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<document id="3438AABB8927B09C84299C339158CBAC" ID-DOI="10.5281/zenodo.189654" ID-GBIF-Dataset="320aec2f-ea3b-427e-a2a7-bdd4969cb030" ID-ISSN="1175-5326" ID-Zenodo-Dep="189654" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461103618203" checkinUser="plazi" docAuthor="Chiba, Hideyuki, Hsu, Yu-Feng, Tsukiyama, Hiroshi, Lo, Philip Yik-Fui, Chen, Chien-Ren &amp; Wang, Shou-Ming" docDate="2009" docId="03CC500DF30D3421FF2CF4F63CAFBCD4" docLanguage="en" docName="zt02202p058.pdf" docOrigin="Zootaxa 2202" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tagiades menaka subsp. hainana Sonan 1938" docType="treatment" docVersion="7" lastPageNumber="57" masterDocId="FFF52875F3043428FFBBF5023E1BB836" masterDocTitle="Jinhaku Sonans skipper type collection deposited at Taiwan Agricultural Research Institute (Lepidoptera: Hesperiidae)" masterLastPageNumber="58" masterPageNumber="48" pageNumber="57" updateTime="1698592197444" updateUser="plazi">
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<mods:title id="D0A75C58866CC1F62842D4EF13E36404">Jinhaku Sonans skipper type collection deposited at Taiwan Agricultural Research Institute (Lepidoptera: Hesperiidae)</mods:title>
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<mods:namePart id="AB0371CE7FA2C5897DA88FB70ABDAB0E">Chiba, Hideyuki</mods:namePart>
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<mods:namePart id="7A39322D8A20F371B59D26A20E0A2CE4">Hsu, Yu-Feng</mods:namePart>
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<mods:namePart id="8ED0935B6285616BEB426B62DF895763">Tsukiyama, Hiroshi</mods:namePart>
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<mods:namePart id="4C56279951046068BA6BF325A1F10D86">Lo, Philip Yik-Fui</mods:namePart>
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<mods:namePart id="4C87D267399D813E2B6FF79372C60532">Chen, Chien-Ren</mods:namePart>
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<mods:namePart id="5F7C96655157DA594BEBE0D5F934760E">Wang, Shou-Ming</mods:namePart>
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<mods:date id="9D8DA4C931998E49D718B2695633144D">2009</mods:date>
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<treatment id="03CC500DF30D3421FF2CF4F63CAFBCD4" ID-DOI="http://doi.org/10.5281/zenodo.6225228" ID-GBIF-Taxon="119622485" ID-Zenodo-Dep="6225228" LSID="urn:lsid:plazi:treatment:03CC500DF30D3421FF2CF4F63CAFBCD4" httpUri="http://treatment.plazi.org/id/03CC500DF30D3421FF2CF4F63CAFBCD4" lastPageNumber="57" pageId="9" pageNumber="57">
<subSubSection id="C37FB290F30D3421FF2CF4F63CB5BA02" pageId="9" pageNumber="57" type="nomenclature">
<paragraph id="8BDAE11BF30D3421FF2CF4F63C68BA3B" blockId="9.[151,686,499,564]" box="[151,627,499,526]" pageId="9" pageNumber="57">
<heading id="D0925677F30D3421FF2CF4F63C68BA3B" bold="true" box="[151,627,499,526]" fontSize="11" level="1" pageId="9" pageNumber="57" reason="1">
<taxonomicName id="4C659A98F30D3421FF2CF4F63C68BA3B" ID-CoL="7BHRN" authority="Sonan, 1938" authorityName="Sonan" authorityYear="1938" box="[151,627,499,526]" class="Insecta" family="Hesperiidae" genus="Tagiades" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="57" phylum="Arthropoda" rank="subSpecies" species="menaka" subSpecies="hainana">
<emphasis id="B9113D09F30D3421FF2CF4F63C68BA3B" bold="true" box="[151,627,499,526]" pageId="9" pageNumber="57">
<emphasis id="B9113D09F30D3421FF2CF4F63FCEBA38" bold="true" box="[151,469,500,526]" italics="true" pageId="9" pageNumber="57">Tagiades menaka hainana</emphasis>
Sonan, 1938
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDAE11BF30D3421FF2CF7183CB5BA02" blockId="9.[151,686,499,564]" box="[151,686,538,564]" pageId="9" pageNumber="57">
Trans. nat. Hist. Soc.
<collectingCountry id="F372A18BF30D3421FE28F7183FE1BA02" box="[403,506,538,564]" name="Taiwan" pageId="9" pageNumber="57">Formosa</collectingCountry>
28: 370, fig. 1.
</paragraph>
</subSubSection>
<subSubSection id="C37FB290F30D3421FF2CF76A3CAFBCD4" pageId="9" pageNumber="57" type="description">
<paragraph id="8BDAE11BF30D3421FF2CF76A3CC4BAA9" blockId="9.[151,1436,616,671]" pageId="9" pageNumber="57">
Labels: “
<taxonomicName id="4C659A98F30D3421FF40F76A3F41BA49" box="[251,346,616,639]" class="Insecta" family="Hesperiidae" genus="Tagiades" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="57" phylum="Arthropoda" rank="genus">Tagiades</taxonomicName>
/
<taxonomicName id="4C659A98F30D3421FEDAF76A3FAEBA49" box="[353,437,616,639]" class="Insecta" family="Hesperiidae" genus="Tagiades" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="57" phylum="Arthropoda" rank="species" species="menaka">menaka</taxonomicName>
/hainana Sonan DET. J. SONAN,” “Ling-sui/Hainan/ 1930/COL. J. SONAN,” “menata,” “hainana/Sonan,” “Holo/
<typeStatus id="54DE5FB9F30D3421FE74F78A3C1DBAA9" box="[463,518,648,671]" pageId="9" pageNumber="57">Type</typeStatus>
[in red],” “HES(3)”
</paragraph>
<paragraph id="8BDAE11BF30D3421FF2CF7CC3CAFBCD4" blockId="9.[151,1437,718,1250]" pageId="9" pageNumber="57">
<bibRefCitation id="EFF49CEAF30D3421FF2CF7CC3F55BADE" author="Sonan" box="[151,334,718,744]" pageId="9" pageNumber="58" refString="Sonan, J. (1938 a) A list of the butterflies of Hainan, with descriptions of two new sub-species of Hesperiidae. Transactions of the Natural History Society of Formosa, 28, 348 - 372, 2 f. [In Japanese]" type="journal article" year="1938" yearSuffix="a">Sonan (1938a)</bibRefCitation>
specified just one male specimen when he described
<taxonomicName id="4C659A98F30D3421FC4EF7CD3B27BADE" box="[1013,1340,719,744]" class="Insecta" family="Hesperiidae" genus="Tagiades" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="57" phylum="Arthropoda" rank="subSpecies" species="menaka" subSpecies="hainana">
<emphasis id="B9113D09F30D3421FC4EF7CD3B27BADE" box="[1013,1340,719,744]" italics="true" pageId="9" pageNumber="57">Tagiades menaka hainana</emphasis>
</taxonomicName>
, thus it represents the
<typeStatus id="54DE5FB9F30D3421FE84F7F73FBEBB39" box="[319,421,757,783]" pageId="9" pageNumber="57" type="holotype">holotype</typeStatus>
of the taxon. The unique
<typeStatus id="54DE5FB9F30D3421FD6AF7F73D18BB39" box="[721,771,757,783]" pageId="9" pageNumber="57">type</typeStatus>
(
<typeStatus id="54DE5FB9F30D3421FCA9F7F73D67BB39" box="[786,892,757,783]" pageId="9" pageNumber="57" type="holotype">holotype</typeStatus>
) of the taxon was found in TARI (
<figureCitation id="135EFD9EF30D3421FAB0F7F73B95BB39" box="[1291,1422,757,783]" captionStart="FIGURES 19 39" captionStartId="4.[151,270,1872,1896]" captionTargetBox="[192,1389,202,1833]" captionTargetId="figure@4.[189,1401,197,1835]" captionTargetPageId="4" captionText="FIGURES 19 39. Types of skipper taxa described by Jinhaku Sonan: 19 21, paratype of Augiades bouddha niitakana, upperside, underside, labels; 22 24, holotype of Parnara kotoshona, upperside, underside, labels; 25 27, holotype of Parnara ranrunna, upperside, underside, labels; 28 30, holotype of Pamara kiraizana, upperside, underside, labels; 31 33, holotype of Gangara thyrsis hainana, upperside, underside, labels; 34 36, holotype of Telicota palmarum hainanum, upperside, upperside, labels; 37 39, holotype of Tagiades menaka hainana, upperside, underside, labels." httpUri="https://zenodo.org/record/189656/files/figure.png" pageId="9" pageNumber="57">Fig. 3739</figureCitation>
