD49AB0F" type="Zenodo-Dep">10150449</mods:identifier>
<mods:identifier id="CEF006D12A5A6BAC7C9078265B1187C0" type="ZooBank">B960FE5C-095B-455B-9639-52ACE8CEFD22</mods:identifier>
</mods:mods>
<treatment id="03EF6024B450CE4810FD33C1FED6FB21" LSID="urn:lsid:plazi:treatment:03EF6024B450CE4810FD33C1FED6FB21" httpUri="http://treatment.plazi.org/id/03EF6024B450CE4810FD33C1FED6FB21" lastPageId="29" lastPageNumber="210" pageId="27" pageNumber="208">
<subSubSection id="C35C82B9B450CE4E10FD33C1FD65FBD8" box="[151,761,1014,1041]" pageId="27" pageNumber="208" type="nomenclature">
<paragraph id="8BF9D132B450CE4E10FD33C1FD65FBD8" blockId="27.[151,761,1014,1077]" box="[151,761,1014,1041]" pageId="27" pageNumber="208">
<heading id="D0B1665EB450CE4E10FD33C1FD65FBD8" bold="true" box="[151,761,1014,1041]" fontSize="11" level="1" pageId="27" pageNumber="208" reason="1">
<emphasis id="B9320D20B450CE4E10FD33C1FD65FBD8" bold="true" box="[151,761,1014,1041]" pageId="27" pageNumber="208">
<taxonomicName id="4C46AAB1B450CE4E10FD33C1FDC7FBD8" authority="Zhao &amp; Ahrens" authorityName="Zhao &amp; Ahrens" authorityYear="2023" box="[151,603,1014,1041]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="208" phylum="Arthropoda" rank="species" species="zhanchaoi" status="sp. nov.">
<emphasis id="B9320D20B450CE4E10FD33C1FE0EFBD9" bold="true" box="[151,402,1014,1041]" italics="true" pageId="27" pageNumber="208">Maladera zhanchaoi</emphasis>
Zhao &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A201B05BB450CE4E120333C0FD65FBD8" box="[617,761,1014,1041]" pageId="27" pageNumber="208" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C82B9B450CE4E10FD342CFED2FBFD" box="[151,334,1051,1077]" pageId="27" pageNumber="208" type="description">
<paragraph id="8BF9D132B450CE4E10FD342CFED2FBFD" blockId="27.[151,761,1014,1077]" box="[151,334,1051,1077]" pageId="27" pageNumber="208">
(
<figureCitation id="137DCDB7B450CE4E10F5342CFE96FBFD" box="[159,266,1051,1077]" captionStart="FIGURE 10" captionStartId="28.[151,250,1634,1658]" captionTargetBox="[166,1431,252,1601]" captionTargetId="figure-49@28.[151,1436,237,1611]" captionTargetPageId="28" captionText="FIGURE 10. Habitus in dorsal view.A, Maladera zhanchaoi Zhao &amp;Ahrens, new species, holotype; B, M.parabikouesis Zhao &amp; Ahrens, new species, holotype; C, M. qianqingtangensis Ahrens, Fabrizi &amp; Liu, 2021, female (Fujian); D, M. qianqingtangensis, male (Fujian); E, M. shikengkongensis Zhao &amp; Ahrens, new species, holotype; F, M. shikengkongensis, female paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10150469" httpUri="https://zenodo.org/record/10150469/files/figure.png" pageId="27" pageNumber="208">Figs 10A</figureCitation>
,
<figureCitation id="137DCDB7B450CE4E117C342CFEDBFBFD" box="[278,327,1051,1077]" captionStart="FIGURE 11" captionStartId="30.[151,250,1336,1360]" captionTargetBox="[164,1424,202,1298]" captionTargetId="figure-18@30.[151,1436,181,1312]" captionTargetPageId="30" captionText="FIGURE 11. Aedeagus. A, Maladera zhanchaoi Zhao &amp; Ahrens, new species, holotype; B, M. parabikouesis Zhao &amp; Ahrens, new species, holotype; C, M. shikengkongensis Zhao &amp; Ahrens, new species, holotype; D, M. qianqingtangensis Ahrens, Fabrizi &amp; Liu, 2021 (Fujian). a, aedeagus in left lateral view; b, parameres in dorsal view; c, aedeagus in right lateral view; d, aedeagus in dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10150471" httpUri="https://zenodo.org/record/10150471/files/figure.png" pageId="27" pageNumber="208">11A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C82B9B450CE4E10FD3454FA19FB0D" pageId="27" pageNumber="208" type="materials_examined">
<paragraph id="8BF9D132B450CE4E10FD3454FA19FB0D" blockId="27.[151,1437,1122,2013]" pageId="27" pageNumber="208">
<emphasis id="B9320D20B450CE4E10FD3454FE5FFBB4" bold="true" box="[151,451,1122,1149]" pageId="27" pageNumber="208">Type material examined.</emphasis>
<materialsCitation id="3B2EDB6FB450CE4E11A63454FC3EFB69" collectingDate="2020-05-23" collectionCode="SCAU" collectorName="Bao-Xiang Zhan" country="China" county="Libo County" elevation="507" location="Wuyanqiao" municipality="Maolan" pageId="27" pageNumber="208" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="B9320D20B450CE4E11A63454FDDCFBB5" bold="true" box="[460,576,1123,1149]" pageId="27" pageNumber="208">
<typeStatus id="54FD6F90B450CE4E11A63454FDA7FBB5" box="[460,571,1123,1149]" pageId="27" pageNumber="208" type="holotype">Holotype</typeStatus>
.
</emphasis>
♁ (
<collectionCode id="ED5749F7B450CE4E12043454FD22FBB5" box="[622,702,1123,1149]" pageId="27" pageNumber="208">SCAU</collectionCode>
), “
<collectingCountry id="F35191A2B450CE4E12B43454FCA1FBB5" box="[734,829,1123,1149]" name="China" pageId="27" pageNumber="208">CHINA</collectingCountry>
:
<collectingRegion id="49821FD0B450CE4E13233454FC30FBB5" box="[841,940,1123,1149]" country="China" name="Guizhou" pageId="27" pageNumber="208">Guizhou</collectingRegion>
,
<collectingCounty id="6298A9BEB450CE4E13D03454FBD7FBB5" box="[954,1099,1123,1149]" pageId="27" pageNumber="208">Libo County</collectingCounty>
,
<collectingMunicipality id="6B9D4B48B450CE4E14323454FB2CFBB5" box="[1112,1200,1123,1149]" pageId="27" pageNumber="208">Maolan</collectingMunicipality>
,
<location id="8E9987E9B450CE4E14D73454FADCFBB5" LSID="urn:lsid:plazi:treatment:03EF6024B450CE4810FD33C1FED6FB21:8E9987E9B450CE4E14D73454FADCFBB5" box="[1213,1344,1123,1149]" country="China" county="Libo County" municipality="Maolan" name="Wuyanqiao" pageId="27" pageNumber="208" stateProvince="Guizhou">Wuyanqiao</location>
,
<quantity id="4CBE7CD7B450CE4E15273455FA0BFBB5" box="[1357,1431,1122,1149]" metricMagnitude="2" metricUnit="m" metricValue="5.07" pageId="27" pageNumber="208" unit="m" value="507.0">
<elevation id="006B3601B450CE4E15273455FA0BFBB5" box="[1357,1431,1122,1149]" metricMagnitude="2" metricUnit="m" metricValue="5.07" pageId="27" pageNumber="208" unit="m" value="507.0">507 m</elevation>
</quantity>
, N 25°17′16″, E 108°4′18″,
<date id="FFF8F7F2B450CE4E11A334B0FDA2FB69" box="[457,574,1159,1185]" pageId="27" pageNumber="208" value="2020-05-23">
<collectingDate id="EFBC0E1AB450CE4E11A334B0FDA2FB69" box="[457,574,1159,1185]" pageId="27" pageNumber="208" value="2020-05-23">2020.V.23</collectingDate>
</date>
, at light,
<collectorName id="26B3B4E4B450CE4E12CD34B0FCFFFB69" box="[679,867,1159,1185]" pageId="27" pageNumber="208">Bao-Xiang Zhan</collectorName>
leg.”
</materialsCitation>
.
<materialsCitation id="3B2EDB6FB450CE4E13C734B0FA1DFB0D" collectingDate="2020-05-23" collectionCode="CZMZ" collectorName="Bao-Xiang Zhan" country="China" county="Libo County" elevation="507" location="Wuyanqiao" municipality="Maolan" pageId="27" pageNumber="208" specimenCount="1" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="B9320D20B450CE4E13C734B0FBBDFB69" bold="true" box="[941,1057,1159,1185]" pageId="27" pageNumber="208">
<typeStatus id="54FD6F90B450CE4E13C734B0FB80FB69" box="[941,1052,1159,1185]" pageId="27" pageNumber="208" type="paratype">Paratype</typeStatus>
.
</emphasis>
1♁ (
<collectionCode id="ED5749F7B450CE4E143034B0FB32FB69" box="[1114,1198,1159,1185]" pageId="27" pageNumber="208">CZMZ</collectionCode>
), “
<collectingCountry id="F35191A2B450CE4E14A134B0FAB6FB69" box="[1227,1322,1159,1185]" name="China" pageId="27" pageNumber="208">CHINA</collectingCountry>
:
<collectingRegion id="49821FD0B450CE4E155F34B0FA04FB69" box="[1333,1432,1159,1185]" country="China" name="Guizhou" pageId="27" pageNumber="208">Guizhou</collectingRegion>
,
<collectingCounty id="6298A9BEB450CE4E10FD349CFEBBFB0D" box="[151,295,1195,1221]" pageId="27" pageNumber="208">Libo County</collectingCounty>
,
<collectingMunicipality id="6B9D4B48B450CE4E1158349CFE16FB0D" box="[306,394,1195,1221]" pageId="27" pageNumber="208">Maolan</collectingMunicipality>
,
<location id="8E9987E9B450CE4E11FF349CFD84FB0D" LSID="urn:lsid:plazi:treatment:03EF6024B450CE4810FD33C1FED6FB21:8E9987E9B450CE4E11FF349CFD84FB0D" box="[405,536,1195,1221]" country="China" county="Libo County" municipality="Maolan" name="Wuyanqiao" pageId="27" pageNumber="208" stateProvince="Guizhou">Wuyanqiao</location>
,
<quantity id="4CBE7CD7B450CE4E1249349DFDF0FB0D" box="[547,620,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="5.07" pageId="27" pageNumber="208" unit="m" value="507.0">
<elevation id="006B3601B450CE4E1249349DFDF0FB0D" box="[547,620,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="5.07" pageId="27" pageNumber="208" unit="m" value="507.0">507 m</elevation>
</quantity>
, N 25°17′16″, E 108°4′18″,
<date id="FFF8F7F2B450CE4E13C3349CFB82FB0D" box="[937,1054,1195,1221]" pageId="27" pageNumber="208" value="2020-05-23">
<collectingDate id="EFBC0E1AB450CE4E13C3349CFB82FB0D" box="[937,1054,1195,1221]" pageId="27" pageNumber="208" value="2020-05-23">2020.V.23</collectingDate>
</date>
, at light,
<collectorName id="26B3B4E4B450CE4E14EC349CFADEFB0D" box="[1158,1346,1195,1221]" pageId="27" pageNumber="208">Bao-Xiang Zhan</collectorName>
leg.”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C82B9B450CE4810AD34F9FDD0FBFD" lastPageId="29" lastPageNumber="210" pageId="27" pageNumber="208" type="description">
<paragraph id="8BF9D132B450CE4E10AD34F9FDB3FA9D" blockId="27.[151,1437,1122,2013]" pageId="27" pageNumber="208">
<emphasis id="B9320D20B450CE4E10AD34F9FD08FB20" bold="true" box="[199,660,1230,1257]" pageId="27" pageNumber="208">
Description of the
<typeStatus id="54FD6F90B450CE4E11D934F8FD80FB21" box="[435,540,1231,1257]" pageId="27" pageNumber="208" type="holotype">holotype</typeStatus>
. General.
</emphasis>
Body ovoid, strongly convex, color generally reddish brown and iridescent, dorsal surface excluding pygidium with greenish toment; elytra, abdomen, and legs iridescent; palpi, antennae and legs reddish brown (excluding pro- and mesofemora, as well as coxae), shiny. Spiniform setae reddish brown, other setae yellowish brown.
</paragraph>
<paragraph id="8BF9D132B450CE4E10AD3568FCF9F951" blockId="27.[151,1437,1122,2013]" pageId="27" pageNumber="208">
<emphasis id="B9320D20B450CE4E10AD3568FE92FAB1" bold="true" box="[199,270,1375,1401]" pageId="27" pageNumber="208">Head.</emphasis>
Labroclypeus subtrapezoidal, ratio of maximum length/ width: 1/ 1.67; lateral margins straight and strongly convergent anteriad, anterior angle moderately curved, anterior margin straight; surface flat and dull, with moderately dense and large punctures on anterior third, otherwise with smaller and sparser punctures bearing a few minute, erect setae, behind anterior margin shiny; frontoclypeal suture finely incised, curved medially; smooth area anterior to eye large and flat, twice as wide as long; ocular canthus triangular (equal to 1/3 of ocular diameter). Frons flat, with scattered and small punctures, on each side with a short erect seta. Eyes small, ratio diameter/interocular width: 0.54. Antenna with ten antennomeres; antennomere 7 transverse and short; club with three antennomeres, slightly shorter than remaining antennomeres combined and straight.
<collectingRegion id="49821FD0B450CE4E13DA366CFB93F9BD" box="[944,1039,1627,1653]" country="Myanmar" name="Chin" pageId="27" pageNumber="208">Mentum</collectingRegion>
strongly and transversely elevated anteriorly. Labrum strongly produced, deeply concave medially.
</paragraph>
<paragraph id="8BF9D132B450CE4E10AD3694FC09F8E1" blockId="27.[151,1437,1122,2013]" pageId="27" pageNumber="208">
<emphasis id="B9320D20B450CE4E10AD3694FEDCF975" bold="true" box="[199,320,1699,1725]" pageId="27" pageNumber="208">Pronotum</emphasis>
subtrapezoidal, widest at base, lateral margins evenly curved and strongly convergent anteriad; anterior angles strongly produced and acute, posterior angles rectangular, rounded at apex; hypomeron carinate at base but not produced ventrally; anterior margin with a fine marginal line interrupted medially; surface with dense and large punctures, lateral margin with sparse and long, erect setae.
</paragraph>
<paragraph id="8BF9D132B450CE4E10AD3705FCFAF8B9" blockId="27.[151,1437,1122,2013]" pageId="27" pageNumber="208">
<emphasis id="B9320D20B450CE4E10AD3705FEA3F884" bold="true" box="[199,319,1842,1868]" pageId="27" pageNumber="208">Scutellum</emphasis>
large, triangular, lateral margin straight, apex obtuse; sparsely punctate basomedially and apically, otherwise with moderately dense and small punctures, glabrous.
</paragraph>
<paragraph id="8BF9D132B450CE4910AD374CFBBFFF1D" blockId="27.[151,1437,1122,2013]" lastBlockId="28.[151,1436,151,213]" lastPageId="28" lastPageNumber="209" pageId="27" pageNumber="208">
<emphasis id="B9320D20B450CE4E10AD374CFE8FF85D" bold="true" box="[199,275,1915,1941]" pageId="27" pageNumber="208">Elytra</emphasis>
oval, widest shortly behind middle. Striae strongly impressed, with dense and large strial punctures, scarcely bearing minute and recumbent setae; intervals weakly convex, with irregularly distributed large punctures concentrated along striae, otherwise with rather sparse punctures. Epipleura wide, ending shortly behind strongly curved external apical angle of elytron, with a row of sparse and short semi-erect setae; apex with a short spine, posterior margin with a very fine membranous rim composed of microtrichomes.
</paragraph>
<caption id="DF3981BAB457CE4910FD3655FE88F92E" ID-DOI="http://doi.org/10.5281/zenodo.10150469" ID-Zenodo-Dep="10150469" httpUri="https://zenodo.org/record/10150469/files/figure.png" pageId="28" pageNumber="209" startId="28.[151,250,1634,1658]" targetBox="[166,1431,252,1601]" targetPageId="28" targetType="figure">
<paragraph id="8BF9D132B457CE4910FD3655FE88F92E" blockId="28.[151,1437,1634,1766]" pageId="28" pageNumber="209">
<emphasis id="B9320D20B457CE4910FD3655FE82F9B2" bold="true" box="[151,286,1634,1658]" pageId="28" pageNumber="209">FIGURE 10.</emphasis>
Habitus in dorsal view.
<emphasis id="B9320D20B457CE4912633655FDBDF9B2" bold="true" box="[521,545,1634,1658]" pageId="28" pageNumber="209">A,</emphasis>
<taxonomicName id="4C46AAB1B457CE49124C3655FC09F9B2" authority="Zhao &amp;Ahrens" authorityName="Zhao &amp; Ahrens" authorityYear="2023" box="[550,917,1634,1659]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="209" phylum="Arthropoda" rank="species" species="zhanchaoi" status="sp. nov.">
<emphasis id="B9320D20B457CE49124C3655FD6AF9B2" box="[550,758,1634,1658]" italics="true" pageId="28" pageNumber="209">Maladera zhanchaoi</emphasis>
Zhao &amp;Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A201B05BB457CE4913F73654FB8BF9B2" box="[925,1047,1634,1658]" pageId="28" pageNumber="209" rank="species">new species</taxonomicNameLabel>
, holotype;
<emphasis id="B9320D20B457CE4914E93654FB06F9B3" bold="true" box="[1155,1178,1635,1659]" pageId="28" pageNumber="209">B,</emphasis>
<taxonomicName id="4C46AAB1B457CE4914F43654FF7FF956" authority="Zhao &amp; Ahrens" authorityName="Zhao &amp; Ahrens" authorityYear="2023" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="209" phylum="Arthropoda" rank="species" species="parabikouesis" status="new species">
<emphasis id="B9320D20B457CE4914F43654FAD0F9B2" box="[1182,1356,1634,1658]" italics="true" pageId="28" pageNumber="209">M. parabikouesis</emphasis>
Zhao &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A201B05BB457CE49108636B0FEFBF956" box="[236,359,1670,1694]" pageId="28" pageNumber="209" rank="species">new species</taxonomicNameLabel>
, holotype;
<emphasis id="B9320D20B457CE4911BE36B1FE71F956" bold="true" box="[468,493,1670,1694]" pageId="28" pageNumber="209">C,</emphasis>
<taxonomicName id="4C46AAB1B457CE49119836B0FC71F957" authority="Ahrens, Fabrizi &amp; Liu, 2021" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[498,1005,1670,1695]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="209" phylum="Arthropoda" rank="species" species="qianqingtangensis">
<emphasis id="B9320D20B457CE49119836B0FD50F956" box="[498,716,1671,1694]" italics="true" pageId="28" pageNumber="209">M. qianqingtangensis</emphasis>
<bibRefCitation id="EFD7ACC3B457CE4912BA36B1FC71F957" author="Fabrizi, S. &amp; Liu, W. G. &amp; Bai, M. &amp; Yang, X. K. &amp; Ahrens, D." box="[720,1005,1670,1695]" pageId="28" pageNumber="209" pagination="1 - 400" refId="ref28212" refString="Fabrizi, S., Liu, W. G., Bai, M., Yang, X. K. &amp; Ahrens, D. (2021) A monograph of the genus Maladera of China (Coleoptera, Scarabaeidae, Sericini). Zootaxa, 4922 (1), 1 - 400. https: // doi. org / 10.11646 / zootaxa. 4922.1.1" type="journal article" year="2021">Ahrens, Fabrizi &amp; Liu, 2021</bibRefCitation>
</taxonomicName>
, female (Fujian);
<emphasis id="B9320D20B457CE4914F436B0FB2AF957" bold="true" box="[1182,1206,1671,1695]" pageId="28" pageNumber="209">D,</emphasis>
<taxonomicName id="4C46AAB1B457CE4914D636B0FA0AF956" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[1212,1430,1671,1694]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="209" phylum="Arthropoda" rank="species" species="qianqingtangensis">
<emphasis id="B9320D20B457CE4914D636B0FA0AF956" box="[1212,1430,1671,1694]" italics="true" pageId="28" pageNumber="209">M. qianqingtangensis</emphasis>
</taxonomicName>
, male (Fujian);
<emphasis id="B9320D20B457CE491144369CFED9F90B" bold="true" box="[302,325,1707,1731]" pageId="28" pageNumber="209">E,</emphasis>
<taxonomicName id="4C46AAB1B457CE491126369CFD59F90A" authority="Zhao &amp; Ahrens" authorityName="Zhao &amp; Ahrens" authorityYear="2023" box="[332,709,1706,1731]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="209" phylum="Arthropoda" rank="species" species="shikengkongensis" status="new species">
<emphasis id="B9320D20B457CE491126369CFD83F90A" box="[332,543,1706,1730]" italics="true" pageId="28" pageNumber="209">M. shikengkongensis</emphasis>
Zhao &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A201B05BB457CE4912A5369CFCD0F90A" box="[719,844,1706,1730]" pageId="28" pageNumber="209" rank="species">new species</taxonomicNameLabel>
, holotype;
<emphasis id="B9320D20B457CE4913D6369CFC4CF90B" bold="true" box="[956,976,1707,1731]" pageId="28" pageNumber="209">F,</emphasis>
<taxonomicName id="4C46AAB1B457CE4913BC369CFB35F90A" authorityName="Zhao &amp; Ahrens" authorityYear="2023" box="[982,1193,1706,1730]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="209" phylum="Arthropoda" rank="species" species="shikengkongensis">
<emphasis id="B9320D20B457CE4913BC369CFB35F90A" box="[982,1193,1706,1730]" italics="true" pageId="28" pageNumber="209">M. shikengkongensis</emphasis>
</taxonomicName>
, female paratype. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8BF9D132B457CE4910AD370EFD1BF82B" blockId="28.[151,1436,1848,2019]" pageId="28" pageNumber="209">
<emphasis id="B9320D20B457CE4910AD370EFE7EF89A" bold="true" box="[199,482,1848,1875]" pageId="28" pageNumber="209">Ventral thoracic surface</emphasis>
. Hypomeron with moderately dense punctures each bearing a short seta; Other portions of ventral thoracic surface including metacoxae with dense and large punctures, mesosternum with recumbent short setae, metacoxa laterally bearing several short spiniform setae. Mesosternum between mesocoxae equal to width of mesofemur, somewhat vertically convex and truncate, with some moderately long and spiniform setae. Ratio of length of metepisternum/ metacoxa: 1/ 1.66.
</paragraph>
<paragraph id="8BF9D132B456CE4810AD30A1FBAFFE89" blockId="29.[151,1437,150,1257]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD30A1FEDFFF78" bold="true" box="[199,323,150,176]" pageId="29" pageNumber="210">Abdomen.</emphasis>
Abdominal ventrites with dense and large punctures, each ventrite with a transverse row of short and spiniform setae which is broadly interrupted medially but complete on penultimate ventrite and absent on ultimate ventrite; ultimate ventrite at middle almost half as long as penultimate one, with sparse and long setae, posterior margin with dense and minute setae. Pygidium weakly convex, posterior margin weakly reflexed; punctate and setose as on ultimate abdominal ventrite, but long setae only present near margins.
</paragraph>
<paragraph id="8BF9D132B456CE4810AD317CFC95FDD1" blockId="29.[151,1437,150,1257]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD317CFEB0FEAD" bold="true" box="[199,300,331,357]" pageId="29" pageNumber="210">Femora.</emphasis>
Profemur dull, with a row of long setae near dorsal margin, another longitudinal row of punctures presented medially, punctures each bearing a short seta. Mesofemur dull, two longitudinal rows of sparse punctures each bearing a short seta, with a ridge near anterior margin. Metafemur shiny and iridescent, with sparse punctures, with two short and spiniform setae preapically; anterior margin sharp, ventral posterior margin carinate and strongly produced at apical half, dorsal posterior margin finely and completely serrate, a sparse row of moderately long and spiniform setae situated between the latter two margins.
</paragraph>
<paragraph id="8BF9D132B456CE4810AD3215FD71FD39" blockId="29.[151,1437,150,1257]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD3215FE86FDF4" bold="true" box="[199,282,546,572]" pageId="29" pageNumber="210">Tibiae.</emphasis>
Protibia stout, bidentate. Metatibia widest at middle, ratio of width/ length: 1/ 2.64; dorsal margin carinate at apical two fifths; lateral face with sparse and small punctures, dorsally with two groups of spiniform setae not touching the dorsal margin, the basal group at two fifths, distal group at two thirds of metatibial length, basal quarter with a short seta; ventral margin convex, serrate at apical two fifth, with three long, spiniform and equidistant setae, basally with several minute setae and a short spiniform seta; medial face glabrous; apex interiorly serrate, near tarsal articulation concavely emarginate.
</paragraph>
<paragraph id="8BF9D132B456CE4810AD32CDFDF9FC49" blockId="29.[151,1437,150,1257]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD32CDFE90FCDC" bold="true" box="[199,268,762,788]" pageId="29" pageNumber="210">Tarsi.</emphasis>
Protarsomeres short, pro- and mesotarsomeres with sparse and minute setae ventrally.All metatarsomeres carinate ventrolaterally, strongly serrate ventrally, first metatarsomere slightly shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protarsal claws symmetric, the distal and basal teeth equal in length, basal tooth slightly wider than distal one.
</paragraph>
<paragraph id="8BF9D132B456CE4810AD33BCFE75FC6D" blockId="29.[151,1437,150,1257]" box="[199,489,906,933]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD33BCFEE5FC6C" bold="true" box="[199,377,906,933]" pageId="29" pageNumber="210">Male genitalia.</emphasis>
<figureCitation id="137DCDB7B456CE48111533BCFE7AFC6D" box="[383,486,907,933]" captionStart="FIGURE 11" captionStartId="30.[151,250,1336,1360]" captionTargetBox="[164,1424,202,1298]" captionTargetId="figure-18@30.[151,1436,181,1312]" captionTargetPageId="30" captionText="FIGURE 11. Aedeagus. A, Maladera zhanchaoi Zhao &amp; Ahrens, new species, holotype; B, M. parabikouesis Zhao &amp; Ahrens, new species, holotype; C, M. shikengkongensis Zhao &amp; Ahrens, new species, holotype; D, M. qianqingtangensis Ahrens, Fabrizi &amp; Liu, 2021 (Fujian). a, aedeagus in left lateral view; b, parameres in dorsal view; c, aedeagus in right lateral view; d, aedeagus in dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10150471" httpUri="https://zenodo.org/record/10150471/files/figure.png" pageId="29" pageNumber="210">Fig. 11A</figureCitation>
.
</paragraph>
<paragraph id="8BF9D132B456CE4810AD3399FC1AFC01" blockId="29.[151,1437,150,1257]" box="[199,902,942,969]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD3399FE15FC01" bold="true" box="[199,393,942,969]" pageId="29" pageNumber="210">Variation. Male.</emphasis>
The only
<typeStatus id="54FD6F90B456CE4811913398FDC7FC01" box="[507,603,943,969]" pageId="29" pageNumber="210" type="paratype">paratype</typeStatus>
is generally more reddish.
</paragraph>
<paragraph id="8BF9D132B456CE4810AD33E4FE3EFC25" blockId="29.[151,1437,150,1257]" box="[199,418,979,1005]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD33E4FEB9FC25" bold="true" box="[199,293,979,1005]" pageId="29" pageNumber="210">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BF9D132B456CE4810AD33C0FDD0FBFD" blockId="29.[151,1437,150,1257]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD33C0FEE2FBD9" bold="true" box="[199,382,1015,1041]" pageId="29" pageNumber="210">Measurements.</emphasis>
Length:
<quantity id="4CBE7CD7B456CE48118933C0FD1AFBD9" box="[483,646,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="1.2399999999999998" metricValueMax="1.26" metricValueMin="1.22" pageId="29" pageNumber="210" unit="mm" value="12.399999999999999" valueMax="12.6" valueMin="12.2">12.2–12.6 mm</quantity>
(
<typeStatus id="54FD6F90B456CE4812FE33C0FD64FBD9" box="[660,760,1015,1041]" pageId="29" pageNumber="210" type="holotype">holotype</typeStatus>
<quantity id="4CBE7CD7B456CE48129533C0FCFBFBD9" box="[767,871,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="29" pageNumber="210" unit="mm" value="12.6">12.6 mm</quantity>
); length of elytra:
<quantity id="4CBE7CD7B456CE48145F33C0FB20FBD9" box="[1077,1212,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="8.95" metricValueMax="9.1" metricValueMin="8.8" pageId="29" pageNumber="210" unit="mm" value="8.95" valueMax="9.1" valueMin="8.8">8.8–9.1 mm</quantity>
(
<typeStatus id="54FD6F90B456CE4814A033C0FAB2FBD9" box="[1226,1326,1015,1041]" pageId="29" pageNumber="210" type="holotype">holotype</typeStatus>
<quantity id="4CBE7CD7B456CE48155F33C0FA0CFBD9" box="[1333,1424,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="29" pageNumber="210" unit="mm" value="9.1">9.1 mm</quantity>
); width: 6.9–7.0 mm (
<typeStatus id="54FD6F90B456CE481111342CFE43FBFD" box="[379,479,1051,1077]" pageId="29" pageNumber="210" type="holotype">holotype</typeStatus>
7.0 mm).
</paragraph>
</subSubSection>
<subSubSection id="C35C82B9B456CE4810AD3409FA88FB69" pageId="29" pageNumber="210" type="diagnosis">
<paragraph id="8BF9D132B456CE4810AD3409FA88FB69" blockId="29.[151,1437,150,1257]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD3409FEDEFB90" bold="true" box="[199,322,1086,1112]" pageId="29" pageNumber="210">Diagnosis.</emphasis>
<taxonomicName id="4C46AAB1B456CE4811203408FD6DFB91" authority="Zhao &amp; Ahrens" authorityName="Zhao &amp; Ahrens" authorityYear="2023" box="[330,753,1087,1113]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="210" phylum="Arthropoda" rank="species" species="zhanchaoi" status="sp. nov.">
<emphasis id="B9320D20B456CE4811203408FDA8FB91" box="[330,564,1087,1113]" italics="true" pageId="29" pageNumber="210">Maladera zhanchaoi</emphasis>
Zhao &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A201B05BB456CE4812973408FC1AFB91" box="[765,902,1087,1113]" pageId="29" pageNumber="210" rank="species">new species</taxonomicNameLabel>
is similar to the species of the
<taxonomicName id="4C46AAB1B456CE4814853408FA00FB91" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[1263,1436,1087,1113]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="210" phylum="Arthropoda" rank="species" species="granuligera">
<emphasis id="B9320D20B456CE4814853408FA00FB91" box="[1263,1436,1087,1113]" italics="true" pageId="29" pageNumber="210">M. granuligera</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EFD7ACC3B456CE4810823454FE6DFBB5" author="Ahrens, D. &amp; Fabrizi, S." box="[232,497,1123,1149]" pageId="29" pageNumber="210" pagination="1 - 355" refId="ref27788" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn zoological Bulletin, 65 (1 - 2), 1 - 355." type="journal article" year="2016">Ahrens &amp; Fabrizi 2016</bibRefCitation>
) whose species possess a dorsal apophysis at apical phallobase. The new species is easily distinguished from the so far known species by the extremely widened parameres (lateral view).
</paragraph>
</subSubSection>
<subSubSection id="C35C82B9B456CE4810AD349CFED6FB21" pageId="29" pageNumber="210" type="etymology">
<paragraph id="8BF9D132B456CE4810AD349CFED6FB21" blockId="29.[151,1437,150,1257]" pageId="29" pageNumber="210">
<emphasis id="B9320D20B456CE4810AD349CFED2FB0D" bold="true" box="[199,334,1195,1221]" pageId="29" pageNumber="210">Etymology.</emphasis>
The species is dedicated to Mr. Chao Zhan, the father of Mr. Bao-Xiang Zhan, who greatly supported his field survey.
</paragraph>
</subSubSection>
</treatment>
</document>
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