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<document id="8589A5925EF9CDAAFA594B36F2DEF022" ID-CLB-Dataset="2947" ID-DOI="10.11646/zootaxa.5082.1.2" ID-GBIF-Dataset="722c160e-2d11-4a58-83fc-d346d227f9e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5783094" ID-ZooBank="795488E3-DE16-4268-8968-628C9D5E3A4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639564243538" checkinUser="plazi" docAuthor="Rueda Martín, Paola A., Gibon, Francois-Marie &amp; Sganga, Julieta V." docDate="2021" docId="03DEE90BFFAF4517FF0AFF18FAFDFA89" docLanguage="en" docName="zootaxa.5082.1.2.pdf" docOrigin="Zootaxa 5082 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrnellus marginalis" docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="FFE79173FFAA4512FF9DFF8FFF9CFFC9" masterDocTitle="New species and records of the Neotropical genus Cyrnellus (Trichoptera: Polycentropodidae) and the re-establishment of C. minimus and C. marginalis" masterLastPageNumber="29" masterPageNumber="15" pageNumber="20" updateTime="1699442671739" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CE7F11ABD753D1356A60362CC6C19C43">New species and records of the Neotropical genus Cyrnellus (Trichoptera: Polycentropodidae) and the re-establishment of C. minimus and C. marginalis</mods:title>
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<mods:namePart id="2045925F282067AB18312EF272256548">Rueda Martín, Paola A.</mods:namePart>
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<mods:namePart id="103C2CD7797AC9AD844EE02E622DCB26">Gibon, Francois-Marie</mods:namePart>
<mods:affiliation id="E743B7F4410C4B332CC6EBFAA78659EA">CBGP, IRD, CIRAD, INRAE, Montpellier SupAgro, Univ. Montpellier, Montpellier, France.</mods:affiliation>
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<mods:namePart id="C7D913BA998A839407D447A7813C8894">Sganga, Julieta V.</mods:namePart>
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<mods:affiliation id="83F80357504588AFFF8584148A6A0DF2">Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales (UBA). CABA, Argentina. jsganga @ gmail. com; https: // orcid. org / 0000 - 0003 - 1210 - 5937</mods:affiliation>
<mods:nameIdentifier id="538820ABCA453BC6EF48276EC072730B" type="email">jsganga@gmail.com</mods:nameIdentifier>
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<treatment id="03DEE90BFFAF4517FF0AFF18FAFDFA89" ID-DOI="http://doi.org/10.5281/zenodo.5783698" ID-GBIF-Taxon="191647276" ID-Zenodo-Dep="5783698" LSID="urn:lsid:plazi:treatment:03DEE90BFFAF4517FF0AFF18FAFDFA89" httpUri="http://treatment.plazi.org/id/03DEE90BFFAF4517FF0AFF18FAFDFA89" lastPageNumber="20" pageId="5" pageNumber="20">
<subSubSection id="C36D0B96FFAF4517FF0AFF18FDDDFF78" box="[151,577,151,178]" pageId="5" pageNumber="20" type="nomenclature">
<paragraph id="8BC8581DFFAF4517FF0AFF18FDDDFF78" blockId="5.[151,577,151,213]" box="[151,577,151,178]" pageId="5" pageNumber="20">
<heading id="D080EF71FFAF4517FF0AFF18FDDDFF78" bold="true" box="[151,577,151,178]" fontSize="11" level="1" pageId="5" pageNumber="20" reason="1">
<taxonomicName id="4C77239EFFAF4517FF0AFF18FDDDFF78" ID-CoL="33JTJ" authority="(Banks 1930)" baseAuthorityName="Banks" baseAuthorityYear="1930" box="[151,577,151,178]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="B903840FFFAF4517FF0AFF18FDDDFF78" bold="true" box="[151,577,151,178]" pageId="5" pageNumber="20">
<emphasis id="B903840FFFAF4517FF0AFF18FE0AFF7B" bold="true" box="[151,406,151,178]" italics="true" pageId="5" pageNumber="20">Cyrnellus marginalis</emphasis>
(
<bibRefCitation id="EFE625ECFFAF4517FE3BFF18FDA5FF78" author="Banks, N." box="[422,569,151,177]" pageId="5" pageNumber="20" pagination="223 - 233" refId="ref8312" refString="Banks, N. (1930) New neuropteroid insects from the United States. Psyche, 37, 223 - 233. https: // doi. org / 10.1155 / 1930 / 12828" type="journal article" year="1930">Banks 1930</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C36D0B96FFAF4517FF0AFF34FEBDFF1C" box="[151,289,187,213]" pageId="5" pageNumber="20" type="description">
<paragraph id="8BC8581DFFAF4517FF0AFF34FEBDFF1C" blockId="5.[151,577,151,213]" box="[151,289,187,213]" pageId="5" pageNumber="20">
<figureCitation id="134C4498FFAF4517FF0AFF34FEBDFF1C" box="[151,289,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1864,1888]" captionTargetBox="[253,1332,189,1829]" captionTargetId="figure-19@6.[242,1345,181,1840]" captionTargetPageId="6" captionText="FIGURE 3. Cyrnellus spp., male genitalia. 3A3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783106" httpUri="https://zenodo.org/record/5783106/files/figure.png" pageId="5" pageNumber="20">Figs 3A3D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFAF4517FF0AFE8FFB04FEBD" pageId="5" pageNumber="20" type="reference_group">
<paragraph id="8BC8581DFFAF4517FF0AFE8FFB7FFED1" blockId="5.[151,1437,256,372]" box="[151,1251,256,280]" pageId="5" pageNumber="20">
<treatmentCitationGroup id="AB677F33FFAF4517FF0AFE8FFB7FFED1" box="[151,1251,256,280]" pageId="5" pageNumber="20">
<treatmentCitation id="0AD67E0CFFAF4517FF0AFE8FFDDEFED1" author="Banks, N." box="[151,578,256,280]" page="231" pageId="5" pageNumber="20" year="1930">
<taxonomicName id="4C77239EFFAF4517FF0AFE8FFDDEFED1" ID-CoL="485SN" authority="Banks 1930: 231" authorityName="Banks" authorityPageNumber="231" authorityYear="1930" box="[151,578,256,280]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="B903840FFFAF4517FF0AFE8FFE0CFED1" box="[151,400,256,280]" italics="true" pageId="5" pageNumber="20">Nyctiophylax marginalis</emphasis>
<bibRefCitation id="EFE625ECFFAF4517FE0BFE8FFDDEFED1" author="Banks, N." box="[406,578,256,280]" pageId="5" pageNumber="20" pagination="223 - 233" refId="ref8312" refString="Banks, N. (1930) New neuropteroid insects from the United States. Psyche, 37, 223 - 233. https: // doi. org / 10.1155 / 1930 / 12828" type="journal article" year="1930">Banks 1930: 231</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation id="0AD67E0CFFAF4517FCB1FE8FFC21FED1" author="Ross, H. H." box="[812,957,256,280]" page="12" pageId="5" pageNumber="20" year="1938">
<bibRefCitation id="EFE625ECFFAF4517FCB1FE8FFC21FED1" author="Ross, H. H." box="[812,957,256,280]" pageId="5" pageNumber="20" pagination="1 - 59" refId="ref9572" refString="Ross, H. H. (1938) Lectotypes of North American caddisflies in the Museum of Comparative Zoology. Psyche, 45, 1 - 59. https: // doi. org / 10.1155 / 1938 / 25928" type="journal article" year="1938">Ross 1938: 12</bibRefCitation>
</treatmentCitation>
(designated male
<typeStatus id="54CCE6BFFFAF4517FBEBFE8FFB44FED1" box="[1142,1240,256,280]" pageId="5" pageNumber="20" type="lectotype">lectotype</typeStatus>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC8581DFFAF4517FF0AFE91FBFFFE9C" blockId="5.[151,1437,256,372]" pageId="5" pageNumber="20">
<treatmentCitationGroup id="AB677F33FFAF4517FF0AFE91FBFFFE9C" pageId="5" pageNumber="20">
<taxonomicName id="4C77239EFFAF4517FF0AFE91FE28FEFF" ID-CoL="33JTJ" authority="Banks" authorityName="Banks" box="[151,436,286,310]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="B903840FFFAF4517FF0AFE91FEF0FEFF" box="[151,364,286,310]" italics="true" pageId="5" pageNumber="20">Cyrnellus marginalis</emphasis>
Banks
</taxonomicName>
;
<treatmentCitation id="0AD67E0CFFAF4517FE23FE90FDD3FEFF" author="Ross, H. H." box="[446,591,286,310]" page="71" pageId="5" pageNumber="20" year="1944">
<bibRefCitation id="EFE625ECFFAF4517FE23FE90FDD3FEFF" author="Ross, H. H." box="[446,591,286,310]" pageId="5" pageNumber="20" pagination="1 - 326" refId="ref9613" refString="Ross, H. H. (1944) The caddisflies or Trichoptera of Illinois. Bulletin of the Illinois Natural History Survey, 23, 1 - 326. https: // doi. org / 10.21900 / j. inhs. v 23.199" type="journal article" year="1944">Ross 1944: 71</bibRefCitation>
</treatmentCitation>
, 72 (descriptions of male, female);
<treatmentCitation id="0AD67E0CFFAF4517FC2FFE91FBD1FEFF" author="Flint, O. S. Jr." box="[946,1101,286,310]" page="469" pageId="5" pageNumber="20" year="1964">
<bibRefCitation id="EFE625ECFFAF4517FC2FFE91FBD1FEFF" author="Flint, O. S. Jr." box="[946,1101,286,310]" pageId="5" pageNumber="20" pagination="467 - 481" refId="ref8850" refString="Flint, O. S. Jr. (1964) Notes on some Neartic Psychomiidae with special reference to their larvae (Trichoptera). Proceedings of the United States National Museum, 115 (3491), 467 - 481. https: // doi. org / 10.5479 / si. 00963801.115 - 3491.467" type="journal article" year="1964">Flint 1964: 469</bibRefCitation>
</treatmentCitation>
[as synonym of
<taxonomicName id="4C77239EFFAF4517FB6EFE91FEE3FE9C" ID-CoL="33JTG" authority="(Banks 1905)" baseAuthorityName="Banks" baseAuthorityYear="1905" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B903840FFFAF4517FB6EFE91FF76FE9C" italics="true" pageId="5" pageNumber="20">Cyrnellus fraternus</emphasis>
(
<bibRefCitation id="EFE625ECFFAF4517FF65FEB2FEEBFE9C" author="Banks, N." box="[248,375,317,341]" pageId="5" pageNumber="20" pagination="1 - 20" refId="ref8253" refString="Banks, N. (1905) Descriptions of new Nearctic neuropteroid insects. Transactions of the American Entomological Society, 32, 1 - 20." type="journal article" year="1905">Banks 1905</bibRefCitation>
)
</taxonomicName>
];
<treatmentCitation id="0AD67E0CFFAF4517FE0DFEB2FD58FE9C" author="Holzenthal, R. W. &amp; Calor, A. R." box="[400,708,317,341]" page="418" pageId="5" pageNumber="20" year="2017">
<bibRefCitation id="EFE625ECFFAF4517FE0DFEB2FD58FE9C" author="Holzenthal, R. W. &amp; Calor, A. R." box="[400,708,317,341]" pageId="5" pageNumber="20" pagination="1 - 566" refId="ref9193" refString="Holzenthal, R. W. &amp; Calor, A. R. (2017) Catalog of the Neotropical Trichoptera (Caddisflies). ZooKeys, 654, 1 - 566. https: // doi. org / 10.3897 / zookeys. 654.9516" type="journal article" year="2017">Holzenthal &amp; Calor 2017: 418</bibRefCitation>
</treatmentCitation>
(catalogue, as synonym of
<taxonomicName id="4C77239EFFAF4517FC47FEB2FBC9FE9C" ID-CoL="33JTG" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[986,1109,317,341]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B903840FFFAF4517FC47FEB2FBC9FE9C" box="[986,1109,317,341]" italics="true" pageId="5" pageNumber="20">C. fraternus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC8581DFFAF4517FF0AFED3FB04FEBD" blockId="5.[151,1437,256,372]" box="[151,1176,348,372]" pageId="5" pageNumber="20">
<treatmentCitationGroup id="AB677F33FFAF4517FF0AFED3FB04FEBD" box="[151,1176,348,372]" pageId="5" pageNumber="20">
