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<document id="401B72E9247B4E3FA107EDFA3EA40C17" ID-DOI="10.5281/zenodo.3673403" ID-GBIF-Dataset="f01c9bd1-7635-40c7-b5ee-5fc57320b880" ID-ISSN="1942-1354" ID-Zenodo-Dep="3673403" ID-ZooBank="08CB0DA0-C83E-438F-AA97-8CD637F044A5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582027555556" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2019" docId="0381879FFF936E52FF1C9EE2FB65F83D" docLanguage="en" docName="InsectaMundi.0706.1-10.pdf.imf" docOrigin="Insecta Mundi 706 (706)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Ochodaeus iniquipes Paulsen 2019, new species" docType="treatment" docVersion="7" lastPageNumber="7" masterDocId="FFB8FFE7FF946E5AFFDC9C2EFFA2FFB4" masterDocTitle="New species of Ochodaeus Dejean from Madagascar II (Coleoptera: Scarabaeoidea: Ochodaeidae)" masterLastPageNumber="10" masterPageNumber="1" pageNumber="6" updateTime="1698718828345" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="CED756BDFBBA2AEC3E8A2D7E4A66BC88" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D261A50FF20A6732C68FD59F520A35EA">
<mods:title id="6721856FF0F79FB1F4AC64DC1AEA298F">New species of Ochodaeus Dejean from Madagascar II (Coleoptera: Scarabaeoidea: Ochodaeidae)</mods:title>
</mods:titleInfo>
<mods:name id="17395EB5DFF29B25496BD18051151E0E" type="personal">
<mods:role id="5CEEC3E2117791D2AB8F6C6DF2AEA857">
<mods:roleTerm id="796A55DCF4CF0E7933B38A444D29898C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7FC72C92075EE157D6EBCDADFD1FFF31">Paulsen, M. J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C78E4C80A5C579D9EC1BC3B21A80F386">text</mods:typeOfResource>
<mods:relatedItem id="7515A7A36A512001771FE18C5C43E41B" type="host">
<mods:titleInfo id="69562B4A3B60F29EBBFAD94087E844F1">
<mods:title id="B646666209CE43DCDBB1A8F0F279DEBB">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="A7BB661E8495E9CA237FBEC831C5270D">
<mods:date id="8C578125B4155CD6468409D95E0FD995">2019</mods:date>
<mods:detail id="AD2AEB9584D60A74F127D6606D7529A2" type="pubDate">
<mods:number id="4DDA189AC69F164F04C1E22F43ADBB93">2019-05-31</mods:number>
</mods:detail>
<mods:detail id="22A120590A35E0CB51592C560A013605" type="volume">
<mods:number id="646A9B7215300DBBD271A12C448DA2C4">706</mods:number>
</mods:detail>
<mods:detail id="8EC3A761589F728A4B3FEA52E32707DF" type="issue">
<mods:number id="DE991171F47D473D9DCD586E3D8E9968">706</mods:number>
</mods:detail>
<mods:extent id="DBD6F817572A32035FD1129583AEA4B6" unit="page">
<mods:start id="93508CE37562494FFF4675B2AC1BDE3A">1</mods:start>
<mods:end id="A9C3035CEC75F49E4012D467541C0B3A">10</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1BA123771DADA866FB3A962955E01902">journal article</mods:classification>
<mods:identifier id="510268797722D5E3CF810A9E12B5935D" type="DOI">10.5281/zenodo.3673403</mods:identifier>
<mods:identifier id="280C0F7BAB7E0BA0E2612477905B146C" type="GBIF-Dataset">f01c9bd1-7635-40c7-b5ee-5fc57320b880</mods:identifier>
<mods:identifier id="BB7AFAE58F5A300CE7C3B55291D2B825" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="F63FDA7DB03DF37165EEB259C538D619" type="Zenodo-Dep">3673403</mods:identifier>
<mods:identifier id="78569A007E701312927B7FAFF98F4445" type="ZooBank">08CB0DA0-C83E-438F-AA97-8CD637F044A5</mods:identifier>
</mods:mods>
<treatment id="0381879FFF936E52FF1C9EE2FB65F83D" ID-DOI="http://doi.org/10.5281/zenodo.3679983" ID-GBIF-Taxon="162161799" ID-Zenodo-Dep="3679983" LSID="urn:lsid:plazi:treatment:0381879FFF936E52FF1C9EE2FB65F83D" httpUri="http://treatment.plazi.org/id/0381879FFF936E52FF1C9EE2FB65F83D" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<subSubSection id="C3326502FF936E5DFF1C9EE2FCA4FD51" box="[192,774,716,741]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="8B973689FF936E5DFF1C9EE2FCA4FD51" blockId="7.[192,774,716,741]" box="[192,774,716,741]" pageId="7" pageNumber="6">
<heading id="D0DF81E5FF936E5DFF1C9EE2FCA4FD51" bold="true" box="[192,774,716,741]" fontSize="10" level="2" pageId="7" pageNumber="6" reason="3">
<emphasis id="B95CEA9BFF936E5DFF1C9EE2FCA4FD51" bold="true" box="[192,774,716,741]" pageId="7" pageNumber="6">
<taxonomicName id="4C284D0AFF936E5DFF1C9EE2FDF7FD51" ID-CoL="48DCQ" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[192,597,716,741]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="iniquipes" status="sp. nov.">
<emphasis id="B95CEA9BFF936E5DFF1C9EE2FE7FFD51" bold="true" box="[192,477,716,741]" italics="true" pageId="7" pageNumber="6">Ochodaeus iniquipes</emphasis>
Paulsen
</taxonomicName>
,
<taxonomicNameLabel id="A26F57E0FF936E5DFDBE9EE3FCA4FD51" box="[610,774,717,741]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3326502FF936E5DFF1C9ED0FA03FB81" pageId="7" pageNumber="6" type="materials_examined">
<paragraph id="8B973689FF936E5DFF1C9ED0FC71FCC1" blockId="7.[192,1441,765,1078]" pageId="7" pageNumber="6">
Type material.
<materialsCitation id="3B403CD4FF936E5DFEA09ED3FC71FCC1" ID-GBIF-Occurrence="2565952363" collectingDate="2009-01-26" collectingDateMax="2009-02-04" collectingDateMin="2009-01-26" collectionCode="CASC, R" collectorName="M. Irwin" country="Madagascar" county="Calif. Acad. of Sciences" elevation="15" latitude="-22.818" location="Tulear" longLatPrecision="12" longitude="43.372665" municipality="Tsifota" pageId="7" pageNumber="6" specimenCount="2" typeStatus="holotype">
<typeStatus id="5493882BFF936E5DFEA09ED3FE48FCA1" box="[380,490,765,789]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED39AE4CFF936E5DFDE19ED3FD29FCA1" box="[573,651,765,789]" pageId="7" pageNumber="6">CASC</collectionCode>
;
<figureCitation id="13132A0CFF936E5DFD4B9ED3FD4FFCA1" box="[663,749,765,789]" captionStart="Figure 3" captionStartId="8.[192,275,1383,1405]" captionTargetBox="[201,1420,250,1299]" captionTargetId="figure@8.[192,1440,236,1346]" captionTargetPageId="8" captionText="Figure 3. Ochodaeus iniquipes Paulsen, new species. a) Dorsal habitus, male holotype. b) Right front leg, dorsal view. Arrow indicating median tooth of anterior margin of profemur. c) Right stridulatory peg, dorsal view (elytra lifted). d) Holotype labels." figureDoi="http://doi.org/10.5281/zenodo.3673409" httpUri="https://zenodo.org/record/3673409/files/figure.png" pageId="7" pageNumber="6">Fig. 3a</figureCitation>
), labeled: a) “
<collectingCountry id="F33F7619FF936E5DFC4F9ED3FBFCFCA1" box="[915,1118,765,789]" name="Madagascar" pageId="7" pageNumber="6">MADAGASCAR</collectingCountry>
:
<location id="8EF76052FF936E5DFBB59ED3FB19FCA1" LSID="urn:lsid:plazi:treatment:0381879FFF936E52FF1C9EE2FB65F83D:8EF76052FF936E5DFBB59ED3FB19FCA1" box="[1129,1211,765,789]" country="Madagascar" county="Calif. Acad. of Sciences" latitude="-22.818" longLatPrecision="12" longitude="43.372665" municipality="Tsifota" name="Tulear" pageId="7" pageNumber="6">Tulear</location>
/
<collectingMunicipality id="6BF3ACF3FF936E5DFB0E9ED3FA88FCA1" box="[1234,1322,765,789]" pageId="7" pageNumber="6">Tsifota</collectingMunicipality>
,
<locationDeviation id="9A252FABFF936E5DFAEA9ED3FE9BFC81" location="Manombo" pageId="7" pageNumber="6">
<quantity id="4CD09B6CFF936E5DFAEA9ED3FA20FCA1" box="[1334,1410,765,789]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="6" unit="km" value="20.0">20 km</quantity>
N Manombo
</locationDeviation>
/
<locationDeviation id="9A252FABFF936E5DFE929F33FDDEFC81" box="[334,636,797,821]" location="Tsifota Village" pageId="7" pageNumber="6">
<quantity id="4CD09B6CFF936E5DFE929F33FE29FC81" box="[334,395,797,821]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="6" unit="km" value="1.0">1 km</quantity>
E of Tsifota Village
</locationDeviation>
/
<date id="FF961049FF936E5DFD4E9F33FCDAFC81" box="[658,888,797,821]" pageId="7" pageNumber="6" value="2009-01-26" valueMax="2009-02-04" valueMin="2009-01-26">
<collectingDate id="EFD2E9A1FF936E5DFD4E9F33FCDAFC81" box="[658,888,797,821]" pageId="7" pageNumber="6" value="2009-01-26" valueMax="2009-02-04" valueMin="2009-01-26">26 Jan–4 Feb 2009</collectingDate>
</date>
/
<geoCoordinate id="EE1C504EFF936E5DFC519F33FBBEFC81" box="[909,1052,796,821]" degrees="22" direction="south" minutes="49.08" orientation="latitude" pageId="7" pageNumber="6" precision="9" value="-22.818">22° 49.08′ S</geoCoordinate>
<geoCoordinate id="EE1C504EFF936E5DFBFF9F33FB16FC81" box="[1059,1204,796,821]" degrees="43" direction="east" minutes="22.36" orientation="longitude" pageId="7" pageNumber="6" precision="9" value="43.372665">43° 22.36′ E</geoCoordinate>
”; b) “
<collectingCounty id="62F64E05FF936E5DFB299F33FE8AFCE1" pageId="7" pageNumber="6">Calif. Acad. of Sciences</collectingCounty>
/ coll:
<collectorName id="26DD535FFF936E5DFEAB9F10FE47FCE1" box="[375,485,829,853]" pageId="7" pageNumber="6">M. Irwin</collectorName>
,
<collectionCode id="ED39AE4CFF936E5DFE2E9F10FDABFCE1" box="[498,521,830,853]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="6">R</collectionCode>
. Harin›Hala /
<collectingMethod id="52694E9EFF936E5DFD6B9F13FCF7FCE1" box="[695,853,829,853]" pageId="7" pageNumber="6">malaise trap</collectingMethod>
, elev
<quantity id="4CD09B6CFF936E5DFC479F13FC7AFCE1" box="[923,984,829,853]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="6" unit="m" value="15.0">
<elevation id="0005D1BAFF936E5DFC479F13FC7AFCE1" box="[923,984,829,853]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="6" unit="m" value="15.0">15 m</elevation>
</quantity>
/ spiny forest MG-48-12”; c) c) on red paper, “
<taxonomicName id="4C284D0AFF936E5DFEC39F73FE06FCC1" authorityName="Dejean" authorityYear="1821" box="[287,420,861,885]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEA9BFF936E5DFEC39F73FE06FCC1" box="[287,420,861,885]" italics="true" pageId="7" pageNumber="6">Ochodaeus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C284D0AFF936E5DFE669F73FD88FCC0" authorityName="Paulsen" authorityYear="2019" box="[442,554,861,884]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="iniquipes">
<emphasis id="B95CEA9BFF936E5DFE669F73FD88FCC0" box="[442,554,861,884]" italics="true" pageId="7" pageNumber="6">iniquipes</emphasis>
</taxonomicName>
/ Paulsen /
<typeStatus id="5493882BFF936E5DFD659F73FCF6FCC1" box="[697,852,861,885]" pageId="7" pageNumber="6" type="holotype">HOLOTYPE</typeStatus>
” (
<figureCitation id="13132A0CFF936E5DFCB29F73FC64FCC1" box="[878,966,861,885]" captionStart="Figure 3" captionStartId="8.[192,275,1383,1405]" captionTargetBox="[201,1420,250,1299]" captionTargetId="figure@8.[192,1440,236,1346]" captionTargetPageId="8" captionText="Figure 3. Ochodaeus iniquipes Paulsen, new species. a) Dorsal habitus, male holotype. b) Right front leg, dorsal view. Arrow indicating median tooth of anterior margin of profemur. c) Right stridulatory peg, dorsal view (elytra lifted). d) Holotype labels." figureDoi="http://doi.org/10.5281/zenodo.3673409" httpUri="https://zenodo.org/record/3673409/files/figure.png" pageId="7" pageNumber="6">Fig. 3d</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B973689FF936E5DFF349F53FED8FBA1" blockId="7.[192,1441,765,1078]" pageId="7" pageNumber="6">
<materialsCitation id="3B403CD4FF936E5DFF349F53FB32FC61" ID-GBIF-Occurrence="2565952375" collectingDate="2003-11-26" collectingDateMax="2003-12-02" collectingDateMin="2003-11-26" collectionCode="R, DKC" collectorName="M. Irwin &amp; S. Murzin &amp; A. Sharnaev" county="Camp Catta" elevation="120" location="Calif. Acad. of Sciences" municipality="Madagascar" pageId="7" pageNumber="6" specimenCount="1" typeStatus="paratype">
Female
<typeStatus id="5493882BFF936E5DFE969F50FE15FC22" box="[330,439,894,918]" pageId="7" pageNumber="6" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED39AE4CFF936E5DFE1A9F53FDB6FC21" box="[454,532,893,917]" pageId="7" pageNumber="6">MJPC</collectionCode>
) labeled: a) “
<collectingCountry id="F33F7619FF936E5DFD6D9F53FCDEFC21" box="[689,892,893,917]" name="Madagascar" pageId="7" pageNumber="6">MADAGASCAR</collectingCountry>
: Tulear / Tsimanampetsotsa National / Park, Mitoho Forest /
<date id="FF961049FF936E5DFE559FB3FD09FC01" box="[393,683,925,949]" pageId="7" pageNumber="6" value="2009-03-25" valueMax="2009-04-02" valueMin="2009-03-25">25 March–2 April 2009</date>
/
<geoCoordinate id="EE1C504EFF936E5DFD1A9FB3FCE9FC01" box="[710,843,924,949]" degrees="24" direction="south" minutes="2.91" orientation="latitude" pageId="7" pageNumber="6" precision="9" value="-24.0485">24° 2.91′ S</geoCoordinate>
<geoCoordinate id="EE1C504EFF936E5DFC899FB3FC4EFC01" box="[853,1004,924,949]" degrees="43" direction="east" minutes="45.14" orientation="longitude" pageId="7" pageNumber="6" precision="9" value="43.752335">43° 45.14′ E</geoCoordinate>
”; b) “
<location id="8EF76052FF936E5DFBEF9FB3FAF6FC01" LSID="urn:lsid:plazi:treatment:0381879FFF936E52FF1C9EE2FB65F83D:8EF76052FF936E5DFBEF9FB3FAF6FC01" box="[1075,1364,925,949]" county="Camp Catta" municipality="Madagascar" name="Calif. Acad. of Sciences" pageId="7" pageNumber="6">Calif. Acad. of Sciences</location>
/ coll:
<collectorName id="26DD535FFF936E5DFF1C9F90FE8CFC61" box="[192,302,957,981]" pageId="7" pageNumber="6">M. Irwin</collectorName>
,
<collectionCode id="ED39AE4CFF936E5DFEE79F90FEF0FC61" box="[315,338,958,981]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="6">R</collectionCode>
.
<location id="8EF76052FF936E5DFE819F93FE04FC61" LSID="urn:lsid:plazi:treatment:0381879FFF936E52FF1C9EE2FB65F83D:8EF76052FF936E5DFE819F93FE04FC61" box="[349,422,957,981]" county="Camp Catta" municipality="Madagascar" name="Harin" pageId="7" pageNumber="6">Harin</location>
›
<location id="8EF76052FF936E5DFE729F93FE48FC61" LSID="urn:lsid:plazi:treatment:0381879FFF936E52FF1C9EE2FB65F83D:8EF76052FF936E5DFE729F93FE48FC61" box="[430,490,957,981]" county="Camp Catta" municipality="Madagascar" name="Hala" pageId="7" pageNumber="6">Hala</location>
/ malaise, transitional forest / elev
<quantity id="4CD09B6CFF936E5DFC419F93FC4BFC61" box="[925,1001,957,981]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="6" unit="m" value="120.0">
<elevation id="0005D1BAFF936E5DFC419F93FC4BFC61" box="[925,1001,957,981]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="6" unit="m" value="120.0">120 m</elevation>
</quantity>
MG-53C-21”.
</materialsCitation>
<materialsCitation id="3B403CD4FF936E5DFB4B9F93FED4FBA1" ID-GBIF-Occurrence="2565952384" collectingDate="2003-11-26" collectingDateMax="2003-12-02" collectingDateMin="2003-11-26" collectionCode="R, DKC" collectorName="M. Irwin &amp; S. Murzin &amp; A. Sharnaev" county="Camp Catta" elevation="120" location="Calif. Acad. of Sciences" municipality="Madagascar" pageId="7" pageNumber="6" specimenCount="1" typeStatus="paratype">
<location id="8EF76052FF936E5DFB4B9F93FB76FC61" LSID="urn:lsid:plazi:treatment:0381879FFF936E52FF1C9EE2FB65F83D:8EF76052FF936E5DFB4B9F93FB76FC61" box="[1175,1236,957,981]" county="Camp Catta" municipality="Madagascar" name="Male" pageId="7" pageNumber="6">Male</location>
<typeStatus id="5493882BFF936E5DFB009F90FAEBFC62" box="[1244,1353,958,982]" pageId="7" pageNumber="6" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED39AE4CFF936E5DFA869F93FA3BFC61" box="[1370,1433,957,981]" pageId="7" pageNumber="6">DKC</collectionCode>
) labeled: a) “
<collectingMunicipality id="6BF3ACF3FF936E5DFE8D9FF3FE44FC41" box="[337,486,989,1013]" pageId="7" pageNumber="6">Madagascar</collectingMunicipality>
/
<collectingCounty id="62F64E05FF936E5DFE219FF3FD31FC41" box="[509,659,989,1013]" pageId="7" pageNumber="6">Camp Catta</collectingCounty>
<locationDeviation id="9A252FABFF936E5DFD479FF3FC30FC41" box="[667,914,989,1013]" location="Ambalavao" pageId="7" pageNumber="6">
<quantity id="4CD09B6CFF936E5DFD479FF3FD45FC41" box="[667,743,989,1013]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="6" unit="km" value="40.0">40 km</quantity>
S Ambalavao
</locationDeviation>
/
<quantity id="4CD09B6CFF936E5DFC749FF3FC54FC41" box="[936,1014,989,1013]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="7" pageNumber="6" unit="m" value="810.0">
<elevation id="0005D1BAFF936E5DFC749FF3FC54FC41" box="[936,1014,989,1013]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="7" pageNumber="6" unit="m" value="810.0">810 m</elevation>
</quantity>
,
<date id="FF961049FF936E5DFBDF9FF3FB6DFC41" box="[1027,1231,989,1013]" pageId="7" pageNumber="6" value="2003-11-26" valueMax="2003-12-02" valueMin="2003-11-26">
<collectingDate id="EFD2E9A1FF936E5DFBDF9FF3FB6DFC41" box="[1027,1231,989,1013]" pageId="7" pageNumber="6" value="2003-11-26" valueMax="2003-12-02" valueMin="2003-11-26">26.XI-2.XII.2003</collectingDate>
</date>
/
<collectorName id="26DD535FFF936E5DFB3A9FF3FAC2FC41" box="[1254,1376,989,1013]" pageId="7" pageNumber="6">S. Murzin</collectorName>
&amp;
<collectorName id="26DD535FFF936E5DFA599FF3FE94FBA1" pageId="7" pageNumber="6">A. Sharnaev</collectorName>
leg.”
</materialsCitation>
.
</paragraph>
<paragraph id="8B973689FF936E5DFF349833FA03FB81" blockId="7.[192,1441,765,1078]" box="[232,1441,1053,1078]" pageId="7" pageNumber="6">
<materialsCitation id="3B403CD4FF936E5DFF349833FA3FFB81" ID-GBIF-Occurrence="2565952383" box="[232,1437,1053,1078]" pageId="7" pageNumber="6" typeStatus="paratype">
Both
<typeStatus id="5493882BFF936E5DFEF69830FE06FB82" box="[298,420,1054,1078]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
(
<emphasis id="B95CEA9BFF936E5DFE689833FE66FB80" box="[436,452,1053,1076]" italics="true" pageId="7" pageNumber="6">n</emphasis>
= 2) with label: on yellow paper, “
<taxonomicName id="4C284D0AFF936E5DFCBD9833FC44FB81" authorityName="Dejean" authorityYear="1821" box="[865,998,1053,1077]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEA9BFF936E5DFCBD9833FC44FB81" box="[865,998,1053,1077]" italics="true" pageId="7" pageNumber="6">Ochodaeus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C284D0AFF936E5DFC209833FBCEFB80" authorityName="Paulsen" authorityYear="2019" box="[1020,1132,1053,1076]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="iniquipes">
<emphasis id="B95CEA9BFF936E5DFC209833FBCEFB80" box="[1020,1132,1053,1076]" italics="true" pageId="7" pageNumber="6">iniquipes</emphasis>
</taxonomicName>
/ Paulsen /
<typeStatus id="5493882BFF936E5DFB269833FA31FB81" box="[1274,1427,1053,1077]" pageId="7" pageNumber="6" type="paratype">PARATYPE</typeStatus>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3326502FF936E5DFF1C9863FBDDF8C1" pageId="7" pageNumber="6" type="description">
<paragraph id="8B973689FF936E5DFF1C9863FAC9F8B1" blockId="7.[192,1441,1101,1797]" pageId="7" pageNumber="6">
<emphasis id="B95CEA9BFF936E5DFF1C9863FEC8FBD1" bold="true" box="[192,362,1101,1125]" pageId="7" pageNumber="6">Description.</emphasis>
<typeStatus id="5493882BFF936E5DFEB39863FE7FFBD1" box="[367,477,1101,1125]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="13132A0CFF936E5DFDF19863FD20FBD1" box="[557,642,1101,1125]" captionStart="Figure 3" captionStartId="8.[192,275,1383,1405]" captionTargetBox="[201,1420,250,1299]" captionTargetId="figure@8.[192,1440,236,1346]" captionTargetPageId="8" captionText="Figure 3. Ochodaeus iniquipes Paulsen, new species. a) Dorsal habitus, male holotype. b) Right front leg, dorsal view. Arrow indicating median tooth of anterior margin of profemur. c) Right stridulatory peg, dorsal view (elytra lifted). d) Holotype labels." figureDoi="http://doi.org/10.5281/zenodo.3673409" httpUri="https://zenodo.org/record/3673409/files/figure.png" pageId="7" pageNumber="6">Fig. 3a</figureCitation>
).
<taxonomicName id="4C284D0AFF936E5DFD4A9863FCBCFBD1" authorityName="Linnaeus" authorityYear="1758" box="[662,798,1101,1125]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
: Scarabaeoidea:
<taxonomicName id="4C284D0AFF936E5DFC3A9863FB25FBD1" authorityName="Mulsant &amp; Rey" authorityYear="1871" box="[998,1159,1101,1125]" class="Insecta" family="Ochodaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="family">Ochodaeidae</taxonomicName>
. Length
<quantity id="4CD09B6CFF936E5DFB319863FAE9FBD1" box="[1261,1355,1101,1125]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="6" unit="mm" value="7.6">7.6 mm</quantity>
. Width
<quantity id="4CD09B6CFF936E5DFF1C9843FEBCFB31" box="[192,286,1133,1157]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="7" pageNumber="6" unit="mm" value="4.1">4.1 mm</quantity>
.
<emphasis id="B95CEA9BFF936E5DFEF49843FEDEFB31" bold="true" box="[296,380,1133,1157]" pageId="7" pageNumber="6">Color:</emphasis>
Everywhere reddish brown, surface shiny.
<emphasis id="B95CEA9BFF936E5DFC569843FC78FB31" bold="true" box="[906,986,1133,1157]" pageId="7" pageNumber="6">Head:</emphasis>
Surface strongly granulate; granules setose, shiny; surface between granules shagreened. Eyes large, globose, lacking ventral projection of canthus. Antennal club moderately large, approximately size of eye in distal view. Mandibles broadly concave, basal angle broadly rounded, apices falcate; left mandible with triangular internal tooth and second tooth behind; right mandible with internal teeth weakly developed. Frons weakly convex. Frontoclypeal suture triangular (projecting posteriorly, not transverse). Clypeus rhomboidal, long (about 1/2 as long as wide), with small horn on posterior margin; anterior margin distinctly beaded throughout, bead slender, bisinuate. Labrum with deep, U-shaped emargination. Mentum slightly wider than long, anteriorly emarginate with broad, subtriangular fovea.
<emphasis id="B95CEA9BFF936E5DFC599943FBB8FA31" bold="true" box="[901,1050,1389,1413]" pageId="7" pageNumber="6">Pronotum:</emphasis>
Form convex. Surface densely granulate everywhere (except furrowed midline in basal fourth); granules shiny, with short seta; surface shagreened between granules.
<emphasis id="B95CEA9BFF936E5DFDA89983FD71FA71" bold="true" box="[628,723,1453,1477]" pageId="7" pageNumber="6">Elytra:</emphasis>
Form convex, distinctly narrower than pronotum, slightly elongate, elytra together as long as wide. Surface with striae moderately impressed, punctate, surface smooth between punctures (except sutural stria with connecting groove); strial punctures large, separated by ~1 puncture diameter, lacking setae. Intervals irregularly tuberculate; tubercles uniformly small, setose; setae slightly longer than diameter of largest strial punctures.
<emphasis id="B95CEA9BFF936E5DFBFB9A03FBCDF9F1" bold="true" box="[1063,1135,1581,1605]" pageId="7" pageNumber="6">Legs:</emphasis>
Protibia broad (
<figureCitation id="13132A0CFF936E5DFAE69A03FA33F9F1" box="[1338,1425,1581,1605]" captionStart="Figure 3" captionStartId="8.[192,275,1383,1405]" captionTargetBox="[201,1420,250,1299]" captionTargetId="figure@8.[192,1440,236,1346]" captionTargetPageId="8" captionText="Figure 3. Ochodaeus iniquipes Paulsen, new species. a) Dorsal habitus, male holotype. b) Right front leg, dorsal view. Arrow indicating median tooth of anterior margin of profemur. c) Right stridulatory peg, dorsal view (elytra lifted). d) Holotype labels." figureDoi="http://doi.org/10.5281/zenodo.3673409" httpUri="https://zenodo.org/record/3673409/files/figure.png" pageId="7" pageNumber="6">Fig. 3b</figureCitation>
), tridentate externally, with moderately long pollex. Profemur with strong, apical and median teeth (
<figureCitation id="13132A0CFF936E5DFAAC9A63FF7DF931" captionStart="Figure 3" captionStartId="8.[192,275,1383,1405]" captionTargetBox="[201,1420,250,1299]" captionTargetId="figure@8.[192,1440,236,1346]" captionTargetPageId="8" captionText="Figure 3. Ochodaeus iniquipes Paulsen, new species. a) Dorsal habitus, male holotype. b) Right front leg, dorsal view. Arrow indicating median tooth of anterior margin of profemur. c) Right stridulatory peg, dorsal view (elytra lifted). d) Holotype labels." figureDoi="http://doi.org/10.5281/zenodo.3673409" httpUri="https://zenodo.org/record/3673409/files/figure.png" pageId="7" pageNumber="6">Fig. 3b</figureCitation>
). Mesofemur and metafemur with apical teeth large, acute. Metatibia flattened, dorsal surface with internal margin blade-like basally.
<emphasis id="B95CEA9BFF936E5DFDAC9AA3FCFCF911" bold="true" box="[624,862,1677,1701]" pageId="7" pageNumber="6">Venter/Abdomen:</emphasis>
Metasternum and abdomen sparsely punctate; punctures large with long, golden setae. Last abdominal segment with erect setae. Pygidium rugopunctate, setose, setae sparse. Stridulatory peg oblong (
<figureCitation id="13132A0CFF936E5DFCC49AE3FCCBF951" box="[792,873,1741,1765]" captionStart="Figure 3" captionStartId="8.[192,275,1383,1405]" captionTargetBox="[201,1420,250,1299]" captionTargetId="figure@8.[192,1440,236,1346]" captionTargetPageId="8" captionText="Figure 3. Ochodaeus iniquipes Paulsen, new species. a) Dorsal habitus, male holotype. b) Right front leg, dorsal view. Arrow indicating median tooth of anterior margin of profemur. c) Right stridulatory peg, dorsal view (elytra lifted). d) Holotype labels." figureDoi="http://doi.org/10.5281/zenodo.3673409" httpUri="https://zenodo.org/record/3673409/files/figure.png" pageId="7" pageNumber="6">Fig. 3c</figureCitation>
).
<emphasis id="B95CEA9BFF936E5DFCA09AE3FBE1F951" bold="true" box="[892,1091,1741,1765]" pageId="7" pageNumber="6">Male genitalia:</emphasis>
Sclerotized patch on internal sac between parameres broadly triangular, apex rounded; internal sac lacking complex armature.
</paragraph>
<paragraph id="8B973689FF936E5DFF1C9B33FBDDF8C1" blockId="7.[192,1440,1821,1910]" pageId="7" pageNumber="6">
<emphasis id="B95CEA9BFF936E5DFF1C9B33FE6FF881" bold="true" box="[192,461,1821,1845]" pageId="7" pageNumber="6">
<typeStatus id="5493882BFF936E5DFF1C9B33FE9DF881" box="[192,319,1821,1845]" pageId="7" pageNumber="6" type="paratype">Paratype</typeStatus>
variation.
</emphasis>
Male (
<emphasis id="B95CEA9BFF936E5DFDFC9B33FD92F880" box="[544,560,1821,1844]" italics="true" pageId="7" pageNumber="6">n</emphasis>
= 1; length
<quantity id="4CD09B6CFF936E5DFD639B33FCBFF881" box="[703,797,1821,1845]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="6" unit="mm" value="7.4">7.4 mm</quantity>
; width 4.0 mm). Female (
<emphasis id="B95CEA9BFF936E5DFB8D9B33FBC3F880" box="[1105,1121,1821,1844]" italics="true" pageId="7" pageNumber="6">n</emphasis>
= 1; length
<quantity id="4CD09B6CFF936E5DFB2D9B33FAECF881" box="[1265,1358,1821,1845]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="7" pageNumber="6" unit="mm" value="8.2">8.2 mm</quantity>
; width
<quantity id="4CD09B6CFF936E5DFF1C9B13FEB9F8E1" box="[192,283,1853,1877]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="6" unit="mm" value="4.5">4.5 mm</quantity>
). The female specimen has much narrower protibiae, the median tooth on the profemur is reduced, and the setae on the last abdominal segment are recumbent rather than erect.
</paragraph>
</subSubSection>
<caption id="DF576601FF9C6E52FF1C9949FE43FA03" ID-DOI="http://doi.org/10.5281/zenodo.3673409" ID-Zenodo-Dep="3673409" httpUri="https://zenodo.org/record/3673409/files/figure.png" pageId="8" pageNumber="7" startId="8.[192,275,1383,1405]" targetBox="[201,1420,250,1299]" targetPageId="8">
<paragraph id="8B973689FF9C6E52FF1C9949FE43FA03" blockId="8.[192,1440,1383,1463]" pageId="8" pageNumber="7">
<emphasis id="B95CEA9BFF9C6E52FF1C9949FE8DFAC8" bold="true" box="[192,303,1383,1405]" pageId="8" pageNumber="7">Figure 3.</emphasis>
<taxonomicName id="4C284D0AFF9C6E52FEEA9949FDDEFAC9" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[310,636,1383,1405]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="iniquipes" status="sp. nov.">
<emphasis id="B95CEA9BFF9C6E52FEEA9949FE0CFAC8" box="[310,430,1383,1404]" italics="true" pageId="8" pageNumber="7">Ochodaeus</emphasis>
<emphasis id="B95CEA9BFF9C6E52FE699946FDB8FAC9" box="[437,538,1384,1405]" italics="true" pageId="8" pageNumber="7">iniquipes</emphasis>
Paulsen
</taxonomicName>
,
<taxonomicNameLabel id="A26F57E0FF9C6E52FD5B9946FCAEFAC9" box="[647,780,1383,1405]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="B95CEA9BFF9C6E52FCCA9949FC8CFAC9" bold="true" box="[790,814,1383,1405]" pageId="8" pageNumber="7">a)</emphasis>
Dorsal habitus, male holotype.
<emphasis id="B95CEA9BFF9C6E52FB519949FB04FAC9" bold="true" box="[1165,1190,1383,1405]" pageId="8" pageNumber="7">b)</emphasis>
Right front leg, dorsal view. Arrow indicating median tooth of anterior margin of profemur.
<emphasis id="B95CEA9BFF9C6E52FC6A99AAFC6EFA2E" bold="true" box="[950,972,1412,1434]" pageId="8" pageNumber="7">c)</emphasis>
Right stridulatory peg, dorsal view (elytra lifted).
<emphasis id="B95CEA9BFF9C6E52FED3998EFE8AFA02" bold="true" box="[271,296,1440,1462]" pageId="8" pageNumber="7">d)</emphasis>
Holotype labels.
</paragraph>
</caption>
<subSubSection id="C3326502FF9C6E52FF1C99DEFE26F95C" pageId="8" pageNumber="7" type="discussion">
<paragraph id="8B973689FF9C6E52FF1C99DEFE26F95C" blockId="8.[192,1440,1520,1768]" pageId="8" pageNumber="7">
<emphasis id="B95CEA9BFF9C6E52FF1C99DEFEE1F9BC" bold="true" box="[192,323,1520,1544]" pageId="8" pageNumber="7">Remarks.</emphasis>
<taxonomicName id="4C284D0AFF9C6E52FE9699DEFDE4F9BC" authorityName="Paulsen" authorityYear="2019" box="[330,582,1520,1544]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="iniquipes">
<emphasis id="B95CEA9BFF9C6E52FE9699DEFDE4F9BC" box="[330,582,1520,1544]" italics="true" pageId="8" pageNumber="7">Ochodaeus iniquipes</emphasis>
</taxonomicName>
has the clypeal armature most strongly developed, clearly referable to as a small horn rather than a shiny tubercle or bump. The frontoclypeal suture is produced posteriorly to the horn, which lies on the suture itself, whereas in
<taxonomicName id="4C284D0AFF9C6E52FCB59A1EFBB5F9FC" authorityName="Paulsen" authorityYear="2019" box="[873,1047,1584,1608]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="umbonulus">
<emphasis id="B95CEA9BFF9C6E52FCB59A1EFBB5F9FC" box="[873,1047,1584,1608]" italics="true" pageId="8" pageNumber="7">O. umbonulus</emphasis>
</taxonomicName>
the suture is straight. The profemur has an anteromedian tooth in all three known specimens, although it is reduced in the female
<typeStatus id="5493882BFF9C6E52FF1C9A5FFE93F93D" box="[192,305,1649,1673]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
. This tooth would presumably be partially or entirely obsolete in smaller individuals, and the largest specimens of
<taxonomicName id="4C284D0AFF9C6E52FE619ABEFDCBF91C" authorityName="Paulsen" authorityYear="2019" box="[445,617,1680,1704]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="umbonulus">
<emphasis id="B95CEA9BFF9C6E52FE619ABEFDCBF91C" box="[445,617,1680,1704]" italics="true" pageId="8" pageNumber="7">O. umbonulus</emphasis>
</taxonomicName>
have the median tooth weakly indicated. In such a case, the clypeal character will better serve to distinguish the two species. The deep U-shaped labrum is also diagnostic for
<taxonomicName id="4C284D0AFF9C6E52FF359AFEFEDEF95C" authorityName="Paulsen" authorityYear="2019" box="[233,380,1744,1768]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="iniquipes">
<emphasis id="B95CEA9BFF9C6E52FF359AFEFEDEF95C" box="[233,380,1744,1768]" italics="true" pageId="8" pageNumber="7">O. iniquipes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3326502FF9C6E52FF1C9B2EFCAEF8EC" pageId="8" pageNumber="7" type="etymology">
<paragraph id="8B973689FF9C6E52FF1C9B2EFCAEF8EC" blockId="8.[192,1440,1792,1880]" pageId="8" pageNumber="7">
<emphasis id="B95CEA9BFF9C6E52FF1C9B2EFEF9F8AC" bold="true" box="[192,347,1792,1816]" pageId="8" pageNumber="7">Etymology.</emphasis>
The name is a Latin noun in the nominative singular derived from ‘iniquus’, dangerous or evil, with “pes”, legs. The evil legs are the heavily armed profemora, with a large apical tooth and also a median tooth on the anterior margin (
<figureCitation id="13132A0CFF9C6E52FD7B9B6EFD5CF8EC" box="[679,766,1856,1880]" captionStart="Figure 3" captionStartId="8.[192,275,1383,1405]" captionTargetBox="[201,1420,250,1299]" captionTargetId="figure@8.[192,1440,236,1346]" captionTargetPageId="8" captionText="Figure 3. Ochodaeus iniquipes Paulsen, new species. a) Dorsal habitus, male holotype. b) Right front leg, dorsal view. Arrow indicating median tooth of anterior margin of profemur. c) Right stridulatory peg, dorsal view (elytra lifted). d) Holotype labels." figureDoi="http://doi.org/10.5281/zenodo.3673409" httpUri="https://zenodo.org/record/3673409/files/figure.png" pageId="8" pageNumber="7">Fig. 3b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3326502FF9C6E52FF1C9B5EFB65F83D" box="[192,1223,1904,1929]" pageId="8" pageNumber="7" type="distribution">
<paragraph id="8B973689FF9C6E52FF1C9B5EFB65F83D" blockId="8.[192,1223,1904,1929]" box="[192,1223,1904,1929]" pageId="8" pageNumber="7">
<emphasis id="B95CEA9BFF9C6E52FF1C9B5EFED1F83C" bold="true" box="[192,371,1904,1928]" pageId="8" pageNumber="7">Distribution.</emphasis>
This species is distributed in the southern third of the island (
<figureCitation id="13132A0CFF9C6E52FBAD9B5EFB1BF83D" box="[1137,1209,1904,1929]" captionStart="Figures 9–10" captionStartId="10.[192,287,1877,1899]" captionTargetBox="[192,1440,864,1839]" captionTargetId="figure@10.[192,1440,864,1839]" captionTargetPageId="10" captionText="Figures 9–10. Distributions of new species of Madagascar Ochodaeus. 9) Blue squares = O. iniquipes Paulsen, new species; orange circle = O. meridialis Paulsen, new species. 10) White circles = O. umbonulus Paulsen, new species." figureDoi="http://doi.org/10.5281/zenodo.3673413" httpUri="https://zenodo.org/record/3673413/files/figure.png" pageId="8" pageNumber="7">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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