394 lines
46 KiB
XML
394 lines
46 KiB
XML
<document id="D968C2A0A079A8842FA46AF25D437783" ID-DOI="10.5281/zenodo.5273151" ID-ISSN="0253-116X" ID-Zenodo-Dep="5273151" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629964482392" checkinUser="marcus" docAuthor="Theischinger, Günther, D, Zacariah & Martin, John" docDate="2018" docId="038187EEFF9DFFA5FF5C7D53E5A364EC" docLanguage="en" docName="LinzerbiolBeitr.2018.50.2.1587-1633.pdf" docOrigin="Linzer biologische Beiträge 50 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Styringomyia bipunctata EDWARDS" docType="treatment" docVersion="4" lastPageNumber="1597" masterDocId="FFB8FF96FF94FFAEFFEE7910E67B654C" masterDocTitle="The genus Styringomyia LOEW in Australia (Diptera: Tipuloidea: Limoniidae)" masterLastPageNumber="1633" masterPageNumber="1587" pageNumber="1596" updateTime="1699131846658" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7937F1AFBC946018E939623B76B293A8">The genus Styringomyia LOEW in Australia (Diptera: Tipuloidea: Limoniidae)</mods:title>
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<mods:namePart id="09F4CC712C07F679285180CD9F1D3FDA">D, Zacariah</mods:namePart>
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EDWARDS
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(
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M a t e r i a l s t u d i e d.
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(see
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(
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;
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, L.
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W by
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(
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.
|
||
</specimenCount>
|
||
off
|
||
<location id="8EF76023FF9DFFA7FD877DF9E48B61B3" LSID="urn:lsid:plazi:treatment:038187EEFF9DFFA5FF5C7D53E5A364EC:8EF76023FF9DFFA7FD877DF9E48B61B3" box="[617,752,1257,1279]" county="Sluice Ck" municipality="Mt Edith Forest Road" name="Danbulla Road" pageId="9" pageNumber="1596" stateProvince="Queensland">Danbulla Road</location>
|
||
,
|
||
<date id="FF961038FF9DFFA7FD177DF9E53061B3" box="[761,843,1257,1279]" pageId="9" pageNumber="1596" value="1967-05-06">
|
||
<collectingDate id="EFD2E9D0FF9DFFA7FD177DF9E53061B3" box="[761,843,1257,1279]" pageId="9" pageNumber="1596" value="1967-05-06">6-v-1967</collectingDate>
|
||
</date>
|
||
, D.H.
|
||
<collectorName id="26DD532EFF9DFFA7FC6B7DF9E5BE61B3" box="[901,965,1257,1279]" pageId="9" pageNumber="1596">Colless</collectorName>
|
||
(
|
||
<collectionCode id="ED39AE3DFF9DFFA7FC3C7DF9E27761B3" box="[978,1036,1257,1279]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="1596">ANIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B403CA5FF9DFFA7FBF27DF9E7E0600B" collectingDate="1967-04-27" collectingDateMax="1967-05-07" collectingDateMin="1967-04-27" collectionCode="ANIC" collectorName="Danbulla Road & D. H. Colless & Yungaburra" county="Mt Edith" elevation="6" location="State Forest" municipality="Wongabel State Forest" pageId="9" pageNumber="1596" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Queensland" typeStatus="holotype">
|
||
<specimenCount id="9D2EFD71FF9DFFA7FBF27DF9E24361B2" box="[1052,1080,1257,1278]" pageId="9" pageNumber="1596" type="female">1♀</specimenCount>
|
||
,
|
||
<collectingCounty id="62F64E74FF9DFFA7FF317C12E74B605B" box="[223,304,1281,1303]" pageId="9" pageNumber="1596">Mt Edith</collectingCounty>
|
||
<quantity id="4CD09B1DFF9DFFA7FED87C11E714605B" box="[310,367,1281,1303]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="9" pageNumber="1596" unit="m" value="5.5" valueMax="7.0" valueMin="4.0">
|
||
<elevation id="0005D1CBFF9DFFA7FED87C11E714605B" box="[310,367,1281,1303]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="9" pageNumber="1596" unit="m" value="5.5" valueMax="7.0" valueMin="4.0">4-7 m</elevation>
|
||
</quantity>
|
||
. off
|
||
<collectorName id="26DD532EFF9DFFA7FE767C11E45B605B" box="[408,544,1281,1303]" pageId="9" pageNumber="1596">Danbulla Road</collectorName>
|
||
,
|
||
<date id="FF961038FF9DFFA7FDC47C11E4F6605B" box="[554,653,1281,1303]" pageId="9" pageNumber="1596" value="1967-04-27">
|
||
<collectingDate id="EFD2E9D0FF9DFFA7FDC47C11E4F6605B" box="[554,653,1281,1303]" pageId="9" pageNumber="1596" value="1967-04-27">27-iv-1967</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26DD532EFF9DFFA7FD797C12E572605B" box="[663,777,1281,1303]" pageId="9" pageNumber="1596">D.H. Colless</collectorName>
|
||
(
|
||
<collectionCode id="ED39AE3DFF9DFFA7FCF87C11E52A605B" box="[790,849,1281,1303]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="1596">ANIC</collectionCode>
|
||
).
