251 lines
24 KiB
XML
251 lines
24 KiB
XML
<document id="A9B6BD6EE5164F1C8714DB90082B15DE" ID-DOI="10.1649/655" ID-ISSN="1938-4394" ID-Zenodo-Dep="5345291" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630199912524" checkinUser="tatiana" docAuthor="Spomer, Stephen M." docDate="2004" docId="D65887D2DA2B441AFE93FEC8BBC1A30E" docLanguage="en" docName="655.pdf" docOrigin="The Coleopterists Bulletin 58 (3)" docSource="http://dx.doi.org/10.1649/655" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Cicindela nevadica subsp. makosika Spomer 2004, new subspecies" docType="treatment" docVersion="3" lastPageNumber="412" masterDocId="2A61FFAADA29441EFF8CFFD2B955A203" masterDocTitle="A New Subspecies of Cicindela nevadica Leconte (Coleoptera: Carabidae: Cicindelinae) from the Badlands of South Dakota" masterLastPageNumber="412" masterPageNumber="409" pageNumber="409" updateTime="1699183386285" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="AB40A19E61921F91FA2D9293C2675555" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="E720BF7D2D27CC9EA7653AD3ABBD783E">
|
||
<mods:title id="8A292E1B7AA6C85985057AA54A79EDB6">A New Subspecies of Cicindela nevadica Leconte (Coleoptera: Carabidae: Cicindelinae) from the Badlands of South Dakota</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="1F9C9F415BD60CEEDB172F7B703001CE" type="personal">
|
||
<mods:role id="61781A48FB23F8D0AE026DA3E7AFD686">
|
||
<mods:roleTerm id="C89C171C519640B8FB393EC028EBE621">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CF676EA02B60FD13EC2D754E7FEAAA75">Spomer, Stephen M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="B6D349DA8C7E468F953EA3C406BA171A">text</mods:typeOfResource>
|
||
<mods:relatedItem id="E34735A46E49CBD12F72C631EDF6AE93" type="host">
|
||
<mods:titleInfo id="939DC4CA5DF84892FD11754BD2741CE2">
|
||
<mods:title id="CA0CB6D1BB50CCA932828A664A0B6C16">The Coleopterists Bulletin</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="BBFC6DD1362F4FBB4E00F3BF649D19C5">
|
||
<mods:date id="02F69A18B08C24A44202191399C98E8A">2004</mods:date>
|
||
<mods:detail id="16286688DA297DC6B77598727EF6BAEE" type="pubDate">
|
||
<mods:number id="49481CDEB21159D5761553B6AC9EE75F">2004-09-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AAB22D5C9997DECAB28DBF296CFF170E" type="volume">
|
||
<mods:number id="A01BC17DCB7A5F1D9B32222929A7B7E2">58</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="0E3C85B1D9590369FD7502E51075BBDD" type="issue">
|
||
<mods:number id="2D3A224948A75913475A46761407A291">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="E8C3F00887B9D4AD79A9A6F83EBB6B8D" unit="page">
|
||
<mods:start id="D019D1163AB1AD12D89401674AA9E4DF">409</mods:start>
|
||
<mods:end id="2783BB2D4F5252884A00A55894404B13">412</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="A1E46C4F5E0C2B6459C740DEC87EBA4D">
|
||
<mods:url id="A669F7659DB72DF1CD893B3322D1825D">http://dx.doi.org/10.1649/655</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="CB95527E16E7659D8AE650ECAA2D09F6">journal article</mods:classification>
|
||
<mods:identifier id="5E17D4A1D2C26FB054440CD39FE28F67" type="DOI">10.1649/655</mods:identifier>
|
||
<mods:identifier id="5F2C9D6578B8820E896C48B63E34BF73" type="ISSN">1938-4394</mods:identifier>
|
||
<mods:identifier id="638734B82D9E9CC3AAA839BCF7CDCD37" type="Zenodo-Dep">5345291</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="D65887D2DA2B441AFE93FEC8BBC1A30E" LSID="urn:lsid:plazi:treatment:D65887D2DA2B441AFE93FEC8BBC1A30E" httpUri="http://treatment.plazi.org/id/D65887D2DA2B441AFE93FEC8BBC1A30E" lastPageId="4" pageId="2">
|
||
<subSubSection id="16EB654FDA2B441CFE93FEC8BA10A32C" box="[287,837,282,303]" pageId="2" type="nomenclature">
|
||
<paragraph id="5E4E36C4DA2B441CFE93FEC8BA10A32C" blockId="2.[287,837,282,330]" box="[287,837,282,303]" pageId="2">
|
||
<heading id="050681A8DA2B441CFE93FEC8BA10A32C" box="[287,837,282,303]" centered="true" fontSize="9" level="2" pageId="2" reason="2">
|
||
<emphasis id="6C85EAD6DA2B441CFE93FEC8BA10A32C" bold="true" box="[287,837,282,303]" pageId="2">
|
||
<taxonomicName id="99F14D47DA2B441CFE93FEC8BBC8A32C" authority="Spomer" authorityName="Spomer" authorityYear="2004" box="[287,669,282,303]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="subSpecies" species="nevadica" status="subsp. nov." subSpecies="makosika">
|
||
<emphasis id="6C85EAD6DA2B441CFE93FEC8BB11A32C" bold="true" box="[287,580,282,303]" italics="true" pageId="2">Cicindela nevadica makosika</emphasis>
|
||
Spomer
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="77B657ADDA2B441CFD26FEC8BA10A32C" box="[682,837,282,303]" pageId="2" rank="subSpecies">new subspecies</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16EB654FDA2B441CFD82FEE7BB03A349" box="[526,598,309,330]" pageId="2" type="description">
|
||
<paragraph id="5E4E36C4DA2B441CFD82FEE7BB03A349" blockId="2.[287,837,282,330]" box="[526,598,309,330]" pageId="2">
|
||
(
|
||
<figureCitation id="C6CA2A41DA2B441CFD99FEE7BB18A349" box="[533,589,309,330]" captionStart="Fig" captionStartId="3.[197,228,1333,1352]" captionTargetBox="[382,798,194,1312]" captionTargetId="figure-96@3.[382,798,194,1312]" captionTargetPageId="3" captionText="Fig. 1. Habitus, holotype of Cicindela nevadica makosika Spomer." figureDoi="http://doi.org/10.5281/zenodo.5345295" httpUri="https://zenodo.org/record/5345295/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16EB654FDA2B441CFF20FE8EBAD1A012" pageId="2" type="materials_examined">
|
||
<paragraph id="5E4E36C4DA2B441CFF20FE8EBAD1A012" blockId="2.[148,977,348,529]" pageId="2">
|
||
<emphasis id="6C85EAD6DA2B441CFF20FE8EB879A372" bold="true" box="[172,300,348,369]" pageId="2">Type Series.</emphasis>
|
||
<materialsCitation id="EE993C99DA2B441CFEB4FE8FBB15A38F" collectionCode="T, R" collectorName="Pennington Co., SD & E Sec. & Indian Creek, D." county="Pennington Co." location="Indian Creek" pageId="2" specimenCode="T3, R12" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="814A8866DA2B441CFEB4FE8FB8C5A371" box="[312,400,349,370]" pageId="2" type="holotype">Holotype</typeStatus>
|
||
(male) labeled ‘‘23: VII: 2002,
|
||
<collectorName id="F3045312DA2B441CFD59FE8FBAC9A371" box="[725,924,349,370]" pageId="2">
|
||
<collectingCounty id="B72F4E48DA2B441CFD59FE8FBA3AA371" box="[725,879,349,370]" pageId="2">Pennington Co.</collectingCounty>
|
||
, SD
|
||
</collectorName>
|
||
,
|
||
<specimenCode id="0E579EBFDA2B441CFC25FE8FBA91A371" box="[937,964,349,370]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2">T3</specimenCode>
|
||
S
|
||
<specimenCode id="0E579EBFDA2B441CFF18FEA5B9EFA38F" box="[148,186,375,396]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2">R12</specimenCode>
|
||
<collectorName id="F3045312DA2B441CFF37FEA5B9AEA38F" box="[187,251,375,396]" pageId="2">E Sec.</collectorName>
|
||
27,
|
||
<collectorName id="F3045312DA2B441CFEABFEA5B89CA38F" box="[295,457,375,396]" pageId="2">
|
||
<location id="5B2E601FDA2B441CFEABFEA5B8F3A38F" LSID="urn:lsid:plazi:treatment:D65887D2DA2B441AFE93FEC8BBC1A30E:5B2E601FDA2B441CFEABFEA5B8F3A38F" box="[295,422,375,396]" county="Pennington Co." name="Indian Creek" pageId="2">Indian Creek</location>
|
||
, D.
|
||
</collectorName>
|
||
Backlund.’’
|
||
</materialsCitation>
|
||
<materialsCitation id="EE993C99DA2B441CFDC4FEA5BA3FA38F" box="[584,874,375,396]" collectionCode="USNM" pageId="2" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="814A8866DA2B441CFDC4FEA5BBF5A38F" box="[584,672,375,396]" pageId="2" type="holotype">Holotype</typeStatus>
|
||
deposited at
|
||
<collectionCode id="38E0AE01DA2B441CFCAEFEA5BA3FA38F" box="[802,874,375,396]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" type="Museum">USNM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="EE993C99DA2B441CFCF9FEA5BAD5A012" collectingDate="2002-07-30" collectionCode="USNM, UNSM, SDSU, CSU" collectorName="D. Backlund & S. Spomer & W. Allgeier. Three & W. Allgeier" county="Museum" location="University" municipality="University" pageId="2" specimenCount="37" specimenCount-female="16" specimenCount-male="16" stateProvince="Nebraska" typeStatus="paratype">
|
||
<typeStatus id="814A8866DA2B441CFCF9FEA5BA84A38F" box="[885,977,375,396]" pageId="2" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<specimenCount id="48F7FD4DDA2B441CFF17FE40B9A6A3A4" box="[155,243,402,423]" pageId="2" type="male">16 males</specimenCount>
|
||
,
|
||
<specimenCount id="48F7FD4DDA2B441CFF72FE40B83DA3A4" box="[254,360,402,423]" pageId="2" type="female">16 females</specimenCount>
|
||
) collected 6 July, 23 July, and
|
||
<date id="2A4F1004DA2B441CFD19FE40BA47A3A4" box="[661,786,402,423]" pageId="2" value="2002-07-30">
|
||
<collectingDate id="3A0BE9ECDA2B441CFD19FE40BA47A3A4" box="[661,786,402,423]" pageId="2" value="2002-07-30">30 July 2002</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="F3045312DA2B441CFCBBFE40BAE6A3A4" box="[823,947,402,423]" pageId="2">D. Backlund</collectorName>
|
||
,
|
||
<collectorName id="F3045312DA2B441CFC31FE40B9B5A3C2" pageId="2">S. Spomer</collectorName>
|
||
, and
|
||
<collectorName id="F3045312DA2B441CFE91FE7EB889A3C2" box="[285,476,428,449]" pageId="2">W. Allgeier. Three</collectorName>
|
||
<typeStatus id="814A8866DA2B441CFE64FE7EBB16A3C2" box="[488,579,428,449]" pageId="2" type="paratype">paratypes</typeStatus>
|
||
deposited at
|
||
<collectionCode id="38E0AE01DA2B441CFD5EFE7EBA4CA3C2" box="[722,793,428,449]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" type="Museum">USNM</collectionCode>
|
||
,
|
||
<specimenCount id="48F7FD4DDA2B441CFCA4FE7EBAE6A3C2" box="[808,947,428,449]" pageId="2" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
to
|
||
<collectionCode id="38E0AE01DA2B441CFF18FE15B98CA3DF" box="[148,217,455,476]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="2">UNSM</collectionCode>
|
||
(Univ.
|
||
<collectingRegion id="9C35F826DA2B441CFEA9FE15B8E1A3DF" box="[293,436,455,476]" country="United States of America" name="Nebraska" pageId="2">Nebraska State</collectingRegion>
|
||
<collectingCounty id="B72F4E48DA2B441CFE30FE15BB46A3DF" box="[444,531,455,476]" pageId="2">Museum</collectingCounty>
|
||
),
|
||
<specimenCount id="48F7FD4DDA2B441CFDA8FE15BBFFA3DF" box="[548,682,455,476]" pageId="2" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
to
|
||
<collectionCode id="38E0AE01DA2B441CFD47FE15BA5DA3DF" box="[715,776,455,476]" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" pageId="2">SDSU</collectionCode>
|
||
(
|
||
<collectingRegion id="9C35F826DA2B441CFC9AFE15BA84A3DF" box="[790,977,455,476]" country="United States of America" name="South Dakota" pageId="2">South Dakota State</collectingRegion>
|
||
<collectingMunicipality id="BE2AACBEDA2B441CFF18FE33B9A9A3F5" box="[148,252,481,502]" pageId="2">University</collectingMunicipality>
|
||
),
|
||
<specimenCount id="48F7FD4DDA2B441CFE82FE33B8C1A3F5" box="[270,404,481,502]" pageId="2" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
to
|
||
<collectionCode id="38E0AE01DA2B441CFE3BFE33B8B0A3F5" box="[439,485,481,502]" country="USA" httpUri="http://grbio.org/cool/hsbf-qixz" name="Colorado State University" pageId="2">CSU</collectionCode>
|
||
(
|
||
<collectingRegion id="9C35F826DA2B441CFE78FE33BBD0A3F5" box="[500,645,481,502]" country="United States of America" name="Colorado" pageId="2">Colorado State</collectingRegion>
|
||
<location id="5B2E601FDA2B441CFD01FE33BBA0A3F5" LSID="urn:lsid:plazi:treatment:D65887D2DA2B441AFE93FEC8BBC1A30E:5B2E601FDA2B441CFD01FE33BBA0A3F5" box="[653,757,481,502]" county="Museum" municipality="University" name="University" pageId="2" stateProvince="Nebraska">University</location>
|
||
),
|
||
<specimenCount id="48F7FD4DDA2B441CFC8BFE33BAC4A3F5" box="[775,913,481,502]" pageId="2" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
|
||
to the author, sixteen
|
||
<typeStatus id="814A8866DA2B441CFEA5FE2EB8D1A012" box="[297,388,508,529]" pageId="2" type="paratype">paratypes</typeStatus>
|
||
to
|
||
<collectorName id="F3045312DA2B441CFE2AFE2EBB76A012" box="[422,547,508,529]" pageId="2">D. Backlund</collectorName>
|
||
, and
|
||
<specimenCount id="48F7FD4DDA2B441CFDD4FE2EBBBFA012" box="[600,746,508,529]" pageId="2" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
|
||
to
|
||
<collectorName id="F3045312DA2B441CFC80FE2EBAD5A012" box="[780,896,508,529]" pageId="2">W. Allgeier</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16EB654FDA2B441CFF25FDF7B854A74D" pageId="2" type="description">
|
||
<paragraph id="5E4E36C4DA2B441CFF25FDF7BAEDA1E4" blockId="2.[148,978,549,999]" pageId="2">
|
||
<emphasis id="6C85EAD6DA2B441CFF25FDF7B842A03B" bold="true" box="[169,279,549,568]" pageId="2">Description.</emphasis>
|
||
Male. Length
|
||
<quantity id="99099B21DA2B441CFE17FDF4B8BDA03A" box="[411,488,550,569]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="2" unit="mm" value="10.5">10.5 mm</quantity>
|
||
. Color coppery with pronounced white maculation.
|
||
<emphasis id="6C85EAD6DA2B441CFC12FDF4BA87A03A" box="[926,978,550,569]" italics="true" pageId="2">Head.</emphasis>
|
||
Color coppery, with greenish reflections, especially along edges of eyes, near scape, and on clypeus and gena. Frons with wrinkles, most distinctive near eyes; middle of frons punctate and bearing many white, decumbent setae. Clypeus also punctate and bearing many white, decumbent setae. Labrum white, outer edge dark, unidentate with blunt tooth. Mandible white on outer edge about one-half way toward tip, teeth and tip of mandible black with green reflectance. Scape with 1 sensory seta and 3 decumbent setae, first 4 segments of antenna coppery with green reflectance, remaining 7 segments tomentose and dark brown. Maxillary palpi light brown with proximal segment dark brown; labial palpi identical in color to maxillary palpi, penultimate segment longest and heavily setose.
|
||
<emphasis id="6C85EAD6DA2B441CFEB2FD2FB82EA113" box="[318,379,765,784]" italics="true" pageId="2">Thorax</emphasis>
|
||
. Pronotum narrower than head across eyes, slightly wider than long; color coppery; median sulcus shallow, anterior sulcus deep with green reflectance, posterior sulcus deep, with slight green reflectance; numerous white setae except at prominent points.
|
||
<emphasis id="6C85EAD6DA2B441CFCEEFCFEBAC3A13C" box="[866,918,812,831]" italics="true" pageId="2">Elytra</emphasis>
|
||
. Color coppery; punctures with blue reflectance except on maculations; humeral edge with green reflectance. Scutellum coppery. Maculation wide, occupying approximately 50% of surface area. Middle band nearly fused with apical lunules; maculation continuous at outer edges of elytra; apical lunules and middle bands traceable but expanded. Microserrations present.
|
||
<emphasis id="6C85EAD6DA2B441CFC86FC5EBAC7A19C" box="[778,914,908,927]" italics="true" pageId="2">Ventral surface.</emphasis>
|
||
Lateral surface coppery and setose; prosternum coppery with green reflectance, glabrous. Metasternum and proepisternum metallic green. Genae coppery and metallic green, heavily setose. Femora coppery green. Tibiae and tarsi coppery with slight green reflectance. Legs with numerous white setae.
|
||
</paragraph>
|
||
<paragraph id="5E4E36C4DA2B441CFF20FC28B854A74D" blockId="2.[148,977,1018,1570]" pageId="2">
|
||
<emphasis id="6C85EAD6DA2B441CFF20FC28B841A60C" bold="true" box="[172,276,1018,1039]" pageId="2">Variation.</emphasis>
|
||
<emphasis id="6C85EAD6DA2B441CFEACFC28B837A60C" box="[288,354,1018,1039]" italics="true" pageId="2">Length</emphasis>
|
||
: male (n
|
||
<emphasis id="6C85EAD6DA2B441CFE35FC28B89EA60D" bold="true" box="[441,459,1018,1038]" italics="true" pageId="2">¼</emphasis>
|
||
17), x
|
||
<emphasis id="6C85EAD6DA2B441CFD86FC28BB49A60D" bold="true" box="[522,540,1018,1038]" italics="true" pageId="2">¼</emphasis>
|
||
<quantity id="99099B21DA2B441CFDACFC28BB22A60C" box="[544,631,1018,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="2" unit="mm" value="10.6">10.6 mm</quantity>
|
||
, ranging from 10.0 mm to
|
||
<quantity id="99099B21DA2B441CFCFAFC28BA98A60C" box="[886,973,1018,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="2" unit="mm" value="11.2">11.2 mm</quantity>
|
||
; female (n
|
||
<emphasis id="6C85EAD6DA2B441CFF7DFBC7B856A62A" bold="true" box="[241,259,1045,1065]" italics="true" pageId="2">¼</emphasis>
|
||
16), x
|
||
<emphasis id="6C85EAD6DA2B441CFECEFBC7B801A62A" bold="true" box="[322,340,1045,1065]" italics="true" pageId="2">¼</emphasis>
|
||
<quantity id="99099B21DA2B441CFED4FBC7B8FAA629" box="[344,431,1045,1066]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="2" unit="mm" value="11.6">11.6 mm</quantity>
|
||
, ranging from
|
||
<quantity id="99099B21DA2B441CFDB7FBC7BB8EA629" box="[571,731,1045,1066]" metricMagnitude="-2" metricUnit="m" metricValue="1.145" metricValueMax="1.22" metricValueMin="1.0699999999999998" pageId="2" unit="mm" value="11.45" valueMax="12.2" valueMin="10.7">10.7 to 12.2 mm</quantity>
|
||
.
|
||
<emphasis id="6C85EAD6DA2B441CFD68FBC7BA4AA629" box="[740,799,1045,1066]" italics="true" pageId="2">Head.</emphasis>
|
||
Color coppery but greenish reflections sometimes absent; greenish reflections often present at posterior vertex. Scape with up to 11 decumbent setae.
|
||
<emphasis id="6C85EAD6DA2B441CFDDCFB98BBC0A65C" box="[592,661,1098,1119]" italics="true" pageId="2">Thorax</emphasis>
|
||
. Anterior sulcus with or without green reflectance, posterior sulcus usually without green reflectance.
|
||
<emphasis id="6C85EAD6DA2B441CFCC2FBB6BADCA67A" box="[846,905,1124,1145]" italics="true" pageId="2">Elytra</emphasis>
|
||
. Color coppery (88%) or coppery with green reflectance (12%); pits with green or blue reflectance except on maculations; humeral edge with blue or green reflectance in most (94%) specimens; scutellum coppery (55%) or with blue or green reflectance (45%); maculation wide to very wide, occupying up to nearly 70% of the surface area. One specimen (3%) with humeral band fused with middle band and
|
||
<specimenCount id="48F7FD4DDA2B441CFD60FB3BBA35A6FD" box="[748,864,1257,1278]" pageId="2" type="generic">7 specimens</specimenCount>
|
||
(21%) with middle band fused with apical lunules.
|
||
<emphasis id="6C85EAD6DA2B441CFDA9FAD6BBE8A71A" box="[549,701,1284,1305]" italics="true" pageId="2">Ventral surface</emphasis>
|
||
. Genae often only metallic green; femora green or coppery green; tibiae and tarsi coppery or coppery with green reflectance.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16EB654FDA2B441AFF20FA81BBC1A30E" lastPageId="4" pageId="2" type="etymology">
|
||
<paragraph id="5E4E36C4DA2B441CFF20FA81BA2AA780" blockId="2.[148,977,1018,1570]" pageId="2">
|
||
<emphasis id="6C85EAD6DA2B441CFF20FA81B875A76B" bold="true" box="[172,288,1363,1384]" pageId="2">Etymology.</emphasis>
|
||
The name
|
||
<emphasis id="6C85EAD6DA2B441CFE16FA81B8A1A76B" box="[410,500,1363,1384]" italics="true" pageId="2">makosika</emphasis>
|
||
is a composite of two Sioux Indian words for ‘‘land’’ and ‘‘bad’’ (
|
||
<emphasis id="6C85EAD6DA2B441CFED2FABCB8D4A780" box="[350,385,1390,1411]" italics="true" pageId="2">i.e.,</emphasis>
|
||
badlands), the area in which this subspecies occurs.
|
||
</paragraph>
|
||
<paragraph id="5E4E36C4DA2B441CFF20FA5AB9A6A421" blockId="2.[148,977,1018,1570]" pageId="2">
|
||
<emphasis id="6C85EAD6DA2B441CFF20FA5AB821A79E" bold="true" box="[172,372,1416,1437]" pageId="2">Similar Subspecies.</emphasis>
|
||
<emphasis id="6C85EAD6DA2B441CFE0CFA5BBB0CA79D" box="[384,601,1417,1438]" italics="true" pageId="2">
|
||
<taxonomicName id="99F14D47DA2B441CFE0CFA5BB889A79D" authorityName="Linnaeus" authorityYear="1758" box="[384,476,1417,1438]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">Cicindela</taxonomicName>
|
||
n. makosika
|
||
</emphasis>
|
||
most closely resembles
|
||
<emphasis id="6C85EAD6DA2B441CFCC6FA5BBA84A79D" box="[842,977,1417,1438]" italics="true" pageId="2">C. n. tubensis</emphasis>
|
||
and the population of
|
||
<taxonomicName id="99F14D47DA2B441CFEE8FA71B883A7BB" box="[356,470,1443,1464]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="nevadica">
|
||
<emphasis id="6C85EAD6DA2B441CFEE8FA71B883A7BB" box="[356,470,1443,1464]" italics="true" pageId="2">C. nevadica</emphasis>
|
||
</taxonomicName>
|
||
in Torrance Co.,
|
||
<collectingRegion id="9C35F826DA2B441CFDF1FA71BBA2A7BB" box="[637,759,1443,1464]" country="United States of America" name="New Mexico" pageId="2">New Mexico</collectingRegion>
|
||
being referred to as
|
||
<emphasis id="6C85EAD6DA2B441CFC37FA71B9A3A7D0" italics="true" pageId="2">C. n. olmosa</emphasis>
|
||
. However,
|
||
<emphasis id="6C85EAD6DA2B441CFEE2FA6CBB53A7D0" box="[366,518,1470,1491]" italics="true" pageId="2">C. n. makosika</emphasis>
|
||
is slightly larger and less reddish than
|
||
<emphasis id="6C85EAD6DA2B441CFC12FA6CB9B7A7EE" italics="true" pageId="2">C. n. tubensis</emphasis>
|
||
and much more coppery than the Torrance Co. population. The most heavily maculated
|
||
<emphasis id="6C85EAD6DA2B441CFF72FA21B82EA40B" box="[254,379,1523,1544]" italics="true" pageId="2">C. n. knausii</emphasis>
|
||
examined was less maculated than the least maculated
|
||
<emphasis id="6C85EAD6DA2B441CFC2CFA21B9BBA421" italics="true" pageId="2">C. n. makosika</emphasis>
|
||
.
|
||
</paragraph>
|
||
<caption id="0A8E664CDA2A441DFF49FAE7BA40A74B" ID-DOI="http://doi.org/10.5281/zenodo.5345295" ID-Zenodo-Dep="5345295" box="[197,789,1333,1352]" httpUri="https://zenodo.org/record/5345295/files/figure.png" pageId="3" startId="3.[197,228,1333,1352]" targetBox="[382,798,194,1312]" targetPageId="3">
|
||
<paragraph id="5E4E36C4DA2A441DFF49FAE7BA40A74B" blockId="3.[197,789,1333,1352]" box="[197,789,1333,1352]" pageId="3">
|
||
<emphasis id="6C85EAD6DA2A441DFF49FAE7B9AAA74B" bold="true" box="[197,255,1333,1352]" pageId="3">Fig. 1.</emphasis>
|
||
Habitus, holotype of
|
||
<taxonomicName id="99F14D47DA2A441DFE47FAE7BA40A74B" authority="Spomer." box="[459,789,1333,1352]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="subSpecies" species="nevadica" subSpecies="makosika">
|
||
<emphasis id="6C85EAD6DA2A441DFE47FAE7BB93A74B" box="[459,710,1333,1352]" italics="true" pageId="3">Cicindela nevadica makosika</emphasis>
|
||
Spomer.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5E4E36C4DA2A441AFF4BFABCBBC1A30E" blockId="3.[175,1005,1390,1570]" lastBlockId="4.[148,977,194,269]" lastPageId="4" pageId="3">
|
||
<emphasis id="6C85EAD6DA2A441DFF4BFABCB849A780" bold="true" box="[199,284,1390,1411]" pageId="3">Habitat.</emphasis>
|
||
Indian Creek is an intermittent stream that probably picks up salinity from seeps through cracks in the underlying Pierre shale. In lower Indian Creek (where
|
||
<emphasis id="6C85EAD6DA2A441DFC32FA5BB85CA7BB" italics="true" pageId="3">C. n. makosika</emphasis>
|
||
occurs), portions of the streambed consist of a light colored, viscous mud overlying the Pierre shale.
|
||
<emphasis id="6C85EAD6DA2A441DFE3EFA6CBBDDA7D0" box="[434,648,1470,1491]" italics="true" pageId="3">
|
||
<taxonomicName id="99F14D47DA2A441DFE3EFA6CBB5BA7D0" authorityName="Linnaeus" authorityYear="1758" box="[434,526,1470,1491]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="genus">Cicindela</taxonomicName>
|
||
n. makosika
|
||
</emphasis>
|
||
was found almost exclusively where this mud occurred and was not found on other badlands streams that lacked this feature of Pierre shale underlying the stream bed. Willis (1967) suggested that selection pressure on tiger beetles in saline habitats has caused certain species to match the color 412 of the soil in the local area and to match the white of the salt. The expanded white maculations on the elytra of
|
||
<emphasis id="6C85EAD6DA2D441AFE29FF0FBB61A2F1" box="[421,564,221,242]" italics="true" pageId="4">C. n. makosika</emphasis>
|
||
seems to correspond with the light colored mud along Indian Creek, thus fitting this hypothesis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |