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<document id="E83E520BACF65E53EB9BB351D6340464" ID-CLB-Dataset="7955" ID-DOI="10.11646/zootaxa.4878.1.6" ID-GBIF-Dataset="1ddcd915-594e-4b9c-8c80-913922c14b26" ID-ISSN="1175-5326" ID-Zenodo-Dep="4424780" ID-ZooBank="ED7F02DC-27B8-4BB8-A4C7-EC1995D5C473" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610035136151" checkinUser="plazi" docAuthor="Soto, Eulogio H., Martín, Guillermo San &amp; Lucas, Yolanda" docDate="2020" docId="039EEC36FFC49B53399AFC38FAD4FE84" docLanguage="en" docName="zootaxa.4878.1.6.pdf" docOrigin="Zootaxa 4878 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Syllis patersoni Soto &amp; Martín &amp; Lucas 2020, new species" docType="treatment" docVersion="8" lastPageNumber="155" masterDocId="FFA7944EFFCC9B58390DFFEAFF9DFFE1" masterDocTitle="Three new species of Syllis Savigny in Lamarck, 1818 (Annelida, Syllidae) from Chilean Patagonia" masterLastPageNumber="158" masterPageNumber="145" pageNumber="153" updateTime="1698923634718" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="5D0EA6C37C73D4869030C26CE48CEE0C">
<mods:title id="84609271264D6C7DA0500F3FA6671679">Three new species of Syllis Savigny in Lamarck, 1818 (Annelida, Syllidae) from Chilean Patagonia</mods:title>
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<mods:name id="E8515F10706B36A685D11B0901E246D1" type="personal">
<mods:role id="E2868E79A4BDA3B194144603B9780D84">
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<mods:namePart id="25B5B21E30AED0141BF11C8435FEE61F">Soto, Eulogio H.</mods:namePart>
<mods:nameIdentifier id="19B0496955E3A876C3CB358B72A2F392" type="ORCID">0000-0003-1056-4494</mods:nameIdentifier>
<mods:affiliation id="B95F1330C306F4A6FD7457244503A80B">Facultad de Ciencias del Mar y de Recursos Naturales, Universidad de Valparaíso. Avenida Borgoño 16344, Viña del Mar. Chile eulogio. soto @ uv. cl; https: // orcid. org / 0000 - 0003 - 1056 - 4494 &amp; Centro de Observación Marino para Estudios de Riesgos del Ambiente Costero (COSTAR), Universidad de Valparaíso, Chile. Corresponding author</mods:affiliation>
<mods:nameIdentifier id="38C33D2EF1CFB8EFDEF3E1C0E2BA4F09" type="email">eulogio.soto@uv.cl</mods:nameIdentifier>
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<mods:role id="F8B0E31A46C21EB4542618D438DE2FD3">
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<mods:namePart id="8E5650C7A8C532AAEA39F2F9299E7808">Martín, Guillermo San</mods:namePart>
<mods:affiliation id="C3A49F2228DCB467CCB8796CC30AFB07">Departamento de Biología (Zoología), Facultad de Ciencias, Universidad Autónoma de Madrid, Canto Blanco, 28049 Madrid, Spain &amp; Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM). Universidad Autónoma de Madrid, Calle Darwin 2, E- 28049 Madrid, Spain.</mods:affiliation>
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<mods:role id="A513AAE6E34C89277EFEE6482069AA9A">
<mods:roleTerm id="A6A518C2D93711D4FD9DA03CF143B649">Author</mods:roleTerm>
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<mods:namePart id="CC625121368176476D954B6AB4718761">Lucas, Yolanda</mods:namePart>
<mods:affiliation id="45F63E0595375BD9042D49D7590EC9BD">Departamento de Biología (Zoología), Facultad de Ciencias, Universidad Autónoma de Madrid, Canto Blanco, 28049 Madrid, Spain &amp; Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM). Universidad Autónoma de Madrid, Calle Darwin 2, E- 28049 Madrid, Spain.</mods:affiliation>
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<mods:title id="7C8A5DFBA15AF9F5E554F3603B1EDF06">Zootaxa</mods:title>
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<mods:part id="68E15E45C82A49B4276D364FA9FDE5F3">
<mods:date id="2F0D951E8FC433EFBB2E543C7E2B4703">2020</mods:date>
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<mods:number id="616F9B45D630D9097925D20E9258BF72">2020-11-12</mods:number>
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<mods:number id="A988FF201FF4703389437664ED6489B2">4878</mods:number>
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<mods:identifier id="34A30F205C5EC8FA09553C4BBBB648B0" type="DOI">10.11646/zootaxa.4878.1.6</mods:identifier>
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<treatment id="039EEC36FFC49B53399AFC38FAD4FE84" ID-DOI="http://doi.org/10.5281/zenodo.4425543" ID-GBIF-Taxon="179983506" ID-Zenodo-Dep="4425543" LSID="urn:lsid:plazi:treatment:039EEC36FFC49B53399AFC38FAD4FE84" httpUri="http://treatment.plazi.org/id/039EEC36FFC49B53399AFC38FAD4FE84" lastPageId="11" lastPageNumber="155" pageId="8" pageNumber="153">
<subSubSection id="C32D0EABFFC49B50399AFC38FE7AFC0D" box="[151,487,978,1005]" pageId="8" pageNumber="153" type="nomenclature">
<paragraph id="8B885D20FFC49B50399AFC38FE7AFC0D" blockId="8.[151,487,978,1041]" box="[151,487,978,1005]" pageId="8" pageNumber="153">
<heading id="D0C0EA4CFFC49B50399AFC38FE7AFC0D" bold="true" box="[151,487,978,1005]" fontSize="11" level="1" pageId="8" pageNumber="153" reason="1">
<emphasis id="B9438132FFC49B50399AFC38FE7AFC0D" bold="true" box="[151,487,978,1005]" pageId="8" pageNumber="153">
<taxonomicName id="4C3726A3FFC49B50399AFC38FECDFC0C" ID-CoL="85S3W" authority="Soto &amp; Martín &amp; Lucas, 2020" authorityName="Soto &amp; Martín &amp; Lucas" authorityYear="2020" box="[151,336,978,1005]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="153" phylum="Annelida" rank="species" species="patersoni" status="sp. nov.">
<emphasis id="B9438132FFC49B50399AFC38FECDFC0C" bold="true" box="[151,336,978,1005]" italics="true" pageId="8" pageNumber="153">Syllis patersoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2703C49FFC49B503855FC39FE7AFC0D" box="[344,487,978,1005]" pageId="8" pageNumber="153" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32D0EABFFC49B50399AFC1DFF68FBF0" box="[151,245,1015,1041]" pageId="8" pageNumber="153" type="description">
<paragraph id="8B885D20FFC49B50399AFC1DFF68FBF0" blockId="8.[151,487,978,1041]" box="[151,245,1015,1041]" pageId="8" pageNumber="153">
<figureCitation id="130C41A5FFC49B50399AFC1DFF68FBF0" box="[151,245,1015,1041]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Figure 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D0EABFFC49B50399AFBD4FBBCFB9C" pageId="8" pageNumber="153" type="materials_examined">
<paragraph id="8B885D20FFC49B50399AFBD4FBBCFB9C" blockId="8.[151,1437,1086,2013]" pageId="8" pageNumber="153">
<emphasis id="B9438132FFC49B50399AFBD4FEE5FBB9" bold="true" box="[151,376,1086,1112]" pageId="8" pageNumber="153">Material examined</emphasis>
.
<materialsCitation id="3B5F577DFFC49B503888FBD5FB81FB9C" ID-GBIF-Occurrence="3044013301" collectingDate="2009-10-19" collectionCode="MNCN" latitude="-50.276943" longLatPrecision="18" longitude="-74.88917" pageId="8" pageNumber="153" specimenCode="MNCN 16.01" specimenCount="2" typeStatus="holotype">
St. 41.
