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<document id="DC6AD6618C60546DBD01711DD35CB561" ID-DOI="10.5281/zenodo.5320566" ID-ISSN="0374-1036" ID-Zenodo-Dep="5320566" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630136727139" checkinUser="felipe" docAuthor="Toledano, Luca" docDate="2009" docId="03EF20294A0C9F57FEBFFCF3373DFDC4" docLanguage="en" docName="ActaEntMusNatPra.2009.49.2.577-598.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Bembidion (Blepharoplataphus) teradai Toledano 2009, sp. nov." docType="treatment" docVersion="4" lastPageNumber="585" masterDocId="FFD658514A099F5FFFEFFFDF3359FF97" masterDocTitle="Notes on the Bembidiina of Taiwan with description of three new species (Coleoptera: Carabidae)" masterLastPageNumber="598" masterPageNumber="577" pageNumber="582" updateTime="1699163666101" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F19414AA23F44BB65A8E93C42220E6C7">Notes on the Bembidiina of Taiwan with description of three new species (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="CE5DF6D3682E90E6C3223AD2BB2F6131">Toledano, Luca</mods:namePart>
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<mods:title id="8BD757DBCE21636C25EB99C1C9F4C4E7">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="97312AE33C90649EF697D48EE1F1DB04">2009</mods:date>
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<mods:number id="605C91CB02956E65EF5D67859B429F33">2009-12-15</mods:number>
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<treatment id="03EF20294A0C9F57FEBFFCF3373DFDC4" ID-DOI="http://doi.org/10.5281/zenodo.5342699" ID-Zenodo-Dep="5342699" LSID="urn:lsid:plazi:treatment:03EF20294A0C9F57FEBFFCF3373DFDC4" httpUri="http://treatment.plazi.org/id/03EF20294A0C9F57FEBFFCF3373DFDC4" lastPageId="8" lastPageNumber="585" pageId="5" pageNumber="582">
<subSubSection id="C35CC2B44A0C9F5AFEBFFCF3303BFCD2" box="[336,866,812,837]" pageId="5" pageNumber="582" type="nomenclature">
<paragraph id="8BF9913F4A0C9F5AFEBFFCF3303BFCD2" blockId="5.[336,866,812,837]" box="[336,866,812,837]" pageId="5" pageNumber="582">
<heading id="D0B126534A0C9F5AFEBFFCF3303BFCD2" bold="true" box="[336,866,812,837]" centered="true" fontSize="8" level="2" pageId="5" pageNumber="582" reason="3">
<emphasis id="B9324D2D4A0C9F5AFEBFFCF3303BFCD2" bold="true" box="[336,866,812,837]" pageId="5" pageNumber="582">
<taxonomicName id="4C46EABC4A0C9F5AFEBFFCF3305FFCD2" ID-CoL="LDQD" authority="Toledano, 2009" authorityName="Toledano" authorityYear="2009" box="[336,774,812,837]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="582" phylum="Arthropoda" rank="species" species="teradai" status="sp. nov." subGenus="Blepharoplataphus">
<emphasis id="B9324D2D4A0C9F5AFEBFFCF33291FCD2" bold="true" box="[336,456,812,837]" italics="true" pageId="5" pageNumber="582">Bembidion</emphasis>
(
<emphasis id="B9324D2D4A0C9F5AFE38FCF331F0FCD2" bold="true" box="[471,681,812,837]" italics="true" pageId="5" pageNumber="582">Blepharoplataphus</emphasis>
)
<emphasis id="B9324D2D4A0C9F5AFD56FCF3305FFCD2" bold="true" box="[697,774,812,837]" italics="true" pageId="5" pageNumber="582">teradai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201F0564A0C9F5AFCE2FCF2303BFCD2" box="[781,866,813,837]" pageId="5" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35CC2B44A0C9F5AFDCAFC9431D4FCC8" box="[549,653,843,863]" pageId="5" pageNumber="582" type="description">
<paragraph id="8BF9913F4A0C9F5AFDCAFC9431D4FCC8" blockId="5.[549,653,843,863]" box="[549,653,843,863]" pageId="5" pageNumber="582">
(
<figureCitation id="137D8DBA4A0C9F5AFDC4FC94313EFCC9" box="[555,615,843,862]" captionStart="Figs" captionStartId="6.[151,190,1518,1537]" captionTargetBox="[210,1085,225,1493]" captionTargetId="figure-10@6.[193,1085,219,1493]" captionTargetPageId="6" captionText="Figs.15. Bembidion spp., habitus. 1 B. (Armatocillenus) sauteri Jedlička, 1954, holotype; 2 B. (Desarmatocillenus) foochowense Lindroth, 1980; 3 B. (Odontium) fusiforme Netolitzky, 1914; 4 B. (Plataphus) takasagonis Habu, 1973; 5 B. (Blepharoplataphus) teradai sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5320570" httpUri="https://zenodo.org/record/5320570/files/figure.png" pageId="5" pageNumber="582">Figs. 5</figureCitation>
,