<treatmentCitation id="0AD67E0CFFAF4517FF0AFED3FE61FEBD" author="Mosely, M. E." box="[151,509,348,372]" page="142" pageId="5" pageNumber="20" year="1934">
<taxonomicName id="4C77239EFFAF4517FF0AFED3FE61FEBD" ID-CoL="33JTQ" authority="Mosely 1934: 142" authorityName="Mosely" authorityPageNumber="142" authorityYear="1934" box="[151,509,348,372]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="zernyi">
<emphasis id="B903840FFFAF4517FF0AFED3FEA2FEBA" box="[151,318,348,372]" italics="true" pageId="5" pageNumber="20">Cyrnellus zernyi</emphasis>
<bibRefCitation id="EFE625ECFFAF4517FED9FED3FE61FEBD" author="Mosely, M. E." box="[324,509,348,372]" pageId="5" pageNumber="20" pagination="139 - 142" refId="ref9307" refString="Mosely, M. E. (1934) Some new exotic Trichoptera. Stylops, 3 (6), 139 - 142. https: // doi. org / 10.1111 / j. 1365 - 3113.1934. tb 01566. x" type="journal article" year="1934">Mosely 1934: 142</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation id="0AD67E0CFFAF4517FD7AFED3FCE4FEBD" author="Ross, H. H." box="[743,888,348,372]" page="13" pageId="5" pageNumber="20" year="1938">
<bibRefCitation id="EFE625ECFFAF4517FD7AFED3FCE4FEBD" author="Ross, H. H." box="[743,888,348,372]" pageId="5" pageNumber="20" pagination="1 - 59" refId="ref9572" refString="Ross, H. H. (1938) Lectotypes of North American caddisflies in the Museum of Comparative Zoology. Psyche, 45, 1 - 59. https: // doi. org / 10.1155 / 1938 / 25928" type="journal article" year="1938">Ross 1938: 13</bibRefCitation>
</treatmentCitation>
(synonym of
<taxonomicName id="4C77239EFFAF4517FB98FED3FB0CFEBD" ID-CoL="33JTJ" authority="(Banks 1930)" box="[1029,1168,348,372]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="B903840FFFAF4517FB98FED3FB0CFEBD" box="[1029,1168,348,372]" italics="true" pageId="5" pageNumber="20">C. marginalis</emphasis>
</taxonomicName>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFAF4517FF0AFE2EFD21FCED" pageId="5" pageNumber="20" type="diagnosis">
<paragraph id="8BC8581DFFAF4517FF0AFE2EFD21FCED" blockId="5.[151,1437,417,1344]" pageId="5" pageNumber="20">
<emphasis id="B903840FFFAF4517FF0AFE2EFE96FE72" bold="true" box="[151,266,417,443]" pageId="5" pageNumber="20">Diagnosis</emphasis>
.
<taxonomicName id="4C77239EFFAF4517FE84FE2DFD9AFE75" box="[281,518,418,444]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="B903840FFFAF4517FE84FE2DFD9AFE75" box="[281,518,418,444]" italics="true" pageId="5" pageNumber="20">Cyrnellus marginalis</emphasis>
</taxonomicName>
is distinguished from other species in the genus by the following character combination: Subapicomesal spine of each inferior appendage conical, wide at base, subapical in position (
<figureCitation id="134C4498FFAF4517FA8DFE49FAFBFE29" box="[1296,1383,454,480]" captionStart="FIGURE 3" captionStartId="6.[151,250,1864,1888]" captionTargetBox="[253,1332,189,1829]" captionTargetId="figure-19@6.[242,1345,181,1840]" captionTargetPageId="6" captionText="FIGURE 3. Cyrnellus spp., male genitalia. 3A3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783106" httpUri="https://zenodo.org/record/5783106/files/figure.png" pageId="5" pageNumber="20">Fig. 3B</figureCitation>
,
<emphasis id="B903840FFFAF4517FAEFFE48FA17FE29" box="[1394,1419,455,480]" italics="true" pageId="5" pageNumber="20">as</emphasis>
); its internal lobe rounded (
<figureCitation id="134C4498FFAF4517FE28FE65FD90FDCD" box="[437,524,490,516]" captionStart="FIGURE 3" captionStartId="6.[151,250,1864,1888]" captionTargetBox="[253,1332,189,1829]" captionTargetId="figure-19@6.[242,1345,181,1840]" captionTargetPageId="6" captionText="FIGURE 3. Cyrnellus spp., male genitalia. 3A3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783106" httpUri="https://zenodo.org/record/5783106/files/figure.png" pageId="5" pageNumber="20">Fig. 3B</figureCitation>
,
<emphasis id="B903840FFFAF4517FD8AFE65FDBBFDCD" box="[535,551,490,516]" italics="true" pageId="5" pageNumber="20">il</emphasis>
); the dorsal phallic sclerite has its anterior arm short (
<figureCitation id="134C4498FFAF4517FBE2FE65FB4AFDCD" box="[1151,1238,490,516]" captionStart="FIGURE 3" captionStartId="6.[151,250,1864,1888]" captionTargetBox="[253,1332,189,1829]" captionTargetId="figure-19@6.[242,1345,181,1840]" captionTargetPageId="6" captionText="FIGURE 3. Cyrnellus spp., male genitalia. 3A3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783106" httpUri="https://zenodo.org/record/5783106/files/figure.png" pageId="5" pageNumber="20">Fig. 3C</figureCitation>
,
<emphasis id="B903840FFFAF4517FB7CFE65FB66FDCD" box="[1249,1274,490,516]" italics="true" pageId="5" pageNumber="20">ds</emphasis>
) and posterior arms short and divergent (
<figureCitation id="134C4498FFAF4517FE2AFD81FD93FDE1" box="[439,527,526,552]" captionStart="FIGURE 3" captionStartId="6.[151,250,1864,1888]" captionTargetBox="[253,1332,189,1829]" captionTargetId="figure-19@6.[242,1345,181,1840]" captionTargetPageId="6" captionText="FIGURE 3. Cyrnellus spp., male genitalia. 3A3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783106" httpUri="https://zenodo.org/record/5783106/files/figure.png" pageId="5" pageNumber="20">Fig. 3D</figureCitation>
,
<emphasis id="B903840FFFAF4517FD87FD81FDAFFDE1" box="[538,563,526,552]" italics="true" pageId="5" pageNumber="20">ds</emphasis>
).
<taxonomicName id="4C77239EFFAF4517FDD4FD81FCAFFDE1" box="[585,819,526,552]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="B903840FFFAF4517FDD4FD81FCAFFDE1" box="[585,819,526,552]" italics="true" pageId="5" pageNumber="20">Cyrnellus marginalis</emphasis>
</taxonomicName>
is considered to be distinct from
<taxonomicName id="4C77239EFFAF4517FB3FFD81FAB5FDE1" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[1186,1321,526,552]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B903840FFFAF4517FB3FFD81FAB5FDE1" box="[1186,1321,526,552]" italics="true" pageId="5" pageNumber="20">C. fraternus</emphasis>
</taxonomicName>
by having the base of each intermediate appendage narrower than the median part whereas in
<taxonomicName id="4C77239EFFAF4517FBAFFDBDFB25FD85" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[1074,1209,562,588]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B903840FFFAF4517FBAFFDBDFB25FD85" box="[1074,1209,562,588]" italics="true" pageId="5" pageNumber="20">C. fraternus</emphasis>
</taxonomicName>
the base is subequal to the median part (
<figureCitation id="134C4498FFAF4517FEF3FDD9FE65FDB9" box="[366,505,598,624]" captionStart="FIGURE 3" captionStartId="6.[151,250,1864,1888]" captionTargetBox="[253,1332,189,1829]" captionTargetId="figure-19@6.[242,1345,181,1840]" captionTargetPageId="6" captionText="FIGURE 3. Cyrnellus spp., male genitalia. 3A3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783106" httpUri="https://zenodo.org/record/5783106/files/figure.png" pageId="5" pageNumber="20">Figs 3A, 3E</figureCitation>
). Another constant character is the ventral margin of each inferior appendage which is angulate in
<taxonomicName id="4C77239EFFAF4517FEAAFDF5FE4CFD5D" box="[311,464,634,660]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="B903840FFFAF4517FEAAFDF5FE4CFD5D" box="[311,464,634,660]" italics="true" pageId="5" pageNumber="20">C. marginalis</emphasis>
</taxonomicName>
and sinuous in
<taxonomicName id="4C77239EFFAF4517FD18FDF5FC91FD5D" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[645,781,634,660]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B903840FFFAF4517FD18FDF5FC91FD5D" box="[645,781,634,660]" italics="true" pageId="5" pageNumber="20">C. fraternus</emphasis>
</taxonomicName>
. An additional remarkable difference between the species is the position of the subapicomesal spine of each inferior appendage: in
<taxonomicName id="4C77239EFFAF4517FC50FD11FBFBFD71" box="[973,1127,670,696]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="B903840FFFAF4517FC50FD11FBFBFD71" box="[973,1127,670,696]" italics="true" pageId="5" pageNumber="20">C. marginalis</emphasis>
</taxonomicName>
this spine is subapical by a distance as long as the width of the apex of the appendage (
<figureCitation id="134C4498FFAF4517FCAFFD4DFC17FD15" box="[818,907,706,732]" captionStart="FIGURE 3" captionStartId="6.[151,250,1864,1888]" captionTargetBox="[253,1332,189,1829]" captionTargetId="figure-19@6.[242,1345,181,1840]" captionTargetPageId="6" captionText="FIGURE 3. Cyrnellus spp., male genitalia. 3A3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783106" httpUri="https://zenodo.org/record/5783106/files/figure.png" pageId="5" pageNumber="20">Fig. 3A</figureCitation>
); in
<taxonomicName id="4C77239EFFAF4517FC20FD4DFBD9FD15" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[957,1093,706,732]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B903840FFFAF4517FC20FD4DFBD9FD15" box="[957,1093,706,732]" italics="true" pageId="5" pageNumber="20">C. fraternus</emphasis>
</taxonomicName>
the spines subapical insertion is more than 1.5 times as far as the width of the apex of the appendage (
<figureCitation id="134C4498FFAF4517FC2DFD69FB98FCC9" box="[944,1028,742,768]" captionStart="FIGURE 3" captionStartId="6.[151,250,1864,1888]" captionTargetBox="[253,1332,189,1829]" captionTargetId="figure-19@6.[242,1345,181,1840]" captionTargetPageId="6" captionText="FIGURE 3. Cyrnellus spp., male genitalia. 3A3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783106" httpUri="https://zenodo.org/record/5783106/files/figure.png" pageId="5" pageNumber="20">Fig. 3F</figureCitation>
); this distance is not constant among individuals of the same species (
<figureCitation id="134C4498FFAF4517FD9FFC85FD2CFCED" box="[514,688,778,804]" captionStart="FIGURE 3" captionStartId="6.[151,250,1864,1888]" captionTargetBox="[253,1332,189,1829]" captionTargetId="figure-19@6.[242,1345,181,1840]" captionTargetPageId="6" captionText="FIGURE 3. Cyrnellus spp., male genitalia. 3A3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783106" httpUri="https://zenodo.org/record/5783106/files/figure.png" pageId="5" pageNumber="20">Figs 3F, 3I3K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFAF4517FF5AFCA2FB88FB45" pageId="5" pageNumber="20" type="materials_examined">
<paragraph id="8BC8581DFFAF4517FF5AFCA2FB88FB45" blockId="5.[151,1437,417,1344]" pageId="5" pageNumber="20">
<emphasis id="B903840FFFAF4517FF5AFCA2FE30FC8E" bold="true" box="[199,428,813,839]" pageId="5" pageNumber="20">Material examined</emphasis>
.
<materialsCitation id="3B1F5240FFAF4517FE20FCA2FF41FCA5" ID-GBIF-Occurrence="3418589328" collectingDate="2006-05-06" country="Bolivia" latitude="-13.263889" location="Bella Vista" longLatPrecision="21" longitude="-63.709167" municipality="Laguna Granja" pageId="5" pageNumber="20" specimenCount="1" specimenCount-male="1">
<emphasis id="B903840FFFAF4517FE20FCA2FDDCFC8E" bold="true" box="[445,576,813,839]" pageId="5" pageNumber="20">
<collectingCountry id="F360188DFFAF4517FE20FCA2FDA7FC8E" box="[445,571,813,839]" name="Bolivia" pageId="5" pageNumber="20">BOLIVIA</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6BACC267FFAF4517FDD7FCA1FD6BFC81" box="[586,759,814,840]" pageId="5" pageNumber="20">Laguna Granja</collectingMunicipality>
,
<location id="8EA80EC6FFAF4517FC98FCA1FC19FC81" LSID="urn:lsid:plazi:treatment:03DEE90BFFAF4517FF0AFF18FAFDFA89:8EA80EC6FFAF4517FC98FCA1FC19FC81" box="[773,901,814,840]" country="Bolivia" latitude="-13.263889" longLatPrecision="21" longitude="-63.709167" municipality="Laguna Granja" name="Bella Vista" pageId="5" pageNumber="20">Bella Vista</location>
,
<geoCoordinate id="EE433EDAFFAF4517FC0EFCA1FB82FC80" box="[915,1054,814,841]" degrees="13" direction="south" minutes="15" orientation="latitude" pageId="5" pageNumber="20" precision="15" seconds="50" value="-13.263889">13°1550S</geoCoordinate>
,
<geoCoordinate id="EE433EDAFFAF4517FBB0FCA1FB5DFC81" box="[1069,1217,814,840]" degrees="63" direction="west" minutes="42" orientation="longitude" pageId="5" pageNumber="20" precision="15" seconds="33" value="-63.709167">63°4233W</geoCoordinate>
,
<date id="FFC97EDDFFAF4517FB52FCA1FAA3FC81" box="[1231,1343,814,840]" pageId="5" pageNumber="20" value="2006-05-06">
<collectingDate id="EF8D8735FFAF4517FB52FCA1FAA3FC81" box="[1231,1343,814,840]" pageId="5" pageNumber="20" value="2006-05-06">06.v.2006</collectingDate>
</date>
,
<specimenCount id="9D719394FFAF4517FAD3FCA1FA00FC81" box="[1358,1436,814,840]" pageId="5" pageNumber="20" type="male">1 male</specimenCount>
(IBN)
</materialsCitation>
;
<materialsCitation id="3B1F5240FFAF4517FF74FCDEFCC1FC59" ID-GBIF-Occurrence="3418589327" collectingDate="1991-11-17" collectorName="Dr Peter" country="Brazil" county="Km" elevation="65" latitude="-10.541667" location="Fazenda Rancho Grande" longLatPrecision="914" longitude="-62.8" municipality="Km" pageId="5" pageNumber="20" specimenCount="3" specimenCount-male="3" stateProvince="Rondonia">
<emphasis id="B903840FFFAF4517FF74FCDEFEC2FCA2" bold="true" box="[233,350,849,875]" pageId="5" pageNumber="20">
<collectingCountry id="F360188DFFAF4517FF74FCDEFEC5FCA2" box="[233,345,849,875]" name="Brazil" pageId="5" pageNumber="20">BRAZIL</collectingCountry>
:
</emphasis>
<collectingRegion id="49B396FFFFAF4517FEFBFCDDFE4AFCA5" box="[358,470,850,876]" country="Brazil" name="Rondonia" pageId="5" pageNumber="20">Rondonia</collectingRegion>
, 62
<collectingCounty id="62A92091FFAF4517FD98FCDDFDB3FCA5" box="[517,559,850,876]" pageId="5" pageNumber="20">Km</collectingCounty>
S de
<locationDeviation id="9A7A413FFFAF4517FDF3FCDDFD70FCA5" box="[622,748,850,876]" pageId="5" pageNumber="20">Ariquemes</locationDeviation>
,
<locationDeviation id="9A7A413FFFAF4517FD6AFCDDFCABFCA5" box="[759,823,850,876]" pageId="5" pageNumber="20">Linea</locationDeviation>
C-20, 7
<collectingMunicipality id="6BACC267FFAF4517FC07FCDDFC58FCA5" box="[922,964,850,876]" pageId="5" pageNumber="20">Km</collectingMunicipality>
E B-
<quantity id="4C8FF5F8FFAF4517FB9FFCDDFBE9FCA5" box="[1026,1141,850,877]" metricMagnitude="1" metricUnit="m" metricValue="6.5165" pageId="5" pageNumber="20" unit="m" value="65.165">
<elevation id="005ABF2EFFAF4517FB9FFCDDFBE9FCA5" box="[1026,1141,850,877]" metricMagnitude="1" metricUnit="m" metricValue="6.5165" pageId="5" pageNumber="20" unit="m" value="65.165">65, 165 m</elevation>
</quantity>
,
<location id="8EA80EC6FFAF4517FB1CFCDDFA04FCA5" LSID="urn:lsid:plazi:treatment:03DEE90BFFAF4517FF0AFF18FAFDFA89:8EA80EC6FFAF4517FB1CFCDDFA04FCA5" box="[1153,1432,850,876]" country="Brazil" county="Km" latitude="-10.541667" longLatPrecision="914" longitude="-62.8" municipality="Km" name="Fazenda Rancho Grande" pageId="5" pageNumber="20" stateProvince="Rondonia">Fazenda Rancho Grande</location>
,
<geoCoordinate id="EE433EDAFFAF4517FF0AFCF9FE97FC58" box="[151,267,886,913]" degrees="10" direction="south" minutes="32,5" orientation="latitude" pageId="5" pageNumber="20" precision="92" value="-10.541667">10°32,5´S</geoCoordinate>
,
<geoCoordinate id="EE433EDAFFAF4517FE84FCF9FE16FC59" box="[281,394,886,912]" degrees="62" direction="west" minutes="48" orientation="longitude" pageId="5" pageNumber="20" precision="925" value="-62.8">62° 48´W</geoCoordinate>
,
<date id="FFC97EDDFFAF4517FE04FCF9FD8FFC59" box="[409,531,886,912]" pageId="5" pageNumber="20" value="1991-11-17">
<collectingDate id="EF8D8735FFAF4517FE04FCF9FD8FFC59" box="[409,531,886,912]" pageId="5" pageNumber="20" value="1991-11-17">17.xi.1991</collectingDate>
</date>
,
<collectorName id="26823DCBFFAF4517FDBCFCF9FD18FC59" box="[545,644,886,912]" pageId="5" pageNumber="20">Dr Peter</collectorName>
,
<specimenCount id="9D719394FFAF4517FD0FFCF9FD77FC59" box="[658,747,886,912]" pageId="5" pageNumber="20" type="male">3 males</specimenCount>
(USNM)
</materialsCitation>
;
<materialsCitation id="3B1F5240FFAF4517FCF6FCFAFC49FC7D" ID-GBIF-Occurrence="3418589311" collectingDate="1982-12-29" collectingDateMax="1995-04-18" collectingDateMin="1982-12-29" collectorName="MV Light &amp; O. S. Flint Jr." country="Guyana" county="Karanambo" latitude="5.6566668" location="Takutu Mountains" longLatPrecision="130" longitude="-57.591667" municipality="Mazarini Potaro district" pageId="5" pageNumber="20" specimenCount="3" specimenCount-male="3">
<emphasis id="B903840FFFAF4517FCF6FCFAFC71FC46" bold="true" box="[875,1005,885,911]" pageId="5" pageNumber="20">
<collectingCountry id="F360188DFFAF4517FCF6FCFAFC74FC46" box="[875,1000,885,911]" name="Guyana" pageId="5" pageNumber="20">GUYANA</collectingCountry>
:
</emphasis>
<collectingCounty id="62A92091FFAF4517FC6AFCF9FBE1FC59" box="[1015,1149,886,912]" pageId="5" pageNumber="20">Karanambo</collectingCounty>
,
<collectorName id="26823DCBFFAF4517FB17FCF9FA82FC59" box="[1162,1310,886,912]" pageId="5" pageNumber="20">Rupununi R.</collectorName>
,
<geoCoordinate id="EE433EDAFFAF4517FAB0FCF9FA04FC58" box="[1325,1432,886,913]" degrees="3" direction="north" minutes="45,1" orientation="latitude" pageId="5" pageNumber="20" precision="92" value="3.7516665">3°45,1´N</geoCoordinate>
,
<geoCoordinate id="EE433EDAFFAF4517FF0AFC15FE8FFC7C" box="[151,275,922,949]" degrees="59" direction="west" minutes="18,6" orientation="longitude" pageId="5" pageNumber="20" precision="92" value="-59.31">59°18,6´W</geoCoordinate>
,
<date id="FFC97EDDFFAF4517FEBFFC15FE48FC7D" box="[290,468,922,948]" pageId="5" pageNumber="20" value="1994-03-31" valueMax="1994-04-02" valueMin="1994-03-31">
<collectingDate id="EF8D8735FFAF4517FEBFFC15FE48FC7D" box="[290,468,922,948]" pageId="5" pageNumber="20" value="1994-03-31" valueMax="1994-04-02" valueMin="1994-03-31">31.iii2.iv.1994</collectingDate>
</date>
,
<collectorName id="26823DCBFFAF4517FE79FC15FD0BFC7D" box="[484,663,922,948]" pageId="5" pageNumber="20">MV Light, O.S</collectorName>
. Flint Jr.,
<specimenCount id="9D719394FFAF4517FC8DFC15FCC3FC7D" box="[784,863,922,948]" pageId="5" pageNumber="20" type="male">1 male</specimenCount>
(USNM);
</materialsCitation>
<materialsCitation id="3B1F5240FFAF4517FC42FC15FAAEFC11" ID-GBIF-Occurrence="3418589310" collectingDate="1982-12-29" collectingDateMax="1995-04-18" collectingDateMin="1982-12-29" collectorName="Lowry &amp; Williams &amp;" country="Guyana" county="Karanambo" latitude="5.6566668" location="Takutu Mountains" longLatPrecision="130" longitude="-57.591667" municipality="Mazarini Potaro district" pageId="5" pageNumber="20" specimenCount="3" specimenCount-male="3">
<collectingMunicipality id="6BACC267FFAF4517FC42FC15FB68FC7D" box="[991,1268,922,948]" pageId="5" pageNumber="20">Mazarini Potaro district</collectingMunicipality>
, Takutu Luber Camp.,
<location id="8EA80EC6FFAF4517FF6CFC31FE5FFC11" LSID="urn:lsid:plazi:treatment:03DEE90BFFAF4517FF0AFF18FAFDFA89:8EA80EC6FFAF4517FF6CFC31FE5FFC11" box="[241,451,958,984]" country="Guyana" county="Karanambo" latitude="5.6566668" longLatPrecision="130" longitude="-57.591667" municipality="Mazarini Potaro district" name="Takutu Mountains" pageId="5" pageNumber="20">Takutu Mountains</location>
,
<geoCoordinate id="EE433EDAFFAF4517FE4CFC31FDB8FC10" box="[465,548,958,985]" degrees="6.25" direction="north" orientation="latitude" pageId="5" pageNumber="20" precision="555" value="6.25">6.25°N</geoCoordinate>
,
<geoCoordinate id="EE433EDAFFAF4517FDAFFC31FD15FC10" box="[562,649,958,985]" degrees="50.0" direction="west" orientation="longitude" pageId="5" pageNumber="20" precision="5555" value="-50.0">50.0°W</geoCoordinate>
,
<date id="FFC97EDDFFAF4517FD0BFC31FC84FC11" box="[662,792,958,984]" pageId="5" pageNumber="20" value="1982-12-29">
<collectingDate id="EF8D8735FFAF4517FD0BFC31FC84FC11" box="[662,792,958,984]" pageId="5" pageNumber="20" value="1982-12-29">29.xii.1982</collectingDate>
</date>
, Steiner,
<collectorName id="26823DCBFFAF4517FC1BFC31FC4DFC11" box="[902,977,958,984]" pageId="5" pageNumber="20">Lowry</collectorName>
&amp;
<collectorName id="26823DCBFFAF4517FC64FC31FBFDFC11" box="[1017,1121,958,984]" pageId="5" pageNumber="20">Williams</collectorName>
,
<specimenCount id="9D719394FFAF4517FBF2FC31FB21FC11" box="[1135,1213,958,984]" pageId="5" pageNumber="20" type="male">1 male</specimenCount>
(USNM);
</materialsCitation>
<materialsCitation id="3B1F5240FFAF4517FAA1FC31FB31FC35" ID-GBIF-Occurrence="3418589318" collectingDate="1982-12-29" collectingDateMax="1995-04-18" collectingDateMin="1982-12-29" collectorName="O. S. Flint Jr." country="Guyana" county="Karanambo" latitude="5.6566668" location="Takutu Mountains" longLatPrecision="130" longitude="-57.591667" municipality="Mazarini Potaro district" pageId="5" pageNumber="20" specimenCount="3" specimenCount-male="3">
Dunulay ranch,
<location id="8EA80EC6FFAF4517FF7EFC6DFE09FC35" LSID="urn:lsid:plazi:treatment:03DEE90BFFAF4517FF0AFF18FAFDFA89:8EA80EC6FFAF4517FF7EFC6DFE09FC35" box="[227,405,994,1020]" country="Guyana" county="Karanambo" latitude="5.6566668" longLatPrecision="130" longitude="-57.591667" municipality="Mazarini Potaro district" name="Aramanti River" pageId="5" pageNumber="20">Aramanti River</location>
,
<geoCoordinate id="EE433EDAFFAF4517FE3FFC6DFD97FC34" box="[418,523,994,1021]" degrees="5" direction="north" minutes="39,4" orientation="latitude" pageId="5" pageNumber="20" precision="92" value="5.6566668">5°39,4´N</geoCoordinate>
,
<geoCoordinate id="EE433EDAFFAF4517FD84FC6DFD09FC34" box="[537,661,994,1021]" degrees="57" direction="west" minutes="35,5" orientation="longitude" pageId="5" pageNumber="20" precision="92" value="-57.591667">57°35,5´W</geoCoordinate>
,
<date id="FFC97EDDFFAF4517FD3CFC6DFCD8FC34" box="[673,836,994,1021]" pageId="5" pageNumber="20" value="1995-04-15" valueMax="1995-04-18" valueMin="1995-04-15">
<collectingDate id="EF8D8735FFAF4517FD3CFC6DFCD8FC34" box="[673,836,994,1021]" pageId="5" pageNumber="20" value="1995-04-15" valueMax="1995-04-18" valueMin="1995-04-15">1518.iv.1995</collectingDate>
</date>
,
<collectorName id="26823DCBFFAF4517FCCCFC6DFC78FC35" box="[849,996,994,1020]" pageId="5" pageNumber="20">O.S. Flint Jr.</collectorName>
,
<specimenCount id="9D719394FFAF4517FC6CFC6DFBA1FC35" box="[1009,1085,994,1020]" pageId="5" pageNumber="20" type="male">1 male</specimenCount>
(USNM)
</materialsCitation>
;
<materialsCitation id="3B1F5240FFAF4517FB27FC6EFE0EFB8D" ID-GBIF-Occurrence="3418589314" collectorName="Dpto &amp; Kochalka" country="Paraguay" location="Concepcion" pageId="5" pageNumber="20" specimenCount="12" specimenCount-male="12" stateProvince="Concepcion">
<emphasis id="B903840FFFAF4517FB27FC6EFAC0FC32" bold="true" box="[1210,1372,993,1019]" pageId="5" pageNumber="20">
<collectingCountry id="F360188DFFAF4517FB27FC6EFAC4FC32" box="[1210,1368,993,1019]" name="Paraguay" pageId="5" pageNumber="20">PARAGUAY</collectingCountry>
:
</emphasis>
<collectorName id="26823DCBFFAF4517FAF9FC6DFA00FC35" box="[1380,1436,994,1020]" pageId="5" pageNumber="20">Dpto</collectorName>
<collectingRegion id="49B396FFFFAF4517FF0AFB89FEBCFBE9" box="[151,288,1030,1056]" country="Paraguay" name="Concepcion" pageId="5" pageNumber="20">Concepción</collectingRegion>
,
<collectingRegion id="49B396FFFFAF4517FEB7FB89FE2FFBE9" box="[298,435,1030,1056]" country="Paraguay" name="Concepcion" pageId="5" pageNumber="20">Concepción</collectingRegion>
,
<collectingMethod id="5236200AFFAF4517FE20FB89FDC6FBE9" box="[445,602,1030,1056]" pageId="5" pageNumber="20">UV Light trap</collectingMethod>
between 5:4512:30 PM, warm night 5 days after heavy rain, J.A.
<collectorName id="26823DCBFFAF4517FADFFB89FF28FB8D" pageId="5" pageNumber="20">Kochalka</collectorName>
,
<specimenCount id="9D719394FFAF4517FF22FBA5FEBFFB8D" box="[191,291,1066,1092]" pageId="5" pageNumber="20" type="male">12 males</specimenCount>
(USNM)
</materialsCitation>
;
<materialsCitation id="3B1F5240FFAF4517FE00FBA6FE8CFBA1" ID-GBIF-Occurrence="3418589307" collectingDate="1959-11-29" country="Uruguay" location="Rio Uruguay" municipality="San Gregorio" pageId="5" pageNumber="20" specimenCount="9" specimenCount-male="9" stateProvince="Artigas">
<emphasis id="B903840FFFAF4517FE00FBA6FDACFB8A" bold="true" box="[413,560,1065,1091]" pageId="5" pageNumber="20">
<collectingCountry id="F360188DFFAF4517FE00FBA6FDB0FB8A" box="[413,556,1065,1091]" name="Uruguay" pageId="5" pageNumber="20">URUGUAY</collectingCountry>
:
</emphasis>
<collectingRegion id="49B396FFFFAF4517FDA8FBA5FD10FB8D" box="[565,652,1066,1092]" country="Uruguay" name="Artigas" pageId="5" pageNumber="20">Artigas</collectingRegion>
:
<collectingMunicipality id="6BACC267FFAF4517FD0BFBA5FCB3FB8D" box="[662,815,1066,1092]" pageId="5" pageNumber="20">San Gregorio</collectingMunicipality>
, orillas del
<location id="8EA80EC6FFAF4517FC2DFBA5FBDEFB8D" LSID="urn:lsid:plazi:treatment:03DEE90BFFAF4517FF0AFF18FAFDFA89:8EA80EC6FFAF4517FC2DFBA5FBDEFB8D" box="[944,1090,1066,1092]" country="Uruguay" municipality="San Gregorio" name="Rio Uruguay" pageId="5" pageNumber="20" stateProvince="Artigas">Río Uruguay</location>
,
<collectingMethod id="5236200AFFAF4517FBD0FBA5FB29FB8D" box="[1101,1205,1066,1092]" pageId="5" pageNumber="20">light trap</collectingMethod>
,
<date id="FFC97EDDFFAF4517FB5DFBA5FAA7FB8C" box="[1216,1339,1066,1093]" pageId="5" pageNumber="20" value="1959-11-29">
<collectingDate id="EF8D8735FFAF4517FB5DFBA5FAA7FB8C" box="[1216,1339,1066,1093]" pageId="5" pageNumber="20" value="1959-11-29">29.xi.1959</collectingDate>
</date>
,
<specimenCount id="9D719394FFAF4517FADBFBA5FA00FB8D" box="[1350,1436,1066,1092]" pageId="5" pageNumber="20" type="male">9 males</specimenCount>
(CFHCM)
</materialsCitation>
;
<materialsCitation id="3B1F5240FFAF4517FE81FBC2FB8CFB45" ID-GBIF-Occurrence="3418589321" collectingDate="1979-02-09" collectingDateMax="1983-01-20" collectingDateMin="1979-02-09" collectorName="C. M. &amp; O. S. Flint Jr." country="Venezuela" county="Fundo Masaguaral" location="Rio Caroni" municipality="Cano Caracol" pageId="5" pageNumber="20" specimenCount="2" specimenCount-male="2">
<emphasis id="B903840FFFAF4517FE81FBC2FE4FFBAE" bold="true" box="[284,467,1101,1127]" pageId="5" pageNumber="20">
<collectingCountry id="F360188DFFAF4517FE81FBC2FE53FBAE" box="[284,463,1101,1127]" name="Venezuela" pageId="5" pageNumber="20">VENEZUELA</collectingCountry>
:
</emphasis>
GU,
<collectingCounty id="62A92091FFAF4517FD8EFBC1FD75FBA1" box="[531,745,1102,1128]" pageId="5" pageNumber="20">Fundo Masaguaral</collectingCounty>
,
<collectingMunicipality id="6BACC267FFAF4517FD68FBC1FC0EFBA1" box="[757,914,1102,1128]" pageId="5" pageNumber="20">Caño Caracol</collectingMunicipality>
,
<date id="FFC97EDDFFAF4517FC03FBC1FBB3FBA1" box="[926,1071,1102,1128]" pageId="5" pageNumber="20" value="1983-01-19" valueMax="1983-01-20" valueMin="1983-01-19">
<collectingDate id="EF8D8735FFAF4517FC03FBC1FBB3FBA1" box="[926,1071,1102,1128]" pageId="5" pageNumber="20" value="1983-01-19" valueMax="1983-01-20" valueMin="1983-01-19">19-20.i.1983</collectingDate>
</date>
, O.S.
<collectorName id="26823DCBFFAF4517FBEBFBC1FB4CFBA1" box="[1142,1232,1102,1128]" pageId="5" pageNumber="20">Flint Jr.</collectorName>
,
<specimenCount id="9D719394FFAF4517FB41FBC1FAB4FBA1" box="[1244,1320,1102,1128]" pageId="5" pageNumber="20" type="male">1 male</specimenCount>
(USNM);
<location id="8EA80EC6FFAF4517FF0AFBFDFE8AFB45" LSID="urn:lsid:plazi:treatment:03DEE90BFFAF4517FF0AFF18FAFDFA89:8EA80EC6FFAF4517FF0AFBFDFE8AFB45" box="[151,278,1138,1164]" country="Venezuela" county="Fundo Masaguaral" municipality="Cano Caracol" name="Rio Caroni" pageId="5" pageNumber="20">Río Caroni</location>
, at
<location id="8EA80EC6FFAF4517FEA1FBFDFE7CFB45" LSID="urn:lsid:plazi:treatment:03DEE90BFFAF4517FF0AFF18FAFDFA89:8EA80EC6FFAF4517FEA1FBFDFE7CFB45" box="[316,480,1138,1164]" country="Venezuela" county="Fundo Masaguaral" municipality="Cano Caracol" name="Paso Caruachi" pageId="5" pageNumber="20">Paso Caruachi</location>
,
<date id="FFC97EDDFFAF4517FE76FBFDFDCCFB45" box="[491,592,1138,1164]" pageId="5" pageNumber="20" value="1979-02-09">
<collectingDate id="EF8D8735FFAF4517FE76FBFDFDCCFB45" box="[491,592,1138,1164]" pageId="5" pageNumber="20" value="1979-02-09">9.ii.1979</collectingDate>
</date>
,
<collectorName id="26823DCBFFAF4517FDC0FBFDFCD7FB45" box="[605,843,1138,1164]" pageId="5" pageNumber="20">C.M. &amp; O.S. Flint Jr.</collectorName>
,
<specimenCount id="9D719394FFAF4517FCCBFBFDFC3DFB45" box="[854,929,1138,1164]" pageId="5" pageNumber="20" type="male">1 male</specimenCount>
(USNM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFAF4517FF5AFB1AFAFDFA89" pageId="5" pageNumber="20" type="distribution">
<paragraph id="8BC8581DFFAF4517FF5AFB1AFAFDFA89" blockId="5.[151,1437,417,1344]" pageId="5" pageNumber="20">
<emphasis id="B903840FFFAF4517FF5AFB1AFEFDFB66" bold="true" box="[199,353,1173,1199]" pageId="5" pageNumber="20">Distribution.</emphasis>
<taxonomicName id="4C77239EFFAF4517FEF4FB19FDCBFB79" box="[361,599,1174,1200]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="marginalis">
<emphasis id="B903840FFFAF4517FEF4FB19FDCBFB79" box="[361,599,1174,1200]" italics="true" pageId="5" pageNumber="20">Cyrnellus marginalis</emphasis>
</taxonomicName>
is distributed in various South American countries but was collected and described for the first time in United States of America by Banks from Put-In-Bay, Ohio. Banks material could not be reviewed; his specimen could belong to
<taxonomicName id="4C77239EFFAF4517FDECFB51FD6BFB31" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[625,759,1246,1272]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B903840FFFAF4517FDECFB51FD6BFB31" box="[625,759,1246,1272]" italics="true" pageId="5" pageNumber="20">C. fraternus</emphasis>
</taxonomicName>
, but because the two species have been considered synonyms since 1964, all existing North American records are suspect and need to be reviewed. It is reported for the first time from Bolivia.
<emphasis id="B903840FFFAF4517FEA8FAAAFD8AFA89" bold="true" box="[309,534,1317,1344]" pageId="5" pageNumber="20">
<collectingCountry id="F360188DFFAF4517FEA8FAAAFE15FAF6" box="[309,393,1317,1343]" name="Bolivia" pageId="5" pageNumber="20">Bolivia</collectingCountry>
new record
</emphasis>
, Brazil, Guyana, Paraguay, Uruguay, United States of America, Venezuela.
</paragraph>
</subSubSection>
</treatment>
</document>