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<document id="F6B3770620FEFD877537C0C389F75B04" ID-CLB-Dataset="21852" ID-DOI="10.11646/zootaxa.4780.3.3" ID-GBIF-Dataset="70713ddc-4f67-4f0a-8625-41aa37abb017" ID-ISSN="1175-5326" ID-Zenodo-Dep="3855256" ID-ZooBank="CA5DA614-D448-4D95-B271-DDC8D56A37F60" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590481028709" checkinUser="plazi" docAuthor="Dias, Fernando Maia Silva, Dolibaina, Diego Rodrigo, Mielke, Olaf Hermann Hendrik &amp; Casagrande, Mirna Martins" docDate="2020" docId="03DD0A61FFE9FFE4AD94B044FB9DFD2F" docLanguage="en" docName="zootaxa.4780.3.3.pdf" docOrigin="Zootaxa 4780 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Symmachia uirassu Dolibaina, Dias &amp; Casagrande 2020, sp. nov." docType="treatment" docUuid="A0AC5F0F-D580-4484-BE4F-C1314694EFA6" docUuidSource="ZooBank" docVersion="8" lastPageNumber="499" masterDocId="FFE47219FFF3FFF8AD03B529FFC2FFE0" masterDocTitle="Taxonomic review of the “ Probetoriformes ” species group of Symmachia Hübner [1819] (Lepidoptera: Riodinidae), with the description of two new species from the Brazilian Atlantic Forest" masterLastPageNumber="507" masterPageNumber="471" pageNumber="497" updateTime="1698841375024" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2F851589337ACE3B6306551ED7DFB0D6">Taxonomic review of the “ Probetoriformes ” species group of Symmachia Hübner [1819] (Lepidoptera: Riodinidae), with the description of two new species from the Brazilian Atlantic Forest</mods:title>
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<mods:namePart id="3983C43AB5494281724DD414758B853D">Dias, Fernando Maia Silva</mods:namePart>
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<mods:namePart id="53FC1638995EE18270E3BE5F06F763F0">Dolibaina, Diego Rodrigo</mods:namePart>
<mods:affiliation id="37F86D01BD246D152516702064B9128F">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil. &amp; dirodrido @ gmail. com; https: // orcid. org / 0000 - 0001 - 5258 - 3787</mods:affiliation>
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<mods:namePart id="A6D6B5B9F1A35EEFFF253E7216EE95AD">Mielke, Olaf Hermann Hendrik</mods:namePart>
<mods:affiliation id="8869A807F2E0F8082F3026AF51DD0DCC">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil. &amp; omhesp @ ufpr. br; https: // orcid. org / 0000 - 0003 - 3655 - 4606</mods:affiliation>
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<mods:namePart id="652253EB97BE0989B8EA626B6D4FBB64">Casagrande, Mirna Martins</mods:namePart>
<mods:affiliation id="300520A6FE344133C7621BFF6D3A85DC">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil. &amp; mibras @ ufpr. br; https: // orcid. org / 0000 - 0002 - 6076 - 8463</mods:affiliation>
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<treatment id="03DD0A61FFE9FFE4AD94B044FB9DFD2F" ID-DOI="http://doi.org/10.5281/zenodo.3857488" ID-GBIF-Taxon="164299928" ID-Zenodo-Dep="3857488" LSID="urn:lsid:zoobank.org:act:A0AC5F0F-D580-4484-BE4F-C1314694EFA6" httpUri="http://treatment.plazi.org/id/03DD0A61FFE9FFE4AD94B044FB9DFD2F" lastPageId="28" lastPageNumber="499" pageId="26" pageNumber="497">
<subSubSection id="C36EE8FCFFE9FFE2AD94B044FCB5FA67" box="[151,887,1389,1416]" pageId="26" pageNumber="497" type="nomenclature">
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<heading id="D0830C1BFFE9FFE2AD94B044FCB5FA67" bold="true" box="[151,887,1389,1416]" fontSize="11" level="1" pageId="26" pageNumber="497" reason="1">
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<taxonomicName id="4C74C0F4FFE9FFE2AD94B044FCCFFA67" authority="Dolibaina, Dias &amp; Casagrande" authorityName="Dolibaina, Dias &amp; Casagrande" authorityYear="2020" box="[151,781,1389,1416]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="uirassu" status="sp. nov.">
<emphasis id="B9006765FFE9FFE2AD94B044FE44FA67" bold="true" box="[151,390,1389,1416]" italics="true" pageId="26" pageNumber="497">Symmachia uirassu</emphasis>
Dolibaina, Dias &amp; Casagrande
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,
<taxonomicNameLabel id="A233DA1EFFE9FFE2AE1AB044FCB5FA67" box="[793,887,1389,1415]" pageId="26" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C36EE8FCFFE9FFE2AD94B0B8FB65F9D0" pageId="26" pageNumber="497" type="description">
<paragraph id="8BCBBB77FFE9FFE2AD94B0B8FBCBFA2F" blockId="26.[151,1033,1389,1487]" pageId="26" pageNumber="497">
(
<figureCitation id="134FA7F2FFE9FFE2AD9CB0B8FEE2FA4B" box="[159,288,1424,1451]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="26" pageNumber="497">Figs 25–26</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2AC2FB0B9FE88FA4B" box="[300,330,1424,1451]" captionStart="FIGURES 39–47" captionStartId="7.[151,264,1432,1457]" captionTargetBox="[151,1436,273,1406]" captionTargetId="figure@7.[151,1436,273,1406]" captionTargetPageId="7" captionText="FIGURES 39–47. Heads of the Symmachia belonging to the “Probetoriformes” species group in anterior view. 39. S. probetor. 40. S. belti stat. rev. 41. S. championi stat. rest. 42. S. estellina. 43. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov.. 44. S. praxila. 45. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 46. S. divisora. 47. S. falcistriga. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3855264" httpUri="https://zenodo.org/record/3855264/files/figure.png" pageId="26" pageNumber="497">45</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2AC55B0B9FEB6FA4B" box="[342,372,1424,1451]" captionStart="FIGURES 48–56" captionStartId="8.[151,264,1515,1540]" captionTargetBox="[151,1436,358,1490]" captionTargetId="figure@8.[151,1436,358,1490]" captionTargetPageId="8" captionText="FIGURES 48–56. Heads of the Symmachia belonging to the “Probetoriformes” species group in lateral view. 48. S. probetor. 49. S. belti stat. rev.. 50. S. championi stat. rest. 51. S. estellina. 52. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov.. 53. S. praxila. 54. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 55. S. divisora. 56. S. falcistriga. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3855266" httpUri="https://zenodo.org/record/3855266/files/figure.png" pageId="26" pageNumber="497">54</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2AC83B0B8FE5CFA4B" box="[384,414,1425,1451]" captionStart="FIGURES 57–66" captionStartId="10.[151,264,1222,1247]" captionTargetBox="[155,1433,672,1192]" captionTargetId="figure@10.[151,1436,669,1197]" captionTargetPageId="10" captionText="FIGURES 57–66. Forewings of males of Symmachia belonging to the “Probetoriformes” species group in dorsal view. 57. S. probetor. 58. S. belti stat. rev. 59. S. championi stat. rest. 60. S. estellina. 61. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov.. 62. S. praxila. 63. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 64. S. divisora. 65. S. falcistriga. 66. S. meyi stat. nov." figureDoi="http://doi.org/10.5281/zenodo.3855268" httpUri="https://zenodo.org/record/3855268/files/figure.png" pageId="26" pageNumber="497">63</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2ACA9B0B8FE30FA4B" box="[426,498,1425,1451]" captionStart="FIGURES 95–102" captionStartId="13.