<figureCitation id="137D8DBA4A0C9F5AFD9EFC9331DEFCC8" box="[625,647,844,863]" captionStart="Figs" captionStartId="14.[151,190,1436,1455]" captionTargetBox="[221,1058,219,1412]" captionTargetId="figure-145@14.[219,1058,219,1417]" captionTargetPageId="14" captionText="Figs. 1728. Bembidion and Sinechostictus spp., median lobe of the aedeagus. 17 B. (Desarmatocillenus) foochowense Lindroth, 1980; 18 B. (Odontium) fusiforme Netolitzky, 1914; 19 B. (Plataphus) takasagonis Habu, 1973; 20 B. (Blepharoplataphus) teradai sp. nov., holotype; 21 B. (Trichoplataphus) miwai Jedlička, 1946; 22 B. (Trichoplataphus) taiwanum Netolitzky, 1939; 23 B. (Trichoplataphus) jelineki sp. nov., holotype; 24 B. (Trichoplataphus) seijii sp. nov., holotype; 25 B. (Ocydromus) chloreum Bates, 1873, specimen from Taiwan; 26 B. (Ocydromus) shunichii Habu, 1973; 27 B. (Ocydromus) peleum Jedlička, 1933, specimen from Taiwan; 28 S. (Pseudolimnaeum) chuji Jedlička, 1951." figureDoi="http://doi.org/10.5281/zenodo.5320580" httpUri="https://zenodo.org/record/5320580/files/figure.png" pageId="5" pageNumber="582">20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35CC2B44A0C9F5AFF9EFCAA3186FB99" pageId="5" pageNumber="582" type="materials_examined">
<paragraph id="8BF9913F4A0C9F5AFF9EFCAA3167FC1F" blockId="5.[113,1089,884,1038]" box="[113,574,884,904]" pageId="5" pageNumber="582">
<emphasis id="B9324D2D4A0C9F5AFF9EFCAA33B3FC1F" bold="true" box="[113,234,884,904]" pageId="5" pageNumber="582">
<typeStatus id="54FD2F9D4A0C9F5AFF9EFCAA33C7FC1F" box="[113,158,885,904]" pageId="5" pageNumber="582">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F351D1AF4A0C9F5AFF00FCAA3269FC1F" box="[239,304,885,904]" name="Taiwan" pageId="5" pageNumber="582">Taiwan</collectingCountry>
, Nansichi, Wulai,
<collectingRegion id="49825FDD4A0C9F5AFE23FCAA3158FC1F" box="[460,513,885,904]" country="Taiwan" name="Taipei" pageId="5" pageNumber="582">Taipei</collectingRegion>
Hsien.
</paragraph>
<paragraph id="8BF9913F4A0C9F5AFF9EFC4F3186FB99" blockId="5.[113,1089,884,1038]" pageId="5" pageNumber="582">
<emphasis id="B9324D2D4A0C9F5AFF9EFC4F33AAFC34" bold="true" box="[113,243,911,931]" pageId="5" pageNumber="582">Type material.</emphasis>
<materialsCitation id="3B2E9B624A0C9F5AFF18FC4F30E6FC34" box="[247,959,907,932]" collectingDate="2001-12-23" country="Taiwan" county="Nansichi" location="Hsien" municipality="Wulai" pageId="5" pageNumber="582" specimenCount="1" stateProvince="Taipei" typeStatus="holotype">
<typeStatus id="54FD2F9D4A0C9F5AFF18FC4F3210FC35" box="[247,329,912,931]" pageId="5" pageNumber="582" type="holotype">HOLOTYPE</typeStatus>
: J,
<collectingCountry id="F351D1AF4A0C9F5AFE80FC4F329EFC34" box="[367,455,912,931]" name="Taiwan" pageId="5" pageNumber="582">TAIWAN</collectingCountry>
,
<date id="FFF8B7FF4A0C9F5AFE21FC4F3168FC34" box="[462,561,912,931]" pageId="5" pageNumber="582" value="2001-12-23">
<collectingDate id="EFBC4E174A0C9F5AFE21FC4F3168FC34" box="[462,561,912,931]" pageId="5" pageNumber="582" value="2001-12-23">23-xii-2001</collectingDate>
</date>
/
<collectingCounty id="6298E9B34A0C9F5AFDD0FC4F31D5FC34" box="[575,652,912,931]" pageId="5" pageNumber="582">Nansichi</collectingCounty>
,
<collectingMunicipality id="6B9D0B454A0C9F5AFD7CFC4F319FFC34" box="[659,710,912,931]" pageId="5" pageNumber="582">Wulai</collectingMunicipality>
/
<collectingRegion id="49825FDD4A0C9F5AFD3CFC4F305EFC34" box="[723,775,912,931]" country="Taiwan" name="Taipei" pageId="5" pageNumber="582">Taipei</collectingRegion>
<location id="8E99C7E44A0C9F5AFCE4FC4F3065FC34" LSID="urn:lsid:plazi:treatment:03EF20294A0C9F57FEBFFCF3373DFDC4:8E99C7E44A0C9F5AFCE4FC4F3065FC34" box="[779,828,912,931]" country="Taiwan" county="Nansichi" municipality="Wulai" name="Hsien" pageId="5" pageNumber="582" stateProvince="Taipei">Hsien</location>
/
<location id="8E99C7E44A0C9F5AFCA6FC4F30DBFC34" LSID="urn:lsid:plazi:treatment:03EF20294A0C9F57FEBFFCF3373DFDC4:8E99C7E44A0C9F5AFCA6FC4F30DBFC34" box="[841,898,912,931]" country="Taiwan" county="Nansichi" municipality="Wulai" name="Terada" pageId="5" pageNumber="582" stateProvince="Taipei">Terada</location>
77 //
</materialsCitation>
<typeStatus id="54FD2F9D4A0C9F5AFC2CFC4F3718FC34" box="[963,1089,912,931]" pageId="5" pageNumber="582" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="4C46EABC4A0C9F5AFF91FC743244FC29" authority="Toledano, 2009" authorityName="Toledano" authorityYear="2009" box="[126,285,939,958]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="582" phylum="Arthropoda" rank="species" species="teradai" status="sp. nov.">Bembidion teradai</taxonomicName>
<taxonomicNameLabel id="A201F0564A0C9F5AFECCFC743215FC29" box="[291,332,939,958]" pageId="5" pageNumber="582" rank="species">sp.n.</taxonomicNameLabel>
/ Det. L. Toledano, 2009 [red] (
<collectionCode id="ED5709FA4A0C9F5AFD87FC7431F3FC29" box="[616,682,939,958]" country="Italy" httpUri="http://grbio.org/cool/vmav-1g60" name="Museo Civico di Storia Naturale di Venezia" pageId="5" pageNumber="582">MSNV</collectionCode>
).
