treatments-xml/data/9D/2A/15/9D2A155EBB27FF8E84F7FD17FB4BF91B.xml
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<document id="F700448584F9FA7F971BF18C6B0F7724" ID-ISSN="2201-4349" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629579987626" checkinUser="felipe" docAuthor="Smith-Vaniz, William F." docDate="2004" docId="9D2A155EBB27FF8E84F7FD17FB4BF91B" docLanguage="en" docName="RecAustMus.56.2.209-224.pdf" docOrigin="Records of the Australian Museum 56 (2)" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Opistognathus verecundus Smith-Vaniz 2004, n.sp." docType="treatment" docVersion="2" lastPageNumber="219" masterDocId="61136D26BB2EFF84876CFF84FFB9FF91" masterDocTitle="Descriptions of Six New Species of Jawfishes (Opistognathidae: Opistognathus) from Australia" masterLastPageNumber="224" masterPageNumber="209" pageNumber="218" updateTime="1699080190703" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="90C83A9E5D795DD030446E952A95E939">Descriptions of Six New Species of Jawfishes (Opistognathidae: Opistognathus) from Australia</mods:title>
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<mods:namePart id="81D8296163600226C42430A26B63FBEF">Smith-Vaniz, William F.</mods:namePart>
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<mods:typeOfResource id="943A988CFAF3E50038ADF438B7F13182">text</mods:typeOfResource>
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<mods:title id="60CC8FEE846D5A532B813DFE6DA212AB">Records of the Australian Museum</mods:title>
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<mods:date id="53609FD91463FD348EBB9561B34F2BA0">2004</mods:date>
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<mods:number id="72BFD236FCC8BC751591B0CA01924F35">56</mods:number>
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<treatment id="9D2A155EBB27FF8E84F7FD17FB4BF91B" LSID="urn:lsid:plazi:treatment:9D2A155EBB27FF8E84F7FD17FB4BF91B" httpUri="http://treatment.plazi.org/id/9D2A155EBB27FF8E84F7FD17FB4BF91B" lastPageId="10" lastPageNumber="219" pageId="9" pageNumber="218">
<subSubSection id="5D99F7C3BB27FF8D84F7FD17FAB5FD3A" box="[923,1292,659,683]" pageId="9" pageNumber="218" type="nomenclature">
<paragraph id="153CA448BB27FF8D84F7FD17FAB5FD3A" blockId="9.[923,1292,659,683]" box="[923,1292,659,683]" pageId="9" pageNumber="218">
<heading id="4E741324BB27FF8D84F7FD17FAB5FD3A" box="[923,1292,659,683]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="218" reason="2">
<emphasis id="27F7785ABB27FF8D84F7FD17FAB5FD3A" bold="true" box="[923,1292,659,683]" pageId="9" pageNumber="218">
<taxonomicName id="D283DFCBBB27FF8D84F7FD17FB77FD3A" ID-CoL="6T2HT" authority="Smith-Vaniz, 2004" authorityName="Smith-Vaniz" authorityYear="2004" box="[923,1230,659,683]" class="Actinopterygii" family="Opistognathidae" genus="Opistognathus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="218" phylum="Chordata" rank="species" species="verecundus" status="sp. nov.">
<emphasis id="27F7785ABB27FF8D84F7FD17FB77FD3A" bold="true" box="[923,1230,659,683]" italics="true" pageId="9" pageNumber="218">Opistognathus verecundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CC4C521BB27FF8D83B9FD17FAB5FD3A" box="[1237,1292,659,683]" pageId="9" pageNumber="218" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5D99F7C3BB27FF8D84C9FD43FC20FCBB" pageId="9" pageNumber="218" type="description">
<paragraph id="153CA448BB27FF8D84C9FD43FB4CFD4F" blockId="9.[933,1269,711,734]" box="[933,1269,711,734]" pageId="9" pageNumber="218">
<figureCitation id="8DB8B8CDBB27FF8D84C9FD43FBBEFD4F" box="[933,1031,711,734]" captionStart="Fig" captionStartId="1.[794,830,1947,1968]" captionTargetBox="[800,1413,1029,1915]" captionTargetId="figure-468@1.[794,1421,1016,1920]" captionTargetPageId="1" captionText="Fig. 2. Maxilla and supramaxilla (shaded): A, Opistognathus alleni; B, O. seminudus; C, O. verecundus; D, O. reticeps. Data as in figure 3; scale bars represent 2 mm." pageId="9" pageNumber="218">Figs. 2C</figureCitation>
,
<figureCitation id="8DB8B8CDBB27FF8D837FFD43FB8CFD4F" box="[1043,1077,711,734]" captionStart="Fig" captionStartId="3.[121,156,1441,1462]" captionTargetBox="[130,1408,164,1414]" captionTargetId="figure-141@3.[121,1422,151,1432]" captionTargetPageId="3" captionText="Fig. 3. Lateral views of left infraorbital bones (3rd infraorbital also in rotated dorsal view), excluding dermosphenotic:A, Opistognathus alleni, ANSP 157594, 54.3 mm SL; B, O. seminudus, ANSP 142950, 55.0 mm SL; C, O. stigmosus, WAM P.29641-001, 62.5 mm SL; D, O. elizabethensis, AMS I.27891-048, 66.6 mm SL; E, O. verecundus, ANSP 167416, 36.9 mm SL; F, O. reticeps, NTM S.10718- 032, 63.7 mm SL. Scale bars represent 1 mm." pageId="9" pageNumber="218">3E</figureCitation>
,
<figureCitation id="8DB8B8CDBB27FF8D832DFD43FBE8FD4F" box="[1089,1105,711,734]" captionStart="Fig" captionStartId="9.[219,255,553,574]" captionTargetBox="[136,1405,177,522]" captionTargetId="figure-385@9.[121,1422,151,541]" captionTargetPageId="9" captionText="Fig. 9. Opistognathus verecundus, holotype, NTM S.10016-003, male, 52.6 mm SL, Cobourg Peninsula, Northern Territory, Australia. Drawn by Jack R. Schroeder." pageId="9" pageNumber="218">9</figureCitation>
,
<figureCitation id="8DB8B8CDBB27FF8D8332FD43FBC4FD4F" box="[1118,1149,711,734]" captionStart="Fig" captionStartId="10.[166,201,817,838]" captionTargetBox="[168,793,151,801]" captionTargetId="figure-737@10.[166,794,151,801]" captionTargetPageId="10" captionText="Fig.10. Semidiagrammatic drawing showing buccal pigmentation in Opistognathus verecundus." pageId="9" pageNumber="218">10</figureCitation>
;
<tableCitation id="580191F3BB27FF8D83E5FD43FB4CFD4F" box="[1161,1269,711,734]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="2.[166,223,172,193]" captionStartId-1="7.[121,178,172,193]" captionStartId-2="7.[121,178,1633,1654]" captionStartId-3="8.[166,223,1660,1681]" captionText-0="Table 1. Frequency distributions for certain fin-ray counts in selectedAustralian species of Opistognathus (holotype values in boldface)." captionText-1="Table 2. Frequency distributions of gill-raker counts in selected Australia species of Opistognathus (values for holotypes in boldface)." captionText-2="Table 3. Frequency distributions of oblique scale rows in longitudinal series in selected Australian species of Opistognathus (values for holotypes in boldface). Asterisk following species name indicates that bilateral counts were made." captionText-3="Table 4. Frequency distributions of lateral-line terminus in relation to total dorsal-fin element position in selected Australian species of Opistognathus (holotype values in boldface). When terminus ended mid-way between two elements, the higher number was used." pageId="9" pageNumber="218">Tables 14</tableCitation>
</paragraph>
<paragraph id="153CA448BB27FF8D8476FD7FFC20FCBB" blockId="9.[794,1397,763,810]" pageId="9" pageNumber="218">
<taxonomicName id="D283DFCBBB27FF8D8476FD7FFC07FC81" box="[794,958,763,784]" class="Actinopterygii" family="Opistognathidae" genus="Opistognathus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="218" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="27F7785ABB27FF8D8476FD7FFC1CFC81" box="[794,933,763,784]" italics="true" pageId="9" pageNumber="218">Opistognathus</emphasis>
sp.
</taxonomicName>
—Larson &amp; Williams, 1997: 366 (listed; Darwin Harbour).
</paragraph>
</subSubSection>
<subSubSection id="5D99F7C3BB27FF8D8476FCC3FAA6FBBA" pageId="9" pageNumber="218" type="materials_examined">
<paragraph id="153CA448BB27FF8D8476FCC3FACAFC60" blockId="9.[794,1421,839,1067]" pageId="9" pageNumber="218">
<emphasis id="27F7785ABB27FF8D8476FCC3FC71FCCE" bold="true" box="[794,968,839,863]" pageId="9" pageNumber="218">Type material</emphasis>
.
<materialsCitation id="A5EBAE15BB27FF8D84B7FCC3FB72FC27" collectingDate="1981-10-18" collectionCode="NTM" collectorName="Cobourg Peninsula, E. &amp; Helen K. Larson &amp; P. Homes" country="Australia" elevation="6" latitude="-11.183333" location="Coral Bay" longLatPrecision="1295" longitude="132.06667" pageId="9" pageNumber="218" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<typeStatus id="CA381AEABB27FF8D84B7FCC3FBECFCCC" box="[987,1109,839,862]" pageId="9" pageNumber="218" type="holotype">HOLOTYPE</typeStatus>
:
<collectionCode id="73923C8DBB27FF8D8305FCC3FB1FFCCF" box="[1129,1190,839,862]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="9" pageNumber="218">NTM</collectionCode>
S.10016-003, male (52.6),
<collectingCountry id="6D94E4D8BB27FF8D8406FCE0FC69FCEA" box="[874,976,868,891]" name="Australia" pageId="9" pageNumber="218">Australia</collectingCountry>
,
<collectingRegion id="D7476AAABB27FF8D84B1FCE0FB10FCEA" box="[989,1193,868,891]" country="Australia" name="Northern Territory" pageId="9" pageNumber="218">Northern Territory</collectingRegion>
,
<collectorName id="B876C19EBB27FF8D83DAFCE0FC88FC08" pageId="9" pageNumber="218">Cobourg Peninsula, E.</collectorName>
side
<location id="105CF293BB27FF8D841FFC06FC5EFC08" LSID="urn:lsid:plazi:treatment:9D2A155EBB27FF8E84F7FD17FB4BF91B:105CF293BB27FF8D841FFC06FC5EFC08" box="[883,999,898,921]" country="Australia" latitude="-11.183333" longLatPrecision="1295" longitude="132.06667" name="Coral Bay" pageId="9" pageNumber="218" stateProvince="Northern Territory">Coral Bay</location>
,
<geoCoordinate id="70B7C28FBB27FF8D849AFC06FBF2FC08" box="[1014,1099,898,921]" degrees="11" direction="south" minutes="11" orientation="latitude" pageId="9" pageNumber="218" precision="925" value="-11.183333">11°11'S</geoCoordinate>
<geoCoordinate id="70B7C28FBB27FF8D833AFC06FB07FC08" box="[1110,1214,898,921]" degrees="132" direction="east" minutes="04" orientation="longitude" pageId="9" pageNumber="218" precision="925" value="132.06667">132°04'E</geoCoordinate>
,
<quantity id="D27B09ADBB27FF8D83A1FC06FAB3FC08" box="[1229,1290,898,921]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="9" pageNumber="218" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">
<elevation id="9EAE437BBB27FF8D83A1FC06FAB3FC08" box="[1229,1290,898,921]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="9" pageNumber="218" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">56 m</elevation>
</quantity>
,
<collectorName id="B876C19EBB27FF8D8276FC06FCD0FC27" pageId="9" pageNumber="218">Helen K. Larson</collectorName>
and
<collectorName id="B876C19EBB27FF8D84C5FC1BFBA2FC27" box="[937,1051,927,950]" pageId="9" pageNumber="218">P. Homes</collectorName>
,
<date id="613D8288BB27FF8D8347FC1BFB72FC27" box="[1067,1227,927,950]" pageId="9" pageNumber="218" value="1981-10-18">
<collectingDate id="71797B60BB27FF8D8347FC1BFB72FC27" box="[1067,1227,927,950]" pageId="9" pageNumber="218" value="1981-10-18">18 Oct. 1981</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="A5EBAE15BB27FF8D83B7FC1BFA39FC42" collectionCode="AMS" country="Australia" pageId="9" pageNumber="218" specimenCount="28" stateProvince="Northern Territory" typeStatus="paratype">
<typeStatus id="CA381AEABB27FF8D83B7FC1BFAE4FC24" box="[1243,1373,927,950]" pageId="9" pageNumber="218" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="03856FC1BB27FF8D821DFC1BFC32FC42" pageId="9" pageNumber="218" type="generic">28 specimens</specimenCount>
,
<quantity id="D27B09ADBB27FF8D84F8FC38FB98FC42" box="[916,1057,956,979]" metricMagnitude="-2" metricUnit="m" metricValue="3.1149999999999998" metricValueMax="4.81" metricValueMin="1.42" pageId="9" pageNumber="218" unit="mm" value="31.15" valueMax="48.1" valueMin="14.2">14.248.1 mm</quantity>
, all taken with the
<typeStatus id="CA381AEABB27FF8D838CFC38FAF9FC42" box="[1248,1344,956,979]" pageId="9" pageNumber="218" type="holotype">holotype</typeStatus>
.