|
||
<specimenCount id="9D2EFD71FF9DFFA7FC8E7C11E506605A" box="[864,893,1281,1302]" pageId="9" pageNumber="1596" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D2EFD71FF9DFFA7FC667C11E5DF605A" box="[904,932,1281,1302]" pageId="9" pageNumber="1596" type="female">1♀</specimenCount>
|
||
, 2 sn,
|
||
<collectingMunicipality id="6BF3AC82FF9DFFA7FC0C7C11E7326063" pageId="9" pageNumber="1596">Wongabel State Forest</collectingMunicipality>
|
||
,
|
||
<date id="FF961038FF9DFFA7FEBF7C09E7D96063" box="[337,418,1305,1327]" pageId="9" pageNumber="1596" value="1967-05-07">
|
||
<collectingDate id="EFD2E9D0FF9DFFA7FEBF7C09E7D96063" box="[337,418,1305,1327]" pageId="9" pageNumber="1596" value="1967-05-07">7-v-1967</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26DD532EFF9DFFA7FE457C0AE4606063" box="[427,539,1305,1327]" pageId="9" pageNumber="1596">D.H. Colless</collectorName>
|
||
(
|
||
<collectionCode id="ED39AE3DFF9DFFA7FDC87C09E41A6063" box="[550,609,1305,1327]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="1596">ANIC</collectionCode>
|
||
).
|
||
<specimenCount id="9D2EFD71FF9DFFA7FD817C09E4F76062" box="[623,652,1305,1326]" pageId="9" pageNumber="1596" type="male">1♂</specimenCount>
|
||
, 1 sn,
|
||
<collectorName id="26DD532EFF9DFFA7FD2D7C09E5576063" box="[707,812,1305,1327]" pageId="9" pageNumber="1596">Yungaburra</collectorName>
|
||
(
|
||
<location id="8EF76023FF9DFFA7FCD97C09E5E46063" LSID="urn:lsid:plazi:treatment:038187EEFF9DFFA5FF5C7D53E5A364EC:8EF76023FF9DFFA7FCD97C09E5E46063" box="[823,927,1305,1327]" county="Mt Edith" municipality="Wongabel State Forest" name="State Forest" pageId="9" pageNumber="1596" stateProvince="Queensland">State Forest</location>
|
||
452),
|
||
<date id="FF961038FF9DFFA7FC397C09E2426063" box="[983,1081,1305,1327]" pageId="9" pageNumber="1596" value="1967-04-29">
|
||
<collectingDate id="EFD2E9D0FF9DFFA7FC397C09E2426063" box="[983,1081,1305,1327]" pageId="9" pageNumber="1596" value="1967-04-29">29-iv-1967</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26DD532EFF9DFFA7FF317C22E734600B" box="[223,335,1329,1351]" pageId="9" pageNumber="1596">D.H. Colless</collectorName>
|
||
(
|
||
<collectionCode id="ED39AE3DFF9DFFA7FEB47C21E7EE600B" box="[346,405,1329,1351]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="1596">ANIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3326573FF9DFFA5FF5F7C4DE5A364EC" lastPageId="11" lastPageNumber="1598" pageId="9" pageNumber="1596" type="description">
|
||
<paragraph id="8B9736F8FF9DFFA7FF5F7C4DE7E0630D" blockId="9.[177,1085,1373,1601]" pageId="9" pageNumber="1596">
|
||
The most valuable diagnostic characters specified in the original Description of the male (
|
||
<bibRefCitation id="EFB94B09FF9DFFA7FF577C6BE72060DD" author="EDWARDS F" box="[185,347,1403,1425]" pageId="9" pageNumber="1596" pagination="265 - 274" refId="ref13530" refString="EDWARDS F. W. (1924): New data concerning Styringomyia (Diptera, Tipulidae). - The Annals and Magazine of Natural History (9) 13: 265 - 274." type="journal article" year="1924">EDWARDS 1924</bibRefCitation>
|
||
) are: "Legs uniformly yellow, except for the extreme tips of femora and tibiae. Wings with the usual dark spots over the crossveins. Hypopygium: Ninth tergite rather short, ending in three little triangular points. Ninth sternite very long, extending beyond the tips of the side-pieces, curled into a tubular shape, truncate and slightly blackened at the tip. Terminal projection of side-pieces rather long and stout, with one rather weak spine and a number of stiff hairs, two or three of which are thickened, black and almost spine-like."
|
||
</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C79CBE51A642A" blockId="10.[178,1085,219,358]" pageId="10" pageNumber="1597">
|
||
From the Description above of the hypopygium and the now available material of
|
||
<taxonomicName id="4C284D7BFF9EFFA4FF5C79E8E7426441" authorityName="LOEW" authorityYear="1845" box="[178,313,248,269]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAEAFF9EFFA4FF5C79E8E7426441" box="[178,313,248,269]" italics="true" pageId="10" pageNumber="1597">Styringomyia</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33F7668FF9EFFA4FE6E79E8E7A36442" box="[384,472,248,270]" name="Australia" pageId="10" pageNumber="1597">Australia</collectingCountry>
|
||
it became obvious that
|
||
<bibRefCitation id="EFB94B09FF9EFFA4FD0B79E8E5E26442" author="EDWARDS F" box="[741,921,248,270]" pageId="10" pageNumber="1597" pagination="265 - 274" refId="ref13530" refString="EDWARDS F. W. (1924): New data concerning Styringomyia (Diptera, Tipulidae). - The Annals and Magazine of Natural History (9) 13: 265 - 274." type="journal article" year="1924">EDWARDS (1924)</bibRefCitation>
|
||
when giving a correct treatment of the dorso-ventral orientation of the male terminalia in
|
||
<taxonomicName id="4C284D7BFF9EFFA4FC5B7806E2476467" authorityName="LOEW" authorityYear="1845" box="[949,1084,278,299]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAEAFF9EFFA4FC5B7806E2476467" box="[949,1084,278,299]" italics="true" pageId="10" pageNumber="1597">Styringomyia</emphasis>
|
||
</taxonomicName>
|
||
did not get it right in
|
||
<emphasis id="B95CEAEAFF9EFFA4FE777823E4536404" box="[409,552,307,328]" italics="true" pageId="10" pageNumber="1597">
|
||
<taxonomicName id="4C284D7BFF9EFFA4FE777823E45E6404" box="[409,549,307,328]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="bipunctata">S. bipunctata</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
If this is taken into account, however, the original description is sufficient for a confident interpretation of the species.
|
||
</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7866E4696782" blockId="10.[178,1085,373,718]" pageId="10" pageNumber="1597">
|
||
<bibRefCitation id="EFB94B09FF9EFFA4FF5C7866E70264C7" author="ALEXANDER C" box="[178,377,373,395]" pageId="10" pageNumber="1597" pagination="221 - 274" refId="ref13282" refString="ALEXANDER C. P. (1925): New or little-known Tipulidae from the Philippines (Diptera). XLVII. - Philippine J ournal of Science 89: 221 - 274." type="journal article" year="1925">ALEXANDER (1925</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB94B09FF9EFFA4FE667865E7C764C7" author="ALEXANDER C" box="[392,444,373,395]" pageId="10" pageNumber="1597" pagination="369 - 377" refId="ref13349" refString="ALEXANDER C. P. (1960): New or little-known Tipulidae from eastern Asia (Diptera). Part III. - Philippine Journal of Science 28: 369 - 377." type="journal article" year="1960">1960</bibRefCitation>
|
||
) mentioned
|
||
<taxonomicName id="4C284D7BFF9EFFA4FDAB7866E4B464C6" box="[581,719,373,395]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="bipunctata">
|
||
<emphasis id="B95CEAEAFF9EFFA4FDAB7866E4B464C6" box="[581,719,373,395]" italics="true" pageId="10" pageNumber="1597">S. bipunctata</emphasis>
|
||
</taxonomicName>
|
||
in comparison with
|
||
<taxonomicName id="4C284D7BFF9EFFA4FC5A7866E70F64E5" authority="ALEXANDER, 1925" authorityName="ALEXANDER" authorityYear="1925" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="mcgregori">
|
||
<emphasis id="B95CEAEAFF9EFFA4FC5A7866E24764C7" box="[948,1084,374,395]" italics="true" pageId="10" pageNumber="1597">S. mcgregori</emphasis>
|
||
<bibRefCitation id="EFB94B09FF9EFFA4FF5C7883E70F64E5" author="ALEXANDER C" box="[178,372,403,425]" pageId="10" pageNumber="1597" pagination="221 - 274" refId="ref13282" refString="ALEXANDER C. P. (1925): New or little-known Tipulidae from the Philippines (Diptera). XLVII. - Philippine J ournal of Science 89: 221 - 274." type="journal article" year="1925">ALEXANDER, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
and with
|
||
<taxonomicName id="4C284D7BFF9EFFA4FE087883E55864E5" authority="ALEXANDER, 1960" authorityName="ALEXANDER" authorityYear="1960" box="[486,803,403,425]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="scalaris">
|
||
<emphasis id="B95CEAEAFF9EFFA4FE087883E42864E4" box="[486,595,403,424]" italics="true" pageId="10" pageNumber="1597">S. scalaris</emphasis>
|
||
<bibRefCitation id="EFB94B09FF9EFFA4FDB07883E55864E5" author="ALEXANDER C" box="[606,803,403,425]" pageId="10" pageNumber="1597" pagination="369 - 377" refId="ref13349" refString="ALEXANDER C. P. (1960): New or little-known Tipulidae from eastern Asia (Diptera). Part III. - Philippine Journal of Science 28: 369 - 377." type="journal article" year="1960">ALEXANDER, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
. The great differences between the male terminalia of these species have nothing to do with the fact that ALEXANDER identified a
|
||
<taxonomicName id="4C284D7BFF9EFFA4FE2B78DEE43764AF" authorityName="LOEW" authorityYear="1845" box="[453,588,462,483]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAEAFF9EFFA4FE2B78DEE43764AF" box="[453,588,462,483]" italics="true" pageId="10" pageNumber="1597">Styringomyia</emphasis>
|
||
</taxonomicName>
|
||
male from Brooklana,
|
||
<collectingRegion id="49ECF81AFF9EFFA4FCBB78DEE26264AF" box="[853,1049,461,483]" country="Australia" name="New South Wales" pageId="10" pageNumber="1597">New South Wales</collectingRegion>
|
||
as
|
||
<emphasis id="B95CEAEAFF9EFFA4FF5C78FBE765674C" box="[178,286,491,512]" italics="true" pageId="10" pageNumber="1597">bipunctata</emphasis>
|
||
(
|
||
<figureCitation id="13132A7DFF9EFFA4FEDF78FBE7DD674D" box="[305,422,491,513]" captionStart="Figs 40-43" captionStartId="36.[178,218,1504,1526]" captionTargetBox="[181,1088,216,1489]" captionTargetId="figure-30@36.[181,1088,216,1489]" captionTargetPageId="36" captionText="Figs 40-43: Styringomyia williamsi nov.sp. (interpreted as Styringomyia bipunctata EDWARDS): (40) wing; (41) terminalia; (42) leg; (43) label." figureDoi="http://doi.org/10.5281/zenodo.5275902" httpUri="https://zenodo.org/record/5275902/files/figure.png" pageId="10" pageNumber="1597">Figs 40-43</figureCitation>
|
||
). This specimen is actually
|
||
<taxonomicName id="4C284D7BFF9EFFA4FD3078FBE521674C" authority="Theischinger & D & Martin, 2018" authorityName="Theischinger & D & Martin" authorityYear="2018" box="[734,858,491,512]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B95CEAEAFF9EFFA4FD3078FBE521674C" box="[734,858,491,512]" italics="true" pageId="10" pageNumber="1597">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26F5791FF9EFFA4FC8B78FBE5D4674C" box="[869,943,491,512]" pageId="10" pageNumber="1597" rank="species">nov.sp.</taxonomicNameLabel>
|
||
as described below. Whereas
|
||
<taxonomicName id="4C284D7BFF9EFFA4FEB17B18E79D6751" box="[351,486,520,541]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="bipunctata">
|
||
<emphasis id="B95CEAEAFF9EFFA4FEB17B18E79D6751" box="[351,486,520,541]" italics="true" pageId="10" pageNumber="1597">S. bipunctata</emphasis>
|
||
</taxonomicName>
|
||
is a tropical species,
|
||
<taxonomicName id="4C284D7BFF9EFFA4FD267B18E53A6751" authorityName="Theischinger & D & Martin" authorityYear="2018" box="[712,833,520,541]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B95CEAEAFF9EFFA4FD267B18E53A6751" box="[712,833,520,541]" italics="true" pageId="10" pageNumber="1597">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
is known to range from Goomburra in
|
||
<collectingRegion id="49ECF81AFF9EFFA4FE837B35E79E6777" box="[365,485,549,571]" country="Australia" name="Queensland" pageId="10" pageNumber="1597">Queensland</collectingRegion>
|
||
(
|
||
<geoCoordinate id="EE1C503FFF9EFFA4FDE87B35E4F16777" box="[518,650,549,571]" degrees="27.978844" direction="south" orientation="latitude" pageId="10" pageNumber="1597" precision="1" value="-27.978844">27.978844°S</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE1C503FFF9EFFA4FD7C7B35E55D6777" box="[658,806,549,571]" degrees="152.293154" direction="east" orientation="longitude" pageId="10" pageNumber="1597" precision="1" value="152.29315">152.293154°E</geoCoordinate>
|
||
) to Minnamurra Falls (
|
||
<geoCoordinate id="EE1C503FFF9EFFA4FF547B53E7776715" box="[186,268,579,601]" degrees="34" direction="south" minutes="38" orientation="latitude" pageId="10" pageNumber="1597" precision="925" value="-34.633335">34°38’S</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE1C503FFF9EFFA4FEFA7B53E70C6715" box="[276,375,579,601]" degrees="152" direction="east" minutes="51" orientation="longitude" pageId="10" pageNumber="1597" precision="925" value="152.85">152°51’E</geoCoordinate>
|
||
) well south of Sydney. The two are apparently a pair of allopatric species occurring north, respectively south, of the Eungella - Paluma gap, a region where ecological and physiographic boundaries coincide with taxonomic discontinuities below the level of closely related species-pairs in Odonata, Plecoptera and Megaloptera (WATSON & THEISCHINGER 1984).
|
||
</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7BF6E70367B7" blockId="10.[178,376,741,763]" box="[178,376,741,763]" pageId="10" pageNumber="1597">D e s c r i p t i o n</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7A1BE710666D" blockId="10.[178,363,779,801]" box="[178,363,779,801]" pageId="10" pageNumber="1597">
|
||
Male (
|
||
<figureCitation id="13132A7DFF9EFFA4FF1D7A1BE719666D" box="[243,354,779,801]" captionStart-0="Fig" captionStart-1="Figs 34-36" captionStartId-0="33.[178,211,867,889]" captionStartId-1="34.[178,218,1553,1575]" captionTargetBox-0="[178,1085,212,847]" captionTargetBox-1="[176,1083,216,1538]" captionTargetId-0="figure-19@33.[178,1085,212,847]" captionTargetId-1="figure-26@34.[176,1083,216,1538]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="Fig. 33: Styringomyia bipunctata EDWARDS, holotype male, habitus, label and terminalia inset." captionText-1="Figs 34-36: Styringomyia bipunctata EDWARDS, terminalia: (34, 35) male: (34) ventral; (35) dorsal; (36) female, lateral." figureDoi-0="http://doi.org/10.5281/zenodo.5273183" figureDoi-1="http://doi.org/10.5281/zenodo.5273185" httpUri-0="https://zenodo.org/record/5273183/files/figure.png" httpUri-1="https://zenodo.org/record/5273185/files/figure.png" pageId="10" pageNumber="1597">Figs 33-35</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7A20E56E6102" blockId="10.[178,1085,816,1102]" pageId="10" pageNumber="1597">Hypopygium. Tergite 9/10 long and wide, largely parallel sided with ear-like lateral expansions in third quarter of its length. Sternite 9 widely trapezoidal, almost hemispherical ending in three narrow lobes, the median apparently with tip triangular, the lateral ones strongly setose; two moderately long and moderately thick subapical setae. Gonocoxites evenly curved, almost conical with basal three fourths markedly expanded and with rather slim apical spine and abundant setae along mesal face, particularly at the apical end of the expansion. Gonostyli a single element represented by a slightly curved lobe triangularly expanded subbasally, with abundant setae and two large apical spines and one subapical spine bearing a secondary spine. What is detectable of the aedeagus appears moderately wide and obtusely conical.</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7D4DE720613F" blockId="10.[178,347,1117,1139]" box="[178,347,1117,1139]" pageId="10" pageNumber="1597">
|
||
Female (
|
||
<figureCitation id="13132A7DFF9EFFA4FEE47D4DE72F613F" box="[266,340,1117,1139]" captionStart="Figs 34-36" captionStartId="34.[178,218,1553,1575]" captionTargetBox="[176,1083,216,1538]" captionTargetId="figure-26@34.[176,1083,216,1538]" captionTargetPageId="34" captionText="Figs 34-36: Styringomyia bipunctata EDWARDS, terminalia: (34, 35) male: (34) ventral; (35) dorsal; (36) female, lateral." figureDoi="http://doi.org/10.5281/zenodo.5273185" httpUri="https://zenodo.org/record/5273185/files/figure.png" pageId="10" pageNumber="1597">Fig. 36</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7D93E40061FA" blockId="10.[178,1085,1155,1206]" pageId="10" pageNumber="1597">Head. Including rostrum and antennae yellowish to pale greyish brown, palps somewhat darker and postocular section greyish brown.</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7DD5E58F61B5" blockId="10.[178,1084,1221,1273]" pageId="10" pageNumber="1597">Thorax. Largely brownish yellow. Legs pale brownish yellow with extreme tip of femora and tibiae slightly darkened and last tarsal segment and claws dark greyish brown.</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7C18E5946052" blockId="10.[178,1007,1288,1310]" box="[178,1007,1288,1310]" pageId="10" pageNumber="1597">Wings. Infumed with pale greyish and with somewhat darker spots on crossveins.</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7C3DE5DF600F" blockId="10.[178,932,1325,1347]" box="[178,932,1325,1347]" pageId="10" pageNumber="1597">Abdomen. Largely dull yellow with tergites almost wholly greyish brown.</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7C43E4BB6024" blockId="10.[178,704,1363,1385]" box="[178,704,1363,1385]" pageId="10" pageNumber="1597">
|
||
Terminalia. As specified under
|
||
<taxonomicName id="4C284D7BFF9EFFA4FE1C7C43E40C6024" box="[498,631,1363,1384]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="bipunctata">
|
||
<emphasis id="B95CEAEAFF9EFFA4FE1C7C43E40C6024" box="[498,631,1363,1384]" italics="true" pageId="10" pageNumber="1597">S. bipunctata</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7C68E44360C2" blockId="10.[178,568,1400,1422]" box="[178,568,1400,1422]" pageId="10" pageNumber="1597">
|
||
Dimensions. Wing length 4.0-
|
||
<quantity id="4CD09B1DFF9EFFA4FE0A7C68E44E60C2" box="[484,565,1400,1422]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="1597" unit="mm" value="4.5">4.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA4FF5C7C8EE26960F8" blockId="10.[178,1042,1437,1460]" box="[178,1042,1437,1460]" pageId="10" pageNumber="1597">
|
||
D i s t r i b u t i o n. North-eastern; known only from tropical
|
||
<collectingRegion id="49ECF81AFF9EFFA4FCD57C8DE5C860FF" box="[827,947,1437,1459]" country="Australia" name="Queensland" pageId="10" pageNumber="1597">Queensland</collectingRegion>
|
||
(
|
||
<figureCitation id="13132A7DFF9EFFA4FC2E7C8EE27E60F8" box="[960,1029,1438,1460]" captionStart="Map 3" captionStartId="46.[178,223,854,875]" captionTargetBox="[299,964,212,834]" captionTargetId="figure-49@46.[299,964,212,834]" captionTargetPageId="46" captionText="Map 3: Distribution of the species of the Styringomyia bipunctata group: S. bipunctata, S. williamsi, S. coronata." figureDoi="http://doi.org/10.5281/zenodo.5275920" httpUri="https://zenodo.org/record/5275920/files/figure.png" pageId="10" pageNumber="1597">Map 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9736F8FF9EFFA5FF5C7CDBE5A364EC" blockId="10.[178,1085,1483,1593]" lastBlockId="11.[178,1086,219,417]" lastPageId="11" lastPageNumber="1598" pageId="10" pageNumber="1597">
|
||
D i s c u s s i o n
|
||
<taxonomicName id="4C284D7BFF9EFFA4FE697CDBE4F160AC" box="[391,650,1483,1504]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="bipunctata">
|
||
<emphasis id="B95CEAEAFF9EFFA4FE697CDBE4F160AC" box="[391,650,1483,1504]" italics="true" pageId="10" pageNumber="1597">Styringomyia bipunctata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C284D7BFF9EFFA4FD4F7CDBE55A60AC" authority="Theischinger & D & Martin, 2018" authorityName="Theischinger & D & Martin" authorityYear="2018" box="[673,801,1483,1504]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B95CEAEAFF9EFFA4FD4F7CDBE55A60AC" box="[673,801,1483,1504]" italics="true" pageId="10" pageNumber="1597">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26F5791FF9EFFA4FCDF7CDBE50160AC" box="[817,890,1483,1504]" pageId="10" pageNumber="1597" rank="species">nov.sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C284D7BFF9EFFA4FC507CDBE24760AC" authority="Theischinger & D & Martin, 2018" authorityName="Theischinger & D & Martin" authorityYear="2018" box="[958,1084,1483,1504]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="species" species="coronata" status="sp. nov.">
|
||
<emphasis id="B95CEAEAFF9EFFA4FC507CDBE24760AC" box="[958,1084,1483,1504]" italics="true" pageId="10" pageNumber="1597">S. coronata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26F5791FF9EFFA4FF5C7CF9E68760B2" box="[178,252,1513,1534]" pageId="10" pageNumber="1597" rank="species">nov.sp.</taxonomicNameLabel>
|
||
are a closely knit group of species with sternite 9 of male ending in three small extensions and tergite 9/10 very long and truncate if the 180 degrees torsion of the male terminalia in
|
||
<taxonomicName id="4C284D7BFF9EFFA4FED47F33E7BA6374" authorityName="LOEW" authorityYear="1845" box="[314,449,1571,1592]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1597" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95CEAEAFF9EFFA4FED47F33E7BA6374" box="[314,449,1571,1592]" italics="true" pageId="10" pageNumber="1597">Styringomyia</emphasis>
|
||
</taxonomicName>
|
||
is correct and consistent (
|
||
<bibRefCitation id="EFB94B09FF9EFFA4FD237F33E50B6374" author="EDWARDS F" box="[717,880,1571,1592]" pageId="10" pageNumber="1597" pagination="265 - 274" refId="ref13530" refString="EDWARDS F. W. (1924): New data concerning Styringomyia (Diptera, Tipulidae). - The Annals and Magazine of Natural History (9) 13: 265 - 274." type="journal article" year="1924">EDWARDS 1924</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB94B09FF9EFFA4FC957F33E2716375" author="RIBEIRO G" box="[891,1034,1571,1593]" pageId="10" pageNumber="1597" pagination="1 - 35" refId="ref13933" refString="RIBEIRO G. C. (2003): Systematics of the Neotropical species of Styringomyia LOEW (Diptera: Tipulomorpha: Limoniidae). - Zootaxa 253: 1 - 35." type="journal article" year="2003">RIBEIRO 2003</bibRefCitation>
|
||
) and if we can assume (based on fresh material of three species) that
|
||
<bibRefCitation id="EFB94B09FF9FFFA5FCAA79CBE58F65BD" author="EDWARDS F" box="[836,1012,219,241]" pageId="11" pageNumber="1598" pagination="265 - 274" refId="ref13530" refString="EDWARDS F. W. (1924): New data concerning Styringomyia (Diptera, Tipulidae). - The Annals and Magazine of Natural History (9) 13: 265 - 274." type="journal article" year="1924">EDWARDS (1924)</bibRefCitation>
|
||
nevertheless got the dorsoventral orientation of the hypopygium wrong in the Description of
|
||
<taxonomicName id="4C284D7BFF9FFFA5FBC479E8E7656466" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="1598" phylum="Arthropoda" rank="species" species="bipunctata">
|
||
<emphasis id="B95CEAEAFF9FFFA5FBC479E8E7656466" italics="true" pageId="11" pageNumber="1598">S. bipunctata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C284D7BFF9FFFA5FEC57806E7CA6466" box="[299,433,277,299]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="1598" phylum="Arthropoda" rank="species" species="bipunctata">
|
||
<emphasis id="B95CEAEAFF9FFFA5FEC57806E7CA6466" box="[299,433,277,299]" italics="true" pageId="11" pageNumber="1598">S. bipunctata</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C284D7BFF9FFFA5FD297806E5456466" authorityName="Theischinger & D & Martin" authorityYear="2018" box="[711,830,277,299]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="1598" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B95CEAEAFF9FFFA5FD297806E5456466" box="[711,830,277,299]" italics="true" pageId="11" pageNumber="1598">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
, its closest ally, possible its sister taxon, by the longer, less flat and narrower median extension of male sternite 9 and by its geographical distribution (more northern, tropical) versus more southern. Both,
|
||
<taxonomicName id="4C284D7BFF9FFFA5FF1C787EE70064CE" box="[242,379,365,387]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="1598" phylum="Arthropoda" rank="species" species="bipunctata">
|
||
<emphasis id="B95CEAEAFF9FFFA5FF1C787EE70064CE" box="[242,379,365,387]" italics="true" pageId="11" pageNumber="1598">S. bipunctata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C284D7BFF9FFFA5FE5F787EE45264CE" authorityName="Theischinger & D & Martin" authorityYear="2018" box="[433,553,365,387]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="1598" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B95CEAEAFF9FFFA5FE5F787EE45264CE" box="[433,553,365,387]" italics="true" pageId="11" pageNumber="1598">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
have the three extensions of sternite 9 shorter and the thinner apical portion of the gonocoxites markedly shorter than
|
||
<taxonomicName id="4C284D7BFF9FFFA5FCB0789BE5A964EC" authorityName="Theischinger & D & Martin" authorityYear="2018" box="[862,978,395,416]" class="Insecta" family="Limoniidae" genus="Styringomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="1598" phylum="Arthropoda" rank="species" species="coronata">
|
||
<emphasis id="B95CEAEAFF9FFFA5FCB0789BE5A964EC" box="[862,978,395,416]" italics="true" pageId="11" pageNumber="1598">S. coronata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |