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<mods:title id="7FF26396F9EB34399C75EE8C827AAF54">Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae)</mods:title>
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: Pseudinae
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In the early Oligocene, 32.3 (25.838.7) Mya, the crown node time of this subfamily, a clade of aquatic and semiaquatic frogs with elongate, calcified intercalary elements in the digits gave rise to the semiaquatic
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in the upper Amazon Basin. The major component of the pseudine clade differentiated into the smaller
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and the larger
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<emphasis id="B937EAA1FF9C121CF2348CA032ACF149" box="[315,402,510,535]" italics="true" pageId="53" pageNumber="54">Pseudis</emphasis>
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in the early Miocene about 21.5 (18.424.6) Mya. Both genera have broad distributions in the
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Region and tropical and subtropical South
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east of the Andes.
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