<collectionCode id="73923C8DBB27FF8D822AFC38FA39FC42" box="[1350,1408,956,979]" pageId="9" pageNumber="218">AMS</collectionCode>
</materialsCitation>
<materialsCitation id="A5EBAE15BB27FF8D8476FC5EFB59FC60" box="[794,1248,986,1009]" collectionCode="ANSP" country="Australia" pageId="9" pageNumber="218" specimenCode="ANSP 167416" specimenCount="1" stateProvince="Northern Territory" typeStatus="paratype">
I.33644-001 (4, 35.239.4),
<specimenCode id="45250C33BB27FF8D832EFC5EFB59FC60" box="[1090,1248,986,1009]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="9" pageNumber="218">ANSP 167416</specimenCode>
</materialsCitation>
(4, 29.548.1,
</paragraph>
<paragraph id="153CA448BB27FF8D8476FC73FAA6FBBA" blockId="9.[794,1421,839,1067]" pageId="9" pageNumber="218">
1 C&amp;S),
<collectionCode id="73923C8DBB27FF8D8419FC73FC09FB9F" box="[885,944,1015,1038]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="9" pageNumber="218">NTM</collectionCode>
S.10016-046 (12, 14.247.2),
<collectionCode id="73923C8DBB27FF8D83B3FC73FA95FB9F" box="[1247,1324,1015,1038]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="218" type="Museum">USNM</collectionCode>
320261 (4, 25.144.8),
<collectionCode id="73923C8DBB27FF8D84DFFB90FC4DFBBA" box="[947,1012,1044,1067]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="9" pageNumber="218" type="Museum">WAM</collectionCode>
P.30593-001 (4, 33.537.3).
</paragraph>
</subSubSection>
<subSubSection id="5D99F7C3BB27FF8D8476FBCCFC50FABC" pageId="9" pageNumber="218" type="diagnosis">
<paragraph id="153CA448BB27FF8D8476FBCCFC50FABC" blockId="9.[794,1421,1096,1325]" pageId="9" pageNumber="218">
<emphasis id="27F7785ABB27FF8D8476FBCCFC30FBF1" bold="true" box="[794,905,1096,1120]" pageId="9" pageNumber="218">Diagnosis</emphasis>
. A species of
<taxonomicName id="D283DFCBBB27FF8D834FFBCCFB79FBCE" authorityName="Smith-Vaniz" authorityYear="2004" box="[1059,1216,1096,1119]" class="Actinopterygii" family="Opistognathidae" genus="Opistognathus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="218" phylum="Chordata" rank="genus">
<emphasis id="27F7785ABB27FF8D834FFBCCFB79FBCE" box="[1059,1216,1096,1119]" italics="true" pageId="9" pageNumber="218">Opistognathus</emphasis>
</taxonomicName>
with dorsal fin X XI, 14 or 15 (typically XI, 14); upper jaw sexually dimorphic with flexible lamina posteriorly, end of maxilla slightly rounded (females) becoming increasingly elongate and scimitar-shaped (adult males); oblique scale rows
<quantity id="D27B09ADBB27FF8D8243FB3AFA39FB44" box="[1327,1408,1214,1237]" metricMagnitude="0" metricUnit="m" metricValue="1.1811" metricValueMax="1.3716" metricValueMin="0.9906" pageId="9" pageNumber="218" unit="in" value="46.5" valueMax="54.0" valueMin="39.0">3954 in</quantity>
longitudinal series; total gill-rakers on first arch 2326; spinous dorsal fin with a single brownish spot, if present, between spines 24.
</paragraph>
</subSubSection>
<subSubSection id="5D99F7C3BB27FF8E8476FACDFC75FBC5" lastPageId="10" lastPageNumber="219" pageId="9" pageNumber="218" type="description">
<paragraph id="153CA448BB27FF8D8476FACDFBA6F9F8" blockId="9.[794,1422,1353,1993]" pageId="9" pageNumber="218">
<emphasis id="27F7785ABB27FF8D8476FACDFC26FAF0" bold="true" box="[794,927,1353,1377]" pageId="9" pageNumber="218">Description</emphasis>
. Dorsal-fin rays XXI, 14 or 15 (typically XI, 14). Anal-fin rays III, 14 (rarely 13 or 15). Pectoral-fin rays 18 or 19. Caudal fin: procurrent rays 45+35, segmented rays 8+8, middle 1214 branched, total elements 2326; hypural 5 absent. Vertebrae: 10+18 (
<quantity id="D27B09ADBB27FF8D83BCFA3BFAB7FA47" box="[1232,1294,1471,1494]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="9" pageNumber="218" unit="in" value="19.0">19 in</quantity>
one of
<specimenCount id="03856FC1BB27FF8D821DFA3BFC36FA62" pageId="9" pageNumber="218" type="generic">29 specimens</specimenCount>
); last pleural rib on vertebra 10; epineural ribs 1112. Supraneurals 1 (based on radiographs but in the only C&amp;S specimen a tiny cartilaginous ball is present in the first interneural space), insertion pattern 0/S/1/1+1/1/. Gill rakers 89+1517 = 2326.
</paragraph>
<paragraph id="153CA448BB27FF8D8454F9EBFA39F950" blockId="9.[794,1422,1353,1993]" pageId="9" pageNumber="218">Scales absent on head, nape, above lateral line, pectoralfin base, breast and anterior ¼ to ½ of belly. Body with about 4454 oblique scale rows in longitudinal series.</paragraph>
<paragraph id="153CA448BB27FF8D8476F943FA38F96A" blockId="9.[794,1422,1353,1993]" pageId="9" pageNumber="218">Lateral-line terminus below verticals from 2nd to 4th segmented dorsal-fin rays (total element position 1315).</paragraph>
<paragraph id="153CA448BB27FF8D8476F886FAE0F858" blockId="9.[794,1422,1353,1993]" pageId="9" pageNumber="218">Lateral line pores numerous, arranged in multiple series above and below embedded lateral-line tubes. Cephalic sensory pores numerous, completely covering most of head, including all of predorsal area except a small area immediately adjacent to dorsal-fin origin; mandibular pore positions 13 occupied by relatively large, single pores, 4th position with 13 (usually 2) pores, 5th with 37 pores.</paragraph>
<caption id="41FCF4C0BB24FF8E87CAFCB5FE73FCF0" pageId="10" pageNumber="219" startId="10.[166,201,817,838]" targetBox="[168,793,151,801]" targetPageId="10">
<paragraph id="153CA448BB24FF8E87CAFCB5FE73FCF0" blockId="10.[166,793,817,865]" pageId="10" pageNumber="219">
Fig. 10. Semidiagrammatic drawing showing buccal pigmentation in
<taxonomicName id="D283DFCBBB24FF8E87ACFCC8FE7DFCF0" authorityName="Smith-Vaniz" authorityYear="2004" box="[192,452,844,865]" class="Actinopterygii" family="Opistognathidae" genus="Opistognathus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="219" phylum="Chordata" rank="species" species="verecundus">
<emphasis id="27F7785ABB24FF8E87ACFCC8FE7DFCF0" box="[192,452,844,865]" italics="true" pageId="10" pageNumber="219">Opistognathus verecundus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="153CA448BB24FF8E87A8FC10FCB3FA11" blockId="10.[166,795,916,1995]" pageId="10" pageNumber="219">Anterior nostril about mid-way between posterior nostril and dorsal margin of upper lip, consisting of a short tube with posterior rim longer, that when depressed does not reach or just reaches margin of posterior nostril; height of tube shorter than to about equal maximum diameter of posterior nostril. Dorsal fin moderately low anteriorly, with profile relatively uniform without change in fin height at junction of last spine and anterior segmented rays. Dorsalfin spines relatively slender and slightly curved distally, with flexible tips; all segmented dorsal- and anal-fin rays branched distally or 1st anal ray unbranched. Outermost segmented pelvic-fin ray not tightly bound to adjacent ray, interradial membrane incised distally. Upper margin of opercle straight and slightly rounded posterodorsally; posterior margin of preopercle indistinct without a free margin. No papillae on inner surface of lips. Fifth cranial nerve passes under A1 section of adductor mandibulae.</paragraph>
<paragraph id="153CA448BB24FF8E87A8FA03FEACF85A" blockId="10.[166,795,916,1995]" pageId="10" pageNumber="219">
Upper jaw sexually dimorphic (longest in adult males) and extending 1.1 to 2.1 eye diameters behind posterior margin of orbit; maxilla widest before end, with flexible lamina posteriorly (
<figureCitation id="8DB8B8CDBB24FF8E861FFA5BFE7DFA67" box="[371,452,1503,1526]" captionStart="Fig" captionStartId="1.[794,830,1947,1968]" captionTargetBox="[800,1413,1029,1915]" captionTargetId="figure-468@1.[794,1421,1016,1920]" captionTargetPageId="1" captionText="Fig. 2. Maxilla and supramaxilla (shaded): A, Opistognathus alleni; B, O. seminudus; C, O. verecundus; D, O. reticeps. Data as in figure 3; scale bars represent 2 mm." pageId="10" pageNumber="219">Fig. 2C</figureCitation>
), scimitar-shaped in adult males; supramaxilla moderately large and subterminally positioned. Jaws subequal, lower slightly included. Premaxilla with an outer row of moderately large, sharply pointed, conical teeth, those near posterior end of jaw noticeably smaller and more closely spaced; 2 or 3 irregular inner rows of much smaller conical teeth present anteriorly, with those adjacent to premaxillary symphysis slightly enlarged. Dentary with an outer row of conical teeth that are blunter anteriorly, those on posterior half of dentary largest and slightly hooked inward; 2 or 3 inner rows of slightly smaller, conical teeth anteriorly, those in innermost row canted backwards. Vomerine teeth absent. Infraorbital bones tubular, with wide openings for sensory canals (
<figureCitation id="8DB8B8CDBB24FF8E8695F8D8FDEAF8E2" box="[505,595,1884,1907]" captionStart="Fig" captionStartId="3.[121,156,1441,1462]" captionTargetBox="[130,1408,164,1414]" captionTargetId="figure-141@3.[121,1422,151,1432]" captionTargetPageId="3" captionText="Fig. 3. Lateral views of left infraorbital bones (3rd infraorbital also in rotated dorsal view), excluding dermosphenotic:A, Opistognathus alleni, ANSP 157594, 54.3 mm SL; B, O. seminudus, ANSP 142950, 55.0 mm SL; C, O. stigmosus, WAM P.29641-001, 62.5 mm SL; D, O. elizabethensis, AMS I.27891-048, 66.6 mm SL; E, O. verecundus, ANSP 167416, 36.9 mm SL; F, O. reticeps, NTM S.10718- 032, 63.7 mm SL. Scale bars represent 1 mm." pageId="10" pageNumber="219">Fig. 3E</figureCitation>
); 3rd infraorbital relatively robust but without a suborbital shelf. Second pharyngobranchial rod-shaped with distal end slightly expanded.
</paragraph>
<paragraph id="153CA448BB24FF8E8409FF1DFAFDFD81" blockId="10.[839,1467,153,528]" pageId="10" pageNumber="219">
Measurements of the
<quantity id="D27B09ADBB24FF8E8302FF1DFB61FF21" box="[1134,1240,153,176]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" pageId="10" pageNumber="219" unit="mm" value="52.6">52.6 mm</quantity>
male
<typeStatus id="CA381AEABB24FF8E8244FF1DFA37FF21" box="[1320,1422,153,176]" pageId="10" pageNumber="219" type="holotype">holotype</typeStatus>
(in parentheses) and
<specimenCount id="03856FC1BB24FF8E836EFF32FB34FF5C" box="[1026,1165,182,205]" pageId="10" pageNumber="219" type="generic" typeStatus="paratypes">15 paratypes</specimenCount>
, 9 ƋƋ
<quantity id="D27B09ADBB24FF8E83B2FF32FAD4FF5C" box="[1246,1389,182,205]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.81" metricValueMin="3.59" pageId="10" pageNumber="219" unit="mm" value="42.0" valueMax="48.1" valueMin="35.9">35.948.1 mm</quantity>
and
<specimenCount id="03856FC1BB24FF8E82CCFF32FCCDFF7A" pageId="10" pageNumber="219" type="female">6 ♀♀</specimenCount>
<quantity id="D27B09ADBB24FF8E8417FF57FBB4FF7B" box="[891,1037,211,234]" metricMagnitude="-2" metricUnit="m" metricValue="4.12" metricValueMax="4.72" metricValueMin="3.52" pageId="10" pageNumber="219" unit="mm" value="41.2" valueMax="47.2" valueMin="35.2">35.247.2 mm</quantity>
, as percentage of SL: predorsal length (31.6) 28.932.0; preanal length (55.1) 52.456.6; dorsalfin base (74.3) 65.076.1; anal-fin base (38.8) 36.340.8; pelvic-fin length (20.0) 20.422.8; caudal-fin length (21.5) 19.822.8; depth at anal-fin origin (15.0) 12.516.2; head length (32.7) 29.832.6; orbit diameter (8.3) 8.19.3; upper jaw length (25.9) 20.424.5 ƋƋ, 19.021.1
<specimenCount id="03856FC1BB24FF8E8230FE07FA30FE0A" box="[1372,1417,387,411]" pageId="10" pageNumber="219" type="female">♀♀</specimenCount>
. As percentage of head length: postorbital head length (68.9) 65.268.4; upper jaw length (79.1) 65.278.9 ƋƋ, 61.6 66.5
<specimenCount id="03856FC1BB24FF8E84EDFE5FFC17FE62" box="[897,942,475,499]" pageId="10" pageNumber="219" type="female">♀♀</specimenCount>
; postorbital jaw length (52.3) 33.945.3 ƋƋ, 29.935.6
<specimenCount id="03856FC1BB24FF8E84D6FE7CFC5EFD81" box="[954,999,504,528]" pageId="10" pageNumber="219" type="female">♀♀</specimenCount>
; orbit diameter (25.3) 25.729.9.
</paragraph>
<paragraph id="153CA448BB24FF8E842BFDA8FC75FBC5" blockId="10.[839,1467,556,1108]" pageId="10" pageNumber="219">
<emphasis id="27F7785ABB24FF8E842BFDA8FB8CFDD5" bold="true" box="[839,1077,556,580]" pageId="10" pageNumber="219">Preserved coloration</emphasis>
. Head and body ground colour light tan, with darker freckling; in larger specimens branchiostegal membranes and pectoral-fin base finely peppered with melanophores; inner lining of maxilla and adjacent membranes with 2 brownish stripes (best developed in adult males) that when mouth is closed are partially visible as lateral streaks on expanded part of the upper jaw; buccal pigmentation (
<figureCitation id="8DB8B8CDBB24FF8E8488FD7EFB8AFC80" box="[996,1075,762,785]" captionStart="Fig" captionStartId="10.[166,201,817,838]" captionTargetBox="[168,793,151,801]" captionTargetId="figure-737@10.[166,794,151,801]" captionTargetPageId="10" captionText="Fig.10. Semidiagrammatic drawing showing buccal pigmentation in Opistognathus verecundus." pageId="10" pageNumber="219">Fig. 10</figureCitation>
) consisting of diffuse dusky area on floor of mouth in front of esophageal opening and a pair of slightly separated dark blotches on roof of mouth behind upper pharyngeal toothplates; in some specimens, a series of small dark blotches present on sides and about 8 equally spaced blotches on dorsum along dorsal-fin base; in a few specimens first 2 blotches relatively dark and extending onto dorsal fin; dorsal and anal fins dusky with blotches and pale spots that that tend to form rows; in some specimens a pair of pale basicaudal spots evident and caudal fin with several indistinct narrow dusky bands; pelvic and pectoral fins immaculate.
</paragraph>
</subSubSection>
<subSubSection id="5D99F7C3BB24FF8E842BFBF4FC53FB53" pageId="10" pageNumber="219" type="etymology">
<paragraph id="153CA448BB24FF8E842BFBF4FC53FB53" blockId="10.[839,1466,1136,1218]" pageId="10" pageNumber="219">
<emphasis id="27F7785ABB24FF8E842BFBF4FC77FB19" bold="true" box="[839,974,1136,1160]" pageId="10" pageNumber="219">Etymology</emphasis>
. The specific epithet is from the Latin
<taxonomicName id="D283DFCBBB24FF8E842BFB0AFC7AFB34" authorityName="Smith-Vaniz" authorityYear="2004" box="[839,963,1166,1189]" class="Actinopterygii" family="Opistognathidae" genus="Opistognathus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="219" phylum="Chordata" rank="species" species="verecundus">
<emphasis id="27F7785ABB24FF8E842BFB0AFC7AFB34" box="[839,963,1166,1189]" italics="true" pageId="10" pageNumber="219">verecundus</emphasis>
</taxonomicName>
(bashful or shy), in allusion to the burrowdwelling habit.
</paragraph>
</subSubSection>
<subSubSection id="5D99F7C3BB24FF8E842BFB5BFC3CFAA0" pageId="10" pageNumber="219" type="distribution">
<paragraph id="153CA448BB24FF8E842BFB5BFC3CFAA0" blockId="10.[839,1466,1247,1329]" pageId="10" pageNumber="219">
<emphasis id="27F7785ABB24FF8E842BFB5BFC65FB66" bold="true" box="[839,988,1247,1271]" pageId="10" pageNumber="219">Distribution</emphasis>
. Known only from the
<typeStatus id="CA381AEABB24FF8E8397FB5BFA95FB67" box="[1275,1324,1247,1270]" pageId="10" pageNumber="219">type</typeStatus>
locality off Cobourg Peninsula, northern
<collectingCountry id="6D94E4D8BB24FF8E83EFFB79FB50FA85" box="[1155,1257,1277,1300]" name="Australia" pageId="10" pageNumber="219">Australia</collectingCountry>
, where collected in
<quantity id="D27B09ADBB24FF8E842BFA9EFC38FAA0" box="[839,897,1306,1329]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="10" pageNumber="219" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">
5
<specimenCount id="03856FC1BB24FF8E8435FA9EFC38FAA0" box="[857,897,1306,1329]" pageId="10" pageNumber="219" type="generic">6 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="5D99F7C3BB24FF8E842BFACAFB4BF91B" pageId="10" pageNumber="219" type="discussion">
<paragraph id="153CA448BB24FF8E842BFACAFA2AF984" blockId="10.[839,1467,1358,1674]" pageId="10" pageNumber="219">
<emphasis id="27F7785ABB24FF8E842BFACAFC09FAF7" bold="true" box="[839,944,1358,1382]" pageId="10" pageNumber="219">Remarks</emphasis>
. This species is apparently most closely related to the allopatric
<taxonomicName id="D283DFCBBB24FF8E8490FAE8FA91FA13" authority="Bleeker 1853" authorityName="Bleeker" authorityYear="1853" box="[1020,1320,1387,1411]" class="Actinopterygii" family="Opistognathidae" genus="Opistognathus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="219" phylum="Chordata" rank="species" species="solorensis">
<emphasis id="27F7785ABB24FF8E8490FAE8FB35FA12" box="[1020,1164,1388,1411]" italics="true" pageId="10" pageNumber="219">O. solorensis</emphasis>
<bibRefCitation id="7112D9B9BB24FF8E83F8FAEFFA91FA13" author="Bleeker, P" box="[1172,1320,1387,1410]" pageId="10" pageNumber="219" pagination="67 - 96" refId="ref13402" refString="Bleeker, P., 1853. Bijdrage tot de Kennis der ichthyologische fauna van Solor. Natuurkundige Tijdschrift voor Nederlandsch-Indie 5: 67 - 96." type="journal article" year="1853">Bleeker 1853</bibRefCitation>
</taxonomicName>
, known from
<collectingCountry id="6D94E4D8BB24FF8E842BFA0DFC0AFA31" box="[839,947,1417,1440]" name="Indonesia" pageId="10" pageNumber="219">Indonesia</collectingCountry>
,
<collectingCountry id="6D94E4D8BB24FF8E84ACFA0DFB83FA31" box="[960,1082,1417,1440]" name="Philippines" pageId="10" pageNumber="219">Philippines</collectingCountry>
and
<collectingCountry id="6D94E4D8BB24FF8E831EFA0DFB7DFA31" box="[1138,1220,1417,1440]" name="Taiwan" pageId="10" pageNumber="219">Taiwan</collectingCountry>
. Both have essentially identical upper jaw structures, dentition, fin ray and vertebral counts and buccal pigmentation.
<taxonomicName id="D283DFCBBB24FF8E83CDFA40FA02FA4A" authorityName="Smith-Vaniz" authorityYear="2004" box="[1185,1467,1476,1499]" class="Actinopterygii" family="Opistognathidae" genus="Opistognathus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="219" phylum="Chordata" rank="species" species="verecundus">
<emphasis id="27F7785ABB24FF8E83CDFA40FA02FA4A" box="[1185,1467,1476,1499]" italics="true" pageId="10" pageNumber="219">Opistognathus verecundus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="D283DFCBBB24FF8E84A2FA65FBE2FA69" authorityName="Bleeker" authorityYear="1853" box="[974,1115,1505,1528]" class="Actinopterygii" family="Opistognathidae" genus="Opistognathus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="219" phylum="Chordata" rank="species" species="solorensis">
<emphasis id="27F7785ABB24FF8E84A2FA65FBE2FA69" box="[974,1115,1505,1528]" italics="true" pageId="10" pageNumber="219">O. solorensis</emphasis>
</taxonomicName>
in having fewer gill-rakers, with 1517 (versus 1820) on the lower limb and a total of 23
</paragraph>
<paragraph id="153CA448BB24FF8E842BF99FFB4BF91B" blockId="10.[839,1467,1358,1674]" pageId="10" pageNumber="219">26 (vs. 2733), only 4454 oblique scale rows in longitudinal series (vs. typically 5869), and only brown spots or markings anteriorly on the spinous dorsal fin, (vs. one or two black spots in this position).</paragraph>
</subSubSection>
</treatment>
</document>