treatments-xml/data/9F/59/0A/9F590A132A9001A571AD92C5BBE0923F.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.85.12288" ID-PMC="PMC5624200" ID-Pensoft-Pub="1314-2003-85-109" ID-Pensoft-UUID="FFAD3505F142FFD38A48014E23541871" ID-PubMed="29033664" ID-Zenodo-Dep="899045" ModsDocID="1314-2003-85-109" checkinTime="1504194362271" checkinUser="pensoft" docAuthor="Brullo, Salvatore, Brullo, Cristian, Cambria, Salvatore, Cristaudo, Antonia &amp; Galdo, Gianpietro Giusso del" docDate="2017" docId="9F590A132A9001A571AD92C5BBE0923F" docLanguage="en" docName="PhytoKeys 85: 109-124" docOrigin="PhytoKeys 85" docPubDate="2017-08-31" docSource="http://dx.doi.org/10.3897/phytokeys.85.12288" docTitle="Bituminaria antiatlantica Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso 2017, sp. nov." docType="treatment" docVersion="7" id="FFAD3505F142FFD38A48014E23541871" lastPageNumber="113" masterDocId="FFAD3505F142FFD38A48014E23541871" masterDocTitle="Bituminaria antiatlantica (Psoraleeae, Fabaceae), a new species from Morocco" masterLastPageNumber="124" masterPageNumber="109" pageNumber="111" updateTime="1668142035405" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Bituminaria antiatlantica (Psoraleeae, Fabaceae), a new species from Morocco</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brullo, Salvatore</mods:namePart>
<mods:affiliation>Department of Biological, Geological and Environmental Sciences, University of Catania, Via A. Longo 19, Catania 95125, Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brullo, Cristian</mods:namePart>
<mods:affiliation>Department of Biological, Geological and Environmental Sciences, University of Catania, Via A. Longo 19, Catania 95125, Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cambria, Salvatore</mods:namePart>
<mods:affiliation>Department of Biological, Geological and Environmental Sciences, University of Catania, Via A. Longo 19, Catania 95125, Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cristaudo, Antonia</mods:namePart>
<mods:affiliation>Department of Biological, Geological and Environmental Sciences, University of Catania, Via A. Longo 19, Catania 95125, Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Galdo, Gianpietro Giusso del</mods:namePart>
<mods:affiliation>Department of Biological, Geological and Environmental Sciences, University of Catania, Via A. Longo 19, Catania 95125, Italy</mods:affiliation>
<mods:nameIdentifier type="email">g.giusso@unict.it</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-08-31</mods:number>
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<mods:detail type="volume">
<mods:number>85</mods:number>
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<mods:start>109</mods:start>
<mods:end>124</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.85.12288</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.85.12288</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-85-109</mods:identifier>
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<treatment ID-GBIF-Taxon="133134951" LSID="urn:lsid:plazi:treatment:9F590A132A9001A571AD92C5BBE0923F" httpUri="http://treatment.plazi.org/id/9F590A132A9001A571AD92C5BBE0923F" lastPageId="4" lastPageNumber="113" pageId="2" pageNumber="111">
<subSubSection pageId="2" pageNumber="111" type="nomenclature">
<paragraph pageId="2" pageNumber="111">
<taxonomicName LSID="9F590A13-2A90-01A5-71AD-92C5BBE0923F" authority="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Fabaceae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="antiatlantica" status="sp. nov.">Bituminaria antiatlantica Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="111">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Diagnostic features regarding the reproductive structures of Bituminaria antiatlantica. A Flower (ventral view) B Flower (dorsal view) C Flower (lateral view) D Bud E Standard F Wings G Keel (lateral view) H Calyx (open) I Staminal tube J Anthers K Pisti L Stigma M Fruiting calyx and pod N Pods O Seed. Illustration by S. Brullo based on living material coming from Mount Tachilla and Djebel Imzi in Morocco (CAT)." figureDoi="10.3897/phytokeys.85.12288.figure1" httpUri="https://binary.pensoft.net/fig/153323" pageId="2" pageNumber="111">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic features regarding the vegetative structures of Bituminaria antiatlantica. A Habit B Leaves C Leaf apex (abaxial side) D Leaf apex (adaxial side) E Stipules. Illustration by S. Brullo based on living material coming from Mount Tachilla and Djebel Imzi in Morocco (CAT)." figureDoi="10.3897/phytokeys.85.12288.figure2" httpUri="https://binary.pensoft.net/fig/153324" pageId="2" pageNumber="111">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="111" type="diagnosis">
<paragraph pageId="2" pageNumber="111">
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria bituminosa" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="bituminosa">Bituminaria bituminosa</taxonomicName>
<emphasis italics="true" pageId="2" pageNumber="111">(L.) C.H. Stirt. affinis, sed habitu suffruticoso, ramis lignosis, foliolis glabris vel sparsim pilosis, subrotundatis vel ovatis, max. 35 mm longis, petiolis usque ad 6 cm longis, inflorescentia laxa, saepe subspicata, 1,5-2 cm longa, 3-10-flora, calice 12-13.5 mm longo, corolla pallida</emphasis>
.
</paragraph>
<caption ID-Zenodo-Dep="991829" doi="10.3897/phytokeys.85.12288.figure1" httpUri="https://binary.pensoft.net/fig/153323" pageId="2" pageNumber="111" start="Figure 1" startId="F1">
<paragraph pageId="2" pageNumber="111">
<emphasis bold="true" pageId="2" pageNumber="111">Figure 1.</emphasis>
Diagnostic features regarding the reproductive structures of 
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">Bituminaria antiatlantica</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="2" pageNumber="111">A</emphasis>
Flower (ventral view) 
<emphasis bold="true" pageId="2" pageNumber="111">B</emphasis>
Flower (dorsal view) 
<emphasis bold="true" pageId="2" pageNumber="111">C</emphasis>
Flower (lateral view) 
<emphasis bold="true" pageId="2" pageNumber="111">D</emphasis>
Bud 
<emphasis bold="true" pageId="2" pageNumber="111">E</emphasis>
Standard 
<emphasis bold="true" pageId="2" pageNumber="111">F</emphasis>
Wings 
<emphasis bold="true" pageId="2" pageNumber="111">G</emphasis>
Keel (lateral view) 
<emphasis bold="true" pageId="2" pageNumber="111">H</emphasis>
Calyx (open) 
<emphasis bold="true" pageId="2" pageNumber="111">I</emphasis>
Staminal tube 
<emphasis bold="true" pageId="2" pageNumber="111">J</emphasis>
Anthers 
<emphasis bold="true" pageId="2" pageNumber="111">K</emphasis>
Pisti 
<emphasis bold="true" pageId="2" pageNumber="111">L</emphasis>
Stigma 
<emphasis bold="true" pageId="2" pageNumber="111">M</emphasis>
Fruiting calyx and pod 
<emphasis bold="true" pageId="2" pageNumber="111">N</emphasis>
Pods 
<emphasis bold="true" pageId="2" pageNumber="111">O</emphasis>
Seed. Illustration by S. Brullo based on living material coming from Mount Tachilla and Djebel Imzi in Morocco (CAT).