<typeStatus id="548CE382FFC49B5038DEFBD5FDA6FBB8" box="[467,571,1087,1113]" pageId="8" pageNumber="153" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DB91F55BFFC49B503B44FBD5FD7FFBBB" box="[585,738,1087,1114]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="8" pageNumber="153" type="Museum">MNCN 16.01</specimenCode>
/18728) and two
<typeStatus id="548CE382FFC49B503A92FBD5FB91FBB8" box="[927,1036,1087,1113]" pageId="8" pageNumber="153" type="paratype">Paratypes</typeStatus>
(
<specimenCode id="DB91F55BFFC49B503D14FBD5FB29FBBB" box="[1049,1204,1087,1114]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="8" pageNumber="153" type="Museum">MNCN 16.01</specimenCode>
/18764). Date of collection:
<date id="FF897BE0FFC49B5039FFFB89FE33FB9C" box="[242,430,1123,1149]" pageId="8" pageNumber="153" value="2009-10-19">
<collectingDate id="EFCD8208FFC49B5039FFFB89FE33FB9C" box="[242,430,1123,1149]" pageId="8" pageNumber="153" value="2009-10-19">19 October 2009</collectingDate>
</date>
(St. 41). Coordinates: St. 41
<geoCoordinate id="EE033BE7FFC49B503BFBFB88FC1FFB9C" box="[758,898,1122,1149]" degrees="50" direction="south" minutes="16" orientation="latitude" pageId="8" pageNumber="153" precision="15" seconds="37" value="-50.276943">50°16’37” S</geoCoordinate>
<geoCoordinate id="EE033BE7FFC49B503A85FB88FB81FB9C" box="[904,1052,1122,1149]" degrees="74" direction="west" minutes="53" orientation="longitude" pageId="8" pageNumber="153" precision="15" seconds="21" value="-74.88917">74°53’21’’W</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D0EABFFC49B5039CAFB6CFC6FF83C" pageId="8" pageNumber="153" type="description">
<paragraph id="8B885D20FFC49B5039CAFB6CFC6FF83C" blockId="8.[151,1437,1086,2013]" pageId="8" pageNumber="153">
<emphasis id="B9438132FFC49B5039CAFB6CFECCFB41" bold="true" box="[199,337,1158,1184]" pageId="8" pageNumber="153">Description</emphasis>
. Body elongated, slender (
<figureCitation id="130C41A5FFC49B503B8DFB6DFD47FB40" box="[640,730,1159,1185]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7A</figureCitation>
), without color markings;
<typeStatus id="548CE382FFC49B503D09FB6DFBFBFB40" box="[1028,1126,1159,1185]" pageId="8" pageNumber="153" type="holotype">holotype</typeStatus>
complete,
<quantity id="4CCFF0C5FFC49B503DEEFB6DFAAFFB40" box="[1251,1330,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="153" unit="mm" value="19.0">19 mm</quantity>
long,
<quantity id="4CCFF0C5FFC49B503C74FB6CFF5EFB24" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="153" unit="mm" value="0.5">0.5 mm</quantity>
wide, 132 chaetigers. Prostomium almost circular, with two pairs of small eyes in open trapezoidal arrangement; median antenna slightly longer than prostomium and palps together, inserted in front of posterior eyes, with about 19 articles; lateral antennae inserted in front of anterior eyes, shorter than median antenna, with 18 articles. Peristomium distinctly shorter than subsequent segments (
<figureCitation id="130C41A5FFC49B503A26FAFDFC1BFAD0" box="[811,902,1303,1329]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7A</figureCitation>
); dorsal tentacular cirri longer than median antenna, with 25 articles; ventral tentacular cirri shorter than dorsal ones, with 13 articles. Dorsal cirri similar in length to body width, longer on anterior segments, slender, elongated, with numerous spiralized glands inside articles (
<figureCitation id="130C41A5FFC49B503C62FAB5FF41FA7C" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7A, B</figureCitation>
); anterior dorsal cirri somewhat longer than remaining, with about 27-14-20-27-17 articles on first five chaetigers; dorsal cirri from proventricular segments alternating long cirri, with 29 articles, and short ones, with 19 articles (
<figureCitation id="130C41A5FFC49B5039ECFA21FEA4FA04" box="[225,313,1483,1509]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7B</figureCitation>
); articles plenty of distinct refringent glands (
<figureCitation id="130C41A5FFC49B503A3BFA21FC32FA04" box="[822,943,1483,1509]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7A, B</figureCitation>
). Parapodia conical; ventral cirri digitiform. Compound chaetae composed by few short, spiniger-like chaetae (
<figureCitation id="130C41A5FFC49B503A77FA05FB92F9E8" box="[890,1039,1519,1545]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7C, E, G</figureCitation>
) and several heterogomph falcigers with elongated, slender, bidentate blades, proximal teeth slightly smaller than distal ones, and short to moderate spines on margin, with dorso-ventral gradation in length (
<figureCitation id="130C41A5FFC49B503A22F9DDFC53F9B0" box="[815,974,1591,1617]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7D, F, H</figureCitation>
), similar throughout. Anterior parapodia each with 2–3 short spiniger-like chaetae, blades
<quantity id="4CCFF0C5FFC49B503BB6F9B1FC9EF994" box="[699,771,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" pageId="8" pageNumber="153" unit="mm" value="69.0">69 µm</quantity>
(
<figureCitation id="130C41A5FFC49B503A1FF9B1FCF5F994" box="[786,872,1627,1653]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7C</figureCitation>
) and 11–12 falcigers with dorso-ventral gradation, blades
<quantity id="4CCFF0C5FFC49B5039E8F995FEB0F978" box="[229,301,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="153" unit="mm" value="36.0">36 µm</quantity>
above,
<quantity id="4CCFF0C5FFC49B503889F995FE51F978" box="[388,460,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="153" unit="mm" value="22.0">22 µm</quantity>
below (
<figureCitation id="130C41A5FFC49B503B2BF995FDE2F978" box="[550,639,1663,1689]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7D</figureCitation>
); midbody parapodia each with two spiniger-like chaetae, blades 75–
<quantity id="4CCFF0C5FFC49B503C8FF994FF20F95C" metricMagnitude="-2" metricUnit="m" metricValue="7.1" pageId="8" pageNumber="153" unit="mm" value="71.0">71 µm</quantity>
long (
<figureCitation id="130C41A5FFC49B50380EF949FECAF95C" box="[259,343,1699,1725]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7E</figureCitation>
) and 6–7 falcigers, blades
<quantity id="4CCFF0C5FFC49B503B73F948FD5BF95C" box="[638,710,1698,1725]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="153" unit="mm" value="35.0">35 µm</quantity>
above,
<quantity id="4CCFF0C5FFC49B503A11F949FCF9F95C" box="[796,868,1699,1725]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="153" unit="mm" value="20.0">20 µm</quantity>
below (
<figureCitation id="130C41A5FFC49B503AB0F949FB8CF95C" box="[957,1041,1699,1725]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7F</figureCitation>
); posterior parapodia each with one spiniger-like, shorter than those of midbody and anterior segments, blades
<quantity id="4CCFF0C5FFC49B503AD5F92CFBBCF900" box="[984,1057,1734,1761]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="153" unit="mm" value="57.0">57 µm</quantity>
long (
<figureCitation id="130C41A5FFC49B503D65F92DFB5CF900" box="[1128,1217,1735,1761]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7G</figureCitation>
), and 5–6 falcigers, blades
<quantity id="4CCFF0C5FFC49B5039EAF901FEACF8E4" box="[231,305,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="153" unit="mm" value="29.0">29 µm</quantity>
above,
<quantity id="4CCFF0C5FFC49B503886F901FE48F8E4" box="[395,469,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="153" unit="mm" value="16.0">16 µm</quantity>
below (
<figureCitation id="130C41A5FFC49B503B3EF901FD13F8E4" box="[563,654,1771,1797]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7H</figureCitation>
). Solitary dorsal capillary chaetae on posterior parapodia, slender bidentate, with minute subdistal serrations (
<figureCitation id="130C41A5FFC49B503B67F8E5FD24F8C8" box="[618,697,1807,1833]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7I</figureCitation>
); ventral simple capillary chaetae on far posterior segments, more distinctly bidentate than dorsal one (
<figureCitation id="130C41A5FFC49B503B21F8D9FDE1F8AC" box="[556,636,1843,1869]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7J</figureCitation>
). Anterior parapodia with four slender, pointed aciculae (
<figureCitation id="130C41A5FFC49B503DFAF8D9FACCF8AC" box="[1271,1361,1843,1869]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7K</figureCitation>
), three on midbody parapodia, distinctly thicker (
<figureCitation id="130C41A5FFC49B503B67F8BDFD5DF890" box="[618,704,1879,1905]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7L</figureCitation>
), and single in posterior parapodia, acuminate (
<figureCitation id="130C41A5FFC49B503DC3F8BDFAB0F890" box="[1230,1325,1879,1905]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7M</figureCitation>
). Pharynx long and slender, through more than 12 segments (everted); pharyngeal tooth conical, on anterior margin (
<figureCitation id="130C41A5FFC49B503C3BF891FA12F874" box="[1334,1423,1915,1941]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7A</figureCitation>
). Proventricle slender and long, shorter than pharynx, through eight segments, with 42 muscle cell rows, without midline (
<figureCitation id="130C41A5FFC49B5039F1F829FECBF83C" box="[252,342,1987,2013]" captionStart="FIGURE 7" captionStartId="9.[151,250,1874,1898]" captionTargetBox="[192,1393,191,1832]" captionTargetId="figure-19@9.[181,1405,181,1850]" captionTargetPageId="9" captionText="FIGURE 7. Syllis patersoni n. sp. (Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="8" pageNumber="153">Fig. 7A</figureCitation>
). Pygidium small, with two long anal cirri, with 25 articles.
</paragraph>
</subSubSection>
<caption id="DF480DA8FFC59B51399AF8B8FBD3F81A" ID-DOI="http://doi.org/10.5281/zenodo.4424798" ID-Zenodo-Dep="4424798" httpUri="https://zenodo.org/record/4424798/files/figure.png" pageId="9" pageNumber="154" startId="9.[151,250,1874,1898]" targetBox="[192,1393,191,1832]" targetPageId="9">
<paragraph id="8B885D20FFC59B51399AF8B8FBD3F81A" blockId="9.[151,1437,1874,2043]" pageId="9" pageNumber="154">
<emphasis id="B9438132FFC59B51399AF8B8FE8BF88B" bold="true" box="[151,278,1874,1898]" pageId="9" pageNumber="154">FIGURE 7.</emphasis>
<taxonomicName id="4C3726A3FFC59B513812F8B8FE22F88B" authority="Soto &amp; Martín &amp; Lucas, 2020" authorityName="Soto &amp; Martín &amp; Lucas" authorityYear="2020" box="[287,447,1874,1898]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="154" phylum="Annelida" rank="species" species="patersoni" status="sp. nov.">
<emphasis id="B9438132FFC59B513812F8B8FE22F88B" box="[287,447,1874,1898]" italics="true" pageId="9" pageNumber="154">Syllis patersoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2703C49FFC59B5138CAF8B9FD9CF88B" box="[455,513,1875,1898]" pageId="9" pageNumber="154" rank="species">n. sp.</taxonomicNameLabel>
(Holotype, MNCN 16.01/18728). A, anterior end, dorsal view. B, dorsal cirri, midbody. C, spiniger-like chaeta, anterior parapodium. D, compound chaetae, anterior parapodium. E, spiniger-like chaetae, midbody parapodium. F, compound chaetae, midbody parapodium. G, spiniger-like chaeta, posterior parapodium. H, compound chaetae, posterior parapodium. I, dorsal simple chaeta. J, ventral simple chaeta. K, aciculae, anterior parapodium. L, aciculae, midbody parapodium. M, aciculum, posterior parapodium. Scale: A, 0.4 mm; B, 0,2 mm; C–M, 20 µm.
</paragraph>
</caption>
<subSubSection id="C32D0EABFFC69B5239CAFF7CFC07FEFC" pageId="10" pageNumber="155" type="distribution">
<paragraph id="8B885D20FFC69B5239CAFF7CFAFBFF50" blockId="10.[151,1437,150,2013]" box="[199,1382,150,177]" pageId="10" pageNumber="155">
<emphasis id="B9438132FFC69B5239CAFF7CFEC4FF51" bold="true" box="[199,345,150,176]" pageId="10" pageNumber="155">Distribution</emphasis>
. Only known from Chilean Patagonia in Concepción channel, Drumond Hay island (
<figureCitation id="130C41A5FFC69B523C1EFF7DFAC8FF50" box="[1299,1365,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1918,1942]" captionTargetBox="[232,1355,1099,1893]" captionTargetId="figure-412@1.[232,1355,1099,1893]" captionTargetPageId="1" captionText="FIGURE 1. Study area showing the sampling stations. Map modified from Google Earth©" figureDoi="http://doi.org/10.5281/zenodo.4424782" httpUri="https://zenodo.org/record/4424782/files/figure.png" pageId="10" pageNumber="155">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B885D20FFC69B5239CAFF50FC07FEFC" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="155">
<emphasis id="B9438132FFC69B5239CAFF50FEBEFF35" bold="true" box="[199,291,186,212]" pageId="10" pageNumber="155">Habitat</emphasis>
. From boulders, sediment bottoms and associated to
<taxonomicName id="4C3726A3FFC69B523A74FF51FC64FF34" authorityName="C. Agardh" authorityYear="1820" baseAuthorityName="Linnaeus" box="[889,1017,187,213]" class="Agaricomycetes" family="Marasmiaceae" genus="Macrocystis" higherTaxonomySource="CoL" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="155" phylum="Basidiomycota" rank="species" species="pyrifera">
<emphasis id="B9438132FFC69B523A74FF51FC64FF34" box="[889,1017,187,213]" italics="true" pageId="10" pageNumber="155">M. pyrifera</emphasis>
</taxonomicName>
holdfasts in fjords, islands and channels from Chilean Patagonia. Salinity: from 30.2 PSU, temperature: 8.9°C. Shallow subtidal at
<quantity id="4CCFF0C5FFC69B523DD4FF35FA89FF18" box="[1241,1300,223,249]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="155" unit="m" value="30.0">30 m</quantity>
depth. This species was not found inside tubes of
<taxonomicName id="4C3726A3FFC69B523B32FEE9FC0AFEFC" baseAuthorityName="Renier" baseAuthorityYear="1804" box="[575,919,259,285]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="variopedatus">
<emphasis id="B9438132FFC69B523B32FEE9FD4BFEFC" box="[575,726,259,285]" italics="true" pageId="10" pageNumber="155">Chaetopterus</emphasis>
cf.
<emphasis id="B9438132FFC69B523A0CFEE9FC0AFEFC" box="[769,919,259,285]" italics="true" pageId="10" pageNumber="155">variopedatus</emphasis>
</taxonomicName>
<emphasis id="B9438132FFC69B523A9BFEE9FC07FEFC" box="[918,922,259,285]" italics="true" pageId="10" pageNumber="155">.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32D0EABFFC69B5239CAFECDFB11FEA3" box="[199,1164,294,322]" pageId="10" pageNumber="155" type="materials_examined">
<paragraph id="8B885D20FFC69B5239CAFECDFB11FEA3" blockId="10.[151,1437,150,2013]" box="[199,1164,294,322]" pageId="10" pageNumber="155">
<materialsCitation id="3B5F577DFFC69B5239CAFECDFB14FEA3" ID-GBIF-Occurrence="3044013302" box="[199,1161,294,322]" country="Chile" location="Concepcion" municipality="Drumond Hay island" pageId="10" pageNumber="155" specimenCount="1" typeStatus="holotype">
<emphasis id="B9438132FFC69B5239CAFECDFEFDFEA1" bold="true" box="[199,352,294,321]" pageId="10" pageNumber="155">
<typeStatus id="548CE382FFC69B5239CAFECDFE9FFEA0" box="[199,258,295,321]" pageId="10" pageNumber="155">Type</typeStatus>
locality
</emphasis>
.
<location id="8EE80BFBFFC69B523863FECDFE69FEA0" LSID="urn:lsid:plazi:treatment:039EEC36FFC49B53399AFC38FAD4FE84:8EE80BFBFFC69B523863FECDFE69FEA0" box="[366,500,295,321]" country="Chile" municipality="Drumond Hay island" name="Concepcion" pageId="10" pageNumber="155">Concepción</location>
channel,
<collectingMunicipality id="6BECC75AFFC69B523B6DFECDFCD3FEA0" box="[608,846,295,321]" pageId="10" pageNumber="155">Drumond Hay island</collectingMunicipality>
, southern
<collectingCountry id="F3201DB0FFC69B523ACDFECDFC60FEA0" box="[960,1021,295,321]" name="Chile" pageId="10" pageNumber="155">Chile</collectingCountry>
(Patagonia)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D0EABFFC69B5239CAFEA1FBC7FAEC" pageId="10" pageNumber="155" type="discussion">
<paragraph id="8B885D20FFC69B5239CAFEA1FDD8FE30" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="155">
<emphasis id="B9438132FFC69B5239CAFEA1FEA9FE84" bold="true" box="[199,308,331,357]" pageId="10" pageNumber="155">Remarks</emphasis>
.
<taxonomicName id="4C3726A3FFC69B52384CFEA1FE73FE85" authority="Soto &amp; Martín &amp; Lucas, 2020" authorityName="Soto &amp; Martín &amp; Lucas" authorityYear="2020" box="[321,494,331,357]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="patersoni" status="sp. nov.">
<emphasis id="B9438132FFC69B52384CFEA1FE73FE85" box="[321,494,331,357]" italics="true" pageId="10" pageNumber="155">Syllis patersoni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2703C49FFC69B5238F6FEA1FDABFE84" box="[507,566,331,357]" pageId="10" pageNumber="155" rank="species">n. sp.</taxonomicNameLabel>
is characterized by its elongated, slender body, long pharynx and proventricle, acuminate posterior acicula, and two
<typeStatus id="548CE382FFC69B523B35FE85FDEEFE68" box="[568,627,367,393]" pageId="10" pageNumber="155">types</typeStatus>
of compound chaetae on each parapodium, relatively short spiniger-like chaetae and falcigers, both bidentate, with proximal teeth well-marked, and distal teeth slightly rounded, and short to moderate straight spines on margin.
</paragraph>
<paragraph id="8B885D20FFC69B5239CAFE31FA17FC0C" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="155">
<taxonomicName id="4C3726A3FFC69B5239CAFE31FDBBFE14" authority="San Martin, 1992" authorityName="San Martin" authorityYear="1992" box="[199,550,475,501]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="alosae">
<emphasis id="B9438132FFC69B5239CAFE31FEC8FE14" box="[199,341,475,501]" italics="true" pageId="10" pageNumber="155">Syllis alosae</emphasis>
<bibRefCitation id="EFA620D1FFC69B523853FE31FDBBFE14" author="San Martin, G." box="[350,550,475,501]" pageId="10" pageNumber="155" pagination="167 - 196" refId="ref8388" refString="San Martin, G. (1992) Syllis Savigny in Lamarck, 1818 (Syllinae: Syllidae: Polychaeta) from Cuba, the Gulf of Mexico, Florida and North Carolina, with a revision of several species described by Verrill. Bulletin of Marine Science, 51 (2), 167 - 196." type="journal article" year="1992">San Martín, 1992</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F3201DB0FFC69B523B7CFE31FD32FE14" box="[625,687,475,501]" name="Cuba" pageId="10" pageNumber="155">Cuba</collectingCountry>
, also has similar falcigers, but the spiniger-like are distally broad, shows a marked color pattern, the dorsal cirri are shorter, and the posterior acicula are straight, protruding out from the parapodial lobes (
<bibRefCitation id="EFA620D1FFC69B523887FDC9FDD3FDDC" author="San Martin, G." box="[394,590,547,573]" pageId="10" pageNumber="155" pagination="167 - 196" refId="ref8388" refString="San Martin, G. (1992) Syllis Savigny in Lamarck, 1818 (Syllinae: Syllidae: Polychaeta) from Cuba, the Gulf of Mexico, Florida and North Carolina, with a revision of several species described by Verrill. Bulletin of Marine Science, 51 (2), 167 - 196." type="journal article" year="1992">San Martín 1992</bibRefCitation>
).
<taxonomicName id="4C3726A3FFC69B523B6FFDC9FC56FDDC" authority="San Martin, 1992" authorityName="San Martin" authorityYear="1992" box="[610,971,547,573]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="maryae">
<emphasis id="B9438132FFC69B523B6FFDC9FD66FDDD" box="[610,763,547,573]" italics="true" pageId="10" pageNumber="155">Syllis maryae</emphasis>
<bibRefCitation id="EFA620D1FFC69B523A0EFDC9FC56FDDC" author="San Martin, G." box="[771,971,547,573]" pageId="10" pageNumber="155" pagination="167 - 196" refId="ref8388" refString="San Martin, G. (1992) Syllis Savigny in Lamarck, 1818 (Syllinae: Syllidae: Polychaeta) from Cuba, the Gulf of Mexico, Florida and North Carolina, with a revision of several species described by Verrill. Bulletin of Marine Science, 51 (2), 167 - 196." type="journal article" year="1992">San Martín, 1992</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F3201DB0FFC69B523D18FDC9FBCDFDDC" box="[1045,1104,547,573]" name="Cuba" pageId="10" pageNumber="155">Cuba</collectingCountry>
and
<collectingCountry id="F3201DB0FFC69B523D84FDC9FB4DFDDC" box="[1161,1232,547,573]" name="Brazil" pageId="10" pageNumber="155">Brazil</collectingCountry>
, has similar body, dorsal cirri, pharynx and proventricle, but the spiniger-like chaetae are proportionally longer, in relation to the falcigers, than those of
<taxonomicName id="4C3726A3FFC69B52388BFD81FDA8FD65" authority="Soto &amp; Martín &amp; Lucas, 2020" authorityName="Soto &amp; Martín &amp; Lucas" authorityYear="2020" box="[390,565,619,645]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="patersoni" status="sp. nov.">
<emphasis id="B9438132FFC69B52388BFD81FDA8FD65" box="[390,565,619,645]" italics="true" pageId="10" pageNumber="155">Syllis patersoni</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2703C49FFC69B523B48FD81FD1FFD64" box="[581,642,619,645]" pageId="10" pageNumber="155" rank="species">n. sp.</taxonomicNameLabel>
The posterior acicula are distally bent at a right angle, and the dorsal simple chaetae are truncate; furthermore, some specimens show some dorsal glands on some post-proventricular segments (
<bibRefCitation id="EFA620D1FFC69B523800FD59FE51FD2C" author="San Martin, G." box="[269,460,691,717]" pageId="10" pageNumber="155" pagination="167 - 196" refId="ref8388" refString="San Martin, G. (1992) Syllis Savigny in Lamarck, 1818 (Syllinae: Syllidae: Polychaeta) from Cuba, the Gulf of Mexico, Florida and North Carolina, with a revision of several species described by Verrill. Bulletin of Marine Science, 51 (2), 167 - 196." type="journal article" year="1992">San Martín 1992</bibRefCitation>
;
<bibRefCitation id="EFA620D1FFC69B5238DAFD59FCBCFD2C" author="Nogueira, J. M. &amp; San Martin, G." box="[471,801,691,717]" pageId="10" pageNumber="155" pagination="57 - 93" refId="ref8241" refString="Nogueira, J. M. &amp; San Martin, G. (2002) Species of Syllis Lamarck, 1818 (Polychaeta: Syllidae) living in corals in the state of Sao Paulo, southeastern Brazil. Beaufortia, 52 (7), 57 - 93." type="journal article" year="2002">Nogueira &amp; San Martín 2002</bibRefCitation>
). Similar differences are shared with
<taxonomicName id="4C3726A3FFC69B523DB1FD59FE08FD10" authority="(Hartmann-Schroder, 1989)" baseAuthorityName="Hartmann-Schroder" baseAuthorityYear="1989" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="edensis">
<emphasis id="B9438132FFC69B523DB1FD59FACCFD2C" box="[1212,1361,691,717]" italics="true" pageId="10" pageNumber="155">Syllis edensis</emphasis>
(
<bibRefCitation id="EFA620D1FFC69B523C52FD59FE10FD10" author="Hartmann-Schroder, G." pageId="10" pageNumber="155" pagination="11 - 63" refId="ref7506" refString="Hartmann-Schroder, G. (1989) Teil 14. Die Polychaeten der antiborealen und subtropisch-tropischen Kuste Sudost-Australien zwischen Lakes Entrance (Victoria) im Suden und Maclean (New South Wales) im Norden. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 86, 11 - 63." type="journal article" year="1989">Hartmann-Schröder, 1989</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F3201DB0FFC69B5238D3FD3DFDD8FD10" box="[478,581,727,753]" name="Australia" pageId="10" pageNumber="155">Australia</collectingCountry>
(
<bibRefCitation id="EFA620D1FFC69B523B5BFD3DFCE1FD10" author="Hartmann-Schroder, G." box="[598,892,727,753]" pageId="10" pageNumber="155" pagination="11 - 63" refId="ref7506" refString="Hartmann-Schroder, G. (1989) Teil 14. Die Polychaeten der antiborealen und subtropisch-tropischen Kuste Sudost-Australien zwischen Lakes Entrance (Victoria) im Suden und Maclean (New South Wales) im Norden. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 86, 11 - 63." type="journal article" year="1989">Hartmann-Schröder 1989</bibRefCitation>
;
<bibRefCitation id="EFA620D1FFC69B523A86FD32FB7EFD10" author="Alvarez-Campos, P. &amp; San Martin, G. &amp; Hutchings, P." box="[907,1251,726,755]" pageId="10" pageNumber="155" pagination="297 - 331" refId="ref6539" refString="Alvarez-Campos, P., San Martin, G. &amp; Hutchings, P. (2015 b) The genus Syllis Lamark, 1818 (Annelida, Syllidae) from Australia. Molecular analysis and re-description of some poorly-know species. Zootaxa, 4052 (2), 297 - 331. https: // doi. org / 10.11646 / zootaxa. 4052.3.2" type="journal article" year="2015">
Álvarez-Campos
<emphasis id="B9438132FFC69B523D5FFD3DFB13FD10" box="[1106,1166,727,753]" italics="true" pageId="10" pageNumber="155">et al.</emphasis>
2015b
</bibRefCitation>
).
<taxonomicName id="4C3726A3FFC69B523DF5FD3DFEA8FCF4" authority="Gravier, 1900" authorityName="Gravier" authorityYear="1900" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="bouvieri">
<emphasis id="B9438132FFC69B523DF5FD3DFA01FD10" box="[1272,1436,727,753]" italics="true" pageId="10" pageNumber="155">Syllis bouvieri</emphasis>
<bibRefCitation id="EFA620D1FFC69B52399AFD11FEA8FCF4" author="Gravier, C." box="[151,309,763,789]" pageId="10" pageNumber="155" pagination="137 - 282" refId="ref7209" refString="Gravier, C. (1900) Contribution a l'etude des Annelides Polychetes de la Mer Rouge. Premiere partie. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 2, 137 - 282." type="journal article" year="1900">Gravier, 1900</bibRefCitation>
</taxonomicName>
, from the Red Sea, also has similar
<typeStatus id="548CE382FFC69B523BC7FD11FD67FCF4" box="[714,762,763,789]" pageId="10" pageNumber="155">type</typeStatus>
of compound chaetae but the longer ones are not so long in proportion with the falcigers, the proximal and distal teeth are more separated, with a curved arc in between, the spines on margin of blades are longer, the dorsal cirri are longer, and the proventricle is shorter and wider (
<bibRefCitation id="EFA620D1FFC69B523C48FCA9FF4FFC60" author="Gravier, C." pageId="10" pageNumber="155" pagination="137 - 282" refId="ref7209" refString="Gravier, C. (1900) Contribution a l'etude des Annelides Polychetes de la Mer Rouge. Premiere partie. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 2, 137 - 282." type="journal article" year="1900">Gravier 1900</bibRefCitation>
;
<bibRefCitation id="EFA620D1FFC69B5239D3FC8DFE38FC60" author="Lucas, Y. &amp; San Martin, G. &amp; Fiege, D." box="[222,421,871,897]" pageId="10" pageNumber="155" pagination="73 - 88" refId="ref7913" refString="Lucas, Y., San Martin, G. &amp; Fiege, D. (2020) New species and records of the genus Syllis Savigny in Lamarck, 1818 (Annelida: Syllidae) from Socotra Archipelago (Indian Ocean). Zootaxa, 4742 (1), 73 - 88. https: // doi. org / 10.11646 / zootaxa. 4742.1.4" type="journal article" year="2020">
Lucas
<emphasis id="B9438132FFC69B523825FC8DFEFCFC60" box="[296,353,871,897]" italics="true" pageId="10" pageNumber="155">et al.</emphasis>
2020
</bibRefCitation>
).
<taxonomicName id="4C3726A3FFC69B5238B5FC8DFC8DFC60" authority="(Licher 1999)" baseAuthorityName="Licher" baseAuthorityYear="1999" box="[440,784,871,897]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="hyllebergi">
<emphasis id="B9438132FFC69B5238B5FC8DFDF6FC60" box="[440,619,871,897]" italics="true" pageId="10" pageNumber="155">Syllis hyllebergi</emphasis>
(
<bibRefCitation id="EFA620D1FFC69B523B76FC8DFC94FC60" author="Licher, F." box="[635,777,871,897]" pageId="10" pageNumber="155" pagination="1 - 336" refId="ref7875" refString="Licher, F. (1999) Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 551, 1 - 336." type="journal article" year="1999">Licher 1999</bibRefCitation>
)
</taxonomicName>
, from the Eastern Mediterranean, Red Sea and
<collectingCountry id="F3201DB0FFC69B523C29FC8DFAF6FC60" box="[1316,1387,871,897]" name="Brazil" pageId="10" pageNumber="155">Brazil</collectingCountry>
, has shorter dorsal cirri, shorter proventricle, and the falcigers are more markedly bidentate, with longer spines on margin (
<bibRefCitation id="EFA620D1FFC69B523992FC45FEB6FC28" author="Licher, F." box="[159,299,943,969]" pageId="10" pageNumber="155" pagination="1 - 336" refId="ref7875" refString="Licher, F. (1999) Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 551, 1 - 336." type="journal article" year="1999">Licher 1999</bibRefCitation>
;
<bibRefCitation id="EFA620D1FFC69B52383AFC45FD1EFC28" author="Nogueira, J. M. &amp; San Martin, G." box="[311,643,943,969]" pageId="10" pageNumber="155" pagination="57 - 93" refId="ref8241" refString="Nogueira, J. M. &amp; San Martin, G. (2002) Species of Syllis Lamarck, 1818 (Polychaeta: Syllidae) living in corals in the state of Sao Paulo, southeastern Brazil. Beaufortia, 52 (7), 57 - 93." type="journal article" year="2002">Nogueira &amp; San Martín 2002</bibRefCitation>
). Several other species also have spiniger-like and falciger chaetae, but the spiniger-like chaetae are much more longer and slender, with filiform blades, in proportion with the falcigers.
</paragraph>
<paragraph id="8B885D20FFC69B5239CAFC1DFE36FB40" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="155">
<taxonomicName id="4C3726A3FFC69B5239CAFC1DFD4FFBF0" authority="(Campoy &amp; Alquezar 1982)" baseAuthorityName="Campoy &amp; Alquezar" baseAuthorityYear="1982" box="[199,722,1015,1041]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="beneliahuae">
<emphasis id="B9438132FFC69B5239CAFC1DFE0EFBF0" box="[199,403,1015,1041]" italics="true" pageId="10" pageNumber="155">Syllis beneliahuae</emphasis>
(
<bibRefCitation id="EFA620D1FFC69B5238ACFC1DFD56FBF0" author="Campoy, A. &amp; Alquezar, E." box="[417,715,1015,1041]" pageId="10" pageNumber="155" pagination="121 - 137" refId="ref6902" refString="Campoy, A. &amp; Alquezar, E. (1982) Anelidos poliquetos de las formaciones de Dendropoma petraeum (Monterosato) de las costas del sureste de Espana. Actas del II Simposio Iberico de Estudios del Bentos Marino, Barcelona, 3, 121 - 137." type="journal article" year="1982">Campoy &amp; Alquézar 1982</bibRefCitation>
)
</taxonomicName>
, widely distributed in tropical and temperate seas, is also a similar species, with slender body, and relatively short spiniger-like chaetae, acuminate posterior acicula and relatively long dorsal cirri, with articles with plenty of refringent glands; however that species has longer spiniger-like chaetae (blades more than twice as long as the longer falciger) and shorter proventricle (
<bibRefCitation id="EFA620D1FFC69B523D07FB89FAAEFB9C" author="Campoy, A. &amp; Alquezar, E." box="[1034,1331,1123,1149]" pageId="10" pageNumber="155" pagination="121 - 137" refId="ref6902" refString="Campoy, A. &amp; Alquezar, E. (1982) Anelidos poliquetos de las formaciones de Dendropoma petraeum (Monterosato) de las costas del sureste de Espana. Actas del II Simposio Iberico de Estudios del Bentos Marino, Barcelona, 3, 121 - 137." type="journal article" year="1982">Campoy &amp; Alquézar 1982</bibRefCitation>
; Campoy 1982;
<bibRefCitation id="EFA620D1FFC69B5239D3FB6DFE02FB40" author="San Martin, G." box="[222,415,1159,1185]" pageId="10" pageNumber="155" refId="ref8446" refString="San Martin, G. (2003) Annelida, Polychaeta II, Syllidae. In: Ramos, M. A., Alba, J., Belles, X., Gosalbez, J., Guerra, A., Macpherson, E., Martin, F., Serrano, J. &amp; Templado, J. (Eds.), Fauna Iberica. Vol. 21. Museo Nacional de Ciencias Naturales, CSIC, Madrid, pp. 1 - 554." type="book" year="2003">San Martín 2003</bibRefCitation>
).
</paragraph>
<paragraph id="8B885D20FFC69B5239CAFB41FBC7FAEC" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="155">
<taxonomicName id="4C3726A3FFC69B5239CAFB41FDBCFB24" authority="Haswell, 1920" authorityName="Haswell" authorityYear="1920" box="[199,545,1195,1221]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="augeneri">
<emphasis id="B9438132FFC69B5239CAFB41FEECFB25" box="[199,369,1195,1221]" italics="true" pageId="10" pageNumber="155">Syllis augeneri</emphasis>
<bibRefCitation id="EFA620D1FFC69B523877FB41FDBCFB24" author="Haswell, W. A." box="[378,545,1195,1221]" pageId="10" pageNumber="155" pagination="90 - 112" refId="ref7557" refString="Haswell, W. A. (1920) Australian Syllidae, Eusyllidae and Autolytidae. Journal of the Linnean Society of London, 24, 90 - 112." type="journal article" year="1920">Haswell, 1920</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F3201DB0FFC69B523B60FB41FD49FB24" box="[621,724,1195,1221]" name="Australia" pageId="10" pageNumber="155">Australia</collectingCountry>
and
<collectingCountry id="F3201DB0FFC69B523A03FB41FCE3FB24" box="[782,894,1195,1221]" name="Indonesia" pageId="10" pageNumber="155">Indonesia</collectingCountry>
, also has acuminate posterior acicula and short spiniger-like chaetae, but the dorsal cirri are much shorter and proportionally thicker, and the spiniger-like chaetae are much shorter than those of
<taxonomicName id="4C3726A3FFC69B5238FFFB19FDE5FAED" authority="Soto &amp; Martín &amp; Lucas, 2020" authorityName="Soto &amp; Martín &amp; Lucas" authorityYear="2020" box="[498,632,1267,1293]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="patersoni" status="sp. nov.">
<emphasis id="B9438132FFC69B5238FFFB19FDE5FAED" box="[498,632,1267,1293]" italics="true" pageId="10" pageNumber="155">S. patersoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2703C49FFC69B523B72FB19FD26FAEC" box="[639,699,1267,1293]" pageId="10" pageNumber="155" rank="species">n. sp.</taxonomicNameLabel>
(
<bibRefCitation id="EFA620D1FFC69B523BC4FB19FCF4FAEC" author="Haswell, W. A." box="[713,873,1267,1293]" pageId="10" pageNumber="155" pagination="90 - 112" refId="ref7557" refString="Haswell, W. A. (1920) Australian Syllidae, Eusyllidae and Autolytidae. Journal of the Linnean Society of London, 24, 90 - 112." type="journal article" year="1920">Haswell 1920</bibRefCitation>
;
<bibRefCitation id="EFA620D1FFC69B523A7EFB19FBD2FAEC" author="Aguado, M. T. &amp; San Martin, G. &amp; ten Hove, H." box="[883,1103,1267,1293]" pageId="10" pageNumber="155" pagination="1 - 48" refId="ref6337" refString="Aguado, M. T., San Martin, G. &amp; ten Hove, H. (2008) Syllidae (Annelida: Polychaeta) from Indonesia collected by the Siboga (1899 - 1900) and Snellius II (1984) expeditions. Zootaxa, 1673 (1), 1 - 48. https: // doi. org / 10.11646 / zootaxa. 1673.1.1" type="journal article" year="2008">
Aguado
<emphasis id="B9438132FFC69B523ADEFB19FB99FAEC" box="[979,1028,1267,1293]" italics="true" pageId="10" pageNumber="155">et al</emphasis>
. 2008
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32D0EABFFC69B5239CAFAFDFF4FFAB4" pageId="10" pageNumber="155" type="etymology">
<paragraph id="8B885D20FFC69B5239CAFAFDFF4FFAB4" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="155">
<emphasis id="B9438132FFC69B5239CAFAFDFED5FAD0" bold="true" box="[199,328,1303,1329]" pageId="10" pageNumber="155">Etymology</emphasis>
. This species is named in honor of Dr. Gordon LJ Paterson, British polychaetologist currently retired.
</paragraph>
</subSubSection>
<subSubSection id="C32D0EABFFC69B5339CAFAB4FAD4FE84" lastPageId="11" lastPageNumber="156" pageId="10" pageNumber="155" type="discussion">
<paragraph id="8B885D20FFC69B5239CAFAB4FD94FA98" blockId="10.[151,1437,150,2013]" box="[199,521,1374,1401]" pageId="10" pageNumber="155">
<emphasis id="B9438132FFC69B5239CAFAB4FE63FA99" bold="true" box="[199,510,1374,1401]" pageId="10" pageNumber="155">Discussion and conclusion</emphasis>
s
</paragraph>
<paragraph id="8B885D20FFC69B5339CAFA69FAD4FE84" blockId="10.[151,1437,150,2013]" lastBlockId="11.[151,1437,151,357]" lastPageId="11" lastPageNumber="156" pageId="10" pageNumber="155">
It is well-known that some invertebrates and biological substrates are important habitats for polychaetes (
<bibRefCitation id="EFA620D1FFC69B523C78FA69FE77FA20" author="Alvarez-Campos, P. &amp; Verdes, A." pageId="10" pageNumber="155" pagination="520 - 531" refId="ref6406" refString="Alvarez-Campos, P. &amp; Verdes, A. (2017) Syllids inhabiting holdfast of Lessonia spicata in Central Chile: diversity, systematics, and description of three new species. Systematics and Biodiversity, 15 (6), 520 - 531. https: // doi. org / 10.1080 / 14772000.2017.1285364" type="journal article" year="2017">Alvarez-Campos &amp; Verdes 2017</bibRefCitation>
;
<bibRefCitation id="EFA620D1FFC69B5238FBFA4DFC94FA20" author="Martin, D. &amp; Britayev, T. A." box="[502,777,1447,1473]" pageId="10" pageNumber="155" pagination="217 - 340" refId="ref7980" refString="Martin, D. &amp; Britayev, T. A. (1998) Symbiotic polychaetes: review of known species. Oceanography and Marine Biology: An Annual Review, 36, 217 - 340." type="journal article" year="1998">Martin &amp; Britayev 1998</bibRefCitation>
).
<taxonomicName id="4C3726A3FFC69B523A11FA4DFCC4FA20" authorityName="Savigny in Lamarck" authorityYear="1818" box="[796,857,1447,1473]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="genus">
<emphasis id="B9438132FFC69B523A11FA4DFCC4FA20" box="[796,857,1447,1473]" italics="true" pageId="10" pageNumber="155">Syllis</emphasis>
</taxonomicName>
species presented on this work were collected from biological substrates such as
<taxonomicName id="4C3726A3FFC69B5238D0FA21FB9AFA04" authority="(Linnaeus) C. Agardh, 1820" authorityName="C. Agardh" authorityYear="1820" baseAuthorityName="Linnaeus" box="[477,1031,1483,1510]" class="Agaricomycetes" family="Marasmiaceae" genus="Macrocystis" higherTaxonomySource="CoL" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="155" phylum="Basidiomycota" rank="species" species="pyrifera">
<emphasis id="B9438132FFC69B5238D0FA21FD5BFA04" box="[477,710,1483,1509]" italics="true" pageId="10" pageNumber="155">Macrocystis pyrifera</emphasis>
(Linnaeus) C.
<bibRefCitation id="EFA620D1FFC69B523A60FA21FB9AFA04" author="Agardh, C. A." box="[877,1031,1483,1509]" pageId="10" pageNumber="155" refId="ref6282" refString="Agardh, C. A. (1820) Species algarum rite cognitae, cum synonymis, differentiis specificis et descriptionibus succinctis. Vol. 1. Part 1. Berling, Lund, [iv] + 168 pp. https: // doi. org / 10.5962 / bhl. title. 45326" type="book" year="1820">Agardh, 1820</bibRefCitation>
</taxonomicName>
kelp holdfasts and
<taxonomicName id="4C3726A3FFC69B523DECFA21FE45F9E8" authority="(Renier, 1804)" baseAuthorityName="Renier" baseAuthorityYear="1804" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="variopedatus">
<emphasis id="B9438132FFC69B523DECFA21FAE5FA04" box="[1249,1400,1483,1509]" italics="true" pageId="10" pageNumber="155">Chaetopterus</emphasis>
cf.
<emphasis id="B9438132FFC69B52399AFA05FEB4F9E8" box="[151,297,1519,1545]" italics="true" pageId="10" pageNumber="155">variopedatus</emphasis>
(
<bibRefCitation id="EFA620D1FFC69B523837FA05FE52F9E8" author="Renier, S. A." box="[314,463,1519,1545]" pageId="10" pageNumber="155" refId="ref8351" refString="Renier, S. A. (1804) Prospetto della Classe dei Vermi, nominati el ordinati secondo il Sistemo de Bosc. Unavailable publication, ICZN Opinion 316, Padua, pp. xv-xxviii." type="book" year="1804">Renier, 1804</bibRefCitation>
)
</taxonomicName>
tubes (
<taxonomicName id="4C3726A3FFC69B523B20FA05FD47F9EB" box="[557,730,1519,1546]" class="Polychaeta" family="Chaetopteridae" kingdom="Animalia" pageId="10" pageNumber="155" phylum="Annelida" rank="family">Chaetopteridae</taxonomicName>
) as well as from pebbles and sandy bottoms. In this research three new
<taxonomicName id="4C3726A3FFC69B523806F9F9FED5F9CC" authorityName="Savigny in Lamarck" authorityYear="1818" box="[267,328,1555,1581]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="genus">
<emphasis id="B9438132FFC69B523806F9F9FED5F9CC" box="[267,328,1555,1581]" italics="true" pageId="10" pageNumber="155">Syllis</emphasis>
</taxonomicName>
species were described; two of them were found associated to
<taxonomicName id="4C3726A3FFC69B523D0FF9F9FB4AF9CC" baseAuthorityName="Renier" baseAuthorityYear="1804" box="[1026,1239,1555,1581]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="10" pageNumber="155" phylum="Annelida" rank="species" species="variopedatus">
<emphasis id="B9438132FFC69B523D0FF9F9FB81F9CC" box="[1026,1052,1555,1581]" italics="true" pageId="10" pageNumber="155">C.</emphasis>
cf.
<emphasis id="B9438132FFC69B523D48F9F9FB4AF9CC" box="[1093,1239,1555,1581]" italics="true" pageId="10" pageNumber="155">variopedatus</emphasis>
</taxonomicName>
and all three species were found living on
<taxonomicName id="4C3726A3FFC69B5238CDF9DDFD37F9B0" authorityName="C. Agardh" authorityYear="1820" baseAuthorityName="Linnaeus" box="[448,682,1591,1617]" class="Agaricomycetes" family="Marasmiaceae" genus="Macrocystis" higherTaxonomySource="CoL" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="155" phylum="Basidiomycota" rank="species" species="pyrifera">
<emphasis id="B9438132FFC69B5238CDF9DDFD37F9B0" box="[448,682,1591,1617]" italics="true" pageId="10" pageNumber="155">Macrocystis pyrifera</emphasis>
</taxonomicName>
holdfasts.
<taxonomicName id="4C3726A3FFC69B523A2BF9DDFC20F9B0" box="[806,957,1591,1617]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="10" pageNumber="155" phylum="Annelida" rank="genus">
<emphasis id="B9438132FFC69B523A2BF9DDFC20F9B0" box="[806,957,1591,1617]" italics="true" pageId="10" pageNumber="155">Chaetopterus</emphasis>
</taxonomicName>
tubes are a new habitat for
<taxonomicName id="4C3726A3FFC69B523DF7F9DDFAAAF9B0" authorityName="Savigny in Lamarck" authorityYear="1818" box="[1274,1335,1591,1617]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="genus">
<emphasis id="B9438132FFC69B523DF7F9DDFAAAF9B0" box="[1274,1335,1591,1617]" italics="true" pageId="10" pageNumber="155">Syllis</emphasis>
</taxonomicName>
possibly as an adaptation to the lack of other suitable habitats due to the presence of extreme environmental conditions and the influence of ice-fields present in the fjords and channels of the Chilean Patagonia.
<taxonomicName id="4C3726A3FFC69B523D8FF995FA84F978" box="[1154,1305,1663,1689]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="10" pageNumber="155" phylum="Annelida" rank="genus">
<emphasis id="B9438132FFC69B523D8FF995FA84F978" box="[1154,1305,1663,1689]" italics="true" pageId="10" pageNumber="155">Chaetopterus</emphasis>
</taxonomicName>
tubes have demonstrated to be a suitable habitat for several syllid species previously recorded at Chilean Patagonia by
<bibRefCitation id="EFA620D1FFC69B523C41F949FEEEF903" author="Soto, E. H. &amp; San Martin, G." pageId="10" pageNumber="155" pagination="1599 - 1607" refId="ref9131" refString="Soto, E. H. &amp; San Martin, G. (2017 a) New reports and a new species of Syllidae (Annelida) from Chilean Patagonia. Journal of the Marine Biological Association of the United Kingdom, 98 (7), 1599 - 1607. https: // doi. org / 10.1017 / S 0025315417001242" type="journal article" year="2017">Soto &amp; San Martin (2017 a</bibRefCitation>
,b). These authors have already found twelve species of syllids hence with the current work the total number of syllid species increases to 15. High abundance and diversity of syllids in association with biological substrates have been previously reported (
<bibRefCitation id="EFA620D1FFC69B523B6AF8E5FCFDF8C8" author="Hernandez, C. E. &amp; Munoz, G. &amp; Rozbaczylo, N." box="[615,864,1807,1833]" pageId="10" pageNumber="155" pagination="99 - 108" refId="ref7587" refString="Hernandez, C. E., Munoz, G. &amp; Rozbaczylo, N. (2001) Poliquetos asociados con Austromegabalanus psittacus (Molina, 1782) (Crustacea: Cirripedia) en Peninsula Gualpen, Chile central: Biodiversidad y efecto del tamano del sustrato biologico. Revista de Biologia Marina y Oceanografia, 36, 99 - 108. https: // doi. org / 10.4067 / S 0718 - 19572001000100009" type="journal article" year="2001">
Hernández
<emphasis id="B9438132FFC69B523BE5F8E5FC85F8C8" box="[744,792,1807,1833]" italics="true" pageId="10" pageNumber="155">et al</emphasis>
. 2001
</bibRefCitation>
;
<bibRefCitation id="EFA620D1FFC69B523A67F8FAFB4BF8C8" author="Alvarez-Campos, P. &amp; Verdes, A." box="[874,1238,1807,1835]" pageId="10" pageNumber="155" pagination="520 - 531" refId="ref6406" refString="Alvarez-Campos, P. &amp; Verdes, A. (2017) Syllids inhabiting holdfast of Lessonia spicata in Central Chile: diversity, systematics, and description of three new species. Systematics and Biodiversity, 15 (6), 520 - 531. https: // doi. org / 10.1080 / 14772000.2017.1285364" type="journal article" year="2017">Álvarez-Campos &amp; Verdes 2017</bibRefCitation>
;
<bibRefCitation id="EFA620D1FFC69B523DEDF8E5FE9EF8AC" author="Soto, E. H. &amp; San Martin, G." pageId="10" pageNumber="155" pagination="1599 - 1607" refId="ref9131" refString="Soto, E. H. &amp; San Martin, G. (2017 a) New reports and a new species of Syllidae (Annelida) from Chilean Patagonia. Journal of the Marine Biological Association of the United Kingdom, 98 (7), 1599 - 1607. https: // doi. org / 10.1017 / S 0025315417001242" type="journal article" year="2017">Soto &amp; San Martin 2017a</bibRefCitation>
, b). Chaetopterid tubes clearly favour symbiotic associations offering a good habitat for diverse species, due to well-protected shelter and continuous water flow, likely bringing easily accessible food particles (
<bibRefCitation id="EFA620D1FFC69B523C37F8BDFEC5F874" author="Britayev, T. A. &amp; Martin, D." pageId="10" pageNumber="155" pagination="1833" refId="ref6783" refString="Britayev, T. A. &amp; Martin, D. (2019) Chapter 5.3. Chaetopteridae Audouin &amp; Milne Edwards, 1833. In: Purschke, G., Boggemann, M. &amp; Westheide, W. (Eds.), Annelida basal groups and Pleistoannelida, Sedentaria. Vol. 1. De Gruyter, Berlin and Boston, pp. 156 - 174." type="book chapter" year="2019">Britayev and Martin 2019</bibRefCitation>
). The great majority of tubes were occupied by one
<taxonomicName id="4C3726A3FFC69B523AABF891FBA0F874" box="[934,1085,1915,1941]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="10" pageNumber="155" phylum="Annelida" rank="genus">
<emphasis id="B9438132FFC69B523AABF891FBA0F874" box="[934,1085,1915,1941]" italics="true" pageId="10" pageNumber="155">Chaetopterus</emphasis>
</taxonomicName>
species although
<taxonomicName id="4C3726A3FFC69B523C0AF891FAD9F874" authorityName="Savigny in Lamarck" authorityYear="1818" box="[1287,1348,1915,1941]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="155" phylum="Annelida" rank="genus">
<emphasis id="B9438132FFC69B523C0AF891FAD9F874" box="[1287,1348,1915,1941]" italics="true" pageId="10" pageNumber="155">Syllis</emphasis>
</taxonomicName>
species were also recorded inside empty tubes and even between the layers that form these tubes. Similarly,
<taxonomicName id="4C3726A3FFC69B523C16F875FA01F858" authorityName="C. Agardh" authorityYear="1820" baseAuthorityName="Linnaeus" box="[1307,1436,1951,1977]" class="Agaricomycetes" family="Marasmiaceae" genus="Macrocystis" higherTaxonomySource="CoL" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="155" phylum="Basidiomycota" rank="species" species="pyrifera">
<emphasis id="B9438132FFC69B523C16F875FA01F858" box="[1307,1436,1951,1977]" italics="true" pageId="10" pageNumber="155">M. pyrifera</emphasis>
</taxonomicName>
holdfasts also allow that several syllid species may inhabit them. Nine species have been recorded by
<bibRefCitation id="EFA620D1FFC69B533C04F829FEA6FF50" author="Soto, E. H. &amp; San Martin, G." lastPageId="11" lastPageNumber="156" pageId="10" pageNumber="155" pagination="1599 - 1607" refId="ref9131" refString="Soto, E. H. &amp; San Martin, G. (2017 a) New reports and a new species of Syllidae (Annelida) from Chilean Patagonia. Journal of the Marine Biological Association of the United Kingdom, 98 (7), 1599 - 1607. https: // doi. org / 10.1017 / S 0025315417001242" type="journal article" year="2017">Soto and San Martin (2017a</bibRefCitation>
, b) for Chilean Patagonia and with this work the total number of syllids increase to 12.
<taxonomicName id="4C3726A3FFC79B533C19FF7DFACCFF50" authorityName="Savigny in Lamarck" authorityYear="1818" box="[1300,1361,151,177]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="156" phylum="Annelida" rank="genus">
<emphasis id="B9438132FFC79B533C19FF7DFACCFF50" box="[1300,1361,151,177]" italics="true" pageId="11" pageNumber="156">Syllis</emphasis>
</taxonomicName>
seems to be a common inhabitant on this
<typeStatus id="548CE382FFC79B533B2EFF51FDCEFF34" box="[547,595,187,213]" pageId="11" pageNumber="156">type</typeStatus>
of habitat because
<bibRefCitation id="EFA620D1FFC79B533A23FF51FB5EFF34" author="Alvarez-Campos, P. &amp; Verdes, A." box="[814,1219,187,213]" pageId="11" pageNumber="156" pagination="520 - 531" refId="ref6406" refString="Alvarez-Campos, P. &amp; Verdes, A. (2017) Syllids inhabiting holdfast of Lessonia spicata in Central Chile: diversity, systematics, and description of three new species. Systematics and Biodiversity, 15 (6), 520 - 531. https: // doi. org / 10.1080 / 14772000.2017.1285364" type="journal article" year="2017">Alvarez-Campos and Verdes (2017)</bibRefCitation>
recorded nine species of this genus and total number of twelve species of
<taxonomicName id="4C3726A3FFC79B533A1EFF35FCF2FF18" authorityName="Grube" authorityYear="1850" box="[787,879,223,249]" class="Polychaeta" family="Syllidae" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="156" phylum="Annelida" rank="family">Syllidae</taxonomicName>
inhabiting holdfasts of the brown algae
<taxonomicName id="4C3726A3FFC79B533C34FF35FF75FEFD" authorityName="Santelices" authorityYear="2012" class="Aves" family="Tyrannidae" genus="Lessonia" kingdom="Animalia" order="Passeriformes" pageId="11" pageNumber="156" phylum="Chordata" rank="species" species="spicata">
<emphasis id="B9438132FFC79B533C34FF35FF75FEFD" italics="true" pageId="11" pageNumber="156">Lessonia spicata</emphasis>
</taxonomicName>
in Central
<collectingCountry id="F3201DB0FFC79B533868FEE9FE38FEFC" box="[357,421,259,285]" name="Chile" pageId="11" pageNumber="156">Chile</collectingCountry>
. These authors highlighted the importance of kelp holdfasts as hotspots for syllid diversity, understanding that kelp-like species play an important ecological role providing food, mechanical shelter against wave impact, refuge from predators, secure spawning areas and nursery grounds (
<bibRefCitation id="EFA620D1FFC79B533D27FEA1FAA0FE84" author="Pabis, K. &amp; Sicinski, J." box="[1066,1341,331,357]" pageId="11" pageNumber="156" pagination="1277 - 1288" refId="ref8291" refString="Pabis, K. &amp; Sicinski, J. (2010) Polychaete fauna associated with holdfasts of the large brown alga Himantothallus grandifolius in Admiralty Bay, King George Island, Antarctic. Polar Biology, 33, 1277 - 1288. https: // doi. org / 10.1007 / s 00300 - 010 - 0816 - x" type="journal article" year="2010">Pabis and Sicinski 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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