[151,264,1478,1503]" captionTargetBox="[152,1436,182,1451]" captionTargetId="figure@13.[151,1436,181,1452]" captionTargetPageId="13" captionText="FIGURES 95–102. Abdomens of males of Symmachia belonging to the “Probetoriformes” species group in dorsal (left) and lateral (right) views. 95–96. S. praxila. 97–98. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 99–100. S. divisora. 101–102. S. falcistriga. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3855282" httpUri="https://zenodo.org/record/3855282/files/figure.png" pageId="26" pageNumber="497">97–98</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2ACFDB0B9FDA2FA4B" box="[510,608,1424,1451]" captionStart="FIGURES 113–122" captionStartId="16.[151,264,1642,1667]" captionTargetBox="[152,1408,187,1605]" captionTargetId="figure@16.[151,1436,181,1616]" captionTargetPageId="16" captionText="FIGURES 113–122. Fourth (left) and fifith (right) tergites of males of Symmachia belonging to the “Probetoriformes” species group. 113–114. S. praxila. 115–116. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 117–118. S. divisora. 119–120. S. falcistriga. 121–122. S. meyi stat. nov.. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3855286" httpUri="https://zenodo.org/record/3855286/files/figure.png" pageId="26" pageNumber="497">115–116</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2AF6FB0B8FD5AFA4B" box="[620,664,1425,1451]" captionStart="FIGURES 128–132" captionStartId="19.[151,264,1154,1179]" captionTargetBox="[155,1432,186,1123]" captionTargetId="figure@19.[151,1436,181,1129]" captionTargetPageId="19" captionText="FIGURES 128–132. Male genitalia of Symmachia belonging to the “Probetoriformes” species group in lateral view, uncus in dorsal view and anterior projection of saccus in ventral view. 128. S. praxila. 129. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 130. S. divisora. 131. S. falcistriga. 132. S. meyi stat. nov.. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3855292" httpUri="https://zenodo.org/record/3855292/files/figure.png" pageId="26" pageNumber="497">129</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2AFA7B0B8FD12FA4B" box="[676,720,1425,1451]" captionStart="FIGURES 138–146" captionStartId="20.[151,264,1576,1601]" captionTargetBox="[155,1432,185,1546]" captionTargetId="figure@20.[151,1436,181,1551]" captionTargetPageId="20" captionText="FIGURES 138–146. Female genitalia of Symmachia belonging to the “Probetoriformes” species group in ventral view and papilla analis in lateral view. 138. S. probetor. 139. S. belti stat. rev. 140. S. championi stat. rest. 141. S. estellina. 142. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov. 143. S. praxila. 144. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 145. S. divisora. 146. S. falcistriga. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3855299" httpUri="https://zenodo.org/record/3855299/files/figure.png" pageId="26" pageNumber="497">144</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2AFDFB0B9FCCAFA4B" box="[732,776,1424,1451]" captionStart="FIGURE 153" captionStartId="28.[151,250,1806,1831]" captionTargetBox="[151,1436,740,1780]" captionTargetId="figure@28.[151,1436,740,1780]" captionTargetPageId="28" captionText="FIGURE 153. Geographical distribution of Symmachia praxila, S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov., S. divisora, S. falcistriga and S. meyi stat. nov. Asterisks indicate records from the literature." figureDoi="http://doi.org/10.5281/zenodo.3855307" httpUri="https://zenodo.org/record/3855307/files/figure.png" pageId="26" pageNumber="497">153</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2AE17B0B9FCFDFA4B" box="[788,831,1424,1451]" captionStart="FIGURES 154–156" captionStartId="29.[151,264,1034,1059]" captionTargetBox="[151,1436,181,1008]" captionTargetId="figure@29.[151,1436,181,1008]" captionTargetPageId="29" captionText="FIGURES 154–156. Habitat of some species of Symmachia belonging to the “Probetoriformes” species group. 154. Hilltop at the Bromélias trail, Reserva Serra Bonita, Camacan, Bahia, Brazil, the arrow indicates the exact location where males of S. atlantica Dias, Dolibaina &amp; Casagrande sp. nov. were collected. 155. Opposite view of the previous figure, arrow indicates the exact location where males of S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. were collected. 156. DRD on a tree at the lightgap of the hilltop in Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil where males of S. probetor, S. estellina, S. divisora and S. falcistriga were collected (photo by Eduardo Carneiro)." figureDoi="http://doi.org/10.5281/zenodo.3855309" httpUri="https://zenodo.org/record/3855309/files/figure.png" pageId="26" pageNumber="497">155</figureCitation>
) http://
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<taxonomicName id="4C74C0F4FFE9FFE2AD94B0D0FECEF9F1" authorityName="Hubner" authorityYear="1819" box="[151,268,1529,1553]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="B9006765FFE9FFE2AD94B0D0FECEF9F1" box="[151,268,1529,1553]" italics="true" pageId="26" pageNumber="497">Symmachia</emphasis>
</taxonomicName>
? sp.;
<bibRefCitation id="EFE5C686FFE9FFE2AC4DB0D0FE30F9F2" author="D'Abrera, B. L." box="[334,498,1529,1554]" pageId="26" pageNumber="497" refId="ref32991" refString="D'Abrera, B. L. (1994) Butterflies of the Neotropical Region. Part VI. Riodinidae. Black Rock, Hill House, Victoria, 217 pp." type="book" year="1994">D’Abrera, 1994</bibRefCitation>
.
<emphasis id="B9006765FFE9FFE2ACF8B0D3FCD3F9F2" bold="true" box="[507,785,1529,1554]" pageId="26" pageNumber="497">Butt. Neot. Reg. IV Riod.,</emphasis>
p. 1044, figs [1–2] (male dorsal and ventral).
</paragraph>
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<taxonomicName id="4C74C0F4FFE9FFE2AD94B331FE99F9D0" authorityName="Westwood" authorityYear="1851" box="[151,347,1560,1584]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="praxila">
<emphasis id="B9006765FFE9FFE2AD94B331FE99F9D0" box="[151,347,1560,1584]" italics="true" pageId="26" pageNumber="497">Symmachia praxila</emphasis>
</taxonomicName>
[
<emphasis id="B9006765FFE9FFE2AC69B33EFDE2F9D0" bold="true" box="[362,544,1559,1584]" pageId="26" pageNumber="497">misidentification</emphasis>
];
<bibRefCitation id="EFE5C686FFE9FFE2AF35B33EFCCFF9D0" author="Hall, J. P. W. &amp; Harvey, D. J." box="[566,781,1559,1585]" pageId="26" pageNumber="497" pagination="7 - 10" refId="ref33740" refString="Hall, J. P. W. &amp; Harvey, D. J. (2007) Three new species of Symmachiini from Panama and Colombia (Lepidoptera: Riodinidae). Tropical Lepidoptera, 16:, 7 - 10." type="journal article" year="2007">Hall &amp; Harvey, 2007</bibRefCitation>
.
<emphasis id="B9006765FFE9FFE2AE14B331FBD1F9D0" bold="true" box="[791,1043,1559,1584]" pageId="26" pageNumber="497">Trop. Lep. Res. 16(1–2)</emphasis>
, p. 7 [
<emphasis id="B9006765FFE9FFE2A957B330FB5AF9D0" box="[1108,1176,1561,1584]" italics="true" pageId="26" pageNumber="497">in part</emphasis>
].
</paragraph>
</subSubSection>
<subSubSection id="C36EE8FCFFE9FFE2AD94B371FF10F83B" pageId="26" pageNumber="497" type="diagnosis">
<paragraph id="8BCBBB77FFE9FFE2AD94B371FF10F83B" blockId="26.[151,1437,1624,2011]" pageId="26" pageNumber="497">
<emphasis id="B9006765FFE9FFE2AD94B371FED0F993" bold="true" box="[151,274,1624,1651]" pageId="26" pageNumber="497">Diagnosis.</emphasis>
<taxonomicName id="4C74C0F4FFE9FFE2AC1AB370FE36F993" authorityName="Dolibaina, Dias &amp; Casagrande" authorityYear="2020" box="[281,500,1625,1651]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="uirassu">
<emphasis id="B9006765FFE9FFE2AC1AB370FE36F993" box="[281,500,1625,1651]" italics="true" pageId="26" pageNumber="497">Symmachia uirassu</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="4C74C0F4FFE9FFE2AFC6B373FC84F993" authorityName="Dias, Dolibaina &amp; Casagrande" authorityYear="2020" box="[709,838,1625,1652]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="B9006765FFE9FFE2AFC6B373FC84F993" box="[709,838,1625,1652]" italics="true" pageId="26" pageNumber="497">S. atlantica</emphasis>
</taxonomicName>
(
<figureCitation id="134FA7F2FFE9FFE2AE56B370FC37F993" box="[853,1013,1624,1651]" captionStart-0="FIGURE 152" captionStart-1="FIGURE 153" captionStartId-0="26.[151,250,1246,1271]" captionStartId-1="28.[151,250,1806,1831]" captionTargetBox-0="[151,1436,181,1221]" captionTargetBox-1="[151,1436,740,1780]" captionTargetId-0="figure@26.[151,1436,181,1221]" captionTargetId-1="figure@28.[151,1436,740,1780]" captionTargetPageId-0="26" captionTargetPageId-1="28" captionText-0="FIGURE 152. Geographical distribution of Symmachia estellina and S. atlantica Dias, Dolibaina &amp; Casagrande sp. nov. Asterisks indicate records from the literature." captionText-1="FIGURE 153. Geographical distribution of Symmachia praxila, S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov., S. divisora, S. falcistriga and S. meyi stat. nov. Asterisks indicate records from the literature." figureDoi-0="http://doi.org/10.5281/zenodo.3855305" figureDoi-1="http://doi.org/10.5281/zenodo.3855307" httpUri-0="https://zenodo.org/record/3855305/files/figure.png" httpUri-1="https://zenodo.org/record/3855307/files/figure.png" pageId="26" pageNumber="497">Figs 152–153</figureCitation>
). Males are most similar to
<taxonomicName id="4C74C0F4FFE9FFE2A82CB373FA5EF993" authorityName="Westwood" authorityYear="1851" box="[1327,1436,1625,1652]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="praxila">
<emphasis id="B9006765FFE9FFE2A82CB373FA5EF993" box="[1327,1436,1625,1652]" italics="true" pageId="26" pageNumber="497">S. praxila</emphasis>
</taxonomicName>
and
<taxonomicName id="4C74C0F4FFE9FFE2ADC4B357FE82F977" authorityName="Dolibaina &amp; Leite" authorityYear="2012" box="[199,320,1661,1688]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="divisora">
<emphasis id="B9006765FFE9FFE2ADC4B357FE82F977" box="[199,320,1661,1688]" italics="true" pageId="26" pageNumber="497">S. divisora</emphasis>
</taxonomicName>
(
<figureCitation id="134FA7F2FFE9FFE2AC4CB354FE12F977" box="[335,464,1660,1687]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="26" pageNumber="497">Figs 25–26</figureCitation>
) and females although unknown likely to be similar to most species of the “Probetoriformes” species group. Males with frons, patagia, dorsal thorax and tegulae as in
<taxonomicName id="4C74C0F4FFE9FFE2A936B38BFB61F95B" authorityName="Westwood" authorityYear="1851" box="[1077,1187,1697,1724]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="praxila">
<emphasis id="B9006765FFE9FFE2A936B38BFB61F95B" box="[1077,1187,1697,1724]" italics="true" pageId="26" pageNumber="497">S. praxila</emphasis>
</taxonomicName>
; FW strongly bulged; bulged area about 38% of the FW length; end of bulge aligned with the origin of M
<subScript id="17F0B932FFE9FFE2A95BB3FEFBA2F905" attach="left" box="[1112,1120,1751,1765]" fontSize="6" pageId="26" pageNumber="497">3</subScript>
(
<figureCitation id="134FA7F2FFE9FFE2A96EB3ECFB07F93F" box="[1133,1221,1732,1759]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="26" pageNumber="497">Figs 25</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2A9D0B3ECFB30F900" box="[1235,1266,1733,1760]" captionStart="FIGURES 57–66" captionStartId="10.[151,264,1222,1247]" captionTargetBox="[155,1433,672,1192]" captionTargetId="figure@10.[151,1436,669,1197]" captionTargetPageId="10" captionText="FIGURES 57–66. Forewings of males of Symmachia belonging to the “Probetoriformes” species group in dorsal view. 57. S. probetor. 58. S. belti stat. rev. 59. S. championi stat. rest. 60. S. estellina. 61. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov.. 62. S. praxila. 63. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 64. S. divisora. 65. S. falcistriga. 66. S. meyi stat. nov." figureDoi="http://doi.org/10.5281/zenodo.3855268" httpUri="https://zenodo.org/record/3855268/files/figure.png" pageId="26" pageNumber="497">63</figureCitation>
); FWD mostly black, with reddish orange spots on the basal area and a dark green sheen on the discal and postdiscal areas, and some light bluish green scaling on 1A (
<figureCitation id="134FA7F2FFE9FFE2AF52B224FD6AF8C7" box="[593,680,1804,1831]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="26" pageNumber="497">Figs 25</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2AFB7B224FD11F8C8" box="[692,723,1805,1832]" captionStart="FIGURES 57–66" captionStartId="10.[151,264,1222,1247]" captionTargetBox="[155,1433,672,1192]" captionTargetId="figure@10.[151,1436,669,1197]" captionTargetPageId="10" captionText="FIGURES 57–66. Forewings of males of Symmachia belonging to the “Probetoriformes” species group in dorsal view. 57. S. probetor. 58. S. belti stat. rev. 59. S. championi stat. rest. 60. S. estellina. 61. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov.. 62. S. praxila. 63. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 64. S. divisora. 65. S. falcistriga. 66. S. meyi stat. nov." figureDoi="http://doi.org/10.5281/zenodo.3855268" httpUri="https://zenodo.org/record/3855268/files/figure.png" pageId="26" pageNumber="497">63</figureCitation>
); FWD with a translucent white and creamy sickle-shaped spot between the discal cell and the costal margin, a small round whitish spots on the postdiscal area in R
<subScript id="17F0B932FFE9FFE2A9E4B26AFB2DF8B1" attach="left" box="[1255,1263,1859,1873]" fontSize="6" pageId="26" pageNumber="497">2</subScript>
–R
<subScript id="17F0B932FFE9FFE2A813B26AFADAF8B1" attach="left" box="[1296,1304,1859,1873]" fontSize="6" pageId="26" pageNumber="497">3</subScript>
and M
<subScript id="17F0B932FFE9FFE2A866B26AFAAFF8B1" attach="left" box="[1381,1389,1859,1873]" fontSize="6" pageId="26" pageNumber="497">1</subScript>
–M
<subScript id="17F0B932FFE9FFE2A897B26AFA5EF8B1" attach="left" box="[1428,1436,1859,1873]" fontSize="6" pageId="26" pageNumber="497">2</subScript>
near the discal cell (
<figureCitation id="134FA7F2FFE9FFE2AC74B27CFE0CF88F" box="[375,462,1876,1903]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="26" pageNumber="497">Figs 25</figureCitation>
,
<figureCitation id="134FA7F2FFE9FFE2ACDAB27CFE3AF890" box="[473,504,1877,1904]" captionStart="FIGURES 57–66" captionStartId="10.[151,264,1222,1247]" captionTargetBox="[155,1433,672,1192]" captionTargetId="figure@10.[151,1436,669,1197]" captionTargetPageId="10" captionText="FIGURES 57–66. Forewings of males of Symmachia belonging to the “Probetoriformes” species group in dorsal view. 57. S. probetor. 58. S. belti stat. rev. 59. S. championi stat. rest. 60. S. estellina. 61. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov.. 62. S. praxila. 63. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 64. S. divisora. 65. S. falcistriga. 66. S. meyi stat. nov." figureDoi="http://doi.org/10.5281/zenodo.3855268" httpUri="https://zenodo.org/record/3855268/files/figure.png" pageId="26" pageNumber="497">63</figureCitation>
); HWD homogeneously dark brown, with some reddish orange scales along the inner margin (
<figureCitation id="134FA7F2FFE9FFE2AC23B250FEB1F873" box="[288,371,1912,1939]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="26" pageNumber="497">Fig. 25</figureCitation>
); abdomen color pattern as in
<taxonomicName id="4C74C0F4FFE9FFE2AFC7B253FCF3F873" authorityName="Westwood" authorityYear="1851" box="[708,817,1913,1940]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="praxila">
<emphasis id="B9006765FFE9FFE2AFC7B253FCF3F873" box="[708,817,1913,1940]" italics="true" pageId="26" pageNumber="497">S. praxila</emphasis>
</taxonomicName>
(
<figureCitation id="134FA7F2FFE9FFE2AE3CB250FC00F873" box="[831,962,1913,1939]" captionStart="FIGURES 95–102" captionStartId="13.[151,264,1478,1503]" captionTargetBox="[152,1436,182,1451]" captionTargetId="figure@13.[151,1436,181,1452]" captionTargetPageId="13" captionText="FIGURES 95–102. Abdomens of males of Symmachia belonging to the “Probetoriformes” species group in dorsal (left) and lateral (right) views. 95–96. S. praxila. 97–98. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 99–100. S. divisora. 101–102. S. falcistriga. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3855282" httpUri="https://zenodo.org/record/3855282/files/figure.png" pageId="26" pageNumber="497">Figs 97–98</figureCitation>
); CAS size and color as in
<taxonomicName id="4C74C0F4FFE9FFE2A9ECB253FA9EF873" authorityName="Westwood" authorityYear="1851" box="[1263,1372,1913,1940]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="praxila">
<emphasis id="B9006765FFE9FFE2A9ECB253FA9EF873" box="[1263,1372,1913,1940]" italics="true" pageId="26" pageNumber="497">S. praxila</emphasis>
</taxonomicName>
(
<figureCitation id="134FA7F2FFE9FFE2A869B250FF3EF857" captionStart="FIGURES 113–122" captionStartId="16.[151,264,1642,1667]" captionTargetBox="[152,1408,187,1605]" captionTargetId="figure@16.[151,1436,181,1616]" captionTargetPageId="16" captionText="FIGURES 113–122. Fourth (left) and fifith (right) tergites of males of Symmachia belonging to the “Probetoriformes” species group. 113–114. S. praxila. 115–116. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 117–118. S. divisora. 119–120. S. falcistriga. 121–122. S. meyi stat. nov.. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3855286" httpUri="https://zenodo.org/record/3855286/files/figure.png" pageId="26" pageNumber="497">Figs 115–116</figureCitation>
); valva larger than in
<taxonomicName id="4C74C0F4FFE9FFE2ACF5B2B7FDB7F857" baseAuthorityName="Stoll" baseAuthorityYear="1782" box="[502,629,1949,1976]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="497" phylum="Arthropoda" rank="species" species="probetor">
<emphasis id="B9006765FFE9FFE2ACF5B2B7FDB7F857" box="[502,629,1949,1976]" italics="true" pageId="26" pageNumber="497">S. probetor</emphasis>
</taxonomicName>
; aedeagus thick and long; vesica with a rounded patch of cornuti (
<figureCitation id="134FA7F2FFE9FFE2A86CB2B4FF07F83B" captionStart="FIGURES 128–132" captionStartId="19.[151,264,1154,1179]" captionTargetBox="[155,1432,186,1123]" captionTargetId="figure@19.[151,1436,181,1129]" captionTargetPageId="19" captionText="FIGURES 128–132. Male genitalia of Symmachia belonging to the “Probetoriformes” species group in lateral view, uncus in dorsal view and anterior projection of saccus in ventral view. 128. S. praxila. 129. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 130. S. divisora. 131. S. falcistriga. 132. S. meyi stat. nov.. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3855292" httpUri="https://zenodo.org/record/3855292/files/figure.png" pageId="26" pageNumber="497">Fig. 129</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36EE8FCFFE8FFE3ADC4B5B1FE5CF903" pageId="27" pageNumber="498" type="description">
<paragraph id="8BCBBB77FFE8FFE3ADC4B5B1FDF4FEA3" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B5B1FE9BFF53" bold="true" box="[199,345,152,179]" pageId="27" pageNumber="498">Description.</emphasis>
Male.
<emphasis id="B9006765FFE8FFE3ACADB5B0FE29FF53" box="[430,491,153,179]" italics="true" pageId="27" pageNumber="498">Head</emphasis>
(
<figureCitation id="134FA7F2FFE8FFE3ACFEB5B0FD95FF53" box="[509,599,152,179]" captionStart="FIGURES 39–47" captionStartId="7.[151,264,1432,1457]" captionTargetBox="[151,1436,273,1406]" captionTargetId="figure@7.[151,1436,273,1406]" captionTargetPageId="7" captionText="FIGURES 39–47. Heads of the Symmachia belonging to the “Probetoriformes” species group in anterior view. 39. S. probetor. 40. S. belti stat. rev. 41. S. championi stat. rest. 42. S. estellina. 43. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov.. 44. S. praxila. 45. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 46. S. divisora. 47. S. falcistriga. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3855264" httpUri="https://zenodo.org/record/3855264/files/figure.png" pageId="27" pageNumber="498">Figs 45</figureCitation>
,
<figureCitation id="134FA7F2FFE8FFE3AF65B5B1FD44FF53" box="[614,646,152,179]" captionStart="FIGURES 48–56" captionStartId="8.[151,264,1515,1540]" captionTargetBox="[151,1436,358,1490]" captionTargetId="figure@8.[151,1436,358,1490]" captionTargetPageId="8" captionText="FIGURES 48–56. Heads of the Symmachia belonging to the “Probetoriformes” species group in lateral view. 48. S. probetor. 49. S. belti stat. rev.. 50. S. championi stat. rest. 51. S. estellina. 52. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov.. 53. S. praxila. 54. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 55. S. divisora. 56. S. falcistriga. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3855266" httpUri="https://zenodo.org/record/3855266/files/figure.png" pageId="27" pageNumber="498">54</figureCitation>
): mostly dark brown and black, with some scattered reddish orange scales on the frons and bordering the compound eyes (some of the orange may have been lost); antenna black, each segment with proximal white scales, antennal club formed by the last 10–12 segments tip yellowish; labial palpus dark brown, short, not extending beyond middle of frons, first segment as long as half of the second segment, third segment about 1/6 as long as second.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B464FF20FE33" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B464FEDAFE87" box="[199,280,333,359]" italics="true" pageId="27" pageNumber="498">Thorax</emphasis>
(
<figureCitation id="134FA7F2FFE8FFE3AC2AB464FE6CFE87" box="[297,430,332,359]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="27" pageNumber="498">Figs 25–26</figureCitation>
): ventrally brown and dark brown; prothorax dorsally with reddish orange scales; mesothorax dorsally black, with a dark bluish sheen and some reddish orange scales; metathorax dorsally with long, dark brown and some reddish orange scales; legs mostly dark brown, each segment ventrally with creamy and white scales.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B4F7FF31FDDF" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B4F7FE66FE17" box="[199,420,477,503]" italics="true" pageId="27" pageNumber="498">FW, shape and size:</emphasis>
average size
<quantity id="4C8C1692FFE8FFE3AF3BB4F4FD52FE17" box="[568,656,477,503]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="27" pageNumber="498" unit="cm" value="1.16">1.16 cm</quantity>
(
<quantity id="4C8C1692FFE8FFE3AF9DB4F4FCDDFE17" box="[670,799,477,503]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="27" pageNumber="498" unit="cm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 cm</quantity>
; n=3); roughly triangular; costal margin sinuous, strongly bulged in the basal area; apex pointed, angled; outer margin almost straight; tornus angled; inner margin almost straight.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B763FE50FCDB" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B763FEC6FD83" box="[199,260,586,611]" italics="true" pageId="27" pageNumber="498">FWD</emphasis>
(
<figureCitation id="134FA7F2FFE8FFE3AC17B760FEAEFD83" box="[276,364,584,611]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="27" pageNumber="498">Figs 25</figureCitation>
,
<figureCitation id="134FA7F2FFE8FFE3AC7AB760FE5BFD84" box="[377,409,585,612]" captionStart="FIGURES 57–66" captionStartId="10.[151,264,1222,1247]" captionTargetBox="[155,1433,672,1192]" captionTargetId="figure@10.[151,1436,669,1197]" captionTargetPageId="10" captionText="FIGURES 57–66. Forewings of males of Symmachia belonging to the “Probetoriformes” species group in dorsal view. 57. S. probetor. 58. S. belti stat. rev. 59. S. championi stat. rest. 60. S. estellina. 61. S. atlantica Dias, Dolibaina &amp; Mielke sp. nov.. 62. S. praxila. 63. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 64. S. divisora. 65. S. falcistriga. 66. S. meyi stat. nov." figureDoi="http://doi.org/10.5281/zenodo.3855268" httpUri="https://zenodo.org/record/3855268/files/figure.png" pageId="27" pageNumber="498">63</figureCitation>
): mostly black with iridescent dark blue and dark green scales, and translucent white, reddish orange and creamy spots; dark blue sheen on the costal margin bulge and on the basal area; dark green sheen roughly on the postbasal, discal and postdiscal areas; irregular reddish orange spots in the basal area of the discal cell, Sc–R, CuA–1A, and 1A and the inner margin; sickle-shaped spot formed by a translucent white spot in the discal cell, and creamy spots between the costal margin and Sc, Sc–R
<subScript id="17F0B932FFE8FFE3AE9DB7C3FC64FD18" attach="left" box="[926,934,746,760]" fontSize="6" pageId="27" pageNumber="498">1</subScript>
, R
<subScript id="17F0B932FFE8FFE3AEC5B7C3FC0CFD18" attach="left" box="[966,974,746,760]" fontSize="6" pageId="27" pageNumber="498">1</subScript>
–R
<subScript id="17F0B932FFE8FFE3AEECB7C3FC35FD18" attach="left" box="[1007,1015,746,760]" fontSize="6" pageId="27" pageNumber="498">2</subScript>
and most of the discal cell width; and further postdiscal spots in R
<subScript id="17F0B932FFE8FFE3ACD5B627FE1CFCFC" attach="left" box="[470,478,782,796]" fontSize="6" pageId="27" pageNumber="498">2</subScript>
–R
<subScript id="17F0B932FFE8FFE3ACFCB627FDC5FCFC" attach="right" box="[511,519,782,796]" fontSize="6" pageId="27" pageNumber="498">3</subScript>
, M
<subScript id="17F0B932FFE8FFE3AF2CB627FDF5FCFC" attach="left" box="[559,567,782,796]" fontSize="6" pageId="27" pageNumber="498">1</subScript>
–M
<subScript id="17F0B932FFE8FFE3AF5DB627FDA4FCFC" attach="left" box="[606,614,782,796]" fontSize="6" pageId="27" pageNumber="498">2</subScript>
and M
<subScript id="17F0B932FFE8FFE3AFB4B627FD7DFCFC" attach="left" box="[695,703,782,796]" fontSize="6" pageId="27" pageNumber="498">2</subScript>
–M
<subScript id="17F0B932FFE8FFE3AFE5B627FD2CFCFC" attach="left" box="[742,750,782,796]" fontSize="6" pageId="27" pageNumber="498">3</subScript>
, nearer to the discal cell, and in R
<subScript id="17F0B932FFE8FFE3A975B627FBBCFCFC" attach="left" box="[1142,1150,782,796]" fontSize="6" pageId="27" pageNumber="498">3</subScript>
–R
<subScript id="17F0B932FFE8FFE3A99CB627FB65FCFC" attach="left" box="[1183,1191,782,796]" fontSize="6" pageId="27" pageNumber="498">4</subScript>
and R
<subScript id="17F0B932FFE8FFE3A9F1B627FB38FCFC" attach="left" box="[1266,1274,782,796]" fontSize="6" pageId="27" pageNumber="498">4</subScript>
–M
<subScript id="17F0B932FFE8FFE3A822B627FAEBFCFC" attach="left" box="[1313,1321,782,796]" fontSize="6" pageId="27" pageNumber="498">1</subScript>
, nearer to the apex; fringe black.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B66FFA83FC47" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B66FFEC2FCBF" box="[199,256,838,863]" italics="true" pageId="27" pageNumber="498">FWV</emphasis>
(
<figureCitation id="134FA7F2FFE8FFE3AC12B66CFEA4FC80" box="[273,358,837,864]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="27" pageNumber="498">Fig. 26</figureCitation>
): mostly brown to dark brown in a more or less checkered pattern, with a white band along the inner margin below 1A and on the postdiscal and submarginal areas of CuA
<subScript id="17F0B932FFE8FFE3AEE1B653FC28FC68" attach="left" box="[994,1002,890,904]" fontSize="6" pageId="27" pageNumber="498">1</subScript>
–1A; sickle-shaped spot and postdiscal spots as on the upper side, and further three fainter postdiscal spots in M
<subScript id="17F0B932FFE8FFE3AEC0B6B7FC09FC4C" attach="left" box="[963,971,926,940]" fontSize="6" pageId="27" pageNumber="498">2</subScript>
–M
<subScript id="17F0B932FFE8FFE3AEF1B6B7FC38FC4C" attach="left" box="[1010,1018,926,940]" fontSize="6" pageId="27" pageNumber="498">3</subScript>
, M
<subScript id="17F0B932FFE8FFE3A922B6B7FBEBFC4C" attach="left" box="[1057,1065,926,940]" fontSize="6" pageId="27" pageNumber="498">3</subScript>
–CuA
<subScript id="17F0B932FFE8FFE3A96FB6B7FBB6FC4C" attach="left" box="[1132,1140,926,940]" fontSize="6" pageId="27" pageNumber="498">1</subScript>
, and CuA
<subScript id="17F0B932FFE8FFE3A9E4B6B7FB2DFC4C" attach="left" box="[1255,1263,926,940]" fontSize="6" pageId="27" pageNumber="498">1</subScript>
–CuA
<subScript id="17F0B932FFE8FFE3A831B6B7FAF8FC4C" attach="left" box="[1330,1338,926,940]" fontSize="6" pageId="27" pageNumber="498">2</subScript>
.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B69BFD65FC0F" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B69BFE8AFC2B" box="[199,328,945,971]" italics="true" pageId="27" pageNumber="498">HW, shape:</emphasis>
roughly triangular; costal margin slightly convex; apex rounded; outer margin straight; tornus pronounced, angled; inner margin slightly convex.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B6D3FE41FBD7" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B6D3FEC5FBF3" box="[199,263,1018,1043]" italics="true" pageId="27" pageNumber="498">HWD</emphasis>
(
<figureCitation id="134FA7F2FFE8FFE3AC14B6D0FEAFFBF3" box="[279,365,1016,1043]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="27" pageNumber="498">Fig. 25</figureCitation>
): mostly black, with long, reddish orange scales in the basal area of CuA
<subScript id="17F0B932FFE8FFE3A9AEB123FB77FBF8" attach="left" box="[1197,1205,1034,1048]" fontSize="6" pageId="27" pageNumber="498">2</subScript>
–1A and along inner margin; fringe black.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B16BFBD0FB43" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B16BFEC1FBBB" box="[199,259,1090,1115]" italics="true" pageId="27" pageNumber="498">HWV</emphasis>
(
<figureCitation id="134FA7F2FFE8FFE3AC10B168FEAAFBBC" box="[275,360,1089,1116]" captionStart="FIGURES 21–38" captionStartId="5.[151,264,1570,1595]" captionTargetBox="[157,1432,185,1538]" captionTargetId="figure@5.[151,1436,181,1545]" captionTargetPageId="5" captionText="FIGURES 21–38. Species of Symmachia belonging to the “Probetoriformes” species group. 21–24. S. praxila. 21–22. Male from Joinville, Santa Catarina, Brazil (DZ 43.995 (DZUP)). 23–24. Female from Rio Natal, São Bento do Sul, Santa Catarina, Brazil (OM 67.541). 25–26. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. male holotype (DZUP). 27–30. S. divisora. 27–28. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.479 (DZUP)). 29–30. Female from Fazenda Rancho Grande, Cacaulândia, Rondônia, Brazil (MGCL). 31–34. S. falcistriga. 31–32. Male from Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil (DZ 43.509 (DZUP)). 33–34. Female, same locality as male (DZ 43.965 (DZUP)). 35–38. S. meyi stat. nov.. 35–36. Male from Chemin de la Sourçe, Cacao, French Guiana (SF). 37–38. Female from Route de l’Este, French Guiana (SF)." figureDoi="http://doi.org/10.5281/zenodo.3855262" httpUri="https://zenodo.org/record/3855262/files/figure.png" pageId="27" pageNumber="498">Fig. 26</figureCitation>
): mostly dark brown, with whitish spots and a white submarginal band; whitish spots in a more or less checkered pattern from the basal to the postdiscal area; submarginal band straight and more or less regular from the apex to the tornus; some reddish orange scales along the inner margin.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B184FB7EFA9B" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B184FEF3FB27" box="[199,305,1197,1223]" italics="true" pageId="27" pageNumber="498">Abdomen</emphasis>
(
<figureCitation id="134FA7F2FFE8FFE3AC40B184FE04FB27" box="[323,454,1197,1223]" captionStart="FIGURES 95–102" captionStartId="13.[151,264,1478,1503]" captionTargetBox="[152,1436,182,1451]" captionTargetId="figure@13.[151,1436,181,1452]" captionTargetPageId="13" captionText="FIGURES 95–102. Abdomens of males of Symmachia belonging to the “Probetoriformes” species group in dorsal (left) and lateral (right) views. 95–96. S. praxila. 97–98. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 99–100. S. divisora. 101–102. S. falcistriga. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3855282" httpUri="https://zenodo.org/record/3855282/files/figure.png" pageId="27" pageNumber="498">Figs 97–98</figureCitation>
,
<figureCitation id="134FA7F2FFE8FFE3ACD6B185FDF8FB27" box="[469,570,1196,1223]" captionStart="FIGURES 113–122" captionStartId="16.[151,264,1642,1667]" captionTargetBox="[152,1408,187,1605]" captionTargetId="figure@16.[151,1436,181,1616]" captionTargetPageId="16" captionText="FIGURES 113–122. Fourth (left) and fifith (right) tergites of males of Symmachia belonging to the “Probetoriformes” species group. 113–114. S. praxila. 115–116. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 117–118. S. divisora. 119–120. S. falcistriga. 121–122. S. meyi stat. nov.. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3855286" httpUri="https://zenodo.org/record/3855286/files/figure.png" pageId="27" pageNumber="498">115–116</figureCitation>
): terga mostly, dorsally and laterally, black, with a dark blue sheen, except segments 5–8 completely reddish orange, segments 3–4 with a narrow reddish orange stripe dorsally but almost completely laterally, and only few reddish orange scales laterally in segment 2; pleura black; sternites ventrally dark brown, with large distal white scales; genitalia surrounded by long scales, dorsally reddish orange, laterally and ventrally black; tergites 4 and 5 with an undivided band of CAS on the posterior margin, as long as 1/9 of the tergites’ length and as wide as half the tergites’ width, not projected anteriorly; CAS scales pale yellow.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B0ACFEDAF95F" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B0ACFEF3FA7F" box="[199,305,1413,1439]" italics="true" pageId="27" pageNumber="498">Genitalia</emphasis>
(
<figureCitation id="134FA7F2FFE8FFE3AC42B0ACFE61FA7F" box="[321,419,1413,1439]" captionStart="FIGURES 128–132" captionStartId="19.[151,264,1154,1179]" captionTargetBox="[155,1432,186,1123]" captionTargetId="figure@19.[151,1436,181,1129]" captionTargetPageId="19" captionText="FIGURES 128–132. Male genitalia of Symmachia belonging to the “Probetoriformes” species group in lateral view, uncus in dorsal view and anterior projection of saccus in ventral view. 128. S. praxila. 129. S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov.. 130. S. divisora. 131. S. falcistriga. 132. S. meyi stat. nov.. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3855292" httpUri="https://zenodo.org/record/3855292/files/figure.png" pageId="27" pageNumber="498">Fig. 129</figureCitation>
): tegumen rectangular; membranous area between the uncus and the tegumen more or less triangular; uncus rectangular, with a medial anterior projection; gnathos C-shaped, angled, ventral part about one third larger than the dorsal, connected to the tegumen by a small dorsal projection; ventral arm of the tegumen thin and obliquely projected, connected ventrally to the wider dorsal arm of the saccus; anterior projection of the saccus wider than long and bilobed anteriorly; valva small, roughly triangular in lateral view, connected ventrally to the saccus and the pedicel, and dorsally to the opposite valva; pedicel posteriorly sclerotized and keeled, connected to the middle of the aedeagus; aedeagus roughly cylindrical, anterior opening wide and on the left side; vesica with an basal patch of very small pointed cornuti of similar size and a apical band of fewer and larger cornuti; subscaphium sclerotized.
</paragraph>
<paragraph id="8BCBBB77FFE8FFE3ADC4B3E0FE5CF903" blockId="27.[151,1437,152,2015]" box="[199,414,1737,1763]" pageId="27" pageNumber="498">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36EE8FCFFE8FFE3ADC4B3C4FABCF893" pageId="27" pageNumber="498" type="etymology">
<paragraph id="8BCBBB77FFE8FFE3ADC4B3C4FABCF893" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B3C4FE88F8E7" bold="true" box="[199,330,1773,1799]" pageId="27" pageNumber="498">Etymology.</emphasis>
This species is dedicated to the Instituto Uiraçu, a non-profit association established in 2001 by the eminent Brazilian lepidopterist Dr. Victor O. Becker to protect the Serra Bonita mountain range in southern
<collectingRegion id="49B07595FFE8FFE3A855B238FA5AF8CB" box="[1366,1432,1809,1835]" country="Brazil" name="Bahia" pageId="27" pageNumber="498">Bahia</collectingRegion>
,
<collectingCountry id="F363FBE7FFE8FFE3AD94B21CFF1EF8AF" box="[151,220,1845,1871]" name="Brazil" pageId="27" pageNumber="498">Brazil</collectingCountry>
, and
<typeStatus id="54CF05D5FFE8FFE3AC17B21CFE81F8AF" box="[276,323,1845,1871]" pageId="27" pageNumber="498">type</typeStatus>
locality of this species. The specific epithet means “Harpy Eagle” in the Brazilian indigenous Tupi language, and is composed of two elements: “uirá” (bird) and “açu” (big). The name is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C36EE8FCFFE8FFE3ADC4B255FE46F83F" pageId="27" pageNumber="498" type="distribution">
<paragraph id="8BCBBB77FFE8FFE3ADC4B255FE46F83F" blockId="27.[151,1437,152,2015]" pageId="27" pageNumber="498">
<emphasis id="B9006765FFE8FFE3ADC4B255FEA3F877" bold="true" box="[199,353,1916,1943]" pageId="27" pageNumber="498">Distribution.</emphasis>
This species is apparently restricted to coastal Atlantic Forest of
<collectingCountry id="F363FBE7FFE8FFE3A930B254FBB5F877" box="[1075,1143,1917,1943]" name="Brazil" pageId="27" pageNumber="498">Brazil</collectingCountry>
(
<collectingRegion id="49B07595FFE8FFE3A986B254FB0AF877" box="[1157,1224,1917,1943]" country="Brazil" name="Bahia" pageId="27" pageNumber="498">Bahia</collectingRegion>
and
<collectingRegion id="49B07595FFE8FFE3A9FEB254FA5EF877" box="[1277,1436,1917,1943]" country="Brazil" name="Espirito Santo" pageId="27" pageNumber="498">Espírito Santo</collectingRegion>
(
<bibRefCitation id="EFE5C686FFE8FFE3AD9CB288FE8EF85B" author="D'Abrera, B. L." box="[159,332,1953,1979]" pageId="27" pageNumber="498" refId="ref32991" refString="D'Abrera, B. L. (1994) Butterflies of the Neotropical Region. Part VI. Riodinidae. Black Rock, Hill House, Victoria, 217 pp." type="book" year="1994">D’Abrera 1994</bibRefCitation>
)) (
<figureCitation id="134FA7F2FFE8FFE3AC6DB288FE13F85B" box="[366,465,1952,1979]" captionStart="FIGURE 153" captionStartId="28.[151,250,1806,1831]" captionTargetBox="[151,1436,740,1780]" captionTargetId="figure@28.[151,1436,740,1780]" captionTargetPageId="28" captionText="FIGURE 153. Geographical distribution of Symmachia praxila, S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov., S. divisora, S. falcistriga and S. meyi stat. nov. Asterisks indicate records from the literature." figureDoi="http://doi.org/10.5281/zenodo.3855307" httpUri="https://zenodo.org/record/3855307/files/figure.png" pageId="27" pageNumber="498">Fig. 153</figureCitation>
). It is not known if this species occurs further south along the Brazilian coast, in sympatry with
<taxonomicName id="4C74C0F4FFE8FFE3AC13B2ECFEBFF83F" authorityName="Westwood" authorityYear="1851" box="[272,381,1989,2015]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="498" phylum="Arthropoda" rank="species" species="praxila">
<emphasis id="B9006765FFE8FFE3AC13B2ECFEBFF83F" box="[272,381,1989,2015]" italics="true" pageId="27" pageNumber="498">S. praxila</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36EE8FCFFEFFFE4ADC4B5B1FC93FDFB" pageId="28" pageNumber="499" type="discussion">
<paragraph id="8BCBBB77FFEFFFE4ADC4B5B1FC35FE6A" blockId="28.[151,1437,152,719]" pageId="28" pageNumber="499">
<emphasis id="B9006765FFEFFFE4ADC4B5B1FE8DFF53" bold="true" box="[199,335,152,179]" pageId="28" pageNumber="499">Comments.</emphasis>
<taxonomicName id="4C74C0F4FFEFFFE4AC55B5B1FDF3FF52" authorityName="Dolibaina, Dias &amp; Casagrande" authorityYear="2020" box="[342,561,152,178]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="499" phylum="Arthropoda" rank="species" species="uirassu">
<emphasis id="B9006765FFEFFFE4AC55B5B1FDF3FF52" box="[342,561,152,178]" italics="true" pageId="28" pageNumber="499">Symmachia uirassu</emphasis>
</taxonomicName>
is described based on three males specimens collected in RPPN Serra Bonita, Camacan,
<collectingRegion id="49B07595FFEFFFE4AC0DB595FE90FF36" box="[270,338,188,214]" country="Brazil" name="Bahia" pageId="28" pageNumber="499">Bahia</collectingRegion>
,
<collectingCountry id="F363FBE7FFEFFFE4AC5DB595FE64FF36" box="[350,422,188,214]" name="Brazil" pageId="28" pageNumber="499">Brazil</collectingCountry>
. The specimen identified as “S
<emphasis id="B9006765FFEFFFE4AE00B595FCB4FF36" box="[771,886,188,214]" italics="true" pageId="28" pageNumber="499">ymmachia</emphasis>
sp.?” illustrated by
<bibRefCitation id="EFE5C686FFEFFFE4A959B595FA94FF37" author="D'Abrera, B. L." box="[1114,1366,188,215]" pageId="28" pageNumber="499" refId="ref32991" refString="D'Abrera, B. L. (1994) Butterflies of the Neotropical Region. Part VI. Riodinidae. Black Rock, Hill House, Victoria, 217 pp." type="book" year="1994">D’Abrera (1994: 1044</bibRefCitation>
[1,2]) from
<collectingRegion id="49B07595FFEFFFE4ADD4B5C9FEBEFF1B" box="[215,380,224,251]" country="Brazil" name="Espirito Santo" pageId="28" pageNumber="499">Espírito Santo</collectingRegion>
,
<collectingCountry id="F363FBE7FFEFFFE4AC8AB5C9FE0FFF1A" box="[393,461,224,250]" name="Brazil" pageId="28" pageNumber="499">Brazil</collectingCountry>
corresponds to
<taxonomicName id="4C74C0F4FFEFFFE4AF85B5C8FD34FF1A" authorityName="Dolibaina, Dias &amp; Casagrande" authorityYear="2020" box="[646,758,225,251]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="499" phylum="Arthropoda" rank="species" species="uirassu">
<emphasis id="B9006765FFEFFFE4AF85B5C8FD34FF1A" box="[646,758,225,251]" italics="true" pageId="28" pageNumber="499">S. uirassu</emphasis>
</taxonomicName>
, and is recognized as an “undescribed taxon (…) probably probably related to
<taxonomicName id="4C74C0F4FFEFFFE4AC71B42CFDC1FEFE" authorityName="Westwood" authorityYear="1851" box="[370,515,261,287]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="499" phylum="Arthropoda" rank="species" species="accusatrix">
<emphasis id="B9006765FFEFFFE4AC71B42CFDC1FEFE" box="[370,515,261,287]" italics="true" pageId="28" pageNumber="499">S. accusatrix</emphasis>
</taxonomicName>
”, and is identified as a male of
<taxonomicName id="4C74C0F4FFEFFFE4AE6BB42CFC17FEFE" authorityName="Westwood" authorityYear="1851" box="[872,981,260,287]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="499" phylum="Arthropoda" rank="species" species="praxila">
<emphasis id="B9006765FFEFFFE4AE6BB42CFC17FEFE" box="[872,981,260,287]" italics="true" pageId="28" pageNumber="499">S. praxila</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE5C686FFEFFFE4AEFCB42DFB39FEFF" author="Hall, J. P. W. &amp; Harvey, D. J." box="[1023,1275,260,287]" pageId="28" pageNumber="499" pagination="7 - 10" refId="ref33740" refString="Hall, J. P. W. &amp; Harvey, D. J. (2007) Three new species of Symmachiini from Panama and Colombia (Lepidoptera: Riodinidae). Tropical Lepidoptera, 16:, 7 - 10." type="journal article" year="2007">Hall &amp; Harvey (2007)</bibRefCitation>
. However, the wings size and shape, the dark ground color, and the absence of the FWD reddish orange submarginal band and postdiscal costal spots clearly indentify this specimen as
<taxonomicName id="4C74C0F4FFEFFFE4AE09B464FCB4FE86" authorityName="Dolibaina, Dias &amp; Casagrande" authorityYear="2020" box="[778,886,333,359]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="499" phylum="Arthropoda" rank="species" species="uirassu">
<emphasis id="B9006765FFEFFFE4AE09B464FCB4FE86" box="[778,886,333,359]" italics="true" pageId="28" pageNumber="499">S. uirassu</emphasis>
</taxonomicName>
. Females still are unknown although it is possible they are misidentified with other species of the group in museum collections.
</paragraph>
<paragraph id="8BCBBB77FFEFFFE4ADC4B4BDFC93FDFB" blockId="28.[151,1437,152,719]" pageId="28" pageNumber="499">
The
<typeStatus id="54CF05D5FFEFFFE4ADFAB4BDFE99FE4E" box="[249,347,404,430]" pageId="28" pageNumber="499" type="holotype">holotype</typeStatus>
and
<typeStatus id="54CF05D5FFEFFFE4AC93B4BCFE39FE4F" box="[400,507,405,431]" pageId="28" pageNumber="499" type="paratype">paratypes</typeStatus>
were all collected in a hilltop at
<quantity id="4C8C1692FFEFFFE4AE66B4BCFC69FE4F" box="[869,939,405,431]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="28" pageNumber="499" unit="m" value="920.0">920 m</quantity>
covered by a dense understory with bamboo and liana where they were flying and resting on leaves of trees up to five meters high (
<figureCitation id="134FA7F2FFEFFFE4A957B491FB32FE33" box="[1108,1264,440,467]" captionStart="FIGURES 154–156" captionStartId="29.[151,264,1034,1059]" captionTargetBox="[151,1436,181,1008]" captionTargetId="figure@29.[151,1436,181,1008]" captionTargetPageId="29" captionText="FIGURES 154–156. Habitat of some species of Symmachia belonging to the “Probetoriformes” species group. 154. Hilltop at the Bromélias trail, Reserva Serra Bonita, Camacan, Bahia, Brazil, the arrow indicates the exact location where males of S. atlantica Dias, Dolibaina &amp; Casagrande sp. nov. were collected. 155. Opposite view of the previous figure, arrow indicates the exact location where males of S. uirassu Dolibaina, Dias &amp; Casagrande sp. nov. were collected. 156. DRD on a tree at the lightgap of the hilltop in Parque Nacional da Serra do Divisor, Mâncio Lima, Acre, Brazil where males of S. probetor, S. estellina, S. divisora and S. falcistriga were collected (photo by Eduardo Carneiro)." figureDoi="http://doi.org/10.5281/zenodo.3855309" httpUri="https://zenodo.org/record/3855309/files/figure.png" pageId="28" pageNumber="499">Figs 154–155</figureCitation>
) at about 11:00 h. Two males were collected in the second and one in the fourth day of collecting at this site occurring simultaneously with other species of
<taxonomicName id="4C74C0F4FFEFFFE4ACC9B729FD89FDFA" authorityName="Hubner" authorityYear="1819" box="[458,587,512,538]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="B9006765FFEFFFE4ACC9B729FD89FDFA" box="[458,587,512,538]" italics="true" pageId="28" pageNumber="499">Symmachia</emphasis>
</taxonomicName>
, including
<taxonomicName id="4C74C0F4FFEFFFE4AFC9B728FC88FDFA" authorityName="Dias, Dolibaina &amp; Casagrande" authorityYear="2020" box="[714,842,512,539]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="499" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="B9006765FFEFFFE4AFC9B728FC88FDFA" box="[714,842,512,539]" italics="true" pageId="28" pageNumber="499">S. atlantica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36EE8FCFFEFFFE4ADC4B70CFB9DFD2F" pageId="28" pageNumber="499" type="materials_examined">
<paragraph id="8BCBBB77FFEFFFE4ADC4B70CFD0CFD4B" blockId="28.[151,1437,152,719]" pageId="28" pageNumber="499">
<emphasis id="B9006765FFEFFFE4ADC4B70CFEB4FDDF" bold="true" box="[199,374,548,575]" pageId="28" pageNumber="499">Type material.</emphasis>
<materialsCitation id="3B1CB12AFFEFFFE4AC7CB70DFA5EFDDF" ID-GBIF-Occurrence="2625548308" box="[383,1436,548,575]" collectionCode="HOLOTY-" location="Male" pageId="28" pageNumber="499" specimenCount="2" typeStatus="holotype">
<location id="8EABEDACFFEFFFE4AC7CB70DFE7BFDDE" LSID="urn:lsid:plazi:treatment:03DD0A61FFE9FFE4AD94B044FB9DFD2F:8EABEDACFFEFFFE4AC7CB70DFE7BFDDE" box="[383,441,548,574]" name="Male" pageId="28" pageNumber="499">Male</location>
<typeStatus id="54CF05D5FFEFFFE4ACC2B70DFDE1FDDE" box="[449,547,548,574]" pageId="28" pageNumber="499" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C74C0F4FFEFFFE4AF48B70DFCE5FDDE" authorityName="Dolibaina, Dias &amp; Casagrande" authorityYear="2020" box="[587,807,548,574]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="499" phylum="Arthropoda" rank="species" species="uirassu">
<emphasis id="B9006765FFEFFFE4AF48B70DFCE5FDDE" box="[587,807,548,574]" italics="true" pageId="28" pageNumber="499">Symmachia uirassu</emphasis>
</taxonomicName>
with the following labels: /
<typeStatus id="54CF05D5FFEFFFE4A967B70CFAC8FDDF" box="[1124,1290,549,575]" pageId="28" pageNumber="499" type="holotype">HOLOTYPUS</typeStatus>
/
<collectionCode id="ED6523B2FFEFFFE4A820B70CFA5EFDDF" box="[1315,1436,549,575]" pageId="28" pageNumber="499">HOLOTY-</collectionCode>
</materialsCitation>
<materialsCitation id="3B1CB12AFFEFFFE4AD94B760FE12FD67" ID-GBIF-Occurrence="2625547484" collectionCode="TRILHA" country="Brazil" location="BAHIA" pageId="28" pageNumber="499" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
PUS
<taxonomicName id="4C74C0F4FFEFFFE4ADD6B761FD49FD82" authority="Dolibaina &amp; Dias" authorityName="Dolibaina &amp; Dias" box="[213,651,584,611]" class="Insecta" family="Lycaenidae" genus="Symmachia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="499" phylum="Arthropoda" rank="species" species="uirassu">
<emphasis id="B9006765FFEFFFE4ADD6B761FE71FD82" box="[213,435,584,610]" italics="true" pageId="28" pageNumber="499">Symmachia uirassu</emphasis>
Dolibaina &amp; Dias
</taxonomicName>
det. 2019/ DZ 43.449 /
<collectingCountry id="F363FBE7FFEFFFE4AEA6B760FBC9FD83" box="[933,1035,585,611]" name="Brazil" pageId="28" pageNumber="499">BRASIL</collectingCountry>
,
<collectingRegion id="49B07595FFEFFFE4A91AB760FBB7FD83" box="[1049,1141,585,611]" country="Brazil" name="Bahia" pageId="28" pageNumber="499">BAHIA</collectingRegion>
,
<collectingRegion id="49B07595FFEFFFE4A980B760FAD7FD83" box="[1155,1301,585,611]" country="Brazil" name="CAMACAN" pageId="28" pageNumber="499">CAMACAN</collectingRegion>
,
<collectingRegion id="49B07595FFEFFFE4A827B760FEA0FD67" country="Brazil" name="RESERVA SERRA BONITA" pageId="28" pageNumber="499">RESERVA SERRA BONITA</collectingRegion>
,
<collectionCode id="ED6523B2FFEFFFE4AC68B744FE12FD67" box="[363,464,621,647]" pageId="28" pageNumber="499">TRILHA</collectionCode>
</materialsCitation>
<materialsCitation id="3B1CB12AFFEFFFE4ACD4B744FB46FD67" ID-GBIF-Occurrence="2625548319" box="[471,1156,620,647]" collectingDate="2016-12-03" collectingDateMax="2016-12-10" collectingDateMin="2016-12-03" collectionCode="BROMELIAS" country="Brazil" elevation="920" latitude="-15.383333" location="S" longLatPrecision="1285" longitude="-39.566666" pageId="28" pageNumber="499" specimenCount="1" stateProvince="S" typeStatus="holotype">
DAS
<collectionCode id="ED6523B2FFEFFFE4AF15B745FD7AFD67" box="[534,696,620,647]" pageId="28" pageNumber="499">BROMÉLIAS</collectionCode>
<geoCoordinate id="EE40DDB0FFEFFFE4AFBCB745FCD8FD67" box="[703,794,620,647]" degrees="15" direction="south" minutes="23" orientation="latitude" pageId="28" pageNumber="499" precision="925" value="-15.383333">
15º23’
<collectingRegion id="49B07595FFEFFFE4AE09B745FCD8FD67" box="[778,794,620,647]" country="Brazil" name="S" pageId="28" pageNumber="499">S</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="EE40DDB0FFEFFFE4AE25B744FC48FD67" box="[806,906,621,647]" degrees="39" direction="west" minutes="34" orientation="longitude" pageId="28" pageNumber="499" precision="925" value="-39.566666">
39º34’
<collectingRegion id="49B07595FFEFFFE4AE6EB744FC48FD67" box="[877,906,621,647]" country="Brazil" name="W" pageId="28" pageNumber="499">W</collectingRegion>
</geoCoordinate>
,
<date id="FFCA9DB7FFEFFFE4AE96B744FBF8FD66" box="[917,1082,620,647]" pageId="28" pageNumber="499" value="2016-12-03" valueMax="2016-12-10" valueMin="2016-12-03">
<collectingDate id="EF8E645FFFEFFFE4AE96B744FBF8FD66" box="[917,1082,620,647]" pageId="28" pageNumber="499" value="2016-12-03" valueMax="2016-12-10" valueMin="2016-12-03">3–10.XII.2016</collectingDate>
</date>
<quantity id="4C8C1692FFEFFFE4A942B744FB46FD67" box="[1089,1156,621,647]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="28" pageNumber="499" unit="m" value="920.0">
<elevation id="00595C44FFEFFFE4A942B744FB46FD67" box="[1089,1156,621,647]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="28" pageNumber="499" unit="m" value="920.0">920m</elevation>
</quantity>
</materialsCitation>
,
<materialsCitation id="3B1CB12AFFEFFFE4A98CB744FDD2FD4B" ID-GBIF-Occurrence="2625548360" collectionCode="DOLIBAINA, SANTOS" country="Brazil" location="MIELKE" pageId="28" pageNumber="499" specimenCount="1" stateProvince="MIELKE" typeStatus="holotype">
<collectingRegion id="49B07595FFEFFFE4A98CB744FB3EFD67" box="[1167,1276,621,647]" country="Brazil" name="MIELKE" pageId="28" pageNumber="499">MIELKE</collectingRegion>
,
<collectingRegion id="49B07595FFEFFFE4A805B744FA5AFD67" box="[1286,1432,621,647]" country="Brazil" name="CARNEIRO" pageId="28" pageNumber="499">CARNEIRO</collectingRegion>
,
<collectingRegion id="49B07595FFEFFFE4AD94B7B8FF19FD4B" box="[151,219,657,683]" country="Brazil" name="DIAS" pageId="28" pageNumber="499">DIAS</collectingRegion>
,
<collectionCode id="ED6523B2FFEFFFE4ADE4B7B8FE43FD4B" box="[231,385,657,683]" pageId="28" pageNumber="499">DOLIBAINA</collectionCode>
&amp;
<collectionCode id="ED6523B2FFEFFFE4ACA7B7B8FDD2FD4B" box="[420,528,657,683]" pageId="28" pageNumber="499">SANTOS</collectionCode>
</materialsCitation>
LEG /. (
<collectionCode id="ED6523B2FFEFFFE4AF70B7B8FD01FD4B" box="[627,707,657,683]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="28" pageNumber="499" type="University or college">DZUP</collectionCode>
).
</paragraph>
<paragraph id="8BCBBB77FFEFFFE4ADC4B79CFB9DFD2F" blockId="28.[151,1437,152,719]" box="[199,1119,692,719]" pageId="28" pageNumber="499">
<materialsCitation id="3B1CB12AFFEFFFE4ADC4B79CFB9DFD2F" ID-GBIF-Occurrence="2625547407" box="[199,1119,692,719]" collectionCode="DZUP" location="Male" pageId="28" pageNumber="499" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B9006765FFEFFFE4ADC4B79CFE84FD2F" bold="true" box="[199,326,693,719]" pageId="28" pageNumber="499">
<typeStatus id="54CF05D5FFEFFFE4ADC4B79CFE83FD2F" box="[199,321,693,719]" pageId="28" pageNumber="499" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D7270FEFFEFFFE4AC4EB79CFE64FD2E" box="[333,422,692,719]" pageId="28" pageNumber="499" type="male">2 males</specimenCount>
, same data as the holotype, DZ 36.622*, DZ 43.459* (
<collectionCode id="ED6523B2FFEFFFE4A909B79CFB95FD2F" box="[1034,1111,693,719]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="28" pageNumber="499" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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