</paragraph>
</caption>
<caption ID-Zenodo-Dep="991831" doi="10.3897/phytokeys.85.12288.figure2" httpUri="https://binary.pensoft.net/fig/153324" pageId="2" pageNumber="111" start="Figure 2" startId="F2">
<paragraph pageId="2" pageNumber="111">
<emphasis bold="true" pageId="2" pageNumber="111">Figure 2.</emphasis>
Diagnostic features regarding the vegetative structures of 
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">Bituminaria antiatlantica</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="2" pageNumber="111">A</emphasis>
Habit 
<emphasis bold="true" pageId="2" pageNumber="111">B</emphasis>
Leaves 
<emphasis bold="true" pageId="2" pageNumber="111">C</emphasis>
Leaf apex (abaxial side) 
<emphasis bold="true" pageId="2" pageNumber="111">D</emphasis>
Leaf apex (adaxial side) 
<emphasis bold="true" pageId="2" pageNumber="111">E</emphasis>
Stipules. Illustration by S. Brullo based on living material coming from Mount Tachilla and Djebel Imzi in Morocco (CAT).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="111" type="synonym">
<paragraph pageId="2" pageNumber="111">Synonym.</paragraph>
<paragraph pageId="2" pageNumber="111">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Psoralea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psoralea bituminosa subsp. var. var. rotundata" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="variety" species="bituminosa" subSpecies="var." variety="rotundata">Psoralea bituminosa L. var. rotundata</taxonomicName>
Maire, Bull. Soc. Hist. Nat. Afr. N. 27(8): 222, 1936.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="111" type="materials_examined">
<paragraph pageId="2" pageNumber="111">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="2" pageNumber="111">
<materialsCitation collectingDate="1935-04-10" collectorName="R. Maire, E. Wilczek, Moris" country="Morocco" county="Anti-Atlantis" elevation="400" location="In" municipality="Mountis Tachilla" specimenCount="1" typeStatus="holotype">
<collectingCountry name="Morocco">Morocco</collectingCountry>
: 
<location LSID="urn:lsid:plazi:treatment:9F590A132A9001A571AD92C5BBE0923F:BE9365C9B27440294A0148C1A341C472" country="Morocco" county="Anti-Atlantis" municipality="Mountis Tachilla" name="In">In</location>
rupibus arenaceis 
<collectingMunicipality>Mountis Tachilla</collectingMunicipality>
ad radices septentr. 
<collectingCounty>Anti-Atlantis</collectingCounty>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</elevation>
</quantity>
, 
<collectingDate value="1935-04-10">10 April 1935</collectingDate>
, 
<emphasis italics="true" pageId="2" pageNumber="111">
<collectorName>R. Maire</collectorName>
&amp; 
<collectorName>E. Wilczek</collectorName>
s.n.
</emphasis>
(
<typeStatus>holotype</typeStatus>
MPU!; isotype RAB!), sub 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Psoralea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psoralea bituminosa subsp. var. var. latifolia" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="variety" species="bituminosa" subSpecies="var." variety="latifolia">Psoralea bituminosa L. var. latifolia</taxonomicName>
<collectorName>Moris</collectorName>
f. 
<taxonomicName class="Magnoliopsida" family="Fabaceae" form="rotundata" genus="Psoralea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="form">rotundata</taxonomicName>
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="111" type="description">
<paragraph pageId="2" pageNumber="111">Description.</paragraph>
<paragraph pageId="2" pageNumber="111">
Perennial, suffruticose, dark green, erect to ascending, up to 60 cm tall. 
<emphasis italics="true" pageId="2" pageNumber="111">Stems</emphasis>
dark green-brown, sparsely hairy, with hairs short and appressed, very branched; branches woody, leafy along entire length. 
<emphasis italics="true" pageId="2" pageNumber="111">Stipules</emphasis>
5-6 mm long, rigid, linear-triangular, adnate to the petiole. 
<emphasis italics="true" pageId="2" pageNumber="111">Leaves</emphasis>
pinnately 3-foliolate, green, with petiole 1.8-6(7) cm long, sparsely hairy; 
<emphasis italics="true" pageId="2" pageNumber="111">leaflets</emphasis>
semi-round to ovate, subglabrous above and sparsely hairy below, 10-35 
<normalizedToken originalValue="×">x</normalizedToken>
8-21 mm, with apex obtuse to acute, ending in a straight mucro 0.3-0.5 mm long. 
<emphasis italics="true" pageId="2" pageNumber="111">Inflorescence</emphasis>
definite, subspicate, lax, 1.5-2 cm long, with 3-10 flowers. 
<emphasis italics="true" pageId="2" pageNumber="111">Peduncle</emphasis>
3.5-14 cm long, overtopping the leaves. 
<emphasis italics="true" pageId="2" pageNumber="111">Bracts</emphasis>
1-3 toothed, 5-8 mm long, subtending 2 or more flowers. 
<emphasis italics="true" pageId="2" pageNumber="111">Flowers</emphasis>
16-17 mm long. 
<emphasis italics="true" pageId="2" pageNumber="111">Calyx</emphasis>
12-13.5 mm long, green, densely hairy, with hairs white mixed to short black hairs; lower teeth 7-8 mm long, laterals shorter, 5.5-7 mm long. 
<emphasis italics="true" pageId="2" pageNumber="111">Corolla</emphasis>
whitish-pink to whitish lilac; standard 16-16.5 
<normalizedToken originalValue="×">x</normalizedToken>
7-7.5 mm, elliptic, striate with lilac in the middle, apex obtuse; wings 14-15 
<normalizedToken originalValue="×">x</normalizedToken>
3-4 mm; keel 10.5-11 
<normalizedToken originalValue="×">x</normalizedToken>
2-2.3 mm, having a macula dark violet in the upper part. 
<emphasis italics="true" pageId="2" pageNumber="111">Staminal tube</emphasis>
11-11.5 mm long, with anthers yellow, 0.7-0.8 
<normalizedToken originalValue="×">x</normalizedToken>
0.3-0.35 mm; vexillary w with filament fused below with the other ones. Pistil 10-10.5 long, ovary hairy, style curved towards the apex, thickened at point of flexure, stigma capitate, penicillate. and ovary hairy. Pod 11-23 mm long (beak included), with beak pubescent, 14-16 mm long. Seed reniform, 6-7 
<normalizedToken originalValue="×">x</normalizedToken>
3.4-4 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="111" type="distribution">
<paragraph pageId="2" pageNumber="111">Distribution and ecology.</paragraph>
<paragraph pageId="2" pageNumber="111">
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">Bituminaria antiatlantica</emphasis>
</taxonomicName>
is a rare and localized species, currently known only from Mount Tachilla and Djebel Imzi in the Anti-Atlas Mountains in southern Morocco, (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of Bituminaria antiatlantica (red dots) based on herbarium specimens." figureDoi="10.3897/phytokeys.85.12288.figure3" httpUri="https://binary.pensoft.net/fig/153325" pageId="2" pageNumber="111">3</figureCitation>
). It grows between 300 and 1500 m of altitude, on steep, north-facing slopes, chiefly constituted of Precambrian quartzite. From the bioclimatic viewpoint, this area falls within the infra- and the thermo-mediterranean type, with semiarid to subhumid ombrotype (Benabid and Cuzin, 1997). This species is an element of a chasmophytic vegetation type characterized by 
<taxonomicName authorityName="Murb" authorityYear="1925" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Celsia longirostris" order="Lamiales" pageId="2" pageNumber="111" phylum="Magnoliophyta" rank="species" species="longirostris">Celsia longirostris</taxonomicName>
Murb. var. 
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="var. antiatlantica" order="Lamiales" pageId="2" pageNumber="111" phylum="Magnoliophyta" rank="variety" species="longirostris" variety="antiatlantica">antiatlantica</taxonomicName>
Emb., 
<taxonomicName class="Dicotyledoneae" family="Lamiaceae" genus="Salvia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salvia taraxacifolia" order="Tubiflorae" pageId="2" pageNumber="111" phylum="Angiospermae" rank="species" species="taraxacifolia">
<emphasis italics="true" pageId="2" pageNumber="111">Salvia taraxacifolia</emphasis>
</taxonomicName>
Coss. &amp; Balansa, 
<taxonomicName genus="Chiladenus" lsidName="Chiladenus hesperius" pageId="2" pageNumber="111" rank="species" species="hesperius">
<emphasis italics="true" pageId="2" pageNumber="111">Chiladenus hesperius</emphasis>
</taxonomicName>
(Maire &amp; Wilczek) Brullo, 
<taxonomicName class="Dicotyledoneae" family="Lamiaceae" genus="Teucrium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Teucrium werneri" order="Tubiflorae" pageId="2" pageNumber="111" phylum="Angiospermae" rank="species" species="werneri">
<emphasis italics="true" pageId="2" pageNumber="111">Teucrium werneri</emphasis>
</taxonomicName>
Emb., 
<taxonomicName class="Magnoliopsida" family="Crassulaceae" genus="Aeonium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Aeonium arboreum" order="Rosales" pageId="2" pageNumber="111" phylum="Magnoliophyta" rank="species" species="arboreum">
<emphasis italics="true" pageId="2" pageNumber="111">Aeonium arboreum</emphasis>
</taxonomicName>
(L.) Webb &amp; Berthel, 
<taxonomicName class="Dicotyledoneae" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Dianthus lusitanus" order="Centrospermae" pageId="2" pageNumber="111" phylum="Angiospermae" rank="species" species="lusitanus">
<emphasis italics="true" pageId="2" pageNumber="111">Dianthus lusitanus</emphasis>
</taxonomicName>
Brot., 
<taxonomicName authorityName="Maire" authorityYear="1916" baseAuthorityName="Briq." class="Dicotyledoneae" family="Lamiaceae" genus="Micromeria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Micromeria hochreutineri" order="Tubiflorae" pageId="2" pageNumber="111" phylum="Angiospermae" rank="species" species="hochreutineri">
<emphasis italics="true" pageId="2" pageNumber="111">Micromeria hochreutineri</emphasis>
</taxonomicName>
(Briq.) Maire, 
<taxonomicName class="Magnoliopsida" family="Asclepiadaceae" genus="Caralluma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Caralluma hesperidum" order="Gentianales" pageId="2" pageNumber="111" phylum="Magnoliophyta" rank="species" species="hesperidum">
<emphasis italics="true" pageId="2" pageNumber="111">Caralluma hesperidum</emphasis>
</taxonomicName>
Maire, 
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Teline" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Teline segonnei" order="Fabales" pageId="2" pageNumber="111" phylum="Magnoliophyta" rank="species" species="segonnei">
<emphasis italics="true" pageId="2" pageNumber="111">Teline segonnei</emphasis>
</taxonomicName>
(Maire) Raynaud, 
<taxonomicName class="Filicopsida" family="Davalliaceae" genus="Davallia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Davallia canariensis" order="Aspidiales" pageId="2" pageNumber="111" phylum="Pteridophyta" rank="species" species="canariensis">
<emphasis italics="true" pageId="2" pageNumber="111">Davallia canariensis</emphasis>
</taxonomicName>
(L.) Sm., 
<taxonomicName class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium aethiopicum" order="Polypodiales" pageId="2" pageNumber="111" phylum="Polypodiopsida" rank="species" species="aethiopicum">
<emphasis italics="true" pageId="2" pageNumber="111">Asplenium aethiopicum</emphasis>
</taxonomicName>
(Burm. fil.) Becherer (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phenological features of Bituminaria antiatlantica A Natural habitat with Bituminaria antiatlantica in Djebel Imzi (Morocco) B Habit of B. antiatlantica in Mount Tachilla (Morocco) C Natural habitat of B. antiatlantica with Dracaena draco L. subsp. ajgal in Djebel Imzi (Morocco) D Inflorescence detail of B. antiatlantica E Fructified inflorescence of B. antiatlantica F Leaf detail of B. antiatlantica (Photos by S. Cambria)." figureDoi="10.3897/phytokeys.85.12288.figure4" httpUri="https://binary.pensoft.net/fig/153326" pageId="2" pageNumber="111">4C</figureCitation>
). Besides, several phanaerophytes occur in these rupestrian habitats such as 
<taxonomicName authorityName="Benabid &amp; Cuzin" authorityYear="1997" class="Monocotyledoneae" family="Liliaceae" genus="Dracaena" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Dracaena draco subsp. ajgal" order="Liliiflorae" pageId="2" pageNumber="111" phylum="Angiospermae" rank="subSpecies" species="draco" subSpecies="ajgal">Dracaena draco L. subsp. ajgal</taxonomicName>
Benabid &amp; Cuzin, 
<taxonomicName class="Dicotyledoneae" family="Lauraceae" genus="Laurus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Laurus azorica" order="Magnoliales" pageId="2" pageNumber="111" phylum="Angiospermae" rank="species" species="azorica">
<emphasis italics="true" pageId="2" pageNumber="111">Laurus azorica</emphasis>
</taxonomicName>
(Seub.) Franco, 
<taxonomicName authorityName="Skeels" authorityYear="1911" class="Magnoliopsida" family="Sapotaceae" genus="Argania" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Argania spinosa" order="Ericales" pageId="2" pageNumber="111" phylum="Magnoliophyta" rank="species" species="spinosa">
<emphasis italics="true" pageId="2" pageNumber="111">Argania spinosa</emphasis>
</taxonomicName>
L., 
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Kleinia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Kleinia anteuphorbium" order="Asterales" pageId="2" pageNumber="111" phylum="Magnoliophyta" rank="species" species="anteuphorbium">
<emphasis italics="true" pageId="2" pageNumber="111">Kleinia anteuphorbium</emphasis>
</taxonomicName>
(L.) Haw., 
<taxonomicName class="Dicotyledoneae" family="Anacardiaceae" genus="Rhus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Rhus tripartita" order="Sapindales" pageId="2" pageNumber="111" phylum="Angiospermae" rank="species" species="tripartita">
<emphasis italics="true" pageId="2" pageNumber="111">Rhus tripartita</emphasis>
</taxonomicName>
(Ucria) Grande, 
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Euphorbia echinus" order="Geraniales" pageId="2" pageNumber="111" phylum="Angiospermae" rank="species" species="echinus">
<emphasis italics="true" pageId="2" pageNumber="111">Euphorbia echinus</emphasis>
</taxonomicName>
Hook.f. &amp; Coss., 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Warionia" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Warionia saharae" order="Asterales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="saharae">
<emphasis italics="true" pageId="2" pageNumber="111">Warionia saharae</emphasis>
</taxonomicName>
Benth. et Coss. Many of these species are rare and endemic to the southern part of Morocco, thus highlighting the relic character of this plant community. In particular, 
<bibRefCitation author="Benabid, A" journalOrPublisher="Sciences de la vie" pageId="5" pageNumber="114" pagination="267 - 277" refId="B3" refString="Benabid, A, Cuzin, F, 1997. Populations de dragonnier (Dracaena draco L. subsp. ajal Benabid et Cuzin) au Maroc: valeurs taxinomique, biogeographique et phytosociologique. C. R. Academie des Sciences Paris. Sciences de la vie 320: 267 - 277" title="Populations de dragonnier (Dracaena draco L. subsp. ajal Benabid et Cuzin) au Maroc: valeurs taxinomique, biogeographique et phytosociologique. C. R. Academie des Sciences Paris." volume="320" year="1997">Benabid and Cuzin (1997)</bibRefCitation>
consider 
<taxonomicName lsidName="B. antiatlantica" pageId="2" pageNumber="111" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">B. antiatlantica</emphasis>
</taxonomicName>
(sub 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Psoralea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psoralea bituminosa subsp. var. var. rotundata" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="variety" species="bituminosa" subSpecies="var." variety="rotundata">Psoralea bituminosa var. rotundata</taxonomicName>
) as characteristic of a very peculiar and remarkable association exclusively occurring in a small area of the Anti Atlas range.
</paragraph>
<caption ID-Zenodo-Dep="991833" doi="10.3897/phytokeys.85.12288.figure3" httpUri="https://binary.pensoft.net/fig/153325" pageId="2" pageNumber="111" start="Figure 3" startId="F3">
<paragraph pageId="2" pageNumber="111">
<emphasis bold="true" pageId="2" pageNumber="111">Figure 3.</emphasis>
Distribution map of 
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">Bituminaria antiatlantica</emphasis>
</taxonomicName>
(red dots) based on herbarium specimens.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="991835" doi="10.3897/phytokeys.85.12288.figure4" httpUri="https://binary.pensoft.net/fig/153326" pageId="2" pageNumber="111" start="Figure 4" startId="F4">
<paragraph pageId="2" pageNumber="111">
<emphasis bold="true" pageId="2" pageNumber="111">Figure 4.</emphasis>
Phenological features of 
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">Bituminaria antiatlantica</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="2" pageNumber="111">A</emphasis>
Natural habitat with 
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">Bituminaria antiatlantica</emphasis>
</taxonomicName>
in Djebel Imzi (Morocco) 
<emphasis bold="true" pageId="2" pageNumber="111">B</emphasis>
Habit of 
<taxonomicName lsidName="B. antiatlantica" pageId="2" pageNumber="111" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">B. antiatlantica</emphasis>
</taxonomicName>
in Mount Tachilla (Morocco) 
<emphasis bold="true" pageId="2" pageNumber="111">C</emphasis>
Natural habitat of 
<taxonomicName lsidName="B. antiatlantica" pageId="2" pageNumber="111" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">B. antiatlantica</emphasis>
</taxonomicName>
with 
<taxonomicName authorityName="Benabid &amp; Cuzin" authorityYear="1997" class="Monocotyledoneae" family="Liliaceae" genus="Dracaena" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Dracaena draco subsp. ajgal" order="Liliiflorae" pageId="2" pageNumber="111" phylum="Angiospermae" rank="subSpecies" species="draco" subSpecies="ajgal">Dracaena draco L. subsp. ajgal</taxonomicName>
in Djebel Imzi (Morocco) 
<emphasis bold="true" pageId="2" pageNumber="111">D</emphasis>
Inflorescence detail of 
<taxonomicName lsidName="B. antiatlantica" pageId="2" pageNumber="111" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">B. antiatlantica</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="2" pageNumber="111">E</emphasis>
Fructified inflorescence of 
<taxonomicName lsidName="B. antiatlantica" pageId="2" pageNumber="111" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">B. antiatlantica</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="2" pageNumber="111">F</emphasis>
Leaf detail of 
<taxonomicName lsidName="B. antiatlantica" pageId="2" pageNumber="111" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">B. antiatlantica</emphasis>
</taxonomicName>
(Photos by S. Cambria).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="111" type="etymology">
<paragraph pageId="2" pageNumber="111">Etymology.</paragraph>
<paragraph pageId="2" pageNumber="111">The specific epithet refers to the Anti-Atlas range, where the species occurs.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="111" type="conservation status">
<paragraph pageId="2" pageNumber="111">Conservation status.</paragraph>
<paragraph pageId="2" pageNumber="111">
Based on current knowledge, 
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="2" pageNumber="111">Bituminaria antiatlantica</emphasis>
</taxonomicName>
seems to have a scattered distribution over an area smaller than 2,000 km2. Therefore, following the IUCN criteria (2014), this species should be classified as 
<normalizedToken originalValue="“Vulnerable”">&quot;Vulnerable&quot;</normalizedToken>
(VU B2). As regards the conservation policy of the growing site, it has been proposed its inclusion in the list of the UNESCO World Heritage Sites for its richness in endemic, rare or important plants, as well as for its breath-taking landscape (see http://whc.unesco.org/en/tentativelists/1180/).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="112" pageId="2" pageNumber="111" type="seed and pod micro-morphology">
<paragraph pageId="2" pageNumber="111">Seed and pod micro-morphology.</paragraph>
<paragraph lastPageId="3" lastPageNumber="112" pageId="2" pageNumber="111">
As emphasized by several authors (
<bibRefCitation DOI="https://doi.org/10.1111/j.1756-1051.1981.tb00704.x" author="Barthlott, W" journalOrPublisher="Nordic Journal of Botany" pageId="5" pageNumber="114" pagination="345 - 355" refId="B2" refString="Barthlott, W, 1981. Epidermal and seed surface character of plants: Systematic applicability and some evolutionary aspects. Nordic Journal of Botany 1: 345 - 355, DOI: https://doi.org/10.1111/j.1756-1051.1981.tb00704.x" title="Epidermal and seed surface character of plants: Systematic applicability and some evolutionary aspects." url="https://doi.org/10.1111/j.1756-1051.1981.tb00704.x" volume="1" year="1981">Barthlott 1981</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.1006/anbo.2000.1197" author="Koul, KK" journalOrPublisher="Annals of Botany" pageId="6" pageNumber="115" pagination="385 - 397" refId="B23" refString="Koul, KK, Nagpal, R, Raina, SN, 2000. Seed coat microsculpturing in Brassica and allied genera (Subtribe Brassicinae, Raphaninae, Moricandinae). Annals of Botany 86: 385 - 397, DOI: https://doi.org/10.1006/anbo.2000.1197" title="Seed coat microsculpturing in Brassica and allied genera (Subtribe Brassicinae, Raphaninae, Moricandinae)." url="https://doi.org/10.1006/anbo.2000.1197" volume="86" year="2000">Koul et al. 2000</bibRefCitation>
, 
<bibRefCitation author="Kirkbride, JH Jr" journalOrPublisher="Technical Bulletin" pageId="6" pageNumber="115" pagination="636 - 1185" refId="B22" refString="Kirkbride, JH Jr, Gunn, CR, Weitzman, AL, 2003. Fruits and seeds of genera in the subfamily Faboideae (Fabaceae), vol. 2. Technical Bulletin 1890: 636 - 1185" title="Fruits and seeds of genera in the subfamily Faboideae (Fabaceae), vol. 2." volume="1890" year="2003">Kirkbride et al. 2003</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.1600/036364412X656563" author="Celep, F" journalOrPublisher="Systematic Botany" pageId="5" pageNumber="114" pagination="893 - 912" refId="B8" refString="Celep, F, Koyuncu, M, Fritsch, RM, Kahraman, A, Dogan, M, 2012. Taxonomic importance of seed morphology in Allium (Amaryllidaceae). Systematic Botany 37: 893 - 912, DOI: https://doi.org/10.1600/036364412X656563" title="Taxonomic importance of seed morphology in Allium (Amaryllidaceae)." url="https://doi.org/10.1600/036364412X656563" volume="37" year="2012">Celep et al. 2012</bibRefCitation>
, 
<bibRefCitation author="Gandhi, D" journalOrPublisher="Environmental and Experimental Biology l" pageId="5" pageNumber="114" pagination="105 - 133" refId="B12" refString="Gandhi, D, Albert, S, Pandya, N, 2011. Morphological and micromorphological characterization of some legume seeds from Gujarat, India. Environmental and Experimental Biology l 9: 105 - 133" title="Morphological and micromorphological characterization of some legume seeds from Gujarat, India." volume="9" year="2011">Gandhi et al. 2011</bibRefCitation>
<bibRefCitation DOI="https://doi.org/10.1111/j.1756-1051.2011.01104.x" author="Brullo, C" journalOrPublisher="Nordic Journal of Botany" pageId="5" pageNumber="114" pagination="518 - 533" refId="B5" refString="Brullo, C, Brullo, S, Giusso, del Galdo G, Minissale, P, Sciandrello, S, 2011. Astragalus raphaelis (Fabaceae), a critical species from Sicily and taxonomic remarks on A. sect. Sesamei. Nordic Journal of Botany 29: 518 - 533, DOI: https://doi.org/10.1111/j.1756-1051.2011.01104.x" title="Astragalus raphaelis (Fabaceae), a critical species from Sicily and taxonomic remarks on A. sect. Sesamei." url="https://doi.org/10.1111/j.1756-1051.2011.01104.x" volume="29" year="2011">
<pageBreakToken pageId="3" pageNumber="112" start="start">Brullo</pageBreakToken>
et al. 2011
</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.5735/085.050.0109" author="Brullo, C" journalOrPublisher="Annales Botanici Fennici" pageId="5" pageNumber="114" pagination="61 - 67" refId="B6" refString="Brullo, C, Brullo, S, Giusso, del Galdo G, Minissale, P, Sciandrello, S, 2013. Astragalus kamarinensis (Fabaceae), a new species from Sicily. Annales Botanici Fennici 50: 61 - 67, DOI: https://doi.org/10.5735/085.050.0109" title="Astragalus kamarinensis (Fabaceae), a new species from Sicily." url="https://doi.org/10.5735/085.050.0109" volume="50" year="2013">2013</bibRefCitation>
) seed coat micro-morphology plays an important role in the taxonomic survey at generic and specific level, especially in those rather critical groups. The seed testa sculptures of 
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria" order="Fabales" pageId="3" pageNumber="112" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="112">Bituminaria</emphasis>
</taxonomicName>
were investigated by 
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.98.1.1" author="Minissale, P" journalOrPublisher="Phytotaxa" pageId="6" pageNumber="115" pagination="1 - 15" refId="B27" refString="Minissale, P, Brullo, C, Brullo, S, Giusso, del Galdo G, Sciandrello, S., 2013. Bituminaria basaltica (Fabaceae), a new species from Italy. Phytotaxa 98 (1): 1 - 15, DOI: https://doi.org/10.11646/phytotaxa.98.1.1" title="Bituminaria basaltica (Fabaceae), a new species from Italy." url="https://doi.org/10.11646/phytotaxa.98.1.1" volume="98" year="2013">Minissale et al. (2013)</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.220.3.6" author="Giusso, del Galdo G" journalOrPublisher="Phytotaxa" pageId="5" pageNumber="114" pagination="277 - 286" refId="B13" refString="Giusso, del Galdo G, Brullo, C, Brullo, S, Cambria, S, Minissale, P, 2015. Bituminaria kyreniae (Fabaceae), a new species from Northern Cyprus. Phytotaxa 220 (3): 277 - 286, DOI: https://doi.org/10.11646/phytotaxa.220.3.6" title="Bituminaria kyreniae (Fabaceae), a new species from Northern Cyprus." url="https://doi.org/10.11646/phytotaxa.220.3.6" volume="220" year="2015">Giusso et al. (2015)</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.266.2.1" author="Brullo, S" journalOrPublisher="Phytotaxa" pageId="5" pageNumber="114" pagination="61 - 79" refId="B7" refString="Brullo, S, Brullo, C, Minissale, P, Salmeri, C, Giusso, del Galdo G, 2016. Taxonomic investigations on Psoralea palaestina (Fabaceae), a critical species of Mediterranean flora. Phytotaxa 266 (2): 61 - 79, DOI: https://doi.org/10.11646/phytotaxa.266.2.1" title="Taxonomic investigations on Psoralea palaestina (Fabaceae), a critical species of Mediterranean flora." url="https://doi.org/10.11646/phytotaxa.266.2.1" volume="266" year="2016">Brullo et al. (2016)</bibRefCitation>
and 
<bibRefCitation DOI="https://doi.org/10.5091/plecevo.2016.1211" author="Bogdanovic, S" journalOrPublisher="Plant Ecology and Evolution" pageId="5" pageNumber="114" pagination="347 - 355" refId="B4" refString="Bogdanovic, S, Brullo, C, Brullo, S, Ljubicic, I, Giusso, del Galdo G, 2016. Bituminaria plumosa (Fabaceae), a crical species of the Croatian flora. Plant Ecology and Evolution 149 (3): 347 - 355, DOI: https://doi.org/10.5091/plecevo.2016.1211" title="Bituminaria plumosa (Fabaceae), a crical species of the Croatian flora." url="https://doi.org/10.5091/plecevo.2016.1211" volume="149" year="2016">
<normalizedToken originalValue="Bogdanović">Bogdanovic</normalizedToken>
et al. (2016)
</bibRefCitation>
, who highlighted the systematic relevance of these features providing additional information in order to discriminate among the allied species. In particular, the species hitherto examined are 
<taxonomicName lsidName="B. bituminosa" pageId="3" pageNumber="112" rank="species" species="bituminosa">
<emphasis italics="true" pageId="3" pageNumber="112">B. bituminosa</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="B. palaestina" pageId="3" pageNumber="112" rank="species" species="palaestina">
<emphasis italics="true" pageId="3" pageNumber="112">B. palaestina</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="B. kyreniae" pageId="3" pageNumber="112" rank="species" species="kyreniae">
<emphasis italics="true" pageId="3" pageNumber="112">B. kyreniae</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="B. basaltica" pageId="3" pageNumber="112" rank="species" species="basaltica">
<emphasis italics="true" pageId="3" pageNumber="112">B. basaltica</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="B. plumosa" pageId="3" pageNumber="112" rank="species" species="plumosa">
<emphasis italics="true" pageId="3" pageNumber="112">B. plumosa</emphasis>
</taxonomicName>
, which are characterized by different patterns of seed testa. In this study SEM investigations were carried out on the seed of 
<taxonomicName lsidName="B. antiatlantica" pageId="3" pageNumber="112" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="3" pageNumber="112">B. antiatlantica</emphasis>
</taxonomicName>
, according to the protocol of 
<bibRefCitation author="Stork, AL" journalOrPublisher="Candollea" pageId="7" pageNumber="116" pagination="421 - 450" refId="B35" refString="Stork, AL, Snogerup, S, Wuest, J, 1980. Seed characters 1 in Brassica section Brassica and some related groups. Candollea 35: 421 - 450" title="Seed characters 1 in Brassica section Brassica and some related groups." volume="35" year="1980">Stork et al. (1980)</bibRefCitation>
using the terminology followed Bartholot (1981) and 
<bibRefCitation DOI="https://doi.org/10.1016/j.flora.2008.01.009" author="Gontcharova, SB" journalOrPublisher="Flora" pageId="6" pageNumber="115" pagination="17 - 24" refId="B14" refString="Gontcharova, SB, Gontcharov, AA, Yakubov, VV, Kondo, K, 2009. Seed surface morphology in some representatives of the genus Rhodiola sect. Rhodiola (Crassulaceae) in the Russian Far East. Flora 204: 17 - 24, DOI: https://doi.org/10.1016/j.flora.2008.01.009" title="Seed surface morphology in some representatives of the genus Rhodiola sect. Rhodiola (Crassulaceae) in the Russian Far East." url="https://doi.org/10.1016/j.flora.2008.01.009" volume="204" year="2009">Gontcharova et al. (2009)</bibRefCitation>
. The results revealed that seed coat sculptures of 
<taxonomicName lsidName="B. antiatlantica" pageId="3" pageNumber="112" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="3" pageNumber="112">B. antiatlantica</emphasis>
</taxonomicName>
differ markedly from the species mentioned above. The seed testa of 
<taxonomicName lsidName="B. antiatlantica" pageId="3" pageNumber="112" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="3" pageNumber="112">B. antiatlantica</emphasis>
</taxonomicName>
is characterized by a fine and inconspicuous reticulum, bordering the cells, which appears irregularly polygonal and 3.5-9(10) 
<normalizedToken originalValue="µm">µm</normalizedToken>
wide. The anticlinal walls are irregularly curved to straight, slightly grooved and smooth, while the periclinal walls are flat with the epidermis smooth to finely rugose (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM micrographs of seed testa (A-C) and pod hairs (D-F) of Bituminaria antiatlantica from Mount Tachilla in Morocco. A Seed at low magnification (x 15) B Seed testa at medium magnification (x 1000) C Seed testa at high magnification (x 2500) D Pods at low magnification (x 10) E Pod hairs at medium magnification (x 250) F Pod hairs at high magnification (x 700)." figureDoi="10.3897/phytokeys.85.12288.figure5" httpUri="https://binary.pensoft.net/fig/153327" pageId="3" pageNumber="112">5B-C</figureCitation>
). The pod corpus is characterized by minutely rugose surfaces, and the indumentum hairs are minutely papillose, with the longitudinal furrow broadly widened at the base (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM micrographs of seed testa (A-C) and pod hairs (D-F) of Bituminaria antiatlantica from Mount Tachilla in Morocco. A Seed at low magnification (x 15) B Seed testa at medium magnification (x 1000) C Seed testa at high magnification (x 2500) D Pods at low magnification (x 10) E Pod hairs at medium magnification (x 250) F Pod hairs at high magnification (x 700)." figureDoi="10.3897/phytokeys.85.12288.figure5" httpUri="https://binary.pensoft.net/fig/153327" pageId="3" pageNumber="112">5E-F</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="991837" doi="10.3897/phytokeys.85.12288.figure5" httpUri="https://binary.pensoft.net/fig/153327" pageId="3" pageNumber="112" start="Figure 5" startId="F5">
<paragraph pageId="3" pageNumber="112">
<emphasis bold="true" pageId="3" pageNumber="112">Figure 5.</emphasis>
SEM micrographs of seed testa (
<emphasis bold="true" pageId="3" pageNumber="112">A-C</emphasis>
) and pod hairs (
<emphasis bold="true" pageId="3" pageNumber="112">D-F</emphasis>
) of 
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="3" pageNumber="112" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="3" pageNumber="112">Bituminaria antiatlantica</emphasis>
</taxonomicName>
from Mount Tachilla in Morocco. 
<emphasis bold="true" pageId="3" pageNumber="112">A</emphasis>
Seed at low magnification (
<normalizedToken originalValue="×">x</normalizedToken>
15) 
<emphasis bold="true" pageId="3" pageNumber="112">B</emphasis>
Seed testa at medium magnification (
<normalizedToken originalValue="×">x</normalizedToken>
1000) 
<emphasis bold="true" pageId="3" pageNumber="112">C</emphasis>
Seed testa at high magnification (
<normalizedToken originalValue="×">x</normalizedToken>
2500) 
<emphasis bold="true" pageId="3" pageNumber="112">D</emphasis>
Pods at low magnification (
<normalizedToken originalValue="×">x</normalizedToken>
10) 
<emphasis bold="true" pageId="3" pageNumber="112">E</emphasis>
Pod hairs at medium magnification (
<normalizedToken originalValue="×">x</normalizedToken>
250) 
<emphasis bold="true" pageId="3" pageNumber="112">F</emphasis>
Pod hairs at high magnification (
<normalizedToken originalValue="×">x</normalizedToken>
700).
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="113" pageId="3" pageNumber="112" type="pollen morphology">
<paragraph pageId="3" pageNumber="112">Pollen morphology.</paragraph>
<paragraph lastPageId="4" lastPageNumber="113" pageId="3" pageNumber="112">
Previous studies of pollen morphology of 
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria" order="Fabales" pageId="3" pageNumber="112" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="112">Bituminaria</emphasis>
</taxonomicName>
included those by La Serna Ramos and 
<normalizedToken originalValue="Gó">Go</normalizedToken>
Mez Ferreras (2006) and 
<bibRefCitation author="Halbritter, H" journalOrPublisher="Australian Systematic Botany" pageId="6" pageNumber="115" publicationUrl="www.paldat.org/pub/Bituminaria_bituminosa/300081" refId="B18" refString="Halbritter, H, Weis, B, 2015. Bituminaria bituminosa. In: PalDat (Ed.) A palynological database. https:/ www.paldat.org/pub/Bituminaria_bituminosa/300081" title="Bituminaria bituminosa. In: PalDat (Ed.) A palynological database. https: /" url="www.paldat.org/pub/Bituminaria_bituminosa/300081" year="2015">Halbritter and Weis (2015)</bibRefCitation>
, who published a SEM picture of 
<taxonomicName lsidName="B. bituminosa" pageId="3" pageNumber="112" rank="species" species="bituminosa">
<emphasis italics="true" pageId="3" pageNumber="112">B. bituminosa</emphasis>
</taxonomicName>
s.l., while 
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.266.2.1" author="Brullo, S" journalOrPublisher="Phytotaxa" pageId="5" pageNumber="114" pagination="61 - 79" refId="B7" refString="Brullo, S, Brullo, C, Minissale, P, Salmeri, C, Giusso, del Galdo G, 2016. Taxonomic investigations on Psoralea palaestina (Fabaceae), a critical species of Mediterranean flora. Phytotaxa 266 (2): 61 - 79, DOI: https://doi.org/10.11646/phytotaxa.266.2.1" title="Taxonomic investigations on Psoralea palaestina (Fabaceae), a critical species of Mediterranean flora." url="https://doi.org/10.11646/phytotaxa.266.2.1" volume="266" year="2016">Brullo et al. (2016)</bibRefCitation>
examined the pollen grains of 
<taxonomicName lsidName="B. bituminosa" pageId="3" pageNumber="112" rank="species" species="bituminosa">
<emphasis italics="true" pageId="3" pageNumber="112">B. bituminosa</emphasis>
</taxonomicName>
s.str. and 
<taxonomicName lsidName="B. palaestina" pageId="3" pageNumber="112" rank="species" species="palaestina">
<emphasis italics="true" pageId="3" pageNumber="112">B. palaestina</emphasis>
</taxonomicName>
, pointing out distinctive morphological differences between the pollen of the two species. In this study pollen grains of 
<taxonomicName lsidName="B. antiatlantica" pageId="3" pageNumber="112" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="3" pageNumber="112">B. antiatlantica</emphasis>
</taxonomicName>
were excised from flower buds in ahydrated state and were examined according to 
<bibRefCitation DOI="https://doi.org/10.2307/2395271" author="Walker, JW" journalOrPublisher="Annales of Missouri Botanical Garden" pageId="7" pageNumber="116" pagination="664 - 723" refId="B37" refString="Walker, JW, Doyle, JA, 1975. The bases of angiosperm phylogeny: palynology. Annales of Missouri Botanical Garden 62: 664 - 723, DOI: https://doi.org/10.2307/2395271" title="The bases of angiosperm phylogeny: palynology." url="https://doi.org/10.2307/2395271" volume="62" year="1975">Walker and Doyle (1975)</bibRefCitation>
, 
<bibRefCitation author="Punt, W" journalOrPublisher="Cahiers Options Me ́ diterrane ́ ennes" pageId="6" pageNumber="115" publicationUrl="http://www.pollen.mtu.edu/glos-gtx/glos-int.htm" refId="B29" refString="Punt, W, Blackmore, S, Nilsson, S, Le Thomas, A, 1994. Glossary of pollen and spore terminology. Lpp Foundation, Utrecht. http://www.pollen.mtu.edu/glos-gtx/glos-int.htm" title="Glossary of pollen and spore terminology. Lpp Foundation, Utrecht." url="http://www.pollen.mtu.edu/glos-gtx/glos-int.htm" year="1994">Punt et al. (1994</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.1016/j.revpalbo.2006.06.008" author="Punt, W" journalOrPublisher="Review of Palaeobotany and Palynology" pageId="6" pageNumber="115" pagination="1 - 81" refId="B30" refString="Punt, W, Hoen, PP, Blackmore, S, Nilsson, S, Le Thomas, A, 2007. Glossary of pollen and spore terminology. Review of Palaeobotany and Palynology 143: 1 - 81, DOI: https://doi.org/10.1016/j.revpalbo.2006.06.008" title="Glossary of pollen and spore terminology." url="https://doi.org/10.1016/j.revpalbo.2006.06.008" volume="143" year="2007">2007</bibRefCitation>
) and 
<bibRefCitation author="Hesse, M" journalOrPublisher="Australian Systematic Botany" pageId="6" pageNumber="115" refId="B19" refString="Hesse, M, Halbritter, H, Zetter, R, Weber, M, Buchner, R, Frosch, Radivo A, Ulrich, S, 2009. Pollen terminology. An illustrated handbook. Springer,Wein - New York." title="Pollen terminology. An illustrated handbook. Springer, Wein - New York." year="2009">Hesse et al. (2009)</bibRefCitation>
. The pollen grains are very similar to that of 
<taxonomicName lsidName="B. bituminosa" pageId="3" pageNumber="112" rank="species" species="bituminosa">
<emphasis italics="true" pageId="3" pageNumber="112">B. bituminosa</emphasis>
</taxonomicName>
sensu stricto from Sicily, which has been examined in detail by 
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.266.2.1" author="Brullo, S" journalOrPublisher="Phytotaxa" pageId="5" pageNumber="114" pagination="61 - 79" refId="B7" refString="Brullo, S, Brullo, C, Minissale, P, Salmeri, C, Giusso, del Galdo G, 2016. Taxonomic investigations on Psoralea palaestina (Fabaceae), a critical species of Mediterranean flora. Phytotaxa 266 (2): 61 - 79, DOI: https://doi.org/10.11646/phytotaxa.266.2.1" title="Taxonomic investigations on Psoralea palaestina (Fabaceae), a critical species of Mediterranean flora." url="https://doi.org/10.11646/phytotaxa.266.2.1" volume="266" year="2016">Brullo et al. (2016)</bibRefCitation>
, although there are differences in size and ornamentations which 
<pageBreakToken pageId="4" pageNumber="113" start="start">distinguishes</pageBreakToken>
clearly the pollens of the two species. Actually, The pollen grain of 
<taxonomicName lsidName="B. bituminosa" pageId="4" pageNumber="113" rank="species" species="bituminosa">
<emphasis italics="true" pageId="4" pageNumber="113">B. bituminosa</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.266.2.1" author="Brullo, S" journalOrPublisher="Phytotaxa" pageId="5" pageNumber="114" pagination="61 - 79" refId="B7" refString="Brullo, S, Brullo, C, Minissale, P, Salmeri, C, Giusso, del Galdo G, 2016. Taxonomic investigations on Psoralea palaestina (Fabaceae), a critical species of Mediterranean flora. Phytotaxa 266 (2): 61 - 79, DOI: https://doi.org/10.11646/phytotaxa.266.2.1" title="Taxonomic investigations on Psoralea palaestina (Fabaceae), a critical species of Mediterranean flora." url="https://doi.org/10.11646/phytotaxa.266.2.1" volume="266" year="2016">Brullo et al., 2016</bibRefCitation>
, Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. SEM micrographs of pollen grains of Bituminaria antiatlantica from Mount Tachilla. A-B Polar view (x 1600) C Equatorial view (x 1600) D Detail of pollen surfaces (x 4000)." figureDoi="10.3897/phytokeys.85.12288.figure6" httpUri="https://binary.pensoft.net/fig/153328" pageId="4" pageNumber="113">6A</figureCitation>
) is smaller with a size of 25-30(34) 
<normalizedToken originalValue="µm">µm</normalizedToken>
, with larger brochi (4-17 
<normalizedToken originalValue="µm">µm</normalizedToken>
) and fewer in number, showing a laxly papillose lumen and less deep (0.5-1 
<normalizedToken originalValue="µm">µm</normalizedToken>
) and narrower (1-1.2 
<normalizedToken originalValue="µm">µm</normalizedToken>
) muri. In 
<taxonomicName lsidName="B. antiatltantica" pageId="4" pageNumber="113" rank="species" species="antiatltantica">
<emphasis italics="true" pageId="4" pageNumber="113">B. antiatltantica</emphasis>
</taxonomicName>
, the pollen grain (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. SEM micrographs of pollen grains of Bituminaria antiatlantica from Mount Tachilla. A-B Polar view (x 1600) C Equatorial view (x 1600) D Detail of pollen surfaces (x 4000)." figureDoi="10.3897/phytokeys.85.12288.figure6" httpUri="https://binary.pensoft.net/fig/153328" pageId="4" pageNumber="113">6</figureCitation>
) is slightly larger (37-38 
<normalizedToken originalValue="µm">µm</normalizedToken>
) with smaller brochi (4.3-11 
<normalizedToken originalValue="µm">µm</normalizedToken>
) and more numerous, with a lumen minutely papillose and muri deeper (1.4-1.8 
<normalizedToken originalValue="µm">µm</normalizedToken>
) and larger (1.4-1.7 
<normalizedToken originalValue="µm">µm</normalizedToken>
).
</paragraph>
<caption ID-Zenodo-Dep="991839" doi="10.3897/phytokeys.85.12288.figure6" httpUri="https://binary.pensoft.net/fig/153328" pageId="4" pageNumber="113" start="Figure 6" startId="F6">
<paragraph pageId="4" pageNumber="113">
<emphasis bold="true" pageId="4" pageNumber="113">Figure 6.</emphasis>
SEM micrographs of pollen grains of 
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="4" pageNumber="113" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="4" pageNumber="113">Bituminaria antiatlantica</emphasis>
</taxonomicName>
from Mount Tachilla. 
<emphasis bold="true" pageId="4" pageNumber="113">A-B</emphasis>
Polar view (
<normalizedToken originalValue="×">x</normalizedToken>
1600) 
<emphasis bold="true" pageId="4" pageNumber="113">C</emphasis>
Equatorial view (
<normalizedToken originalValue="×">x</normalizedToken>
1600) 
<emphasis bold="true" pageId="4" pageNumber="113">D</emphasis>
Detail of pollen surfaces (
<normalizedToken originalValue="×">x</normalizedToken>
4000).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="113" type="discussion">
<paragraph pageId="4" pageNumber="113">Discussion.</paragraph>
<paragraph pageId="4" pageNumber="113">
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="4" pageNumber="113" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="4" pageNumber="113">Bituminaria antiatlantica</emphasis>
</taxonomicName>
shares some ecological and morphological characteristics with 
<taxonomicName lsidName="B. flaccida" pageId="4" pageNumber="113" rank="species" species="flaccida">
<emphasis italics="true" pageId="4" pageNumber="113">B. flaccida</emphasis>
</taxonomicName>
, a very rare species occurring in the semidesert countries of Jordan and Sinai in the Middle East; e.g., reduced leaflets, the size and few-flowered inflorescences, and flower colour. However, the latter differs from 
<taxonomicName lsidName="B. antiatlantica" pageId="4" pageNumber="113" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="4" pageNumber="113">B. antiatlantica</emphasis>
</taxonomicName>
in several significant features (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Main diacritic features of Bituminaria antiatlantica and allied species." httpUri="http://table.plazi.org/id/295A09370499666D269BDDA0EC1A3A94" pageId="4" pageNumber="113" tableUuid="295A09370499666D269BDDA0EC1A3A94">1</tableCitation>
), such as its herbaceous habit, greyish-glaucous stems and leaves, obovate to linear-lanceolate and densely pubescent cauline leaflets, shorter calyx, longer and slightly retuse standard, shorter staminal tube, and feature of pod and seed.
</paragraph>
<caption ID-Table-UUID="295A09370499666D269BDDA0EC1A3A94" httpUri="http://table.plazi.org/id/295A09370499666D269BDDA0EC1A3A94" pageId="4" pageNumber="113" start="Table 1" startId="T1">
<paragraph pageId="4" pageNumber="113">
<emphasis bold="true" pageId="4" pageNumber="113">Table 1.</emphasis>
Main diacritic features of 
<taxonomicName authorityName="Brullo, C. Brullo, Cambria, Cristaudo &amp; Giusso" authorityYear="2017" class="Magnoliopsida" family="Leguminosae" genus="Bituminaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bituminaria antiatlantica" order="Fabales" pageId="4" pageNumber="113" phylum="Tracheophyta" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="4" pageNumber="113">Bituminaria antiatlantica</emphasis>
</taxonomicName>
and allied species.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="113">
<table pageId="4" pageNumber="113">
<tr pageId="4" pageNumber="113">
<th colspan="1" isEmpty="true" pageId="4" pageNumber="113" rowspan="1">-</th>
<th colspan="1" pageId="4" pageNumber="113" rowspan="1">
<taxonomicName lsidName="B. antiatlantica" pageId="4" pageNumber="113" rank="species" species="antiatlantica">
<emphasis italics="true" pageId="4" pageNumber="113">B. antiatlantica</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="113" rowspan="1">
<taxonomicName lsidName="B. bituminosa" pageId="4" pageNumber="113" rank="species" species="bituminosa">
<emphasis italics="true" pageId="4" pageNumber="113">B. bituminosa</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="113" rowspan="1">
<taxonomicName lsidName="B. tunetana" pageId="4" pageNumber="113" rank="species" species="tunetana">
<emphasis italics="true" pageId="4" pageNumber="113">B. tunetana</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="113" rowspan="1">
<taxonomicName lsidName="B. basaltica" pageId="4" pageNumber="113" rank="species" species="basaltica">
<emphasis italics="true" pageId="4" pageNumber="113">B. basaltica</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="113" rowspan="1">
<taxonomicName lsidName="B. flaccida" pageId="4" pageNumber="113" rank="species" species="flaccida">
<emphasis italics="true" pageId="4" pageNumber="113">B. flaccida</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="113" rowspan="1">
<taxonomicName lsidName="B. palaestina" pageId="4" pageNumber="113" rank="species" species="palaestina">
<emphasis italics="true" pageId="4" pageNumber="113">B. palaestina</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="113" rowspan="1">
<taxonomicName lsidName="B. morisiana" pageId="4" pageNumber="113" rank="species" species="morisiana">
<emphasis italics="true" pageId="4" pageNumber="113">B. morisiana</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="113" rowspan="1">
<taxonomicName lsidName="B. kyreniae" pageId="4" pageNumber="113" rank="species" species="kyreniae">
<emphasis italics="true" pageId="4" pageNumber="113">B. kyreniae</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="4" pageNumber="113" rowspan="1">
<taxonomicName lsidName="B. plumosa" pageId="4" pageNumber="113" rank="species" species="plumosa">
<emphasis italics="true" pageId="4" pageNumber="113">B. plumosa</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Stem habit</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">erect to ascending</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">erect (rar. prostrate)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">erect to ascending</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">erect to ascending-erect</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">erect-ascending</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">erect</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">erect-ascending</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">erect-ascending</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">erect-ascending</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Stem tallness (cm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">up to 60</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">up to 150</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">up to 50</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">up to 60</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">up to 40</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">100-200</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">up to 60</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">up to 50</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">up to 150</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Stipule length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-15</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-15</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">8-11</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-15</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Leaf indumentum (abaxial side)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">sparsely hair</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">hirsute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">sparsely hairy</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">hirsute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">hirsute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">hirsute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">sparsely hairy</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">sparsely hairy</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">densely villous</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Leaf indumentum (adaxial side)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">glabrous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">hirsute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">sparsely hairy</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">glabrous to subglabrous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">hirsute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">hirsute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">glabrous to subglabrous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">glabrous to subglabrous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">densely villous</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Leaf petiole length (cm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.8-6(7)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.5-15</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1-7.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.5-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.5-20</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.5-6(8)</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Basal leaflet shape</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">rounded to ovate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">rounded-elliptical to lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">lanceolate to linear-lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">rounded-elliptical to linear-lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">suborbicular to obovate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">widely ovate-subcordate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate-lanceolate to elliptical</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate to lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Cauline leaflet shape</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">rounded to ovate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">elliptical to lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">lanceolate to linear-lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">linear</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">obovate to linear-lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate-lanceolate to lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate-lanceolate to lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate to lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate-lanceolate</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Leaflet apex</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">obtuse to acute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">obtuse to retuse</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">rounded to acute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">rounded, apiculate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">rounded to acute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">obtuse to acute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">obtuse to acute</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">retuse to obtuse</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">rounded to acute</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Leaflet mucro (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">0.3-0.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">0.3-0.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">0.5-1.2</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">0.5-0.8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">0.2-1</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">0.5-1</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">0.4-0.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">0.3-0.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1-1.5</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Leaflet length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-35</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-90</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">20-70</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">8-55</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-30</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">20-55</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">27-42</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-60</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-65</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Leaflet width (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">8-21</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-30</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-20</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-15</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-16</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">14-45</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-20</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-20</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-28</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Peduncle raceme length (cm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3.5-14</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">8-22</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">8-14</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-16</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">14-24</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-20</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">(6)8-21</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Raceme shape</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">subspicate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">capitate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">subcapitate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">capitate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">capitate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">subspicate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">capitate to ovoid</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">sub-capitate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">capitate</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Raceme lenght (cm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.5-2</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-2.8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.5-2</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1-1.6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.5-2</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2.5-5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2.5-4.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-2.8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.8-3</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Raceme (number of flowers)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">15-30</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-12 (16)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-16</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-25</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">(10)15-25(30)</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Bract length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-15</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-15</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-9</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-15</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Calyx length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-13.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">14-18</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">11-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-13</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-16</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">15-18</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-16</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">15-16</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Calyx tube length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5.5-6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-5.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">(5.5)6-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-7</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Calyx lower tooth length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-9</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">(5.5)6-7.5(8.5)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-11</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">8-9</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Calyx lateral teeth length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5.5-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-9</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5.5-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5.5-6(7.5)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5.5-9</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-7</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Corolla (colour)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">whitish-pink to whitish-lilac</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">blue-violet</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">pinkish-lilac</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">white</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">whitish-pink</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">pale violet</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">white-violet</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">blue-violet to violet</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">purplish-pink</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Corolla/calyx ratio</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">longer</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">longer</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">longer</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">subequalling</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">longer</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">longer</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">longer</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">longer</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">longer</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Standard shape</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">elliptical</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate-elliptical</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">spathulate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">sphatulate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate-elliptical</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">oblanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">ovate-lanceolate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">oblanceolate-spathulate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">elliptical</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Standard apex</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">obtuse</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">emarginate</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">slightly retuse</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">rounded to obtuse</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">slightly retuse</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">retuse</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">obtuse</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">usually rounded</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">emarginate</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Standard length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">16-16.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">15-20</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">13-14</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">11-13</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">16-19</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">(17)19-21(24)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">18-23</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">16-24</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">19-20</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Standard width (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-7.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-7.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-8(9)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-8.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-7.5</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Wing length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">14-15</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">14-18</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-12.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-11</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">15.5-16.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">17.5-19</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">16-18</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">14-19</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">18-18.5</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Wing limb width (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-4</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-3</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-3.2</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2.5-3</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-4.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3.4-3.7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-4</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2.8-4</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4-4.2</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Keel length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10.5-11</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-14</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7.5-8.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-10.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-14</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">11-14</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">11.5-16</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-13</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Keel limb width (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-2.3</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.8-2.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.8-2</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">1.5-1.8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-2.2</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2.4-2.6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-2.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-2.6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-2.2</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Staminal tube (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">11-11.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-13.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">8.5-9</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">7-8</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9.5-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">11.5-12.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-15</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">11-12</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Pistil length (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-10.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9-12</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">8-8.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9.5-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-14</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9-13</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">13-14</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Pod length incl. beak (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">21-23</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">13-26</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-15</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">15-16</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-14(16)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">18-26</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">16-22</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">16-18</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Pod beak lenght (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">14-16</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-19</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5.5-6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">9-10</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-8(10)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">12-19</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">11-17</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">10-12</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Pod beak indumentum</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">pubescent</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">pubescent</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">subglabrous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">glabrous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">subglabrous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">glabrous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">pubescent</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">glabrous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">sparsely hairy</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Seed lenght (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3.5-4.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3.5-4</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5.5-6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">6.5-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5-7</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4.5-5.5</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">5.5-6</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Seed width (mm)</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3.4-4</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-4</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2.8-3</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2-2.2</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2.7-3</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3.7-4.2</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">3-4</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">2.4-2.6</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">4.5-4.8</td>
</tr>
<tr pageId="4" pageNumber="113">
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">Ecology</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">chasmophilous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">terricolous xerophilous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">terricolous xerophilous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">terricolous xerophilous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">chasmophilous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">terricolous subhygrophilous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">chasmophilous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">chasmophilous</td>
<td colspan="1" pageId="4" pageNumber="113" rowspan="1">terricolous</td>
</tr>
</table>
</paragraph>
<paragraph pageId="4" pageNumber="113">
<emphasis bold="true" pageId="4" pageNumber="113">Other specimens examined (paratypes).</emphasis>
Morocco: Sulle rupi di quarzite arenacea del Jebel Tachilla a circa 200 m di altitudine, 16 June 2015, S. Cambria (CAT!); Sulle rupi di quarzite arenacea in una gola di Jebel Imzi, 300-400 m di altitudine, 19 June 2015, S. Cambria (CAT!); Sulle rupi di quarzite arenacea del Jebel Imzi a 1450 m di altitudine, 18 June 2015, S. Cambria (CAT!).
</paragraph>
</subSubSection>
</treatment>
</document>