<materialsCitation id="3B2E9B624A0C9F5AFD56FC7432BBFC4F" collectingDate="2001-12-23" collectionCode="NTUC, CTHJ, CTVR" country="Taiwan" county="Nansichi" location="Hsien" municipality="Wulai" pageId="5" pageNumber="582" specimenCount="9" specimenCount-female="9" stateProvince="Taipei" typeStatus="paratype">
<typeStatus id="54FD2F9D4A0C9F5AFD56FC743048FC2B" box="[697,785,939,958]" pageId="5" pageNumber="582" type="paratype">PARATYPES</typeStatus>
: 6 JJ
<specimenCount id="9D405AB64A0C9F5AFCBCFC7430DAFC29" box="[851,899,933,958]" pageId="5" pageNumber="582" type="female">9 ♀♀</specimenCount>
, same date and locality as the holotype (
<collectionCode id="ED5709FA4A0C9F5AFEF4FC1A3201FC4F" box="[283,344,965,984]" pageId="5" pageNumber="582">NTUC</collectionCode>
,
<collectionCode id="ED5709FA4A0C9F5AFE8DFC1A32CEFC4F" box="[354,407,965,984]" pageId="5" pageNumber="582">CTHJ</collectionCode>
,
<collectionCode id="ED5709FA4A0C9F5AFE4FFC1A3284FC4F" box="[416,477,965,984]" pageId="5" pageNumber="582">CTVR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2E9B624A0C9F5AFE04FC1A33BDFC64" collectingDate="2001-12-04" collectionCode="CMTJ" collectorName="K. Terada" country="Taiwan" county="Wulai" location="Taiwan" municipality="Hsien" pageId="5" pageNumber="582" specimenCount="6" specimenCount-female="6" stateProvince="Taipei" typeStatus="paratype">
2 JJ
<specimenCount id="9D405AB64A0C9F5AFDCEFC1A3108FC4F" box="[545,593,959,984]" pageId="5" pageNumber="582" type="female">6 ♀♀</specimenCount>
,
<collectingCounty id="6298E9B34A0C9F5AFDB0FC1A31CFFC4F" box="[607,662,965,984]" pageId="5" pageNumber="582">Wulai</collectingCounty>
/
<collectingRegion id="49825FDD4A0C9F5AFD48FC1A3185FC4F" box="[679,732,965,984]" country="Taiwan" name="Taipei" pageId="5" pageNumber="582">Taipei</collectingRegion>
<collectingMunicipality id="6B9D0B454A0C9F5AFD0DFC1A304DFC4F" box="[738,788,965,984]" pageId="5" pageNumber="582">Hsien</collectingMunicipality>
/
<location id="8E99C7E44A0C9F5AFCCBFC1A303BFC4F" LSID="urn:lsid:plazi:treatment:03EF20294A0C9F57FEBFFCF3373DFDC4:8E99C7E44A0C9F5AFCCBFC1A303BFC4F" box="[804,866,965,984]" country="Taiwan" county="Wulai" municipality="Hsien" name="Taiwan" pageId="5" pageNumber="582" stateProvince="Taipei">Taiwan</location>
//
<date id="FFF8B7FF4A0C9F5AFC95FC1A3082FC4F" box="[890,987,965,984]" pageId="5" pageNumber="582" value="2001-12-04">
<collectingDate id="EFBC4E174A0C9F5AFC95FC1A3082FC4F" box="[890,987,965,984]" pageId="5" pageNumber="582" value="2001-12-04">4-XII-2001</collectingDate>
</date>
/
<collectorName id="26B3F4E94A0C9F5AFC02FC193718FC4F" box="[1005,1089,965,985]" pageId="5" pageNumber="582">K. Terada</collectorName>
leg. (
<collectionCode id="ED5709FA4A0C9F5AFF4CFC3F3387FC64" box="[163,222,992,1011]" pageId="5" pageNumber="582">CMTJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2E9B624A0C9F5AFF01FC3F3718FC64" box="[238,1089,986,1012]" collectingDate="2001-12-23" collectionCode="CMTJ, CTVR" collectorName="K. Terada &amp; All" country="Taiwan" location="Wulai Taiwan" pageId="5" pageNumber="582" specimenCount="1" specimenCount-female="1" stateProvince="Taipei" typeStatus="paratype">
1 J
<specimenCount id="9D405AB64A0C9F5AFEFAFC3F326CFC64" box="[277,309,986,1011]" pageId="5" pageNumber="582" type="female">1 ♀</specimenCount>
,
<location id="8E99C7E44A0C9F5AFEABFC3F32E6FC64" LSID="urn:lsid:plazi:treatment:03EF20294A0C9F57FEBFFCF3373DFDC4:8E99C7E44A0C9F5AFEABFC3F32E6FC64" box="[324,447,992,1011]" country="Taiwan" name="Wulai Taiwan" pageId="5" pageNumber="582" stateProvince="Taipei">Wulai Taiwan</location>
/
<date id="FFF8B7FF4A0C9F5AFE3CFC3F3167FC64" box="[467,574,992,1011]" pageId="5" pageNumber="582" value="2001-12-23">
<collectingDate id="EFBC4E174A0C9F5AFE3CFC3F3167FC64" box="[467,574,992,1011]" pageId="5" pageNumber="582" value="2001-12-23">23-XII-2001</collectingDate>
</date>
/
<collectorName id="26B3F4E94A0C9F5AFDBEFC3F31FFFC64" box="[593,678,992,1011]" pageId="5" pageNumber="582">K. Terada</collectorName>
leg. (
<collectionCode id="ED5709FA4A0C9F5AFD32FC3F304EFC64" box="[733,791,992,1011]" pageId="5" pageNumber="582">CMTJ</collectionCode>
,
<collectionCode id="ED5709FA4A0C9F5AFCCEFC3F3007FC64" box="[801,862,992,1011]" pageId="5" pageNumber="582">CTVR</collectionCode>
).
