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<document id="3DADF837822B2FF8315D3319D7D43616" ID-CLB-Dataset="21162" ID-DOI="10.1080/00222933.2015.1009514" ID-GBIF-Dataset="8c6c97a7-1c63-41ba-b75a-217ce1a3411d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3997986" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598292250840" checkinUser="carolina" docAuthor="Rota, Emilia" docDate="2015" docId="039AAD67FFE14504FE79FC9AFEE60BA9" docLanguage="en" docName="JNATHIST.49.33-34.1987-2020.pdf" docOrigin="Journal of Natural History 49 (33)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Achaeta giustii Rota 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="1999" masterDocId="FFA3D51FFFE84508FF92FFAFFFB6091F" masterDocTitle="Five new species of Enchytraeidae (Annelida: Clitellata) from Mediterranean woodlands of Italy and reaffirmed validity of Achaeta etrusca, Fridericia bulbosa and F. miraflores" masterLastPageNumber="2020" masterPageNumber="1987" pageNumber="1996" updateTime="1698845298462" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="51962206AD49037910A84A8A0804BC9E">Five new species of Enchytraeidae (Annelida: Clitellata) from Mediterranean woodlands of Italy and reaffirmed validity of Achaeta etrusca, Fridericia bulbosa and F. miraflores</mods:title>
</mods:titleInfo>
<mods:name id="7014A0BF2DA7FBDB7A79FCB153F38FC2" type="personal">
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<mods:namePart id="2A946525266D379A717C2159B7D2EAEC">Rota, Emilia</mods:namePart>
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<mods:title id="AD6CFB788F74FA4A63396F36B963033B">Journal of Natural History</mods:title>
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<mods:part id="64A7548302313C110BC0AA5A7A4819D2">
<mods:date id="E062B75C86411448F6D8D9CB6AECE16A">2015</mods:date>
<mods:detail id="A8002805B94DA3F76F63EBCA088CC51A" type="pubDate">
<mods:number id="4CFC484A712440A0100BACAFC5239DC3">2015-02-28</mods:number>
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<mods:detail id="1D1F310EEE9C8BEFF779648364514FBA" type="volume">
<mods:number id="6B4B0EED8ED288853773C189E2AFCF2E">49</mods:number>
</mods:detail>
<mods:detail id="8063F610904FBA5ADF1EBDB0FCEAB11D" type="issue">
<mods:number id="2D155723DB06E1961BBD46FE1901B2B3">33</mods:number>
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<mods:start id="7FC18659A85CA09E6221427B9F504191">1987</mods:start>
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<mods:classification id="6681D007AF855916DE5E56622C273C5E">journal article</mods:classification>
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<mods:identifier id="21A0414F90D1E26FF725AD9B568C7420" type="DOI">10.1080/00222933.2015.1009514</mods:identifier>
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<mods:identifier id="181334BF05057C143BF22D88C7B5DA76" type="ISSN">1464-5262</mods:identifier>
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<treatment id="039AAD67FFE14504FE79FC9AFEE60BA9" ID-DOI="http://doi.org/10.5281/zenodo.4330413" ID-GBIF-Taxon="167293503" ID-Zenodo-Dep="4330413" LSID="urn:lsid:zoobank.org:act:AC645870-CFF4-48E7-B934-5A741629DCA8" httpUri="http://treatment.plazi.org/id/039AAD67FFE14504FE79FC9AFEE60BA9" lastPageId="12" lastPageNumber="1999" pageId="9" pageNumber="1996">
<subSubSection id="C3294FFAFFE14501FE79FC9AFD420A51" box="[491,756,821,846]" pageId="9" pageNumber="1996" type="nomenclature">
<paragraph id="8B8C1C71FFE14501FE79FC9AFD420A51" blockId="9.[491,756,821,878]" box="[491,756,821,846]" pageId="9" pageNumber="1996">
<heading id="D0C4AB1DFFE14501FE79FC9AFD420A51" box="[491,756,821,846]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="1996" reason="2">
<taxonomicName id="4C3367F2FFE14501FE79FC9AFD240A52" authority="Rota, 2015" authorityName="Rota" authorityYear="2015" box="[491,658,821,845]" class="Magnoliopsida" family="Poaceae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="9" pageNumber="1996" phylum="Tracheophyta" rank="species" species="giustii" status="sp. nov.">
<emphasis id="B947C063FFE14501FE79FC9AFD240A52" bold="true" box="[491,658,821,845]" italics="true" pageId="9" pageNumber="1996">Achaeta giustii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2747D18FFE14501FD08FC98FD420A51" box="[666,756,823,846]" pageId="9" pageNumber="1996" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFE14501FDA5FCF9FD1F0A71" box="[567,681,854,878]" pageId="9" pageNumber="1996" type="description">
<paragraph id="8B8C1C71FFE14501FDA5FCF9FD1F0A71" blockId="9.[491,756,821,878]" box="[567,681,854,878]" pageId="9" pageNumber="1996">
(
<figureCitation id="130800F4FFE14501FDADFCF9FD290A71" box="[575,671,854,878]" captionStart="Figure 3" captionStartId="11.[144,211,1354,1376]" captionTargetBox="[144,1104,177,1317]" captionTargetId="figure@11.[144,1104,168,1323]" captionTargetPageId="11" captionText="Figure 3. Achaeta giustii sp. nov. (A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="9" pageNumber="1996">Figure 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFE14501FF02FC39FC390D12" pageId="9" pageNumber="1996" type="reference_group">
<paragraph id="8B8C1C71FFE14501FF02FC39FED50AD2" blockId="9.[144,1103,917,1037]" pageId="9" pageNumber="1996">
<taxonomicName id="4C3367F2FFE14501FF05FC39FC010AB2" authority="(Vejdovsky, 1879 a) sensu Nielsen and Christ. 1959" authorityName="sensu Nielsen and Christensen" authorityYear="1959" baseAuthorityName="Vejdovsky" baseAuthorityYear="1879" box="[151,951,917,942]" class="Magnoliopsida" family="Poaceae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="9" pageNumber="1996" phylum="Tracheophyta" rank="species" species="bohemica">
<emphasis id="B947C063FFE14501FF05FC39FED20AB1" box="[151,356,918,942]" italics="true" pageId="9" pageNumber="1996">Achaeta bohemica</emphasis>
(
<bibRefCitation id="EFA26180FFE14501FEE7FC3AFDF60AB2" author="Vejdovsky F" box="[373,576,917,941]" pageId="9" pageNumber="1996" pagination="183 - 185" refId="ref17273" refString="Vejdovsky F. 1879 a. Vorlaufige Mittheilungen uber die fortgesetzten Oligochaetenstudien. Zool Anz. 2: 183 - 185." type="journal article" year="1879">Vejdovský, 1879a</bibRefCitation>
) sensu Nielsen and Christ. 1959
</taxonomicName>
,
<bibRefCitation id="EFA26180FFE14501FC42FC39FBFE0AB2" author="Rota E" box="[976,1096,917,941]" pageId="9" pageNumber="1996" pagination="183 - 231" refId="ref16775" refString="Rota E. 1995. Italian Enchytraeidae (Oligochaeta). I. Boll Zool. 62: 183 - 231." type="journal article" year="1995">Rota 1995</bibRefCitation>
, p. 196 (partim).
</paragraph>
<paragraph id="8B8C1C71FFE14501FF02FC7AFC390D12" blockId="9.[144,1103,917,1037]" pageId="9" pageNumber="1996">
<taxonomicName id="4C3367F2FFE14501FF05FC79FCBD0AF2" authority="sensu Nielsen and Christ. 1959" authorityName="sensu Nielsen and Christensen" authorityYear="1959" baseAuthorityName="Vejdovsky" baseAuthorityYear="1879" box="[151,779,981,1006]" class="Magnoliopsida" family="Poaceae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="9" pageNumber="1996" phylum="Tracheophyta" rank="species" species="bohemica">
<emphasis id="B947C063FFE14501FF05FC79FF440AF1" box="[151,242,982,1006]" italics="true" pageId="9" pageNumber="1996">Achaeta</emphasis>
cf.
<emphasis id="B947C063FFE14501FEB5FC79FE390AF1" box="[295,399,982,1006]" italics="true" pageId="9" pageNumber="1996">bohemica</emphasis>
sensu Nielsen and Christ. 1959
</taxonomicName>
,
<bibRefCitation id="EFA26180FFE14501FCB4FC79FC5C0AF2" author="Rota E &amp; Caruso T &amp; Monaci F &amp; Baldantoni D &amp; De Nicola &amp; Iovieno P &amp; Bargagli R" box="[806,1002,981,1005]" pageId="9" pageNumber="1996" pagination="268 - 276" refId="ref16835" refString="Rota E, Caruso T, Monaci F, Baldantoni D, De Nicola F, Iovieno P, Bargagli R. 2013. Effects of soil pollutants, biogeochemistry and microbiology on the distribution and composition of enchytraeid communities in urban and suburban holm oak stands. Environ Poll. 179: 268 - 276." type="journal article" year="2013">Rota et al. 2013</bibRefCitation>
, table 1 (species s2);
<bibRefCitation id="EFA26180FFE14501FEDEFC59FDB20D12" author="Rota E &amp; Caruso T &amp; Bargagli R" box="[332,516,1013,1037]" pageId="9" pageNumber="1996" pagination="83 - 91" refId="ref16797" refString="Rota E, Caruso T, Bargagli R. 2014. Community structure, diversity and spatial organization of enchytraeids in Mediterranean urban holm oak stands. Eur J Soil Biol. 62: 83 - 91." type="journal article" year="2014">Rota et al. 2014</bibRefCitation>
, tables 1, 2 and Suppl. 1 (partim).
</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFE14501FF02FBF5FD8A0F84" pageId="9" pageNumber="1996" type="materials_examined">
<paragraph id="8B8C1C71FFE14501FF02FBF5FE9A0D6E" blockId="9.[144,1018,1113,1177]" box="[144,300,1113,1137]" pageId="9" pageNumber="1996">
<heading id="D0C4AB1DFFE14501FF02FBF5FE9A0D6E" box="[144,300,1113,1137]" fontSize="10" level="3" pageId="9" pageNumber="1996" reason="5">
<emphasis id="B947C063FFE14501FF02FBF5FE9A0D6E" box="[144,300,1113,1137]" italics="true" pageId="9" pageNumber="1996">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFE14501FF02FB2EFC4C0D87" blockId="9.[144,1018,1113,1177]" box="[144,1018,1153,1177]" pageId="9" pageNumber="1996">
<materialsCitation id="3B5B162CFFE14501FF02FB2EFC4C0D87" ID-GBIF-Occurrence="2848394324" box="[144,1018,1153,1177]" collectionCode="MCZR" pageId="9" pageNumber="1996" specimenCount="1" typeStatus="holotype">
<emphasis id="B947C063FFE14501FF02FB2EFF480D86" box="[144,254,1153,1177]" italics="true" pageId="9" pageNumber="1996">
<typeStatus id="5488A2D3FFE14501FF02FB2EFF4C0D86" box="[144,250,1153,1177]" pageId="9" pageNumber="1996" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED2284B4FFE14501FE94FB2EFEEC0D86" box="[262,346,1153,1177]" country="Italy" httpUri="http://grbio.org/cool/mmbj-pfva" name="Museo Civico di Zoologia" pageId="9" pageNumber="1996">MCZR</collectionCode>
<taxonomicName id="4C3367F2FFE14501FEF1FB2EFE5C0D86" box="[355,490,1153,1177]" class="Oligochaeta" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="1996" phylum="Magnoliopsida" rank="class">Oligochaeta</taxonomicName>
0176, whole-mounted specimen, fully mature.
</materialsCitation>
</paragraph>
<paragraph id="8B8C1C71FFE14501FF02FB49FE940DE2" blockId="9.[144,1103,1253,1412]" box="[144,290,1253,1277]" pageId="9" pageNumber="1996">
<heading id="D0C4AB1DFFE14501FF02FB49FE940DE2" box="[144,290,1253,1277]" fontSize="10" level="3" pageId="9" pageNumber="1996" reason="5">
<emphasis id="B947C063FFE14501FF02FB49FE940DE2" box="[144,290,1253,1277]" italics="true" pageId="9" pageNumber="1996">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFE14501FF02FAA3FE1A0C9B" blockId="9.[144,1103,1253,1412]" pageId="9" pageNumber="1996">
<collectingCountry id="F3245CE1FFE14501FF02FAA3FF710C3B" box="[144,199,1292,1316]" name="Italy" pageId="9" pageNumber="1996">Italy</collectingCountry>
,
<collectingRegion id="49F7D293FFE14501FF46FAA2FE850C3A" box="[212,307,1293,1317]" country="Italy" name="Toscana" pageId="9" pageNumber="1996">Tuscany</collectingRegion>
(Tu-1), La Verna (Arezzo), centuries-old
<taxonomicName id="4C3367F2FFE14501FC87FAA2FC3D0C3A" box="[789,907,1293,1317]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="1996" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="B947C063FFE14501FC87FAA2FC3D0C3A" box="[789,907,1293,1317]" italics="true" pageId="9" pageNumber="1996">Abies alba</emphasis>
</taxonomicName>
and beech forest with young maple,
<taxonomicName id="4C3367F2FFE14501FEFBFA83FE560C5B" box="[361,480,1324,1348]" class="Magnoliopsida" family="Ranunculaceae" genus="Helleborus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ranunculales" pageId="9" pageNumber="1996" phylum="Tracheophyta" rank="genus">
<emphasis id="B947C063FFE14501FEFBFA83FE560C5B" box="[361,480,1324,1348]" italics="true" pageId="9" pageNumber="1996">Helleborus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3367F2FFE14501FD88FA83FDE00C5B" box="[538,598,1324,1348]" class="Magnoliopsida" family="Violaceae" genus="Viola" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="9" pageNumber="1996" phylum="Tracheophyta" rank="genus">
<emphasis id="B947C063FFE14501FD88FA83FDE00C5B" box="[538,598,1324,1348]" italics="true" pageId="9" pageNumber="1996">Viola</emphasis>
</taxonomicName>
on bryozoal limestone (
<geoCoordinate id="EE077AB6FFE14501FCF5FA83FC550C5B" box="[871,995,1324,1348]" degrees="43.7065" direction="north" orientation="latitude" pageId="9" pageNumber="1996" precision="5" value="43.7065">43.7065°N</geoCoordinate>
,
<geoCoordinate id="EE077AB6FFE14501FC7CFA83FF120C7B" degrees="11.9319" direction="east" orientation="longitude" pageId="9" pageNumber="1996" precision="5" value="11.9319">11.9319° E</geoCoordinate>
,
<quantity id="4CCBB194FFE14501FF22FAE3FEB40C7B" box="[176,258,1356,1380]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="9" pageNumber="1996" unit="m" value="1120.0">1120 m</quantity>
asl). Brown sandy humus and wood litter under rotten logs, moist, pH 5.9,
<date id="FF8D3AB1FFE14501FF02FAC3FEBA0C9B" box="[144,268,1388,1412]" pageId="9" pageNumber="1996" value="1996-02-05">02.05.1996</date>
,
<collectionCode id="ED2284B4FFE14501FE8AFAC2FE9B0C9B" box="[280,301,1389,1412]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="9" pageNumber="1996" type="Herbarium">E</collectionCode>
. Rota coll.
</paragraph>
<paragraph id="8B8C1C71FFE14501FF02FA16FD760F10" blockId="9.[144,1104,1464,1551]" pageId="9" pageNumber="1996">
<materialsCitation id="3B5B162CFFE14501FF02FA16FDA00CF0" ID-GBIF-Occurrence="2848394316" collectingDate="1994-05-22" collectionCode="MCZR" country="Italy" location="Latium" pageId="9" pageNumber="1996" specimenCount="1" stateProvince="Lazio" typeStatus="paratype">
<emphasis id="B947C063FFE14501FF02FA16FEB00CCF" box="[144,262,1465,1488]" italics="true" pageId="9" pageNumber="1996">
<typeStatus id="5488A2D3FFE14501FF02FA16FEB50CCF" box="[144,259,1465,1488]" pageId="9" pageNumber="1996" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED2284B4FFE14501FE84FA17FEDF0CCF" box="[278,361,1464,1488]" country="Italy" httpUri="http://grbio.org/cool/mmbj-pfva" name="Museo Civico di Zoologia" pageId="9" pageNumber="1996">MCZR</collectionCode>
<taxonomicName id="4C3367F2FFE14501FEEBFA17FDB60CCF" box="[377,512,1464,1488]" class="Oligochaeta" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="1996" phylum="Magnoliopsida" rank="class">Oligochaeta</taxonomicName>
01770178, two whole-mounted specimens from
<collectingCountry id="F3245CE1FFE14501FF02FA78FF710CF0" box="[144,199,1495,1519]" name="Italy" pageId="9" pageNumber="1996">Italy</collectingCountry>
,
<collectingRegion id="49F7D293FFE14501FF48FA78FE9B0CF0" box="[218,301,1495,1519]" country="Italy" name="Lazio" pageId="9" pageNumber="1996">Latium</collectingRegion>
(La-1),
<date id="FF8D3AB1FFE14501FE08FA78FDA00CF0" box="[410,534,1495,1519]" pageId="9" pageNumber="1996" value="1994-05-22">
<collectingDate id="EFC9C359FFE14501FE08FA78FDA00CF0" box="[410,534,1495,1519]" pageId="9" pageNumber="1996" value="1994-05-22">22.05.1994</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B5B162CFFE14501FDBBFA78FD0A0F10" ID-GBIF-Occurrence="2848394306" collectingDate="1992-03-25" collectionCode="MCZR" country="Italy" location="Latium" pageId="9" pageNumber="1996" specimenCount="1" stateProvince="Lazio" typeStatus="paratype">
SMNH-Types 87218722, two whole-mounted specimens from
<collectingCountry id="F3245CE1FFE14501FEDBFA58FE360F10" box="[329,384,1527,1551]" name="Italy" pageId="9" pageNumber="1996">Italy</collectingCountry>
,
<collectingRegion id="49F7D293FFE14501FE1FFA58FE560F10" box="[397,480,1527,1551]" country="Italy" name="Lazio" pageId="9" pageNumber="1996">Latium</collectingRegion>
(La-2),
<date id="FF8D3AB1FFE14501FDADFA58FD0A0F10" box="[575,700,1527,1551]" pageId="9" pageNumber="1996" value="1992-03-25">
<collectingDate id="EFC9C359FFE14501FDADFA58FD0A0F10" box="[575,700,1527,1551]" pageId="9" pageNumber="1996" value="1992-03-25">25.03.1992</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8C1C71FFE14501FF02F9EBFD8A0F84" blockId="9.[144,1104,1603,1691]" pageId="9" pageNumber="1996">
<emphasis id="B947C063FFE14501FF02F9EBFEFC0F43" box="[144,330,1604,1628]" italics="true" pageId="9" pageNumber="1996">Other material.</emphasis>
<materialsCitation id="3B5B162CFFE14501FEF3F9ECFD810F84" ID-GBIF-Occurrence="2848394326" collectingDate="1993-10-29" collectingDateMax="2009-05-14" collectingDateMin="1993-10-29" country="Italy" location="One" pageId="9" pageNumber="1996" specimenCount="1" stateProvince="Toscana">
<location id="8EEC4AAAFFE14501FEF3F9ECFE390F44" LSID="urn:lsid:plazi:treatment:039AAD67FFE14504FE79FC9AFEE60BA9:8EEC4AAAFFE14501FEF3F9ECFE390F44" box="[353,399,1603,1627]" country="Italy" name="One" pageId="9" pageNumber="1996" stateProvince="Toscana">One</location>
whole-mounted specimen from
<collectingCountry id="F3245CE1FFE14501FCABF9ECFCC60F44" box="[825,880,1603,1627]" name="Italy" pageId="9" pageNumber="1996">Italy</collectingCountry>
,
<collectingRegion id="49F7D293FFE14501FC19F9EBFC5C0F43" box="[907,1002,1604,1628]" country="Italy" name="Toscana" pageId="9" pageNumber="1996">Tuscany</collectingRegion>
(Tu-2),
<date id="FF8D3AB1FFE14501FF02F9CCFEBA0F64" box="[144,268,1635,1659]" pageId="9" pageNumber="1996" value="1993-10-29">
<collectingDate id="EFC9C359FFE14501FF02F9CCFEBA0F64" box="[144,268,1635,1659]" pageId="9" pageNumber="1996" value="1993-10-29">29.10.1993</collectingDate>
</date>
. One alcohol-preserved specimen from
<collectingCountry id="F3245CE1FFE14501FC8CF9CCFCE30F64" box="[798,853,1635,1659]" name="Italy" pageId="9" pageNumber="1996">Italy</collectingCountry>
,
<collectingRegion id="49F7D293FFE14501FCE1F9CCFC5E0F64" box="[883,1000,1635,1659]" country="Italy" name="Campania" pageId="9" pageNumber="1996">Campania</collectingRegion>
(Ca-1),
<date id="FF8D3AB1FFE14501FF02F92CFEBD0F84" box="[144,267,1667,1691]" pageId="9" pageNumber="1996" value="2009-05-14">
<collectingDate id="EFC9C359FFE14501FF02F92CFEBD0F84" box="[144,267,1667,1691]" pageId="9" pageNumber="1996" value="2009-05-14">14.05.2009</collectingDate>
</date>
, in the author s collection
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFE24502FF46FF07FEFB0837" pageId="10" pageNumber="1997" type="etymology">
<paragraph id="8B8C1C71FFE24502FF46FF07FEFB09DF" blockId="10.[212,1172,168,296]" box="[212,333,168,192]" pageId="10" pageNumber="1997">
<heading id="D0C4AB1DFFE24502FF46FF07FEFB09DF" box="[212,333,168,192]" fontSize="10" level="3" pageId="10" pageNumber="1997" reason="5">
<emphasis id="B947C063FFE24502FF46FF07FEFB09DF" box="[212,333,168,192]" italics="true" pageId="10" pageNumber="1997">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFE24502FF46FF7FFEFB0837" blockId="10.[212,1172,168,296]" pageId="10" pageNumber="1997">The new species is named for Prof. Folco Giusti, outstanding malacologist and dedicated zoogeographer, for his contributions to the knowledge of Mediterranean endemism.</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFE24502FF46FED5FD100B27" pageId="10" pageNumber="1997" type="diagnosis">
<paragraph id="8B8C1C71FFE24502FF46FED5FEF6088E" blockId="10.[212,1172,378,568]" box="[212,320,378,401]" pageId="10" pageNumber="1997">
<heading id="D0C4AB1DFFE24502FF46FED5FEF6088E" box="[212,320,378,401]" fontSize="10" level="3" pageId="10" pageNumber="1997" reason="5">
<emphasis id="B947C063FFE24502FF46FED5FEF6088E" box="[212,320,378,401]" italics="true" pageId="10" pageNumber="1997">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFE24502FF46FE0EFD100B27" blockId="10.[212,1172,378,568]" pageId="10" pageNumber="1997">Medium-sized species, with large flask-shaped glands occurring dorsally from II to tail, including XII; clitellum over XII1/2XIII, absent middorsally, hyaline and granular gland cells forming dorsolaterally a reticulate pattern; male pores in XII, penial bulbs compact, preclitellar nephridia two pairs at 6/77/8, spermathecae opening ventrally, ampullae reaching VIIIX.</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFE24504FF46FD2FFB2209FF" lastPageId="12" lastPageNumber="1999" pageId="10" pageNumber="1997" type="description">
<paragraph id="8B8C1C71FFE24502FF46FD2FFEE50B88" blockId="10.[212,1172,640,1691]" box="[212,339,640,663]" pageId="10" pageNumber="1997">
<heading id="D0C4AB1DFFE24502FF46FD2FFEE50B88" box="[212,339,640,663]" fontSize="10" level="3" pageId="10" pageNumber="1997" reason="5">
<emphasis id="B947C063FFE24502FF46FD2FFEE50B88" box="[212,339,640,663]" italics="true" pageId="10" pageNumber="1997">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFE24502FF46FD08FC610AC1" blockId="10.[212,1172,640,1691]" pageId="10" pageNumber="1997">
Live body length
<quantity id="4CCBB194FFE24502FE26FD08FDAD0BA1" box="[436,539,679,703]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="10" pageNumber="1997" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
, width
<quantity id="4CCBB194FFE24502FDEDFD08FC900BA1" box="[639,806,679,703]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="3.5999999999999996" metricValueMin="2.8" pageId="10" pageNumber="1997" unit="mm" value="0.32" valueMax="0.36" valueMin="0.28">0.280.36 mm</quantity>
at XII; after fixation, length
<quantity id="4CCBB194FFE24502FF46FD68FEED0BC1" box="[212,347,711,735]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.8" metricValueMin="4.2" pageId="10" pageNumber="1997" unit="mm" value="5.0" valueMax="5.8" valueMin="4.2">4.25.8 mm</quantity>
, width
<quantity id="4CCBB194FFE24502FE3DFD68FDF80BC1" box="[431,590,711,735]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="3.0" metricValueMin="2.3" pageId="10" pageNumber="1997" unit="mm" value="0.265" valueMax="0.3" valueMin="0.23">0.230.30 mm</quantity>
at XII. Segments 2837. Large flask-shaped glands dorsally paired from II to tail, including XII (
<figureCitation id="130800F4FFE24502FD4EFD49FCC70BE1" box="[732,881,742,766]" captionStart="Figure 3" captionStartId="11.[144,211,1354,1376]" captionTargetBox="[144,1104,177,1317]" captionTargetId="figure@11.[144,1104,168,1323]" captionTargetPageId="11" captionText="Figure 3. Achaeta giustii sp. nov. (A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="10" pageNumber="1997">Figure 3A, F</figureCitation>
), absent ventrally;
<emphasis id="B947C063FFE24502FBD9FD47FB220BE0" box="[1099,1172,744,767]" italics="true" pageId="10" pageNumber="1997">in vivo</emphasis>
measured length throughout up to 120 μm, fixed 85100 μm, glands smaller (5075 μm) in II; dorsal distance between left and right glands 100120 μm (fixed). Knob-like glands and lentiform glands absent. Clitellum in XII1/2 XIII, dorsally interrupted (gap 90 μm wide), elsewhere continuous; dorsolateral sides made of polygonal granular and not much larger hyaline cells forming a reticulate pattern; dorsal edge consisting of granular cells (
<figureCitation id="130800F4FFE24502FD24FC09FC980AA1" box="[694,814,934,958]" captionStart="Figure 3" captionStartId="11.[144,211,1354,1376]" captionTargetBox="[144,1104,177,1317]" captionTargetId="figure@11.[144,1104,168,1323]" captionTargetPageId="11" captionText="Figure 3. Achaeta giustii sp. nov. (A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="10" pageNumber="1997">Figure 3C</figureCitation>
); only granular cells ventrally (
<figureCitation id="130800F4FFE24502FF4EFC69FEC60AC2" box="[220,368,966,990]" captionStart="Figure 3" captionStartId="11.[144,211,1354,1376]" captionTargetBox="[144,1104,177,1317]" captionTargetId="figure@11.[144,1104,168,1323]" captionTargetPageId="11" captionText="Figure 3. Achaeta giustii sp. nov. (A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="10" pageNumber="1997">Figure 3E, F</figureCitation>
); thickness of clitellum at midpoint 1632 μm (fixed).
</paragraph>
<paragraph id="8B8C1C71FFE24502FF6CFC49FE020D22" blockId="10.[212,1172,640,1691]" pageId="10" pageNumber="1997">Head pore on prostomium. Spermathecal pores ventral at 4/5, 6570 μm (fix) distant from one another. Male pores in XII, 5070 μm apart, somewhat closer than spermathecal pores.</paragraph>
<paragraph id="8B8C1C71FFE24502FF6CFBEAFC110DE2" blockId="10.[212,1172,640,1691]" pageId="10" pageNumber="1997">
Cuticle at least 2.5 μm thick, in places reaching 6.0 μm, larger dorsally than ventrally. Brain about 150 μm long in vivo, 125135 μm when fixed. Oesophageal outer ridge dorsal on IIIV, inconspicuous. Pharyngeal glands three primary pairs at 4/56/7, each merging dorsally, no secondary lobes (
<figureCitation id="130800F4FFE24502FCA5FB0AFC1D0DA2" box="[823,939,1189,1213]" captionStart="Figure 3" captionStartId="11.[144,211,1354,1376]" captionTargetBox="[144,1104,177,1317]" captionTargetId="figure@11.[144,1104,168,1323]" captionTargetPageId="11" captionText="Figure 3. Achaeta giustii sp. nov. (A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="10" pageNumber="1997">Figure 3B</figureCitation>
). Two pairs of preclitellar nephridia at 6/77/8 constricted by septum, without swollen terminal vesicle; nephridia generally absent from first five postclitellar segments.
</paragraph>
<paragraph id="8B8C1C71FFE24502FF6CFAABFE380CA3" blockId="10.[212,1172,640,1691]" pageId="10" pageNumber="1997">
Coelomocytes of various size, with grooved but not granular cytoplasm, roundish, often with one to five marginal prominences, brownish when accumulated. Gut linear, without loops. Oesophagus gradually expanding into intestine at 7/8. Intestinal inner ridge extending over three segments between XVIIIXXIII (ventral intestinal ridge). Chloragogenous cells filled with fine granules. Dorsal blood vessel arising in VII (
<figureCitation id="130800F4FFE24502FE9EFA0BFEC90CA3" box="[268,383,1444,1468]" captionStart="Figure 3" captionStartId="11.[144,211,1354,1376]" captionTargetBox="[144,1104,177,1317]" captionTargetId="figure@11.[144,1104,168,1323]" captionTargetPageId="11" captionText="Figure 3. Achaeta giustii sp. nov. (A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="10" pageNumber="1997">Figure 3B</figureCitation>
).
</paragraph>
<paragraph id="8B8C1C71FFE24504FF6CFA6BFB2209FF" blockId="10.[212,1172,640,1691]" lastBlockId="12.[212,1172,168,224]" lastPageId="12" lastPageNumber="1999" pageId="10" pageNumber="1997">
Seminal vesicle absent or small. Sperm funnels elongate equal to or shorter than body width, broadest point slightly below collar, length:width 34:1 (180230 by 5080 μm
<emphasis id="B947C063FFE24502FBEDFA4AFF490F04" italics="true" pageId="10" pageNumber="1997">in vivo</emphasis>
), generally bent at midlength; collar distinct, as wide or narrower than funnel (
<figureCitation id="130800F4FFE24502FF4EF98CFEE40F24" box="[220,338,1571,1595]" captionStart="Figure 3" captionStartId="11.[144,211,1354,1376]" captionTargetBox="[144,1104,177,1317]" captionTargetId="figure@11.[144,1104,168,1323]" captionTargetPageId="11" captionText="Figure 3. Achaeta giustii sp. nov. (A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="10" pageNumber="1997">Figure 3H</figureCitation>
). Spermatozoa 50 μm long, heads 20 μm long. Vasa deferentia long and narrow (810 μm), tightly coiled. Penial bulbs small, compact, enclosed in a muscular sheath, 5364 μm long in fixed worms (
<figureCitation id="130800F4FFE24502FD03F9CCFC950F64" box="[657,803,1635,1659]" captionStart="Figure 3" captionStartId="11.[144,211,1354,1376]" captionTargetBox="[144,1104,177,1317]" captionTargetId="figure@11.[144,1104,168,1323]" captionTargetPageId="11" captionText="Figure 3. Achaeta giustii sp. nov. (A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="10" pageNumber="1997">Figure 3E, F</figureCitation>
). One egg mature. Spermathecal ampullae reaching to VIIIX, ectal ducts almost parallel to long body axis in vivo, more contracted on one side (thus bent at an obtuse angle) in fixed specimens (
<figureCitation id="130800F4FFE44504FC4EFF07FBFA09DF" box="[988,1100,168,192]" captionStart="Figure 3" captionStartId="11.[144,211,1354,1376]" captionTargetBox="[144,1104,177,1317]" captionTargetId="figure@11.[144,1104,168,1323]" captionTargetPageId="11" captionText="Figure 3. Achaeta giustii sp. nov. (A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="12" pageNumber="1999">Figure 3G</figureCitation>
); ectal ducts not tapering towards pore, rather appearing glandular at junction with body wall.
</paragraph>
</subSubSection>
<caption id="DF4C4CF9FFE34503FF02FAE5FD9F0F32" ID-DOI="http://doi.org/10.5281/zenodo.3997992" ID-Zenodo-Dep="3997992" httpUri="https://zenodo.org/record/3997992/files/figure.png" pageId="11" pageNumber="1998" startId="11.[144,211,1354,1376]" targetBox="[144,1104,177,1317]" targetPageId="11">
<paragraph id="8B8C1C71FFE34503FF02FAE5FD9F0F32" blockId="11.[144,1105,1354,1581]" pageId="11" pageNumber="1998">
Figure 3.
<taxonomicName id="4C3367F2FFE34503FF6BFAE5FE260C7F" authority="Rota, 2015" authorityName="Rota" authorityYear="2015" box="[249,400,1354,1376]" class="Magnoliopsida" family="Poaceae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="11" pageNumber="1998" phylum="Tracheophyta" rank="species" species="giustii" status="sp. nov.">
<emphasis id="B947C063FFE34503FF6BFAE5FE260C7F" bold="true" box="[249,400,1354,1376]" italics="true" pageId="11" pageNumber="1998">Achaeta giustii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2747D18FFE34503FE0AFAE4FE5C0C7F" box="[408,490,1355,1376]" pageId="11" pageNumber="1998" rank="species">sp. nov.</taxonomicNameLabel>
(A) Cephalic region (lateral view); (B) dorsolateral view of segments IVIII, showing the well-developed pharyngeal glands; (C) dorsal view of clitellum. Note the granular cells bordering the middorsal interruption (dg); (D) shallow lateral view of body wall (segment IX), revealing the lozenge pattern of the longitudinal muscle fibres; (E) lateral view of clitellum, showing the reticulate pattern of the gland cells; (F) lateral view of same clitellum in a deeper optical section, documenting its midventral continuity and middorsal gap; (G) ectal portion of the spermatheca; (H) sperm funnel. All from permanent wholemounted specimens, anterior to the left.
</paragraph>
</caption>
<subSubSection id="C3294FFAFFE44504FF46FE83FE290B14" pageId="12" pageNumber="1999" type="discussion">
<paragraph id="8B8C1C71FFE44504FF46FE83FE80085B" blockId="12.[212,1172,300,523]" box="[212,310,300,324]" pageId="12" pageNumber="1999">
<heading id="D0C4AB1DFFE44504FF46FE83FE80085B" box="[212,310,300,324]" fontSize="10" level="3" pageId="12" pageNumber="1999" reason="5">
<emphasis id="B947C063FFE44504FF46FE83FE80085B" box="[212,310,300,324]" italics="true" pageId="12" pageNumber="1999">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFE44504FF46FEFBFE290B14" blockId="12.[212,1172,300,523]" pageId="12" pageNumber="1999">
This species belongs to the group of
<taxonomicName id="4C3367F2FFE44504FDFBFEFBFD720873" authorityName="Vejdovsky" authorityYear="1878" box="[617,708,340,364]" class="Magnoliopsida" family="Poaceae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="12" pageNumber="1999" phylum="Tracheophyta" rank="genus">
<emphasis id="B947C063FFE44504FDFBFEFBFD720873" box="[617,708,340,364]" italics="true" pageId="12" pageNumber="1999">Achaeta</emphasis>
</taxonomicName>
with only dorsal flask-shaped glands and spermathecae opening ventrally. While alive it looks similar to
<taxonomicName id="4C3367F2FFE44504FC57FEDBFDDD08B4" authority="sensu Nielsen and Christensen, 1959" authorityName="sensu Nielsen and Christensen" authorityYear="1959" baseAuthorityName="Vejdovsky" baseAuthorityYear="1879" class="Magnoliopsida" family="Poaceae" genus="Achaeta" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="12" pageNumber="1999" phylum="Tracheophyta" rank="species" species="bohemica">
<emphasis id="B947C063FFE44504FC57FEDBFB250893" box="[965,1171,372,396]" italics="true" pageId="12" pageNumber="1999">Achaeta bohemica</emphasis>
sensu
<bibRefCitation id="EFA26180FFE44504FE8AFE3CFDDD08B4" author="Nielsen CO &amp; Christensen B" box="[280,619,403,427]" pageId="12" pageNumber="1999" pagination="1 - 160" refId="ref16681" refString="Nielsen CO, Christensen B. 1959. The Enchytraeidae. Critical revision and taxonomy of European species. Natura Jutl. 8 - 9: 1 - 160." type="journal article" year="1959">Nielsen and Christensen, 1959</bibRefCitation>
</taxonomicName>
, it can be recognized (more easily after fixation) by having a complete series of dorsal flask-shaped glands (i.e. from segment II to tail) and clitellum longer and reticulate, with granular gland cells bordering the dorsal edges (see above).
</paragraph>
</subSubSection>
<subSubSection id="C3294FFAFFE44504FF46FDF8FEE60BA9" pageId="12" pageNumber="1999" type="distribution">
<paragraph id="8B8C1C71FFE44504FF46FDF8FEEE0B70" blockId="12.[212,1172,599,694]" box="[212,344,599,623]" pageId="12" pageNumber="1999">
<heading id="D0C4AB1DFFE44504FF46FDF8FEEE0B70" box="[212,344,599,623]" fontSize="10" level="3" pageId="12" pageNumber="1999" reason="5">
<emphasis id="B947C063FFE44504FF46FDF8FEEE0B70" box="[212,344,599,623]" italics="true" pageId="12" pageNumber="1999">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C1C71FFE44504FF46FDD0FEE60BA9" blockId="12.[212,1172,599,694]" pageId="12" pageNumber="1999">
<materialsCitation id="3B5B162CFFE44504FF46FDD0FBC30B88" ID-GBIF-Occurrence="2848394320" box="[212,1141,639,663]" country="Italy" location="Mediterranean region" pageId="12" pageNumber="1999" specimenCount="1" stateProvince="Toscana">
Apparently endemic to the
<location id="8EEC4AAAFFE44504FD99FDD0FCB30B88" LSID="urn:lsid:plazi:treatment:039AAD67FFE14504FE79FC9AFEE60BA9:8EEC4AAAFFE44504FD99FDD0FCB30B88" box="[523,773,639,663]" country="Italy" name="Mediterranean region" pageId="12" pageNumber="1999" stateProvince="Toscana">Mediterranean region</location>
, recorded in
<collectingCountry id="F3245CE1FFE44504FC08FDD0FC780B88" box="[922,974,639,663]" name="Italy" pageId="12" pageNumber="1999">Italy</collectingCountry>
from
<collectingRegion id="49F7D293FFE44504FB84FDD0FBC30B88" box="[1046,1141,639,663]" country="Italy" name="Toscana" pageId="12" pageNumber="1999">Tuscany</collectingRegion>
</materialsCitation>
to
<collectingRegion id="49F7D293FFE44504FF46FD31FEFB0BA9" box="[212,333,670,694]" country="Italy" name="Campania" pageId="12" pageNumber="1999">Campania</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>