274 lines
34 KiB
XML
274 lines
34 KiB
XML
<document id="6740F29C8E9228A98276102189195A39" ID-DOI="10.1080/00222933.2017.1324050" ID-ISSN="1464-5262" ID-Zenodo-Dep="5181888" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628698566541" checkinUser="felipe" docAuthor="Schmitter-Soto, Juan J." docDate="2017" docId="03806F39C963FFD1FE2EFA34D224FC06" docLanguage="en" docName="JNatHist.51.23-24.1331-1424.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 51 (21 - 24)" docSource="http://dx.doi.org/10.1080/00222933.2017.1324050" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Astyanax dorioni Schmitter-Soto 2017, comb. nov." docType="treatment" docVersion="4" lastPageNumber="1384" masterDocId="FFB91741C957FFE7FFBEFFB5D316FFA7" masterDocTitle="A revision of Astyanax (Characiformes: Characidae) in Central and North America, with the description of nine new species" masterLastPageNumber="1424" masterPageNumber="1331" pageNumber="1382" updateTime="1699050015946" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3BDFFAC8158FCF2DE53E0BA6D1D194E6">A revision of Astyanax (Characiformes: Characidae) in Central and North America, with the description of nine new species</mods:title>
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<mods:namePart id="8532BCC995AD493F0AD2B39F8D36E4C2">Schmitter-Soto, Juan J.</mods:namePart>
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<mods:title id="948D0153AAEE8C08651F7EA1CC3E2593">Journal of Natural History</mods:title>
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<mods:part id="7C325B6D539743F07DAD7B01EC36369A">
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<mods:date id="FB659A0B2B2DD509C33CFACFC0A0027D">2017</mods:date>
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<mods:title id="A93FC9D12751A8B76D616E475EFB846F">J. Nat. Hist.</mods:title>
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<mods:number id="5B68756C565080FC506634D9B2175581">2017-07-03</mods:number>
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<mods:number id="E1A789D665189C2B8A4F757DB83BE8F0">51</mods:number>
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<mods:number id="541667C7BDE668713DB9844E9E9964ED">21 - 24</mods:number>
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<mods:url id="8702D6D433FBFE8B6C00CC382F402BD5">http://dx.doi.org/10.1080/00222933.2017.1324050</mods:url>
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<treatment id="03806F39C963FFD1FE2EFA34D224FC06" ID-DOI="http://doi.org/10.5281/zenodo.5191156" ID-Zenodo-Dep="5191156" LSID="urn:lsid:plazi:treatment:03806F39C963FFD1FE2EFA34D224FC06" httpUri="http://treatment.plazi.org/id/03806F39C963FFD1FE2EFA34D224FC06" lastPageId="54" lastPageNumber="1384" pageId="52" pageNumber="1382">
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<subSubSection id="C3338DA4C963FFD3FE2EFA34D082FA3C" box="[400,916,1409,1435]" pageId="52" pageNumber="1382" type="nomenclature">
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<paragraph id="8B96DE2FC963FFD3FE2EFA34D082FA3C" blockId="52.[285,1030,1409,1470]" box="[400,916,1409,1435]" pageId="52" pageNumber="1382">
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<heading id="D0DE6943C963FFD3FE2EFA34D082FA3C" box="[400,916,1409,1435]" centered="true" fontSize="11" level="2" pageId="52" pageNumber="1382" reason="2">
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<taxonomicName id="4C29A5ACC963FFD3FE2EFA34D010FA3C" authority="(Rosen 1970)" authorityName="Schmitter-Soto" authorityYear="2017" baseAuthorityName="Rosen" baseAuthorityYear="1970" box="[400,774,1409,1435]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="52" pageNumber="1382" phylum="Chordata" rank="species" species="dorioni" status="comb. nov.">
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<emphasis id="B95D023DC963FFD3FE2EFA34D175FA3C" bold="true" box="[400,611,1409,1435]" italics="true" pageId="52" pageNumber="1382">Astyanax dorioni</emphasis>
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(
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<bibRefCitation id="EFB8A3DEC963FFD3FDCCFA37D1EBFA3C" author="Rosen DE" box="[626,765,1410,1435]" pageId="52" pageNumber="1382" pagination="1 - 17" refId="ref48940" refString="Rosen DE. 1970. A new tetragonopterine characid fish from Guatemala. Am Mus Nov. 2435: 1 - 17." type="journal article" year="1970">Rosen 1970</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A26EBF46C963FFD3FCB1FA37D082FA3C" box="[783,916,1410,1435]" pageId="52" pageNumber="1382" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3338DA4C963FFD3FEA3FA10D710FA19" box="[285,1030,1445,1470]" pageId="52" pageNumber="1382" type="description">
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<paragraph id="8B96DE2FC963FFD3FEA3FA10D710FA19" blockId="52.[285,1030,1409,1470]" box="[285,1030,1445,1470]" pageId="52" pageNumber="1382">
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(
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<figureCitation id="1312C2AAC963FFD3FE9BFA10D280FA19" box="[293,406,1445,1470]" captionStart="Figure 24" captionStartId="37.[160,225,1645,1668]" captionTargetBox="[163,1151,941,1611]" captionTargetId="figure-183@37.[163,1153,941,1614]" captionTargetPageId="37" captionText="Figure 24. Map of northern Central America, showing distribution localities of Astyanax baileyi, A. belizianus, A. brevimanus, A. cubilhuitz sp. nov., A. dorioni and A. petenensis. A larger symbol denotes the type locality." figureDoi="http://doi.org/10.5281/zenodo.5181937" httpUri="https://zenodo.org/record/5181937/files/figure.png" pageId="52" pageNumber="1382">Figure 24</figureCitation>
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; see original description for illustration of
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<typeStatus id="5492608DC963FFD3FC2BFA10D716FA19" box="[917,1024,1445,1470]" pageId="52" pageNumber="1382" type="holotype">holotype</typeStatus>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3338DA4C963FFD3FF1EFA5AD177F98D" pageId="52" pageNumber="1382" type="reference_group">
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<paragraph id="8B96DE2FC963FFD3FF1EFA5AD1A5F9A0" blockId="52.[160,691,1519,1578]" box="[160,691,1519,1543]" pageId="52" pageNumber="1382">
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<treatmentCitationGroup id="AB39F901C963FFD3FF1EFA5AD1A5F9A0" box="[160,691,1519,1543]" pageId="52" pageNumber="1382">
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<treatmentCitation id="0A88F83EC963FFD3FF1EFA5AD1B9F9A0" author="Rosen DE" box="[160,687,1519,1543]" page="3" pageId="52" pageNumber="1382" year="1970">
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<taxonomicName id="4C29A5ACC963FFD3FF1EFA5AD1B9F9A0" ID-CoL="N6FF" authority="Rosen 1970: 3" authorityName="Rosen" authorityPageNumber="3" authorityYear="1970" box="[160,687,1519,1543]" class="Actinopterygii" family="Characidae" genus="Bramocharax" kingdom="Animalia" order="Characiformes" pageId="52" pageNumber="1382" phylum="Chordata" rank="subSpecies" species="bransfordii" subSpecies="dorioni">
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<emphasis id="B95D023DC963FFD3FF1EFA5AD113F9A0" box="[160,517,1519,1543]" italics="true" pageId="52" pageNumber="1382">Bramocharax bransfordii dorioni</emphasis>
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<bibRefCitation id="EFB8A3DEC963FFD3FDB3FA5AD1B9F9A0" author="Rosen DE" box="[525,687,1519,1543]" pageId="52" pageNumber="1382" pagination="1 - 17" refId="ref48940" refString="Rosen DE. 1970. A new tetragonopterine characid fish from Guatemala. Am Mus Nov. 2435: 1 - 17." type="journal article" year="1970">Rosen 1970: 3</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B96DE2FC963FFD3FF1EF9A7D177F98D" blockId="52.[160,691,1519,1578]" box="[160,609,1554,1578]" pageId="52" pageNumber="1382">
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<taxonomicName id="4C29A5ACC963FFD3FF1EF9A7D105F98D" ID-CoL="N6FF" authority="Rosen 1970" authorityName="Rosen" authorityYear="1970" box="[160,531,1554,1578]" class="Actinopterygii" family="Characidae" genus="Bramocharax" kingdom="Animalia" order="Characiformes" pageId="52" pageNumber="1382" phylum="Chordata" rank="species" species="dorioni">
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<emphasis id="B95D023DC963FFD3FF1EF9A7D291F98D" box="[160,391,1554,1578]" italics="true" pageId="52" pageNumber="1382">Bramocharax dorioni</emphasis>
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<bibRefCitation id="EFB8A3DEC963FFD3FE2EF9A7D105F98D" author="Rosen DE" box="[400,531,1554,1578]" pageId="52" pageNumber="1382" pagination="1 - 17" refId="ref48940" refString="Rosen DE. 1970. A new tetragonopterine characid fish from Guatemala. Am Mus Nov. 2435: 1 - 17." type="journal article" year="1970">Rosen 1970</bibRefCitation>
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</taxonomicName>
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, auctt.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3338DA4C963FFD2FF1EF9E3D185FEF3" lastPageId="53" lastPageNumber="1383" pageId="52" pageNumber="1382" type="diagnosis">
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<paragraph id="8B96DE2FC963FFD3FF1EF9E3D202F9C8" blockId="52.[160,1155,1622,1717]" box="[160,276,1622,1647]" pageId="52" pageNumber="1382">