<collectorName id="26B3F4E94A0C9F5AFC84FC3F30DFFC64" box="[875,902,992,1011]" pageId="5" pageNumber="582">All</collectorName>
<typeStatus id="54FD2F9D4A0C9F5AFC62FC3F3086FC64" box="[909,991,992,1011]" pageId="5" pageNumber="582" type="paratype">paratypes</typeStatus>
labelled as
</materialsCitation>
<typeStatus id="54FD2F9D4A0C9F5AFF9BFC2433A9FB99" box="[116,240,1019,1038]" pageId="5" pageNumber="582" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="4C46EABC4A0C9F5AFEEEFC2432C6FB99" authority="Toledano, 2009" authorityName="Toledano" authorityYear="2009" box="[257,415,1019,1038]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="582" phylum="Arthropoda" rank="species" species="teradai" status="sp. nov.">Bembidion teradai</taxonomicName>
<taxonomicNameLabel id="A201F0564A0C9F5AFE4AFC243297FB99" box="[421,462,1019,1038]" pageId="5" pageNumber="582" rank="species">sp.n.</taxonomicNameLabel>
/ Det. L. Toledano, 2009 [red].
</paragraph>
</subSubSection>
<subSubSection id="C35CC2B44A0C9F5AFF9EFBF9320DFB0A" pageId="5" pageNumber="582" type="diagnosis">
<paragraph id="8BF9913F4A0C9F5AFF9EFBF9320DFB0A" blockId="5.[113,1089,1062,1598]" pageId="5" pageNumber="582">
<emphasis id="B9324D2D4A0C9F5AFF9EFBF933BFFBA9" bold="true" box="[113,230,1062,1086]" pageId="5" pageNumber="582">Diagnosis.</emphasis>
A
<taxonomicName id="4C46EABC4A0C9F5AFEE5FBF8328FFBA9" authorityName="Netolitzky" authorityYear="1920" box="[266,470,1063,1086]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="597" phylum="Arthropoda" rank="subGenus" subGenus="Blepharoplataphus">
<emphasis id="B9324D2D4A0C9F5AFEE5FBF8328FFBA9" box="[266,470,1063,1086]" italics="true" pageId="5" pageNumber="582">Blepharoplataphus</emphasis>
</taxonomicName>
species with very wide elytra and humeri, very similar to
<taxonomicName id="4C46EABC4A0C9F5AFF60FB983154FBC9" authority="Morita, 2008" authorityName="Morita" authorityYear="2008" box="[143,525,1094,1118]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="582" phylum="Arthropoda" rank="species" species="nirasawai">
<emphasis id="B9324D2D4A0C9F5AFF60FB98322EFBC9" box="[143,375,1095,1118]" italics="true" pageId="5" pageNumber="582">Bembidion nirasawai</emphasis>
<bibRefCitation id="EFD7ECCE4A0C9F5AFE6FFB993154FBC9" author="MORITA S." box="[384,525,1094,1118]" pageId="5" pageNumber="582" pagination="273 - 277" refId="ref11042" refString="MORITA S. 2008: Notes on the Bembidiinae (Coleoptera: Carabidae) of Japan XIV. Two new species of the Subgenus Blepharoplataphus. Entomological Revue of Japan 62: 273 - 277." type="journal article" year="2008">Morita, 2008</bibRefCitation>
</taxonomicName>
from southwestern
<collectingCountry id="F351D1AF4A0C9F5AFD05FB993070FBCA" box="[746,809,1094,1117]" name="Japan" pageId="5" pageNumber="582">Japan</collectingCountry>
, but distinguishable from this last for the apex of the aedeagus, rounded in
<taxonomicName id="4C46EABC4A0C9F5AFD6FFBB830F4FBE9" authority="(MORITA 2008)" baseAuthorityName="MORITA" baseAuthorityYear="2008" box="[640,941,1126,1150]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="582" phylum="Arthropoda" rank="species" species="nirasawai">
<emphasis id="B9324D2D4A0C9F5AFD6FFBB83050FBE9" box="[640,777,1127,1150]" italics="true" pageId="5" pageNumber="582">B. nirasawai</emphasis>
(
<bibRefCitation id="EFD7ECCE4A0C9F5AFCF8FBB930FFFBE9" author="MORITA S." box="[791,934,1126,1150]" pageId="5" pageNumber="582" pagination="273 - 277" refId="ref11042" refString="MORITA S. 2008: Notes on the Bembidiinae (Coleoptera: Carabidae) of Japan XIV. Two new species of the Subgenus Blepharoplataphus. Entomological Revue of Japan 62: 273 - 277." type="journal article" year="2008">MORITA 2008</bibRefCitation>
)
</taxonomicName>
and ventrally hooked in
<taxonomicName id="4C46EABC4A0C9F5AFF0DFB583214FB09" authorityName="Toledano" authorityYear="2009" box="[226,333,1159,1182]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="582" phylum="Arthropoda" rank="species" species="teradai">
<emphasis id="B9324D2D4A0C9F5AFF0DFB583214FB09" box="[226,333,1159,1182]" italics="true" pageId="5" pageNumber="582">B. teradai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CC2B44A0C9F57FF9EFB79307EFEE4" lastPageId="8" lastPageNumber="585" pageId="5" pageNumber="582" type="description">
<paragraph id="8BF9913F4A0C9F5AFF9EFB7932AAFB49" blockId="5.[113,1089,1062,1598]" pageId="5" pageNumber="582">
<emphasis id="B9324D2D4A0C9F5AFF9EFB7933A5FB29" bold="true" box="[113,252,1190,1214]" pageId="5" pageNumber="582">Description.</emphasis>
Body length
<quantity id="4CBE3CDA4A0C9F5AFE68FB79317AFB2A" box="[391,547,1190,1214]" metricMagnitude="-3" metricUnit="m" metricValue="5.609999999999999" metricValueMax="5.84" metricValueMin="5.38" pageId="5" pageNumber="582" unit="mm" value="5.609999999999999" valueMax="5.84" valueMin="5.38">5.385.84 mm</quantity>
. Black with faint metallic greenish reflections. Legs and antennae piceous black (
<figureCitation id="137D8DBA4A0C9F5AFE4BFB1932BCFB49" box="[420,485,1222,1246]" captionStart="Figs" captionStartId="6.[151,190,1518,1537]" captionTargetBox="[210,1085,225,1493]" captionTargetId="figure-10@6.[193,1085,219,1493]" captionTargetPageId="6" captionText="Figs.15. Bembidion spp., habitus. 1 B. (Armatocillenus) sauteri Jedlička, 1954, holotype; 2 B. (Desarmatocillenus) foochowense Lindroth, 1980; 3 B. (Odontium) fusiforme Netolitzky, 1914; 4 B. (Plataphus) takasagonis Habu, 1973; 5 B. (Blepharoplataphus) teradai sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5320570" httpUri="https://zenodo.org/record/5320570/files/figure.png" pageId="5" pageNumber="582">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BF9913F4A0C9F5AFF7FFB39317EFA89" blockId="5.[113,1089,1062,1598]" pageId="5" pageNumber="582">Head with deep, wide and parallel frontal furrows, not extending on clypeus. Eyes convex, antennae rather short (bl/al = 1.982.09).</paragraph>
<paragraph id="8BF9913F4A0C9F5AFF7FFAF930ECFA29" blockId="5.[113,1089,1062,1598]" pageId="5" pageNumber="582">Pronotum transverse (pw/pl = 1.421.48), rather narrow in relation to elytra (ew/pw = 1.751.76). Sides rounded, briefly sinuate before the short, slightly obtuse hind angles. Lateral channel very wide, anterior transverse impression short, in some specimens with some punctures, median line superficial, basal foveae somewhat square, very rugose, laterally delimited by a carina. Posterior margin straight in middle and oblique at sides.</paragraph>
<paragraph id="8BF9913F4A0C9F5AFF7FFA193203F9A9" blockId="5.[113,1089,1062,1598]" pageId="5" pageNumber="582">Elytra (el/ew = 1.401.43) oval, very wide and convex, with humeri wide and rather rounded. Elytral striae 17 complete, visibly impressed, punctate-sulcate, scutellar stria long, apical stria long and connected with apical end of stria 5. Two discal elytral pores in interval 3 near stria 3.</paragraph>
<caption id="DF39C1B74A0F9F59FF78FA31318CF9A0" ID-DOI="http://doi.org/10.5281/zenodo.5320570" ID-Zenodo-Dep="5320570" httpUri="https://zenodo.org/record/5320570/files/figure.png" pageId="6" pageNumber="583" startId="6.[151,190,1518,1537]" targetBox="[210,1085,225,1493]" targetPageId="6">
<paragraph id="8BF9913F4A0F9F59FF78FA31318CF9A0" blockId="6.[151,1129,1518,1591]" pageId="6" pageNumber="583">
Figs.15.
<taxonomicName id="4C46EABC4A0F9F59FF00FA30322CF995" box="[239,373,1519,1538]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="583" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9324D2D4A0F9F59FF00FA303212F996" box="[239,331,1519,1537]" italics="true" pageId="6" pageNumber="583">Bembidion</emphasis>
spp.
</taxonomicName>
, habitus. 1
<taxonomicName id="4C46EABC4A0F9F59FE0FFA30301EF996" authority="Jedlicka, 1954" authorityName="Jedlicka" authorityYear="1954" box="[480,839,1518,1538]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="583" phylum="Arthropoda" rank="species" species="sauteri" subGenus="Armatocillenus">
<emphasis id="B9324D2D4A0F9F59FE0FFA3032ADF996" box="[480,500,1519,1537]" italics="true" pageId="6" pageNumber="583">B.</emphasis>
(
<emphasis id="B9324D2D4A0F9F59FE11FA3031D9F996" box="[510,640,1519,1537]" italics="true" pageId="6" pageNumber="583">Armatocillenus</emphasis>
)
<emphasis id="B9324D2D4A0F9F59FD64FA30319FF996" box="[651,710,1519,1537]" italics="true" pageId="6" pageNumber="583">sauteri</emphasis>
<bibRefCitation id="EFD7ECCE4A0F9F59FD25FA31301EF996" author="JEDLICKA A." box="[714,839,1518,1538]" pageId="6" pageNumber="583" pagination="225 - 229" refId="ref10600" refString="JEDLICKA A. 1954: Neue Carabiden aus der Sammlung des Ungarischen Naturwissenschaftlichen Museums in Budapest (Coleoptera). Annales Historico-Naturales Musei Nationalis Hungarici (N. S.) 5: 225 - 229." type="journal article" year="1954">Jedlička, 1954</bibRefCitation>
</taxonomicName>
, holotype; 2
<taxonomicName id="4C46EABC4A0F9F59FC2FFA3032E3F98B" authority="Lindroth, 1980" authorityName="Lindroth" authorityYear="1980" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="583" phylum="Arthropoda" rank="species" species="foochowense" subGenus="Desarmatocillenus">
<emphasis id="B9324D2D4A0F9F59FC2FFA30308AF996" box="[960,979,1519,1537]" italics="true" pageId="6" pageNumber="583">B.</emphasis>
(
<emphasis id="B9324D2D4A0F9F59FC31FA3033ECF98B" italics="true" pageId="6" pageNumber="583">Desarmatocillenus</emphasis>
)
<emphasis id="B9324D2D4A0F9F59FF2DF9D53269F98B" box="[194,304,1546,1564]" italics="true" pageId="6" pageNumber="583">foochowense</emphasis>
<bibRefCitation id="EFD7ECCE4A0F9F59FED8F9D632E3F98B" author="LINDROTH C. H." box="[311,442,1545,1564]" pageId="6" pageNumber="583" pagination="179 - 205" refId="ref10828" refString="LINDROTH C. H. 1980: A revisionary study of the taxon Cillenus Samouelle, 1819 and related forms (Coleoptera: Carabidae: Bembidiini). Entomologica Scandinavica 11: 179 - 205." type="journal article" year="1980">Lindroth, 1980</bibRefCitation>
</taxonomicName>
; 3
<taxonomicName id="4C46EABC4A0F9F59FE08F9D5300BF98B" authority="Netolitzky, 1914" authorityName=": NETOLITZKY" authorityYear="1914" box="[487,850,1545,1564]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="583" phylum="Arthropoda" rank="species" species="fusiforme" subGenus="Odontium">
<emphasis id="B9324D2D4A0F9F59FE08F9D532A3F98B" box="[487,506,1546,1564]" italics="true" pageId="6" pageNumber="583">B.</emphasis>
(
<emphasis id="B9324D2D4A0F9F59FDE8F9D53105F98B" box="[519,604,1546,1564]" italics="true" pageId="6" pageNumber="583">Odontium</emphasis>
)
<emphasis id="B9324D2D4A0F9F59FD86F9D531E3F98B" box="[617,698,1546,1564]" italics="true" pageId="6" pageNumber="583">fusiforme</emphasis>
<bibRefCitation id="EFD7ECCE4A0F9F59FD2EF9D6300BF98B" author="NETOLITZKY F." box="[705,850,1545,1564]" pageId="6" pageNumber="583" pagination="168 - 169" refId="ref11078" refString="NETOLITZKY F. 1914: H. Sauter's Formosa-Ausbeute. Entomologische Mitteilungen 2: 168 - 169." type="journal article" year="1914">Netolitzky, 1914</bibRefCitation>
</taxonomicName>
; 4
<taxonomicName id="4C46EABC4A0F9F59FC91F9D533A4F9A0" authority="Habu, 1973" authorityName="Habu" authorityYear="1973" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="583" phylum="Arthropoda" rank="species" species="takasagonis" subGenus="Plataphus">
<emphasis id="B9324D2D4A0F9F59FC91F9D530CBF98B" box="[894,914,1546,1564]" italics="true" pageId="6" pageNumber="583">B.</emphasis>
(
<emphasis id="B9324D2D4A0F9F59FC71F9D530ACF98B" box="[926,1013,1546,1564]" italics="true" pageId="6" pageNumber="583">Plataphus</emphasis>
)
<emphasis id="B9324D2D4A0F9F59FBEDF9D53731F98B" box="[1026,1128,1546,1564]" italics="true" pageId="6" pageNumber="583">takasagonis</emphasis>
<bibRefCitation id="EFD7ECCE4A0F9F59FF78F9FB33A4F9A0" author="HABU A." box="[151,253,1572,1591]" pageId="6" pageNumber="583" pagination="107 - 116" refId="ref10453" refString="HABU A. 1973: Notes and descriptions of Formosan Carabidae taken by Dr. S. - I. Ueno in 1961 (Coleoptera: Carabidae) III. Bembidion. Transactions of the Shikoku Entomological Society 11: 107 - 116." type="journal article" year="1973">Habu, 1973</bibRefCitation>
</taxonomicName>
; 5
<taxonomicName id="4C46EABC4A0F9F59FEC9F9FA316BF9A0" authority="Toledano, 2009" authorityName="Toledano" authorityYear="2009" box="[294,562,1572,1591]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="583" phylum="Arthropoda" rank="species" species="teradai" status="sp. nov." subGenus="Blepharoplataphus">
<emphasis id="B9324D2D4A0F9F59FEC9F9FA3260F9A0" box="[294,313,1573,1591]" italics="true" pageId="6" pageNumber="583">B.</emphasis>
(
<emphasis id="B9324D2D4A0F9F59FEAAF9FA32B1F9A0" box="[325,488,1573,1591]" italics="true" pageId="6" pageNumber="583">Blepharoplataphus</emphasis>
)
<emphasis id="B9324D2D4A0F9F59FE1BF9FA316BF9A0" box="[500,562,1573,1591]" italics="true" pageId="6" pageNumber="583">teradai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201F0564A0F9F59FDD8F9FB3125F9A0" box="[567,636,1572,1591]" pageId="6" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
, holotype.
</paragraph>
</caption>
<caption id="DF39C1B74A0E9F58FF9EF9C633A8F9F5" ID-DOI="http://doi.org/10.5281/zenodo.5320572" ID-Zenodo-Dep="5320572" httpUri="https://zenodo.org/record/5320572/files/figure.png" pageId="7" pageNumber="584" startId="7.[113,153,1561,1580]" targetBox="[224,947,219,1511]" targetPageId="7">
<paragraph id="8BF9913F4A0E9F58FF9EF9C633A8F9F5" blockId="7.[113,1090,1561,1634]" pageId="7" pageNumber="584">
Figs. 69.
<taxonomicName id="4C46EABC4A0E9F58FF3DF9C53207F9BA" box="[210,350,1562,1581]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="584" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9324D2D4A0E9F58FF3DF9C53276F9BB" box="[210,303,1562,1580]" italics="true" pageId="7" pageNumber="584">Bembidion</emphasis>
spp.
</taxonomicName>
, habitus. 6
<taxonomicName id="4C46EABC4A0E9F58FE34F9C53000F9BB" authority="Jedlicka, 1946" authorityName="Jedlicka" authorityYear="1946" box="[475,857,1561,1580]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="584" phylum="Arthropoda" rank="species" species="miwai" subGenus="Trichoplataphus">
<emphasis id="B9324D2D4A0E9F58FE34F9C532B7F9BB" box="[475,494,1562,1580]" italics="true" pageId="7" pageNumber="584">B.</emphasis>
(
<emphasis id="B9324D2D4A0E9F58FE13F9C531D3F9BB" box="[508,650,1562,1580]" italics="true" pageId="7" pageNumber="584">Trichoplataphus</emphasis>
)
<emphasis id="B9324D2D4A0E9F58FD76F9C53197F9BB" box="[665,718,1562,1580]" italics="true" pageId="7" pageNumber="584">miwai</emphasis>
<bibRefCitation id="EFD7ECCE4A0E9F58FD39F9C63000F9BB" author="JEDLICKA A." box="[726,857,1561,1580]" pageId="7" pageNumber="584" refId="ref10557" refString="JEDLICKA A. 1946: Descriptions des Carabidae nouvelles de l'Asie orientale. A. Jedlicka, Praha, 13 pp." type="book" year="1946">Jedlička, 1946</bibRefCitation>
</taxonomicName>
; 7
<taxonomicName id="4C46EABC4A0E9F58FC65F9C53202F9D0" authority="Netolitzky, 1939" authorityName=": NETOLITZKY" authorityYear="1939" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="584" phylum="Arthropoda" rank="species" species="taiwanum" subGenus="Trichoplataphus">
<emphasis id="B9324D2D4A0E9F58FC65F9C530C4F9BB" box="[906,925,1562,1580]" italics="true" pageId="7" pageNumber="584">B.</emphasis>
(
<emphasis id="B9324D2D4A0E9F58FC43F9C53763F9BB" box="[940,1082,1562,1580]" italics="true" pageId="7" pageNumber="584">Trichoplataphus</emphasis>
)
<emphasis id="B9324D2D4A0E9F58FF9EF9EA339CF9D0" box="[113,197,1589,1607]" italics="true" pageId="7" pageNumber="584">taiwanum</emphasis>
<bibRefCitation id="EFD7ECCE4A0E9F58FF24F9EB3202F9D0" author="NETOLITZKY F." box="[203,347,1588,1607]" pageId="7" pageNumber="584" pagination="49 - 51" refId="ref11096" refString="NETOLITZKY F. 1939: Beitrag zur Kenntnis der Bembidion-Arten des Fernen Ostens (Japan, Korea, Ostsibirien) (Coleoptera). 3. Mitteilung. Proceedings of the Royal Entomological Society of London (Series B) 8: 49 - 51." type="journal article" year="1939">Netolitzky, 1939</bibRefCitation>
</taxonomicName>
; 8
<taxonomicName id="4C46EABC4A0E9F58FE6BF9EA3122F9D0" authority="Toledano, 2009" authorityName="Toledano" authorityYear="2009" box="[388,635,1588,1607]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="584" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov." subGenus="Trichoplataphus">
<emphasis id="B9324D2D4A0E9F58FE6BF9EA32C1F9D0" box="[388,408,1589,1607]" italics="true" pageId="7" pageNumber="584">B.</emphasis>
(
<emphasis id="B9324D2D4A0E9F58FE4CF9EA3176F9D0" box="[419,559,1589,1607]" italics="true" pageId="7" pageNumber="584">Trichoplataphus</emphasis>
)
<emphasis id="B9324D2D4A0E9F58FDD3F9EA3122F9D0" box="[572,635,1589,1607]" italics="true" pageId="7" pageNumber="584">jelineki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201F0564A0E9F58FD6FF9EB319CF9D0" box="[640,709,1588,1607]" pageId="7" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
, holotype; 9
<taxonomicName id="4C46EABC4A0E9F58FCADF9EA377BF9D0" authority="Toledano, 2009" authorityName="Toledano" authorityYear="2009" box="[834,1058,1588,1607]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="584" phylum="Arthropoda" rank="species" species="seijii" status="sp. nov." subGenus="Trichoplataphus">
<emphasis id="B9324D2D4A0E9F58FCADF9EA300FF9D0" box="[834,854,1589,1607]" italics="true" pageId="7" pageNumber="584">B.</emphasis>
(
<emphasis id="B9324D2D4A0E9F58FC8EF9EA30B4F9D0" box="[865,1005,1589,1607]" italics="true" pageId="7" pageNumber="584">Trichoplataphus</emphasis>
)
<emphasis id="B9324D2D4A0E9F58FC16F9EA377BF9D0" box="[1017,1058,1589,1607]" italics="true" pageId="7" pageNumber="584">seijii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201F0564A0E9F58FBC7F9EB33C0F9F5" pageId="7" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
, holotype.
</paragraph>
</caption>
<paragraph id="8BF9913F4A019F57FF5AFF04304AFE84" blockId="8.[151,1127,219,596]" pageId="8" pageNumber="585">Microsculpture distinct, consisting of small isodiametric, convex cells on head and pronotum and of transverse, rather irregular flat cells on elytra.</paragraph>
<paragraph id="8BF9913F4A019F57FF5AFEC4307EFEE4" blockId="8.[151,1127,219,596]" pageId="8" pageNumber="585">
Male genitalia (
<figureCitation id="137D8DBA4A019F57FEB2FEC432F5FEA3" box="[349,428,283,308]" captionStart="Figs" captionStartId="14.[151,190,1436,1455]" captionTargetBox="[221,1058,219,1412]" captionTargetId="figure-145@14.[219,1058,219,1417]" captionTargetPageId="14" captionText="Figs. 1728. Bembidion and Sinechostictus spp., median lobe of the aedeagus. 17 B. (Desarmatocillenus) foochowense Lindroth, 1980; 18 B. (Odontium) fusiforme Netolitzky, 1914; 19 B. (Plataphus) takasagonis Habu, 1973; 20 B. (Blepharoplataphus) teradai sp. nov., holotype; 21 B. (Trichoplataphus) miwai Jedlička, 1946; 22 B. (Trichoplataphus) taiwanum Netolitzky, 1939; 23 B. (Trichoplataphus) jelineki sp. nov., holotype; 24 B. (Trichoplataphus) seijii sp. nov., holotype; 25 B. (Ocydromus) chloreum Bates, 1873, specimen from Taiwan; 26 B. (Ocydromus) shunichii Habu, 1973; 27 B. (Ocydromus) peleum Jedlička, 1933, specimen from Taiwan; 28 S. (Pseudolimnaeum) chuji Jedlička, 1951." figureDoi="http://doi.org/10.5281/zenodo.5320580" httpUri="https://zenodo.org/record/5320580/files/figure.png" pageId="8" pageNumber="585">Fig. 20</figureCitation>
). Apex of median lobe ventrally hooked. For the other characters, typical shape and endophallus of the subgenus
<taxonomicName id="4C46EABC4A019F57FD7AFEE23039FEC3" authorityName="Netolitzky" authorityYear="1920" box="[661,864,317,340]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="585" phylum="Arthropoda" rank="subGenus" subGenus="Blepharoplataphus">
<emphasis id="B9324D2D4A019F57FD7AFEE23039FEC3" box="[661,864,317,340]" italics="true" pageId="8" pageNumber="585">Blepharoplataphus</emphasis>
</taxonomicName>
. In particular, the endophallus shares the structure of the sclerites with
<taxonomicName id="4C46EABC4A019F57FD77FE823079FEE3" baseAuthorityName="MORITA" baseAuthorityYear="2008" box="[664,800,349,372]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="585" phylum="Arthropoda" rank="species" species="nirasawai">
<emphasis id="B9324D2D4A019F57FD77FE823079FEE3" box="[664,800,349,372]" italics="true" pageId="8" pageNumber="585">B. nirasawai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CC2B44A019F57FF78FEA33045FE44" pageId="8" pageNumber="585" type="etymology">
<paragraph id="8BF9913F4A019F57FF78FEA33045FE44" blockId="8.[151,1127,219,596]" pageId="8" pageNumber="585">
<emphasis id="B9324D2D4A019F57FF78FEA33241FE03" bold="true" box="[151,280,380,404]" pageId="8" pageNumber="585">Etymology.</emphasis>
Dedicated to Katsuyuki Terada, an accomplished specialist in Japanese and Taiwanese
<taxonomicName id="4C46EABC4A019F57FEE2FE443225FE24" box="[269,380,411,435]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="585" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
, who gave me the opportunity to study the interesting fauna of
<collectingCountry id="F351D1AF4A019F57FBF7FE44373FFE24" box="[1048,1126,411,435]" name="Taiwan" pageId="8" pageNumber="585">Taiwan</collectingCountry>
by kindly sending me the specimens dealt with in this paper.
</paragraph>
</subSubSection>
<subSubSection id="C35CC2B44A019F57FF78FE0431B1FE64" box="[151,744,475,500]" pageId="8" pageNumber="585" type="distribution">
<paragraph id="8BF9913F4A019F57FF78FE0431B1FE64" blockId="8.[151,1127,219,596]" box="[151,744,475,500]" pageId="8" pageNumber="585">
<emphasis id="B9324D2D4A019F57FF78FE043273FE64" bold="true" box="[151,298,475,499]" pageId="8" pageNumber="585">Distribution.</emphasis>
So far known only from the
<typeStatus id="54FD2F9D4A019F57FDB1FE0331D5FE63" box="[606,652,476,500]" pageId="8" pageNumber="585">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C35CC2B44A019F57FF78FE23373DFDC4" pageId="8" pageNumber="585" type="discussion">
<paragraph id="8BF9913F4A019F57FF78FE23373DFDC4" blockId="8.[151,1127,219,596]" pageId="8" pageNumber="585">
<emphasis id="B9324D2D4A019F57FF78FE23325FFD83" bold="true" box="[151,262,508,532]" pageId="8" pageNumber="585">Remarks.</emphasis>
The morphological diversity within
<taxonomicName id="4C46EABC4A019F57FD6CFE223017FD83" authorityName="Netolitzky" authorityYear="1920" box="[643,846,509,532]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="597" phylum="Arthropoda" rank="subGenus" subGenus="Blepharoplataphus">
<emphasis id="B9324D2D4A019F57FD6CFE223017FD83" box="[643,846,509,532]" italics="true" pageId="8" pageNumber="585">Blepharoplataphus</emphasis>
</taxonomicName>
is very low: the male genitalia and habitus of all species are very similar;
<taxonomicName id="4C46EABC4A019F57FD65FDC231AAFDA3" authority="Toledano, 2009" authorityName="Toledano" authorityYear="2009" box="[650,755,541,564]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="585" phylum="Arthropoda" rank="species" species="teradai" status="sp. nov.">
<emphasis id="B9324D2D4A019F57FD65FDC231AAFDA3" box="[650,755,541,564]" italics="true" pageId="8" pageNumber="585">B. teradai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201F0564A019F57FD16FDC33013FDA4" box="[761,842,540,563]" pageId="8" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
seems closely related to
<taxonomicName id="4C46EABC4A019F57FBA0FDC2325BFDC3" baseAuthorityName="MORITA" baseAuthorityYear="2008" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="585" phylum="Arthropoda" rank="species" species="nirasawai">
<emphasis id="B9324D2D4A019F57FBA0FDC2325BFDC3" italics="true" pageId="8" pageNumber="585">B. nirasawai</emphasis>
</taxonomicName>
, with which it shares almost identical male genitalia, except for the apical shape.
</paragraph>
</subSubSection>
</treatment>
</document>