), and the specimen conforms to the original description and associated figure in
<bibRefCitation id="EFF49CEAF30D3421FBF4F61E3AE4BB00" author="Sonan" box="[1103,1279,796,822]" pageId="9" pageNumber="58" refString="Sonan, J. (1938 a) A list of the butterflies of Hainan, with descriptions of two new sub-species of Hesperiidae. Transactions of the Natural History Society of Formosa, 28, 348 - 372, 2 f. [In Japanese]" type="journal article" year="1938" yearSuffix="a">Sonan (1938a)</bibRefCitation>
although the collecting year was “1908” in the original description but “1930” on the label.
<bibRefCitation id="EFF49CEAF30D3421FBF8F6403AF8BB6A" author="Evans" box="[1091,1251,834,860]" pageId="9" pageNumber="58" refString="Evans, W. H. (1949) A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum (Natural History). The British Museum (Natural History), London, xix + 502 pp." type="book" year="1949">Evans (1949)</bibRefCitation>
intended to list
<emphasis id="B9113D09F30D3421FF2CF66B3EECBBB4" box="[151,247,873,898]" italics="true" pageId="9" pageNumber="57">hainana</emphasis>
as a synonym of
<taxonomicName id="4C659A98F30D3421FE7DF66B3DB2BBB5" authority="Moore, 1865" authorityName="Moore" authorityYear="1865" box="[454,937,873,899]" class="Insecta" family="Hesperiidae" genus="Tagiades" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="57" phylum="Arthropoda" rank="subSpecies" species="menaka" subSpecies="menaka">
<emphasis id="B9113D09F30D3421FE7DF66B3CEFBBB4" box="[454,756,873,898]" italics="true" pageId="9" pageNumber="57">Tagiades menaka menaka</emphasis>
(Moore, 1865)
</taxonomicName>
, but the name
<emphasis id="B9113D09F30D3421FBEFF66B3AAFBBB4" box="[1108,1204,873,898]" italics="true" pageId="9" pageNumber="57">hainana</emphasis>
was in the index of his catalogue and not in the text. The characteristics of the
<typeStatus id="54DE5FB9F30D3421FCE1F6923DDEBB9C" box="[858,965,912,938]" pageId="9" pageNumber="57" type="holotype">holotype</typeStatus>
, however, do not agree with those of
<taxonomicName id="4C659A98F30D3421FA3DF6923EEFBBE6" class="Insecta" family="Hesperiidae" genus="Tagiades" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="57" phylum="Arthropoda" rank="species" species="menaka">
<emphasis id="B9113D09F30D3421FA3DF6923EEFBBE6" italics="true" pageId="9" pageNumber="57">T. menaka</emphasis>
</taxonomicName>
. Instead, the
<typeStatus id="54DE5FB9F30D3421FE23F6B43C1ABBE6" box="[408,513,950,976]" pageId="9" pageNumber="57" type="holotype">holotype</typeStatus>
possesses diagnostic characters of
<taxonomicName id="4C659A98F30D3421FC17F6B53AD9BBE6" authority="Stoll, 1782" authorityName="Stoll" authorityYear="1782" box="[940,1218,950,976]" class="Insecta" family="Hesperiidae" genus="Tagiades" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="57" phylum="Arthropoda" rank="species" species="japetus">
<emphasis id="B9113D09F30D3421FC17F6B53A3ABBE6" box="[940,1057,951,976]" italics="true" pageId="9" pageNumber="57">T. japetus</emphasis>
(Stoll, 1782)
</taxonomicName>
defined by
<bibRefCitation id="EFF49CEAF30D3421FAE8F6B43EF3BBC1" author="Evans" pageId="9" pageNumber="58" refString="Evans, W. H. (1949) A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum (Natural History). The British Museum (Natural History), London, xix + 502 pp." type="book" year="1949">Evans (1949)</bibRefCitation>
and
<bibRefCitation id="EFF49CEAF30D3421FE9EF6DF3C1FBBC1" author="Tsukiyama" box="[293,516,989,1015]" pageId="9" pageNumber="58" refString="Tsukiyama, H. (1980) Miscellaneous notes on skippers of the Southeast Asia (2). Gekkan-Mushi, 118, 11 - 16. [In Japanese]" type="journal article" year="1980">Tsukiyama (1980)</bibRefCitation>
as it has no hyaline spot in front of spots at the end of discoidal cell and a prominent hyaline spot in cell M3 and Cu1 of the forewing (
<figureCitation id="135EFD9EF30D3421FCE3F1063DC5BC28" box="[856,990,1028,1054]" captionStart="FIGURES 19 39" captionStartId="4.[151,270,1872,1896]" captionTargetBox="[192,1389,202,1833]" captionTargetId="figure@4.[189,1401,197,1835]" captionTargetPageId="4" captionText="FIGURES 19 39. Types of skipper taxa described by Jinhaku Sonan: 19 21, paratype of Augiades bouddha niitakana, upperside, underside, labels; 22 24, holotype of Parnara kotoshona, upperside, underside, labels; 25 27, holotype of Parnara ranrunna, upperside, underside, labels; 28 30, holotype of Pamara kiraizana, upperside, underside, labels; 31 33, holotype of Gangara thyrsis hainana, upperside, underside, labels; 34 36, holotype of Telicota palmarum hainanum, upperside, upperside, labels; 37 39, holotype of Tagiades menaka hainana, upperside, underside, labels." httpUri="https://zenodo.org/record/189656/files/figure.png" pageId="9" pageNumber="57">Fig. 3738</figureCitation>
). The wing markings of the specimen are generally similar to those of the wet season form of
<emphasis id="B9113D09F30D3421FC9AF12C3D9ABC71" box="[801,897,1070,1095]" italics="true" pageId="9" pageNumber="57">T. j. ravi</emphasis>
Moore, 1865, but differ from it by possessing a white hindwing fringe instead of brown. Although it is difficult to accept the fact that this rather common skipper is represented by a single specimen, there is no specific evidence that it was mislabelled. Thus the name
<emphasis id="B9113D09F30D3421FF59F1A33F5CBC8C" box="[226,327,1185,1210]" italics="true" pageId="9" pageNumber="57">hainana</emphasis>
may be used as the valid subspecific name for populations of
<taxonomicName id="4C659A98F30D3421FBE9F1A33AD0BC8C" box="[1106,1227,1185,1210]" class="Insecta" family="Hesperiidae" genus="Tagiades" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="57" phylum="Arthropoda" rank="species" species="japetus">
<emphasis id="B9113D09F30D3421FBE9F1A33AD0BC8C" box="[1106,1227,1185,1210]" italics="true" pageId="9" pageNumber="57">T. japetus</emphasis>
</taxonomicName>
(
<emphasis id="B9113D09F30D3421FB65F1A33B8ABC8D" bold="true" box="[1246,1425,1185,1211]" pageId="9" pageNumber="57">status revised</emphasis>
) inhabiting the island pending further research.
</paragraph>
</subSubSection>
</treatment>
</document>