349 lines
56 KiB
XML
349 lines
56 KiB
XML
<document id="B5579138C0CDCD9E82293F0861A53690" ID-CLB-Dataset="21439" ID-DOI="10.11646/zootaxa.4816.4.2" ID-GBIF-Dataset="361f5fc5-8f69-4803-a3b5-3c9ad44e3f7a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954577" ID-ZooBank="1F9A8E2B-B063-40B2-9CCB-9038D0B69DBF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595340130676" checkinUser="plazi" docAuthor="Shahdadi, Adnan & Schubart, Christoph D." docDate="2020" docId="03A212449A6E324EFF42F38BFCE8EEFD" docLanguage="en" docName="zootaxa.4816.4.2.pdf" docOrigin="Zootaxa 4816 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptarma dispersum Shahdadi & Schubart 2020, n. sp." docType="treatment" docVersion="7" lastPageNumber="445" masterDocId="FF9B6A3C9A6D3248FFD5F006FFC3ED36" masterDocTitle="Two new species of Leptarma (Decapoda, Brachyura, Sesarmidae) from the Western Indian Ocean with notes on their phylogeny" masterLastPageNumber="456" masterPageNumber="439" pageNumber="442" updateTime="1698843807735" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5C2F18938D8A39F96CB70D1483EF4F7F">Two new species of Leptarma (Decapoda, Brachyura, Sesarmidae) from the Western Indian Ocean with notes on their phylogeny</mods:title>
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<mods:name id="5FB7B3B289C9AF4971C3725BCFFCB51D" type="personal">
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<mods:namePart id="A58E9CE929DD18749A5CB6F0F3B7EB52">Shahdadi, Adnan</mods:namePart>
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<mods:affiliation id="427AA5182FA2E4185C0D8EACEB3A44BF">Department of Marine Biology, Faculty of Marine Sciences and Technology, University of Hormozgan, Bandar Abbas, Iran. adnan 1361 @ gmail. com; https: // orcid. org / 0000 - 0001 - 5131 - 7700</mods:affiliation>
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<mods:namePart id="3CEBCC7CA83D718F2D69C9B2E3A971CF">Schubart, Christoph D.</mods:namePart>
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<mods:affiliation id="62B51E9809733CF149F7AD0F996971F1">Zoology & Evolution, University of Regensburg, 93040 Regensburg, Germany.</mods:affiliation>
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<taxonomicName id="4C0BD8D19A6E324BFF42F38BFE53EE9E" ID-CoL="84GLR" authority="Shahdadi & Schubart, 2020" authorityName="Shahdadi & Schubart" authorityYear="2020" box="[151,400,909,936]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="dispersum" status="sp. nov.">
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<emphasis id="B97F7F409A6E324BFF42F38BFE53EE9E" bold="true" box="[151,400,909,936]" italics="true" pageId="3" pageNumber="442">Leptarma dispersum</emphasis>
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<taxonomicNameLabel id="A24CC23B9A6E324BFE42F38BFE1AEE91" box="[407,473,909,935]" pageId="3" pageNumber="442" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C311F0D99A6E324BFF42F3B7FC0EEED9" pageId="3" pageNumber="442" type="description">
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<paragraph id="8BB4A3529A6E324BFF42F3B7FE43EEFD" blockId="3.[151,973,909,1007]" box="[151,384,945,971]" pageId="3" pageNumber="442">
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(
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<figureCitation id="1330BFD79A6E324BFF4AF3B7FF2CEEFD" box="[159,239,945,971]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1878]" captionTargetBox="[162,1422,357,1810]" captionTargetId="figure@4.[147,1439,345,1827]" captionTargetPageId="4" captionText="FIGURE 2. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954581" httpUri="https://zenodo.org/record/3954581/files/figure.png" pageId="3" pageNumber="442">Figs. 2</figureCitation>
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,
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<figureCitation id="1330BFD79A6E324BFF2EF3B7FEC9EEFD" box="[251,266,945,971]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="3" pageNumber="442">3</figureCitation>
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,
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<figureCitation id="1330BFD79A6E324BFEC2F3B7FEFFEEFD" box="[279,316,945,971]" captionStart="FIGURE 7" captionStartId="10.[151,250,1866,1891]" captionTargetBox="[151,1431,190,1835]" captionTargetId="figure@10.[150,1436,181,1841]" captionTargetPageId="10" captionText="FIGURE 7. Comparisons of carapace and ambulatory legs, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="3" pageNumber="442">7A</figureCitation>
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,
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<figureCitation id="1330BFD79A6E324BFE92F3B7FEBBEEFD" box="[327,376,945,971]" captionStart="FIGURE 10" captionStartId="13.[151,250,1672,1697]" captionTargetBox="[155,1432,181,1646]" captionTargetId="figure@13.[155,1432,181,1646]" captionTargetPageId="13" captionText="FIGURE 10. Comparisons of G1, apical part, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); H, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="3" pageNumber="442">10A</figureCitation>
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)
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urn:lsid:zoobank.org:act:
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<emphasis id="B97F7F409A6E324BFF42F41CFE9FE907" box="[151,348,1049,1073]" italics="true" pageId="3" pageNumber="442">Parasesarma lenzii</emphasis>
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</taxonomicName>
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—
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Poupin
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<emphasis id="B97F7F409A6E324BFE16F41CFE33E907" box="[451,496,1049,1073]" italics="true" pageId="3" pageNumber="442">et al</emphasis>
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. (2012: 13
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,
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<figureCitation id="1330BFD79A6E324BFDBFF41EFD78E907" box="[618,699,1048,1073]" captionStart="FIGURE 9" captionStartId="12.[151,250,1898,1923]" captionTargetBox="[408,1179,181,1873]" captionTargetId="figure@12.[408,1179,181,1873]" captionTargetPageId="12" captionText="FIGURE 9. Comparisons of male chelar dactylus and tuberculation patterns, dorsal views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954597" httpUri="https://zenodo.org/record/3954597/files/figure.png" pageId="3" pageNumber="442">Fig. 9A</figureCitation>
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); not
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<taxonomicName id="4C0BD8D19A6E324BFD21F41CFBA6E907" ID-CoL="7XX37" authority="(De Man, 1895)" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[756,1125,1048,1073]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="lenzii">
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<emphasis id="B97F7F409A6E324BFD21F41CFC7AE907" box="[756,953,1049,1073]" italics="true" pageId="3" pageNumber="442">Parasesarma lenzii</emphasis>
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(De
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<bibRefCitation id="EF9ADEA39A6E324BFC3EF41CFB9FE907" author="Man, J. G. de" box="[1003,1116,1048,1073]" pageId="3" pageNumber="442" pagination="75 - 218" refId="ref8738" refString="Man, J. G. de (1895 - 1898) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 9, 75 - 218. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">Man, 1895</bibRefCitation>
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)
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</taxonomicName>
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.
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<emphasis id="B97F7F409A6E324BFF42F459FEBFE94C" bold="true" box="[151,380,1119,1146]" pageId="3" pageNumber="442">Material examined</emphasis>
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.
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<materialsCitation id="3B63A90F9A6E324BFE58F459FB02E9F4" ID-GBIF-Occurrence="2816753302" collectingDate="2011-11-11" collectorName="Channel & Poupin J. & Gravier-Bonnet N. & Zubia M. & Tisne N." country="Mozambique" county="Recifs Coralliens des Iles Eparses" location="Europa Island" municipality="Conservation des Recifs Coralliens" pageId="3" pageNumber="442" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B01DF09A6E324BFE58F459FE39E94F" box="[397,506,1119,1145]" pageId="3" pageNumber="442" type="holotype">Holotype</typeStatus>
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: male (12.2×10.1) (MNHN-IU-2012-660),
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, coll.
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<collectorName id="26FEC6849A6E324BFF1AF485FEFAE9A8" box="[207,313,1155,1182]" pageId="3" pageNumber="442">Poupin J.</collectorName>
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,
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<collectorName id="26FEC6849A6E324BFDF1F485FD4DE9A8" box="[548,654,1155,1182]" pageId="3" pageNumber="442">Zubia M.</collectorName>
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,
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<collectorName id="26FEC6849A6E324BFD4CF485FD39E9A8" box="[665,762,1155,1182]" pageId="3" pageNumber="442">Tisné N.</collectorName>
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(
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,
|
||
<location id="8ED4F5899A6E324BFBEEF482FB7AE9A8" LSID="urn:lsid:plazi:treatment:03A212449A6E324EFF42F38BFCE8EEFD:8ED4F5899A6E324BFBEEF482FB7AE9A8" box="[1083,1209,1156,1182]" country="Mozambique" county="Recifs Coralliens des Iles Eparses" municipality="Conservation des Recifs Coralliens" name="Ressources" pageId="3" pageNumber="442">Ressources</location>
|
||
et
|
||
<collectingMunicipality id="6BD039289A6E324BFB0EF485FE9AE9F7" pageId="3" pageNumber="442">
|
||
Conservation des
|
||
<collectingCounty id="62D5DBDE9A6E324BFF42F4A1FE9AE9F7" box="[151,345,1191,1217]" pageId="3" pageNumber="442">Récifs Coralliens</collectingCounty>
|
||
</collectingMunicipality>
|
||
des Îles Éparses (BIORECIFE) program, station 14) (see
|
||
<bibRefCitation id="EF9ADEA39A6E324BFC36F4A1FB76E9F4" author="Poupin, J. & Zubia, M. & Chabanet, P. & Malay, M." box="[995,1205,1191,1218]" pageId="3" pageNumber="442" pagination="1 - 25" refId="ref8911" refString="Poupin, J., Zubia, M., Chabanet, P. & Malay, M. (2012) Illustrated checklist of the Decapoda at Europa Island. WIO Journal of Marine Science, 11, 1 - 25." type="journal article" year="2012">
|
||
Poupin
|
||
<emphasis id="B97F7F409A6E324BFBECF4AEFBA9E9F4" box="[1081,1130,1192,1218]" italics="true" pageId="3" pageNumber="442">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311F0D99A6E324BFF12F4CDFCA2E8D7" pageId="3" pageNumber="442" type="diagnosis">
|
||
<paragraph id="8BB4A3529A6E324BFF12F4CDFCA2E8D7" blockId="3.[151,1437,1119,2010]" pageId="3" pageNumber="442">
|
||
<emphasis id="B97F7F409A6E324BFF12F4CDFEF9E9D0" bold="true" box="[199,314,1227,1254]" pageId="3" pageNumber="442">Diagnosis</emphasis>
|
||
. Carapace slightly broader than long, front deflexed, slightly sinuous in dorsal view, median postfrontal lobes slightly wider than lateral ones, carapace lateral margin almost straight; eye with eyestalk short, stout, wider than long, corneal part slightly wider; male chela large and robust, upper surface of palm with one oblique pectinated crest, dorsal surface of dactylus bearing 12 low, transversely oval, asymmetric tubercles, with longer distal slope, proximal tubercles positioned at middle part of upper face of palm; ambulatory legs flattened dorsoventrally, long, third pair (P4) longest, total length (ischium-dactylus) ca. 2.0 times carapace width; Male pleon triangular, somite 2 medially longer than lateral edges; G1 long, straight, apical corneous process long, tip rounded, bent at angle of about 50° to vertical axis, aperture subterminal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311F0D99A6E324EFF12F5EDFBF1EC29" lastPageId="6" lastPageNumber="445" pageId="3" pageNumber="442" type="description">
|
||
<paragraph id="8BB4A3529A6E324BFF12F5EDFE20EBE8" blockId="3.[151,1437,1119,2010]" pageId="3" pageNumber="442">
|
||
<emphasis id="B97F7F409A6E324BFF12F5EDFE92EB30" bold="true" box="[199,337,1515,1542]" pageId="3" pageNumber="442">Description</emphasis>
|
||
. Carapace subrectangular to slightly trapezoidal, slightly broader than long (cw/cl = 1.21), greatest width between exorbital angles; carapace surface smooth; front ca. 0.62 times carapace width, markedly deflexed, slightly sinuous in dorsal view; post-frontal lobes prominent, median lobes slightly wider than lateral ones, separated by shallow furrow; dorsal carapace regions well indicated, gastric region demarcated, cardiac region separated from intestinal region, lateral branchial ridges prominent; anterolateral margin with sharp exorbital angle, lateral margin almost straight, edged with row of short setae. Eye with eyestalk short, stout, wider than long, corneal part slightly wider (
|
||
<figureCitation id="1330BFD79A6E324BFE94F6C5FE66EBE8" box="[321,421,1731,1758]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1878]" captionTargetBox="[162,1422,357,1810]" captionTargetId="figure@4.[147,1439,345,1827]" captionTargetPageId="4" captionText="FIGURE 2. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954581" httpUri="https://zenodo.org/record/3954581/files/figure.png" pageId="3" pageNumber="442">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="1330BFD79A6E324BFE64F6C2FE15EBE8" box="[433,470,1732,1758]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="3" pageNumber="442">3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB4A3529A6E324CFF12F6E1FD54EC75" blockId="3.[151,1437,1119,2010]" lastBlockId="4.[151,1437,153,323]" lastPageId="4" lastPageNumber="443" pageId="3" pageNumber="442">
|
||
Male chela large (palm length/cw = 0.54) (
|
||
<figureCitation id="1330BFD79A6E324BFD7AF6E1FCCBEA34" box="[687,776,1767,1794]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1878]" captionTargetBox="[162,1422,357,1810]" captionTargetId="figure@4.[147,1439,345,1827]" captionTargetPageId="4" captionText="FIGURE 2. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954581" httpUri="https://zenodo.org/record/3954581/files/figure.png" pageId="3" pageNumber="442">Fig. 2B</figureCitation>
|
||
), robust (palm width/length = 0.48) (
|
||
<figureCitation id="1330BFD79A6E324BFB7BF6E1FAC4EA34" box="[1198,1287,1767,1794]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="3" pageNumber="442">Fig. 3B</figureCitation>
|
||
); merus with granulate dorsal border, no distinct subdistal spine, ventral border granulate, anterior border granulate, with distinct subdistal spine, accompanied by several spines distally, inner face smooth with longitudinal row of setae, anterior face smooth, outer face with rows of fine granules; upper surface of palm with 1 oblique pectinated crest and two rows of fine granules proximal to the crest, that is composed of 17 well developed teeth, and terminates at inner end in two granules and several granules on outer side (
|
||
<figureCitation id="1330BFD79A6E324BFCDAF79DFC49EA80" box="[783,906,1947,1974]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="3" pageNumber="442">Fig. 3C, D</figureCitation>
|
||
); outer surface of palm with short rows of fine granules except for smooth, punctate fixed finger; inner surface of palm finely granular except area facing carpus and distal half of finger; no vertical ridge; both sides of palm with scattered tufts of setae; dactylus slightly curved downward and inward, dorsal surface bearing 12 low but distinct, transversely oval tubercles, distinct to tip, tubercles asymmetric, with distal slope longer, proximal tubercles positioned at middle part of upper face of palm; small pointed granules on inner side (
|
||
<figureCitation id="1330BFD79A69324CFDEEF103FD77EC29" box="[571,692,261,287]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="4" pageNumber="443">Fig. 3C, D</figureCitation>
|
||
); scattered tufts of setae on inner side, cutting edge of both fingers with a series of variably sized teeth (
|
||
<figureCitation id="1330BFD79A69324CFDE7F12FFD4AEC75" box="[562,649,297,323]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="4" pageNumber="443">Fig. 3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF74F3DA9A69324CFF42F73BFAABEA60" ID-DOI="http://doi.org/10.5281/zenodo.3954581" ID-Zenodo-Dep="3954581" box="[151,1384,1853,1878]" httpUri="https://zenodo.org/record/3954581/files/figure.png" pageId="4" pageNumber="443" startId="4.[151,250,1853,1878]" targetBox="[162,1422,357,1810]" targetPageId="4">
|
||
<paragraph id="8BB4A3529A69324CFF42F73BFAABEA60" blockId="4.[151,1384,1853,1878]" box="[151,1384,1853,1878]" pageId="4" pageNumber="443">
|
||
<emphasis id="B97F7F409A69324CFF42F73BFED0EA60" bold="true" box="[151,275,1853,1878]" pageId="4" pageNumber="443">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C0BD8D19A69324CFECCF738FE5FEA63" authority="Shahdadi & Schubart, 2020" authorityName="Shahdadi & Schubart" authorityYear="2020" box="[281,412,1853,1877]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="dispersum" status="sp. nov.">
|
||
<emphasis id="B97F7F409A69324CFECCF738FE5FEA63" box="[281,412,1853,1877]" italics="true" pageId="4" pageNumber="443">L. dispersum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97F7F409A69324CFE76F73BFE1FEA63" bold="true" box="[419,476,1853,1877]" pageId="4" pageNumber="443">
|
||
<taxonomicNameLabel id="A24CC23B9A69324CFE76F73BFE1FEA63" box="[419,476,1853,1877]" pageId="4" pageNumber="443" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype, male (12.2×10.1) (MNHN-IU-2012-660). A, dorsal habitus; B, ventral habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB4A3529A69324DFF12F7AEFBF4EDE1" blockId="4.[151,1436,1960,2023]" lastBlockId="5.[151,1436,153,323]" lastPageId="5" lastPageNumber="444" pageId="4" pageNumber="443">
|
||
Ambulatory legs flattened dorso-ventrally, relatively long, third pair (P4) longest (
|
||
<figureCitation id="1330BFD79A69324CFB83F7AEFB5AEAF4" box="[1110,1177,1960,1986]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1878]" captionTargetBox="[162,1422,357,1810]" captionTargetId="figure@4.[147,1439,345,1827]" captionTargetPageId="4" captionText="FIGURE 2. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954581" httpUri="https://zenodo.org/record/3954581/files/figure.png" pageId="4" pageNumber="443">Fig. 2</figureCitation>
|
||
), total length (ischiumdactylus) ca. 2.0 times carapace width, cluster of setae on coxae of P3, P4, tufts of long setae scattered along the legs, more on propodi and dactyli, merus of P4 ca. 2.37 times as long as wide, with anterior margin crenulated, propodus ca. 3.5 times as long as wide, dactylus ca. 0.67 times length of propodus.
|
||
</paragraph>
|
||
<paragraph id="8BB4A3529A68324DFF12F0E7FCE2EC75" blockId="5.[151,1436,153,323]" pageId="5" pageNumber="444">
|
||
Male pleon triangular (
|
||
<figureCitation id="1330BFD79A68324DFE1EF0E7FDEDEDCD" box="[459,558,225,251]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1878]" captionTargetBox="[162,1422,357,1810]" captionTargetId="figure@4.[147,1439,345,1827]" captionTargetPageId="4" captionText="FIGURE 2. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954581" httpUri="https://zenodo.org/record/3954581/files/figure.png" pageId="5" pageNumber="444">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="1330BFD79A68324DFDEFF0E7FD98EDCD" box="[570,603,225,251]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="5" pageNumber="444">3E</figureCitation>
|
||
), with rounded telson, slightly wider than long (basal width/length = 1.1), almost as long as somite 6; somite 6 longer than others; somites 4 and 5 trapezoidal, somite 3 widest, laterally convex, somite 2 medially longer than lateral edges (
|
||
<figureCitation id="1330BFD79A68324DFD6AF12FFCD7EC75" box="[703,788,297,323]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="5" pageNumber="444">Fig. 3F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF74F3DA9A68324DFF42F693FEEBEA2F" ID-DOI="http://doi.org/10.5281/zenodo.3954583" ID-Zenodo-Dep="3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="5" pageNumber="444" startId="5.[151,250,1685,1710]" targetBox="[152,1435,345,1660]" targetPageId="5">
|
||
<paragraph id="8BB4A3529A68324DFF42F693FEEBEA2F" blockId="5.[151,1436,1685,1817]" pageId="5" pageNumber="444">
|
||
<emphasis id="B97F7F409A68324DFF42F693FED1EB98" bold="true" box="[151,274,1685,1710]" pageId="5" pageNumber="444">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C0BD8D19A68324DFECCF690FE58EB9B" authority="Shahdadi & Schubart, 2020" authorityName="Shahdadi & Schubart" authorityYear="2020" box="[281,411,1685,1709]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="dispersum" status="sp. nov.">
|
||
<emphasis id="B97F7F409A68324DFECCF690FE58EB9B" box="[281,411,1685,1709]" italics="true" pageId="5" pageNumber="444">L. dispersum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97F7F409A68324DFE74F693FE19EB9B" bold="true" box="[417,474,1685,1709]" pageId="5" pageNumber="444">
|
||
<taxonomicNameLabel id="A24CC23B9A68324DFE74F693FE19EB9B" box="[417,474,1685,1709]" pageId="5" pageNumber="444" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype, male (12.2×10.1) (MNHN-IU-2012-660). A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB4A3529A68324DFF12F76AFC96EA9C" blockId="5.[151,1437,1899,2034]" pageId="5" pageNumber="444">
|
||
G1 relatively long, straight; apical corneous process long, tip rounded, bent at angle of about 50° to vertical axis, arched in cross section, aperture subterminal (
|
||
<figureCitation id="1330BFD79A68324DFD00F796FCD5EA9C" box="[725,790,1936,1962]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="5" pageNumber="444">Fig. 3</figureCitation>
|
||
G–J).
|
||
</paragraph>
|
||
<paragraph id="8BB4A3529A68324DFF12F7B2FD02EAF8" blockId="5.[151,1437,1899,2034]" box="[199,705,1972,1998]" pageId="5" pageNumber="444">
|
||
<emphasis id="B97F7F409A68324DFF12F7B2FEDDEAF8" bold="true" box="[199,286,1972,1998]" pageId="5" pageNumber="444">Female</emphasis>
|
||
. No female specimen was examined.
|
||
</paragraph>
|
||
<paragraph id="8BB4A3529A68324EFF12F7D1FECCEDE1" blockId="5.[151,1437,1899,2034]" lastBlockId="6.[151,1437,153,971]" lastPageId="6" lastPageNumber="445" pageId="5" pageNumber="444">
|
||
<emphasis id="B97F7F409A68324DFF12F7D1FEC8EAC4" bold="true" box="[199,267,2007,2034]" pageId="5" pageNumber="444">Color</emphasis>
|
||
. The specimen preserved in ethanol does not show any coloration, but according to
|
||
<bibRefCitation id="EF9ADEA39A68324DFB68F7DEFA5AEAC4" author="Poupin, J. & Zubia, M. & Chabanet, P. & Malay, M." box="[1213,1433,2008,2034]" pageId="5" pageNumber="444" pagination="1 - 25" refId="ref8911" refString="Poupin, J., Zubia, M., Chabanet, P. & Malay, M. (2012) Illustrated checklist of the Decapoda at Europa Island. WIO Journal of Marine Science, 11, 1 - 25." type="journal article" year="2012">
|
||
Poupin
|
||
<emphasis id="B97F7F409A68324DFAC1F7DEFA86EAC4" box="[1300,1349,2008,2034]" italics="true" pageId="5" pageNumber="444">et al</emphasis>
|
||
. (2012
|
||
</bibRefCitation>
|
||
:
|
||
<figureCitation id="1330BFD79A6B324EFF42F09FFEC8ED85" box="[151,267,153,179]" captionStart="FIGURE 9" captionStartId="12.[151,250,1898,1923]" captionTargetBox="[408,1179,181,1873]" captionTargetId="figure@12.[408,1179,181,1873]" captionTargetPageId="12" captionText="FIGURE 9. Comparisons of male chelar dactylus and tuberculation patterns, dorsal views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954597" httpUri="https://zenodo.org/record/3954597/files/figure.png" pageId="6" pageNumber="445">Fig. 9B, p</figureCitation>
|
||
. 22), the carapace seems brownish, ambulatory legs with scattered black stains dorsally and chelae with orange tip.
|
||
</paragraph>
|
||
<paragraph id="8BB4A3529A6B324EFF12F0E6FBF1EC29" blockId="6.[151,1437,153,971]" pageId="6" pageNumber="445">
|
||
<emphasis id="B97F7F409A6B324EFF12F0E6FE28EDCD" bold="true" box="[199,491,224,251]" pageId="6" pageNumber="445">Habitat and distribution</emphasis>
|
||
. The only specimen known for this species was collected from Europa Island, Grand Lagoon with mangroves and dead coral blocks (
|
||
<bibRefCitation id="EF9ADEA39A6B324EFD65F103FC40EC29" author="Poupin, J. & Zubia, M. & Chabanet, P. & Malay, M." box="[688,899,261,287]" pageId="6" pageNumber="445" pagination="1 - 25" refId="ref8911" refString="Poupin, J., Zubia, M., Chabanet, P. & Malay, M. (2012) Illustrated checklist of the Decapoda at Europa Island. WIO Journal of Marine Science, 11, 1 - 25." type="journal article" year="2012">
|
||
Poupin
|
||
<emphasis id="B97F7F409A6B324EFCDDF100FCF9EC29" box="[776,826,261,287]" italics="true" pageId="6" pageNumber="445">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
: st.
|
||
<quantity id="4CF30EB79A6B324EFC72F103FC22EC29" box="[935,993,261,287]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="6" pageNumber="445" unit="in" value="14.0">14 in</quantity>
|
||
<figureCitation id="1330BFD79A6B324EFC3DF103FBE2EC29" box="[1000,1057,261,287]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1823]" captionTargetBox="[220,1376,684,1749]" captionTargetId="figure@2.[189,1398,669,1773]" captionTargetPageId="2" captionText="FIGURE 1. A, Phylogenetic consensus tree constructed with Maximum Likelihood (ML) and Bayesian Inference (BI) of the COX1 gene, focusing on Leptarma species; four species of Parasesarma were added as outgroups (only confidence values higher than 50% for ML (bootstrap value, bt) and 90% for BI (posterior probabilities, pp) are shown in the trees as bt/pp; B, C, maximum parsimony haplotype network of COX1 gene constructed with PopART for specimens of B, L. dispersum n. sp. and allied species; C, L. reunionense n. sp. and allied species." figureDoi="http://doi.org/10.5281/zenodo.3954579" httpUri="https://zenodo.org/record/3954579/files/figure.png" pageId="6" pageNumber="445">Fig 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311F0D99A6B324EFF12F12FFF35ECBD" pageId="6" pageNumber="445" type="etymology">
|
||
<paragraph id="8BB4A3529A6B324EFF12F12FFF35ECBD" blockId="6.[151,1437,153,971]" pageId="6" pageNumber="445">
|
||
<emphasis id="B97F7F409A6B324EFF12F12FFE8BEC75" bold="true" box="[199,328,297,323]" pageId="6" pageNumber="445">Etymology</emphasis>
|
||
. The name is derived from the Latin verb “dispergere” (dispersed or scattered) and refer to its isolated position. It furthermore is a direct translation of the French name for the archipelago where it occurs: Îles Éparses.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311F0D99A6B324EFF12F193FCE8EEFD" pageId="6" pageNumber="445" type="discussion">
|
||
<paragraph id="8BB4A3529A6B324EFF12F193FD29EFB1" blockId="6.[151,1437,153,971]" pageId="6" pageNumber="445">
|
||
<emphasis id="B97F7F409A6B324EFF12F193FEF7EC99" bold="true" box="[199,308,405,431]" pageId="6" pageNumber="445">Remarks</emphasis>
|
||
. The specimen resembles
|
||
<taxonomicName id="4C0BD8D19A6B324EFD88F190FD7BEC99" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[605,696,405,431]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="lenzii">
|
||
<emphasis id="B97F7F409A6B324EFD88F190FD7BEC99" box="[605,696,405,431]" italics="true" pageId="6" pageNumber="445">L. lenzii</emphasis>
|
||
</taxonomicName>
|
||
in the morphology of chelar dactylar tubercles and G1 (
|
||
<figureCitation id="1330BFD79A6B324EFAE5F193FF69ECE5" captionStart="FIGURE 10" captionStartId="13.[151,250,1672,1697]" captionTargetBox="[155,1432,181,1646]" captionTargetId="figure@13.[155,1432,181,1646]" captionTargetPageId="13" captionText="FIGURE 10. Comparisons of G1, apical part, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); H, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="6" pageNumber="445">Fig. 10A, B</figureCitation>
|
||
) and was first identified as such by
|
||
<bibRefCitation id="EF9ADEA39A6B324EFDE3F1BFFCD5ECE5" author="Poupin, J. & Zubia, M. & Chabanet, P. & Malay, M." box="[566,790,441,467]" pageId="6" pageNumber="445" pagination="1 - 25" refId="ref8911" refString="Poupin, J., Zubia, M., Chabanet, P. & Malay, M. (2012) Illustrated checklist of the Decapoda at Europa Island. WIO Journal of Marine Science, 11, 1 - 25." type="journal article" year="2012">
|
||
Poupin
|
||
<emphasis id="B97F7F409A6B324EFD5EF1BCFD7FECE5" box="[651,700,441,467]" italics="true" pageId="6" pageNumber="445">et al</emphasis>
|
||
. (2012)
|
||
</bibRefCitation>
|
||
. However, the specimen differ from
|
||
<taxonomicName id="4C0BD8D19A6B324EFB7FF1BCFAC7ECE5" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[1194,1284,441,467]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="lenzii">
|
||
<emphasis id="B97F7F409A6B324EFB7FF1BCFAC7ECE5" box="[1194,1284,441,467]" italics="true" pageId="6" pageNumber="445">L. lenzii</emphasis>
|
||
</taxonomicName>
|
||
in some morphological aspects. Most conspicuous is that the new species has proportionally longer ambulatory legs compared to
|
||
<taxonomicName id="4C0BD8D19A6B324EFF42F204FF32EF2D" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[151,241,513,539]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="lenzii">
|
||
<emphasis id="B97F7F409A6B324EFF42F204FF32EF2D" box="[151,241,513,539]" italics="true" pageId="6" pageNumber="445">L. lenzii</emphasis>
|
||
</taxonomicName>
|
||
, e.g. P4 (ischium-dactylus)/cw ca. 2.0 in the new species, while it is ca.
|
||
<quantity id="4CF30EB79A6B324EFBC3F207FB96EF2D" box="[1046,1109,513,539]" metricMagnitude="-2" metricUnit="m" metricValue="4.572" pageId="6" pageNumber="445" unit="in" value="1.8">1.8 in</quantity>
|
||
<taxonomicName id="4C0BD8D19A6B324EFB89F204FB75EF2D" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[1116,1206,513,539]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="lenzii">
|
||
<emphasis id="B97F7F409A6B324EFB89F204FB75EF2D" box="[1116,1206,513,539]" italics="true" pageId="6" pageNumber="445">L. lenzii</emphasis>
|
||
</taxonomicName>
|
||
; merus length/width ca.
|
||
<quantity id="4CF30EB79A6B324EFF6BF223FF3CEF09" box="[190,255,549,575]" metricMagnitude="-2" metricUnit="m" metricValue="5.842" pageId="6" pageNumber="445" unit="in" value="2.3">2.3 in</quantity>
|
||
the new species, while this is ca.
|
||
<quantity id="4CF30EB79A6B324EFDAEF223FD78EF09" box="[635,699,549,575]" metricMagnitude="-2" metricUnit="m" metricValue="5.334" pageId="6" pageNumber="445" unit="in" value="2.1">2.1 in</quantity>
|
||
<taxonomicName id="4C0BD8D19A6B324EFD16F220FCDEEF09" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[707,797,549,575]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="lenzii">
|
||
<emphasis id="B97F7F409A6B324EFD16F220FCDEEF09" box="[707,797,549,575]" italics="true" pageId="6" pageNumber="445">L. lenzii</emphasis>
|
||
</taxonomicName>
|
||
; propodus length/width ca.
|
||
<quantity id="4CF30EB79A6B324EFB8DF222FB5BEF09" box="[1112,1176,548,575]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" pageId="6" pageNumber="445" unit="in" value="3.5">3.5 in</quantity>
|
||
the new species, while it is ca.
|
||
<quantity id="4CF30EB79A6B324EFF3BF24FFEEDEF55" box="[238,302,585,611]" metricMagnitude="-2" metricUnit="m" metricValue="7.366" pageId="6" pageNumber="445" unit="in" value="2.9">2.9 in</quantity>
|
||
<taxonomicName id="4C0BD8D19A6B324EFEE0F24CFE53EF55" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[309,400,585,611]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="lenzii">
|
||
<emphasis id="B97F7F409A6B324EFEE0F24CFE53EF55" box="[309,400,585,611]" italics="true" pageId="6" pageNumber="445">L. lenzii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1330BFD79A6B324EFE4AF24FFDDBEF55" box="[415,536,585,611]" captionStart="FIGURE 7" captionStartId="10.[151,250,1866,1891]" captionTargetBox="[151,1431,190,1835]" captionTargetId="figure@10.[150,1436,181,1841]" captionTargetPageId="10" captionText="FIGURE 7. Comparisons of carapace and ambulatory legs, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="6" pageNumber="445">Fig. 7A, B</figureCitation>
|
||
). The new species differs slightly from
|
||
<taxonomicName id="4C0BD8D19A6B324EFC00F24CFBECEF55" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[981,1071,585,611]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="lenzii">
|
||
<emphasis id="B97F7F409A6B324EFC00F24CFBECEF55" box="[981,1071,585,611]" italics="true" pageId="6" pageNumber="445">L. lenzii</emphasis>
|
||
</taxonomicName>
|
||
in G1 morphology by having an apical corneous process with wider tip (
|
||
<figureCitation id="1330BFD79A6B324EFD80F26BFD1FEFB1" box="[597,732,621,647]" captionStart="FIGURE 10" captionStartId="13.[151,250,1672,1697]" captionTargetBox="[155,1432,181,1646]" captionTargetId="figure@13.[155,1432,181,1646]" captionTargetPageId="13" captionText="FIGURE 10. Comparisons of G1, apical part, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); H, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="6" pageNumber="445">Fig. 10A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB4A3529A6B324EFF12F297FD4CEE0D" blockId="6.[151,1437,153,971]" pageId="6" pageNumber="445">
|
||
The new species differs from the East African
|
||
<taxonomicName id="4C0BD8D19A6B324EFD04F294FCDEEF9D" box="[721,797,658,683]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="gazi">
|
||
<emphasis id="B97F7F409A6B324EFD04F294FCDEEF9D" box="[721,797,658,683]" italics="true" pageId="6" pageNumber="445">L. gazi</emphasis>
|
||
</taxonomicName>
|
||
in several morphological aspects as well as length of the ambulatory legs (
|
||
<figureCitation id="1330BFD79A6B324EFE88F2B3FE1BEFF9" box="[349,472,693,719]" captionStart="FIGURE 7" captionStartId="10.[151,250,1866,1891]" captionTargetBox="[151,1431,190,1835]" captionTargetId="figure@10.[150,1436,181,1841]" captionTargetPageId="10" captionText="FIGURE 7. Comparisons of carapace and ambulatory legs, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="6" pageNumber="445">Fig. 7A, C</figureCitation>
|
||
) and G1 morphology. P4 (ischium-dactylus) / cw ca. 2.0 in the new species, while it is ca.
|
||
<quantity id="4CF30EB79A6B324EFF0CF2DFFEDAEFC5" box="[217,281,729,755]" metricMagnitude="-2" metricUnit="m" metricValue="4.826" pageId="6" pageNumber="445" unit="in" value="1.9">1.9 in</quantity>
|
||
<taxonomicName id="4C0BD8D19A6B324EFEF5F2DCFEADEFC5" box="[288,366,730,755]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="gazi">
|
||
<emphasis id="B97F7F409A6B324EFEF5F2DCFEADEFC5" box="[288,366,730,755]" italics="true" pageId="6" pageNumber="445">L. gazi</emphasis>
|
||
</taxonomicName>
|
||
. The G
|
||
<quantity id="4CF30EB79A6B324EFE13F2DFFE2DEFC5" box="[454,494,729,755]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="445" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C0BD8D19A6B324EFE20F2DCFD81EFC5" box="[501,578,730,755]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="gazi">
|
||
<emphasis id="B97F7F409A6B324EFE20F2DCFD81EFC5" box="[501,578,730,755]" italics="true" pageId="6" pageNumber="445">L. gazi</emphasis>
|
||
</taxonomicName>
|
||
is curved with a long apical corneous process, bent at an angle of about 10° with vertical axis, while the G
|
||
<quantity id="4CF30EB79A6B324EFE3BF2FBFDD5EE21" box="[494,534,765,791]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="445" unit="in" value="1.0">1 in</quantity>
|
||
the new species is straight, with wider apical corneous process, bent at an angle of about 50° with vertical axis (
|
||
<figureCitation id="1330BFD79A6B324EFE2CF327FD41EE0D" box="[505,642,801,827]" captionStart="FIGURE 10" captionStartId="13.[151,250,1672,1697]" captionTargetBox="[155,1432,181,1646]" captionTargetId="figure@13.[155,1432,181,1646]" captionTargetPageId="13" captionText="FIGURE 10. Comparisons of G1, apical part, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); H, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="6" pageNumber="445">Fig. 10A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB4A3529A6B324EFF12F340FCE8EEFD" blockId="6.[151,1437,153,971]" pageId="6" pageNumber="445">
|
||
<taxonomicName id="4C0BD8D19A6B324EFF12F340FE6DEE69" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[199,430,837,863]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="leptosoma">
|
||
<emphasis id="B97F7F409A6B324EFF12F340FE6DEE69" box="[199,430,837,863]" italics="true" pageId="6" pageNumber="445">Leptarma leptosoma</emphasis>
|
||
</taxonomicName>
|
||
is the more common East African congener in the area. This species is also different from the new species in many features. For example the new species has a wider carapace (cw/cl ca. 1.2) and a G1 with long apical corneous process, while
|
||
<taxonomicName id="4C0BD8D19A6B324EFDE6F388FD06EE91" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[563,709,909,935]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="leptosoma">
|
||
<emphasis id="B97F7F409A6B324EFDE6F388FD06EE91" box="[563,709,909,935]" italics="true" pageId="6" pageNumber="445">L. leptosoma</emphasis>
|
||
</taxonomicName>
|
||
has a more squarish carapace (cw/cl less than 1.1) (
|
||
<figureCitation id="1330BFD79A6B324EFACDF38BFA57EE91" box="[1304,1428,909,935]" captionStart="FIGURE 7" captionStartId="10.[151,250,1866,1891]" captionTargetBox="[151,1431,190,1835]" captionTargetId="figure@10.[150,1436,181,1841]" captionTargetPageId="10" captionText="FIGURE 7. Comparisons of carapace and ambulatory legs, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="6" pageNumber="445">Fig. 7A, D</figureCitation>
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) and a G1 with short apical corneous process (
|
||
<figureCitation id="1330BFD79A6B324EFD41F3B7FCDCEEFD" box="[660,799,945,971]" captionStart="FIGURE 10" captionStartId="13.[151,250,1672,1697]" captionTargetBox="[155,1432,181,1646]" captionTargetId="figure@13.[155,1432,181,1646]" captionTargetPageId="13" captionText="FIGURE 10. Comparisons of G1, apical part, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); H, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="6" pageNumber="445">Fig. 10A, D</figureCitation>
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||
).
|
||
</paragraph>
|
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</subSubSection>
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</treatment>
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</document> |