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<emphasis id="B95D023DC963FFD3FF1EF9E3D202F9C8" bold="true" box="[160,276,1622,1647]" italics="true" pageId="52" pageNumber="1382">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8B96DE2FC963FFD2FF1EF9CFD185FEF3" blockId="52.[160,1155,1622,1717]" lastBlockId="53.[160,1156,142,340]" lastPageId="53" lastPageNumber="1383" pageId="52" pageNumber="1382">
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Diagnosed from other
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<taxonomicName id="4C29A5ACC963FFD3FE0AF9CFD160F935" box="[436,630,1658,1682]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="52" pageNumber="1382" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B95D023DC963FFD3FE0AF9CFD101F935" box="[436,535,1658,1682]" italics="true" pageId="52" pageNumber="1382">Astyanax</emphasis>
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species
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</taxonomicName>
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in central-eastern
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<collectingCountry id="F33E9EBFC963FFD3FCE3F9CFD0C0F935" box="[861,982,1658,1682]" name="Guatemala" pageId="52" pageNumber="1382">Guatemala</collectingCountry>
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and
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<collectingCountry id="F33E9EBFC963FFD3FBA3F9CFD748F935" box="[1053,1118,1658,1682]" name="Belize" pageId="52" pageNumber="1382">Belize</collectingCountry>
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as follows: interorbital space, shorter than eye diameter (vs longer, except in
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<taxonomicName id="4C29A5ACC963FFD3FBAEF928D760F912" authorityName="Schmitter-Soto" authorityYear="2017" baseAuthorityName="Rosen" baseAuthorityYear="1972" box="[1040,1142,1693,1717]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="52" pageNumber="1382" phylum="Chordata" rank="species" species="baileyi">
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<emphasis id="B95D023DC963FFD3FBAEF928D760F912" box="[1040,1142,1693,1717]" italics="true" pageId="52" pageNumber="1382">A. baileyi</emphasis>
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</taxonomicName>
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); postorbital length, mean 17% SL (vs mean less tha 15%); pelvic-fin length, 15% SL or more (15% or less, in
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<taxonomicName id="4C29A5ACC962FFD2FE1EFF04D113FF6E" authorityName="Schmitter-Soto" authorityYear="2017" baseAuthorityName="Rosen" baseAuthorityYear="1972" box="[416,517,177,201]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="baileyi">
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<emphasis id="B95D023DC962FFD2FE1EFF04D113FF6E" box="[416,517,177,201]" italics="true" pageId="53" pageNumber="1383">A. baileyi</emphasis>
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</taxonomicName>
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); supraoccipital in dorsal view short, wide based (long, narrow-based, in
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<taxonomicName id="4C29A5ACC962FFD2FEDBFF61D2DEFF4B" authorityName="Schmitter-Soto" authorityYear="2017" baseAuthorityName="Rosen" baseAuthorityYear="1972" box="[357,456,212,236]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="baileyi">
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<emphasis id="B95D023DC962FFD2FEDBFF61D2DEFF4B" box="[357,456,212,236]" italics="true" pageId="53" pageNumber="1383">A. baileyi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C29A5ACC962FFD2FDBDFF61D19DFF4B" baseAuthorityName="Bocourt" baseAuthorityYear="1868" box="[515,651,212,236]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="belizianus">
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<emphasis id="B95D023DC962FFD2FDBDFF61D19DFF4B" box="[515,651,212,236]" italics="true" pageId="53" pageNumber="1383">A. belizianus</emphasis>
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</taxonomicName>
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); maxillary teeth, more than 9 (4 or fewer in
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<emphasis id="B95D023DC962FFD2FF1EFF43D179FEA9" box="[160,623,246,270]" italics="true" pageId="53" pageNumber="1383">
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<taxonomicName id="4C29A5ACC962FFD2FF1EFF43D23AFEA9" baseAuthorityName="Bocourt" baseAuthorityYear="1868" box="[160,300,246,270]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="belizianus">A. belizianus</taxonomicName>
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,
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<taxonomicName id="4C29A5ACC962FFD2FE84FF43D2CFFEA9" authorityName="Gunther" authorityYear="1864" box="[314,473,246,270]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="brevimanus">A. brevimanus</taxonomicName>
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,
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<taxonomicName id="4C29A5ACC962FFD2FE59FF43D179FEA9" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[487,623,246,270]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="cubilhuitz" status="sp. nov.">A. cubilhuitz</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A26EBF46C962FFD2FDC4FF43D1C0FEA9" box="[634,726,246,270]" pageId="53" pageNumber="1383" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="4C29A5ACC962FFD2FD5CFF43D052FEA9" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[738,836,246,270]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="macal" status="sp. nov.">
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<emphasis id="B95D023DC962FFD2FD5CFF43D052FEA9" box="[738,836,246,270]" italics="true" pageId="53" pageNumber="1383">A. macal</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26EBF46C962FFD2FCF0FF43D0BEFEA9" box="[846,936,246,270]" pageId="53" pageNumber="1383" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C29A5ACC962FFD2FC58FF43D760FEA9" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[998,1142,246,270]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="petenensis">
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<emphasis id="B95D023DC962FFD2FC58FF43D760FEA9" box="[998,1142,246,270]" italics="true" pageId="53" pageNumber="1383">A. petenensis</emphasis>
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</taxonomicName>
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); caudad process of postcleithrum, square with no ventral concavity (no such concavity in
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<taxonomicName id="4C29A5ACC962FFD2FF1EFE89D231FEF3" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[160,295,316,340]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="cubilhuitz" status="sp. nov.">
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<emphasis id="B95D023DC962FFD2FF1EFE89D231FEF3" box="[160,295,316,340]" italics="true" pageId="53" pageNumber="1383">A. cubilhuitz</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26EBF46C962FFD2FE8EFE89D29EFEF3" box="[304,392,316,340]" pageId="53" pageNumber="1383" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C29A5ACC962FFD2FE7AFE89D132FEF3" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[452,548,316,340]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="species" species="macal" status="sp. nov.">
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<emphasis id="B95D023DC962FFD2FE7AFE89D132FEF3" box="[452,548,316,340]" italics="true" pageId="53" pageNumber="1383">A. macal</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26EBF46C962FFD2FD93FE89D19EFEF3" box="[557,648,316,340]" pageId="53" pageNumber="1383" rank="species">sp. nov.</taxonomicNameLabel>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3338DA4C962FFD1FF1EFE35D706FE1B" lastPageId="54" lastPageNumber="1384" pageId="53" pageNumber="1383" type="description">
|
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<paragraph id="8B96DE2FC962FFD2FF1EFE35D256FE3E" blockId="53.[160,1157,384,1726]" box="[160,320,384,409]" pageId="53" pageNumber="1383">
|
||
<emphasis id="B95D023DC962FFD2FF1EFE35D256FE3E" bold="true" box="[160,320,384,409]" italics="true" pageId="53" pageNumber="1383">Redescription</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC962FFD2FF1EFE11D793FE1B" blockId="53.[160,1157,384,1726]" box="[160,1157,420,444]" pageId="53" pageNumber="1383">
|
||
A species of
|
||
<taxonomicName id="4C29A5ACC962FFD2FE92FE11D299FE1B" authorityName="Baird and Girard" authorityYear="1854" box="[300,399,420,444]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95D023DC962FFD2FE92FE11D299FE1B" box="[300,399,420,444]" italics="true" pageId="53" pageNumber="1383">Astyanax</emphasis>
|
||
</taxonomicName>
|
||
, subgenus
|
||
<taxonomicName id="4C29A5ACC962FFD2FDB1FE11D164FE1B" authorityName="Baird and Girard" authorityYear="1854" box="[527,626,420,444]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95D023DC962FFD2FDB1FE11D164FE1B" box="[527,626,420,444]" italics="true" pageId="53" pageNumber="1383">Astyanax</emphasis>
|
||
</taxonomicName>
|
||
(i.e. with a complete predorsal series of scales).
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC962FFD2FF7EFE73D0F5FD83" blockId="53.[160,1157,384,1726]" pageId="53" pageNumber="1383">
|
||
Head profile straight to strongly concave; snout elongated (
|
||
<taxonomicName id="4C29A5ACC962FFD2FC3AFE73D703FE79" authorityName="Gill" authorityYear="1877" box="[900,1045,454,478]" class="Actinopterygii" family="Characidae" genus="Bramocharax" kingdom="Animalia" order="Characiformes" pageId="53" pageNumber="1383" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95D023DC962FFD2FC3AFE73D703FE79" box="[900,1045,454,478]" italics="true" pageId="53" pageNumber="1383">Bramocharax</emphasis>
|
||
</taxonomicName>
|
||
-like); lips even, mouth terminal. Pectoral fins almost always reach pelvic fin origin; anal and dorsal fins usually overlap. Lobes of caudal fin, subequal. See also
|
||
<bibRefCitation id="EFB8A3DEC962FFD2FCF2FDB9D0F6FD83" author="Rosen DE" box="[844,992,524,548]" pageId="53" pageNumber="1383" pagination="1 - 17" refId="ref48940" refString="Rosen DE. 1970. A new tetragonopterine characid fish from Guatemala. Am Mus Nov. 2435: 1 - 17." type="journal article" year="1970">Rosen (1970)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC962FFD2FF7EFD9BD2EBFCD9" blockId="53.[160,1157,384,1726]" pageId="53" pageNumber="1383">
|
||
D. 10–11; A. 23–28 (the original description gives 25–30, including the first small spiniform rays), modally 24; pect. 12–17. Procurrent unsegmented dorsal rays on caudal fin, 9 or fewer. Gill rakers on first arch, 20–23, modally 22; on lower limb, 12–15, modally 13. Scales on lateral line, 34–37, modally 35 (the original description gives 37–40, probably including small scales on caudal-fin base); predorsal scales, 10–14, modally 12; scale rows from lateral line to base of first dorsal-fin ray, 6.5–8, modally 7; to base of pelvic fin, 5–6.5; to base of pectoral fin, 3.5–4; circumpeduncular scales, 13–15, mean 14.5. Scaly sheath on base of anal-fin, short, with imbricated scales. Nuptial tubercles, simple. Total vertebrae 32–33, 18–19 caudal. Detailed frequencies are given in
|
||
<tableCitation id="C6ABEB94C962FFD2FE1CFCD3D2E1FCD9" box="[418,503,870,894]" captionStart="Table 3" captionStartId="16.[160,216,142,165]" captionText="Table 3. Meristics of Central and North American species in the genus Astyanax. Numbers are" pageId="53" pageNumber="1383">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC962FFD2FF7EFC3CD00EFC41" blockId="53.[160,1157,384,1726]" pageId="53" pageNumber="1383">
|
||
Largest examined specimen,
|
||
<quantity id="4CD173CAC962FFD2FDA8FC3CD16AFC06" box="[534,636,905,929]" metricMagnitude="-2" metricUnit="m" metricValue="9.95" pageId="53" pageNumber="1383" unit="mm" value="99.5">99.5 mm</quantity>
|
||
SL. Body slender, 30–32% SL. Head length, 30–34% SL; orbital diameter, 27–29% HL; interorbital distance narrow, 6.1–7.8% SL, mean 7.0% SL (further morphometric data appear in
|
||
<tableCitation id="C6ABEB94C962FFD2FD06FC7BD01AFC41" box="[696,780,974,998]" captionStart="Table 4" captionStartId="19.[160,216,137,160]" captionText="Table 4. Selected morphometric proportions of Central and North American species in the genus Astyanax (abbreviated by first letters of specific epithet). Specimens measured are those marked ‘dig’ in Appendix. All measurements given as percentage of standard length. Data are means; whole range in parentheses." pageId="53" pageNumber="1383">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC962FFD1FF7EFC44D21AFF4B" blockId="53.[160,1157,384,1726]" lastBlockId="54.[160,1156,142,444]" lastPageId="54" lastPageNumber="1384" pageId="53" pageNumber="1383">Anterior fontanel long, straight-sided, sharp-tipped. Supraoccipital process in dorsal view, short, wide-based; slightly concave, in lateral view. Vomer, rostrally undulate. Dentigerous arm of premaxilla, much longer; 3–4 teeth. Highest tooth on dentary, first or third; posterior teeth, not abruptly smaller. Dorsal edge of longer articular arm, variable. Maxillary, with a convex anteroinferior edge; 9–13 teeth. Quadrate, dorsal process expanded. Arms of metapterygoid, subequal; 2 dorsorostral projections. Infraorbital II, triangular with an angled base; infraorbital III, inferoposteriorly semicircular; infraorbital IV, rectangular, with a projection; contact between infraorbitals II and III, wide. Urohyal rostral end turned up, blunt; its ventrorostral edge convex, its ventral apex almost at caudal end; ceratohyal foramen, drop-shaped; rostral vertices, round, ventral side, straight. Epibranchial III, insertion of uncinate process angled, the distal segment of the main body straight. Upper pharyngeal bones, S-shaped; lower pharyngeal plate single, its caudal side concave. Dorsal side of hyomandibular, convex. Opercle, sides of dorsal half, parallel; posterior edge, dorsally concave, ventrally straight-convex; ventral tip, rounded. Interopercle, posterior edge, straight-convex, with a spine. Preopercular anterodorsal edge, with a median convexity; ventral rim, straight; posteroventral edge, curved; 2 canals at angle. Five predorsal bony elements; rostral edge of first pterygiophore curved. Coracoid with 2–3 interdigitations in suture to cleithrum, a concave caudal edge, a single posteroinferior spine. Caudad process of postcleithrum, digitiform. Dorsal tip of scaphium, truncate; caudal tip, spiny; dorsoposterior edge, slightly concave. Neural spines under dorsal fin, straight. Postanal element, short. Sixth to eighth caudal vertebra from tail, with a haemal spine displaced caudad. Largest hypuric plate, rostral edge straight. Epuric plate on last neural spine, roundish.</paragraph>
|
||
<paragraph id="8B96DE2FC961FFD1FF7EFF43D706FE1B" blockId="54.[160,1156,142,444]" pageId="54" pageNumber="1384">
|
||
Humeral spot, indistinct to P-shaped. Pigment uniformly distributed on anal fin, only the anteriormost distal edge lighter. Caudal spot, both on peduncle and on fin rays. In life, the species ‘is light amber, the gill cover has silvery reflections, and the fins of large adults, particularly the pelvic, anal, and caudal fins, are rosy. The pectoral and dorsal fins are clear or pale yellow. In half-grown and juvenile
|
||
<taxonomicName id="4C29A5ACC961FFD1FD4AFE34D057FE3E" authorityName="Rosen" authorityYear="1970" box="[756,833,385,409]" class="Actinopterygii" family="Characidae" genus="Bramocharax" kingdom="Animalia" order="Characiformes" pageId="54" pageNumber="1384" phylum="Chordata" rank="species" species="dorioni">
|
||
<emphasis id="B95D023DC961FFD1FD4AFE34D057FE3E" box="[756,833,385,409]" italics="true" pageId="54" pageNumber="1384">dorioni</emphasis>
|
||
</taxonomicName>
|
||
the fins usually are clear or pale yellow; the caudal fin sometimes has a pinkish wash’ (
|
||
<bibRefCitation id="EFB8A3DEC961FFD1FCFAFE11D0DFFE1B" author="Rosen DE" box="[836,969,420,444]" pageId="54" pageNumber="1384" pagination="1 - 17" refId="ref48940" refString="Rosen DE. 1970. A new tetragonopterine characid fish from Guatemala. Am Mus Nov. 2435: 1 - 17." type="journal article" year="1970">Rosen 1970</bibRefCitation>
|
||
, p. 7).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C961FFD1FF1EFE5DD2E6FD09" pageId="54" pageNumber="1384" type="materials_examined">
|
||
<paragraph id="8B96DE2FC961FFD1FF1EFE5DD2E2FDA6" blockId="54.[160,1156,488,686]" box="[160,500,488,513]" pageId="54" pageNumber="1384">
|
||
<emphasis id="B95D023DC961FFD1FF1EFE5DD2E2FDA6" bold="true" box="[160,500,488,513]" italics="true" pageId="54" pageNumber="1384">Type material and depositor</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC961FFD1FF1EFDB9D2E6FD09" blockId="54.[160,1156,488,686]" pageId="54" pageNumber="1384">
|
||
<materialsCitation id="3B41D472C961FFD1FF1EFDB9D796FDE1" collectionCode="R" collectorName="D. E. Rosen" country="Guatemala" county="Usumacinta River" location="Rio Semococh" municipality="Sebol'" pageId="54" pageNumber="1384" specimenCode="AMNH 29411" specimenCount="1" stateProvince="Alta Verapaz" typeStatus="holotype">
|
||
<typeStatus id="5492608DC961FFD1FF1EFDB9D211FD83" box="[160,263,524,548]" pageId="54" pageNumber="1384" type="holotype">Holotype</typeStatus>
|
||
<specimenCode id="DB8F7654C961FFD1FEACFDB9D2BBFD83" box="[274,429,524,548]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="54" pageNumber="1384">AMNH 29411</specimenCode>
|
||
,
|
||
<quantity id="4CD173CAC961FFD1FE02FDB9D134FD83" box="[444,546,524,548]" metricMagnitude="-2" metricUnit="m" metricValue="9.95" pageId="54" pageNumber="1384" unit="mm" value="99.5">99.5 mm</quantity>
|
||
SL,
|
||
<location id="8EF688F4C961FFD1FDE6FDB9D1E8FD83" LSID="urn:lsid:plazi:treatment:03806F39C963FFD1FE2EFA34D224FC06:8EF688F4C961FFD1FDE6FDB9D1E8FD83" box="[600,766,524,548]" country="Guatemala" county="Usumacinta River" municipality="Sebol'" name="Rio Semococh" pageId="54" pageNumber="1384" stateProvince="Alta Verapaz">Río Semococh</location>
|
||
, ‘
|
||
<quantity id="4CD173CAC961FFD1FCACFDB9D04AFD83" box="[786,860,524,548]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="54" pageNumber="1384" unit="km" value="15.0">15 km</quantity>
|
||
by road south of
|
||
<collectingMunicipality id="6BF24455C961FFD1FB84FDB9D76BFD84" box="[1082,1149,523,548]" pageId="54" pageNumber="1384">Sebol’</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62F7A6A3C961FFD1FF1EFD9BD270FDE1" box="[160,358,558,582]" pageId="54" pageNumber="1384">Usumacinta River</collectingCounty>
|
||
basin,
|
||
<collectingRegion id="49ED10CDC961FFD1FE03FD9BD146FDE1" box="[445,592,558,582]" country="Guatemala" name="Alta Verapaz" pageId="54" pageNumber="1384">Alta Verapaz</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33E9EBFC961FFD1FDE3FD9BD1CFFDE1" box="[605,729,558,582]" name="Guatemala" pageId="54" pageNumber="1384">Guatemala</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26DCBBF9C961FFD1FCA0FD9BD081FDE1" box="[798,919,558,582]" pageId="54" pageNumber="1384">D.E. Rosen</collectorName>
|
||
and
|
||
<collectionCode id="ED3846EAC961FFD1FC68FD9BD0F1FDE1" box="[982,999,558,582]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="54" pageNumber="1384">R</collectionCode>
|
||
. Bailey, 1971
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B41D472C961FFD1FF1EFDE4D0CEFD2C" accessionNumber="MCZ 48687" collectionCode="MCZ" country="Guatemala" pageId="54" pageNumber="1384" specimenCode="AMNH 29412, AMNH 27816, UANL 5705, UMMZ 187926" specimenCount="49" stateProvince="Alta Verapaz" typeStatus="paratype">
|
||
<typeStatus id="5492608DC961FFD1FF1EFDE4D206FDCE" box="[160,272,593,617]" pageId="54" pageNumber="1384" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCode id="DB8F7654C961FFD1FEA5FDE4D2B9FDCE" box="[283,431,593,617]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="54" pageNumber="1384">AMNH 29412</specimenCode>
|
||
(
|
||
<specimenCount id="9D2F15A6C961FFD1FE00FDE4D148FDCE" box="[446,606,593,617]" pageId="54" pageNumber="1384" type="generic">49 specimens</specimenCount>
|
||
), 29413 (22), 29414 (6),
|
||
<specimenCode id="DB8F7654C961FFD1FCD7FDE4D0EBFDCE" box="[873,1021,593,617]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="54" pageNumber="1384">AMNH 27816</specimenCode>
|
||
SD (3);
|
||
<collectionCode id="ED3846EAC961FFD1FBF1FDE4D795FDCE" box="[1103,1155,593,617]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="54" pageNumber="1384" type="Museum">MCZ</collectionCode>
|
||
48687 (2);
|
||
<specimenCode id="DB8F7654C961FFD1FEA3FDC6D288FD2C" box="[285,414,627,651]" collectionCode="UANL" country="Mexico" httpUri="http://grbio.org/cool/cw6z-rzgt" name="Universidad Autonoma de Nuevo Leon" pageId="54" pageNumber="1384">UANL 5705</specimenCode>
|
||
(4);
|
||
<specimenCode id="DB8F7654C961FFD1FE68FDC6D168FD2C" box="[470,638,627,651]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="54" pageNumber="1384" type="University or college">UMMZ 187926</specimenCode>
|
||
(23), 187930 (11), 187944 (8)
|
||
</materialsCitation>
|
||
. For collection data, see original description.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C961FFD1FF1EFD6ED1DEFCB1" box="[160,712,731,790]" pageId="54" pageNumber="1384" type="distribution">
|
||
<paragraph id="8B96DE2FC961FFD1FF1EFD6ED23AFD53" blockId="54.[160,712,731,790]" box="[160,300,731,756]" pageId="54" pageNumber="1384">
|
||
<emphasis id="B95D023DC961FFD1FF1EFD6ED23AFD53" bold="true" box="[160,300,731,756]" italics="true" pageId="54" pageNumber="1384">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC961FFD1FF1EFD4BD1DEFCB1" blockId="54.[160,712,731,790]" box="[160,712,766,790]" pageId="54" pageNumber="1384">
|
||
Endemic to
|
||
<collectingRegion id="49ED10CDC961FFD1FE94FD4BD2ADFCB1" box="[298,443,766,790]" country="Guatemala" name="Alta Verapaz" pageId="54" pageNumber="1384">Alta Verapaz</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33E9EBFC961FFD1FE78FD4BD129FCB1" box="[454,575,766,790]" name="Guatemala" pageId="54" pageNumber="1384">Guatemala</collectingCountry>
|
||
(
|
||
<figureCitation id="1312C2AAC961FFD1FDEEFD4BD1ADFCB1" box="[592,699,766,790]" captionStart="Figure 24" captionStartId="37.[160,225,1645,1668]" captionTargetBox="[163,1151,941,1611]" captionTargetId="figure-183@37.[163,1153,941,1614]" captionTargetPageId="37" captionText="Figure 24. Map of northern Central America, showing distribution localities of Astyanax baileyi, A. belizianus, A. brevimanus, A. cubilhuitz sp. nov., A. dorioni and A. petenensis. A larger symbol denotes the type locality." figureDoi="http://doi.org/10.5281/zenodo.5181937" httpUri="https://zenodo.org/record/5181937/files/figure.png" pageId="54" pageNumber="1384">Figure 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C961FFD1FF1EFCF6D224FC06" pageId="54" pageNumber="1384" type="discussion">
|
||
<paragraph id="8B96DE2FC961FFD1FF1EFCF6D210FCFB" blockId="54.[160,1155,835,929]" box="[160,262,835,860]" pageId="54" pageNumber="1384">
|
||
<emphasis id="B95D023DC961FFD1FF1EFCF6D210FCFB" bold="true" box="[160,262,835,860]" italics="true" pageId="54" pageNumber="1384">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC961FFD1FF1EFCD3D224FC06" blockId="54.[160,1155,835,929]" pageId="54" pageNumber="1384">
|
||
The species corresponds to clade Ic of
|
||
<bibRefCitation id="EFB8A3DEC961FFD1FDE1FCD3D085FCD9" author="Ornelas-Garcia CP & Dominguez-Dominguez O & Doadrio I" box="[607,915,870,894]" pageId="54" pageNumber="1384" pagination="340" refId="ref48693" refString="Ornelas-Garcia CP, Dominguez-Dominguez O, Doadrio I. 2008. Evolutionary history of the fish genus Astyanax Baird and Girard 1854 (Actinopterygii, Characidae) in Mesoamerica reveals multiple morphological homoplasies. BMC Evol Biol. 8: 340." type="journal article" year="2008">Ornelas-García et al. (2008)</bibRefCitation>
|
||
, in part. See remarks for
|
||
<taxonomicName id="4C29A5ACC961FFD1FF76FC3CD23DFC06" authorityName="Schmitter-Soto" authorityYear="2017" baseAuthorityName="Rosen" baseAuthorityYear="1972" box="[200,299,905,929]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="54" pageNumber="1384" phylum="Chordata" rank="species" species="baileyi">
|
||
<emphasis id="B95D023DC961FFD1FF76FC3CD23DFC06" box="[200,299,905,929]" italics="true" pageId="54" pageNumber="1384">A. baileyi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |