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<document id="F10768EBAA17C5A58DBA33DDC4A3BEFF" ID-DOI="10.11646/zootaxa.3635.4.4" ID-GBIF-Dataset="5b27dcc4-7605-425c-847d-41f2a8dd86d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="219563" ID-ZooBank="2426930B-7131-49DA-B1BF-682BE3826928" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460586800093" checkinUser="plazi" docAuthor="Jia, Fenglong, Gentili, Elio &amp; Fikáček, Martin" docDate="2013" docId="165F8783FFE7FFE5FF2E0FAD4EE2FCEF" docLanguage="en" docName="zt03635p418.pdf" docOrigin="Zootaxa 3635 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccobius (Microlaccobius) formosus Gentili 1979" docType="treatment" docVersion="8" lastPageNumber="416" masterDocId="EA66FFFBFFECFFEBFFB9094B4F71FFFB" masterDocTitle="The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae)" masterLastPageNumber="418" masterPageNumber="402" pageNumber="413" updateTime="1698325284964" updateUser="plazi">
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<mods:title id="3CB73F5F477027C784AE324BE13CFB20">The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae)</mods:title>
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<mods:name id="DF7FDBFBEEDD19B03D24AED629FABB7D" type="personal">
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<mods:namePart id="C943D01CB8CC9BDFD0DB1C2B9442C904">Jia, Fenglong</mods:namePart>
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<mods:namePart id="9746C7F841BA8073804698AC50F6B806">Gentili, Elio</mods:namePart>
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<mods:namePart id="6F63A5D3C3966CE60897E20278A70E99">Fikáček, Martin</mods:namePart>
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<mods:title id="3B85DFA7D89EDBA40A46221B94F35C14">Zootaxa</mods:title>
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<mods:part id="79B23F13779DA3F8D1BC276222FFBA8F">
<mods:date id="2272995E9FA597BE9ED9A4C6BDA71F35">2013</mods:date>
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<treatment id="165F8783FFE7FFE5FF2E0FAD4EE2FCEF" ID-DOI="http://doi.org/10.5281/zenodo.6164603" ID-GBIF-Taxon="119564696" ID-Zenodo-Dep="6164603" LSID="urn:lsid:plazi:treatment:165F8783FFE7FFE5FF2E0FAD4EE2FCEF" httpUri="http://treatment.plazi.org/id/165F8783FFE7FFE5FF2E0FAD4EE2FCEF" lastPageId="14" lastPageNumber="416" pageId="11" pageNumber="413">
<subSubSection id="D6EC651EFFE7FFE0FF2E0FAD4F88F8DA" pageId="11" pageNumber="413" type="nomenclature">
<paragraph id="9E493695FFE7FFE0FF2E0FAD4C77F8FB" blockId="11.[151,774,1766,1825]" box="[151,774,1766,1792]" pageId="11" pageNumber="413">
<heading id="C50181F9FFE7FFE0FF2E0FAD4C77F8FB" bold="true" box="[151,774,1766,1792]" fontSize="11" level="1" pageId="11" pageNumber="413" reason="1">
<taxonomicName id="59F64D16FFE7FFE0FF2E0FAD4C77F8FB" ID-CoL="3RLQ5" authority="Gentili, 1979" authorityName="Gentili" authorityYear="1979" box="[151,774,1766,1792]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="413" phylum="Arthropoda" rank="species" species="formosus" subGenus="Microlaccobius">
<emphasis id="AC82EA87FFE7FFE0FF2E0FAD4C77F8FB" bold="true" box="[151,774,1766,1792]" pageId="11" pageNumber="413">
<emphasis id="AC82EA87FFE7FFE0FF2E0FAD4E62F8FB" bold="true" box="[151,275,1766,1792]" italics="true" pageId="11" pageNumber="413">Laccobius</emphasis>
(
<emphasis id="AC82EA87FFE7FFE0FE9D0FAD4EAEF8FB" bold="true" box="[292,479,1766,1792]" italics="true" pageId="11" pageNumber="413">Microlaccobius</emphasis>
)
<emphasis id="AC82EA87FFE7FFE0FE490FAD4D11F8FB" bold="true" box="[496,608,1766,1792]" italics="true" pageId="11" pageNumber="413">formosus</emphasis>
Gentili, 1979
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="9E493695FFE7FFE0FF2E0E424F88F8DA" blockId="11.[151,774,1766,1825]" box="[151,249,1801,1825]" pageId="11" pageNumber="413">
(
<figureCitation id="06CD2A10FFE7FFE0FF260E424F80F8DA" box="[159,241,1801,1825]" captionStart="FIGURES 17 19" captionStartId="8.[151,264,1725,1748]" captionTargetBox="[153,1433,527,1673]" captionTargetId="figure@8.[151,1436,527,1706]" captionTargetPageId="8" captionText="FIGURES 17 19. Aedeagi, dorsal views. 17: Laccobius hingstoni. 18: L. inopinus. 19: L. formosus." httpUri="https://zenodo.org/record/219566/files/figure.png" pageId="11" pageNumber="413">Fig. 19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFE7FFE7FF2E0E1A4D84FDEC" lastPageId="12" lastPageNumber="414" pageId="11" pageNumber="413" type="materials_examined">
<paragraph id="9E493695FFE7FFE7FF2E0E1A4D84FDEC" blockId="11.[151,1436,1873,2006]" lastBlockId="12.[151,1436,151,895]" lastPageId="12" lastPageNumber="414" pageId="11" pageNumber="413">
<emphasis id="AC82EA87FFE7FFE0FF2E0E1A4D57F891" bold="true" box="[151,550,1873,1898]" pageId="11" pageNumber="413">Material examined. Guangdong:</emphasis>
<specimenCount id="88F0FD1CFFE7FFE0FD8A0E1A4DF4F892" box="[563,645,1873,1897]" pageId="11" pageNumber="413" type="male">1 male</specimenCount>
,
<specimenCount id="88F0FD1CFFE7FFE0FD2F0E1A4D8DF892" box="[662,764,1873,1897]" pageId="11" pageNumber="413" type="female">1 female</specimenCount>
(SYSU): Xinyi,
<date id="EA481055FFE7FFE0FC700E1A4B35F892" box="[969,1092,1873,1897]" pageId="11" pageNumber="413" value="1960-06-17">17.vi.1960</date>
, Zhelong Pu lgt.;
<specimenCount id="88F0FD1CFFE7FFE0FAA60E1A4A0CF892" box="[1311,1405,1873,1897]" pageId="11" pageNumber="413" type="male">2 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE7FFE0FA370E1A4F9FF876" pageId="11" pageNumber="413" type="female">2 females</specimenCount>
(SYSU): Guangzhou, without additional data;
<specimenCount id="88F0FD1CFFE7FFE0FC980E3E4C03F876" box="[801,882,1909,1933]" pageId="11" pageNumber="413" type="male">1 male</specimenCount>
(SYSU): Fengkai, Heishiding,
<date id="EA481055FFE7FFE0FB520E3E4AE9F876" box="[1259,1432,1909,1933]" pageId="11" pageNumber="413" value="2007-07-20" valueMax="2007-07-22" valueMin="2007-07-20">2022.vii.2007</date>
, Fenglong Jia lgt.
<emphasis id="AC82EA87FFE7FFE0FED20ED24EAAF849" bold="true" box="[363,475,1945,1970]" pageId="11" pageNumber="413">Guangxi:</emphasis>
<specimenCount id="88F0FD1CFFE7FFE0FE5E0ED24D4AF84A" box="[487,571,1945,1969]" pageId="11" pageNumber="413" type="male">1 male</specimenCount>
,
<specimenCount id="88F0FD1CFFE7FFE0FDF40ED24DCFF84A" box="[589,702,1945,1969]" pageId="11" pageNumber="413" type="female">2 females</specimenCount>
(SYSU): Hechi,
<date id="EA481055FFE7FFE0FC280ED24C8CF84A" box="[913,1021,1945,1969]" pageId="11" pageNumber="413" value="1941-11-04">4.xi.1941</date>
, Zhelong Pu lgt.;
<specimenCount id="88F0FD1CFFE7FFE0FB650ED24A5CF84A" box="[1244,1325,1945,1969]" pageId="11" pageNumber="413" type="male">1 male</specimenCount>
(SYSU): Yangshuo, 1985, Shoujian Chen lgt.
<emphasis id="AC82EA87FFE7FFE0FDFF0EF64DD9F82D" bold="true" box="[582,680,1981,2006]" pageId="11" pageNumber="413">Hainan:</emphasis>
<specimenCount id="88F0FD1CFFE7FFE0FD0D0EF64C69F82E" box="[692,792,1981,2005]" pageId="11" pageNumber="413" type="female">1 female</specimenCount>
(SYSU): Jianfengling Mts., Tianchi,
<date id="EA481055FFE7FFE0FB6B0EF64A3AF82E" box="[1234,1355,1981,2005]" pageId="11" pageNumber="413" value="1983-11-11">11.xi.1983</date>
, Zhihe Huang lgt.
<emphasis id="AC82EA87FFE0FFE7FEAF09DC4E19FF4B" bold="true" box="[278,360,151,176]" pageId="12" pageNumber="414">Hubei:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FEC909DC4ECEFF54" box="[368,447,151,175]" pageId="12" pageNumber="414" type="male">1 male</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FE7509DC4D5FFF54" box="[460,558,151,175]" pageId="12" pageNumber="414" type="female">1 female</specimenCount>
(SYSU): Zigui, Jiutouling,
<quantity id="590E9B70FFE0FFE7FCD209DC4CC0FF54" box="[875,945,151,175]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="414" unit="m" value="110.0">110 m</quantity>
a.s.l., at light,
<date id="EA481055FFE0FFE7FBE209DC4BB9FF54" box="[1115,1224,151,175]" pageId="12" pageNumber="414" value="1994-09-05">5.ix.1994</date>
, Fasheng Li lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FA3709DC4FAAFF2F" pageId="12" pageNumber="414" type="male">2 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FF5009F74E26FF2F" box="[233,343,188,212]" pageId="12" pageNumber="414" type="female">5 females</specimenCount>
(NMPC, MSNV): Wutienshan Mts., Yingshan,
<date id="EA481055FFE0FFE7FC3909F74C92FF2F" box="[896,995,188,212]" pageId="12" pageNumber="414" value="2005-05-05">5.v.2005</date>
, V. Ryjáček lgt.
<emphasis id="AC82EA87FFE0FFE7FB1D09F74B8EFF2F" bold="true" box="[1188,1279,188,212]" pageId="12" pageNumber="414">Hunan:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FAB009F74A03FF2F" box="[1289,1394,188,212]" pageId="12" pageNumber="414" type="male">17 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FA3909F74F9FFF0C" pageId="12" pageNumber="414" type="female">17 females</specimenCount>
(SYSU): Nanyue, Hengshan Mt.,
<date id="EA481055FFE0FFE7FD2C09944C70FF0C" box="[661,769,223,247]" pageId="12" pageNumber="414" value="1941-09-04">4.ix.1941</date>
, Zhelong Pu lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FC5E09944B4CFF0C" box="[999,1085,223,247]" pageId="12" pageNumber="414" type="male">1 male</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FBE909944BC8FF0C" box="[1104,1209,223,247]" pageId="12" pageNumber="414" type="female">1 female</specimenCount>
(SYSU): Yizhang,
<date id="EA481055FFE0FFE7FF2E084F4E45FEE7" box="[151,308,260,284]" pageId="12" pageNumber="414" value="1941-02-10" valueMax="1941-02-11" valueMin="1941-02-10">1011.ii.1941</date>
, Zhelong Pu lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FE45084E4D24FEE7" box="[508,597,260,284]" pageId="12" pageNumber="414" type="male">7 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FDD8084F4DBCFEE7" box="[609,717,260,284]" pageId="12" pageNumber="414" type="female">4 females</specimenCount>
(SYSU): Liyuan,
<date id="EA481055FFE0FFE7FC22084F4B78FEE7" box="[923,1033,260,284]" pageId="12" pageNumber="414" value="1941-03-06">6.iii.1941</date>
, Zhelong Pu lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FB68084F4A43FEE7" box="[1233,1330,260,284]" pageId="12" pageNumber="414" type="female">1 female</specimenCount>
(SYSU): Huaihua, Yushuwan,
<date id="EA481055FFE0FFE7FE3E086C4D70FEC4" box="[391,513,295,319]" pageId="12" pageNumber="414" value="1965-06-17">17.vi.1965</date>
, Zhenyao Chen lgt.
<emphasis id="AC82EA87FFE0FFE7FD5D086C4C34FEBB" bold="true" box="[740,837,295,320]" pageId="12" pageNumber="414">Jiangxi:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FCF5086C4CD4FEC4" box="[844,933,295,319]" pageId="12" pageNumber="414" type="male">2 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FC08086C4B6CFEC4" box="[945,1053,295,319]" pageId="12" pageNumber="414" type="female">2 females</specimenCount>
(SYSU): Shangrao, Sanqingshan,
<date id="EA481055FFE0FFE7FF2E08074E4BFE9F" box="[151,314,332,356]" pageId="12" pageNumber="414" value="2007-04-15" valueMax="2007-04-20" valueMin="2007-04-15">1520.iv.2007</date>
, Fenglong Jia lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FD8C08074DE6FE9F" box="[565,663,332,356]" pageId="12" pageNumber="414" type="male">4 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FD1508074C50FE9F" box="[684,801,332,356]" pageId="12" pageNumber="414" type="female">2 females</specimenCount>
(SYSU): Jinggangshan Mts., Xiangzhou, 26°35.5ʹN 114°16.0ʹE,
<quantity id="590E9B70FFE0FFE7FE9A08244E1DFE7C" box="[291,364,367,391]" metricMagnitude="2" metricUnit="m" metricValue="3.7399999999999998" pageId="12" pageNumber="414" unit="m" value="374.0">374 m</quantity>
a.s.l.,
<date id="EA481055FFE0FFE7FE0108244D41FE7C" box="[440,560,367,391]" pageId="12" pageNumber="414" value="2011-04-26">26.iv.2011</date>
, Fenglong Jia &amp; Shuang Zhao lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FC7308244B6BFE7C" box="[970,1050,367,391]" pageId="12" pageNumber="414" type="male">1 male</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FB9E08244BFBFE7C" box="[1063,1162,367,391]" pageId="12" pageNumber="414" type="female">1 female</specimenCount>
(NMPC): same locality and date, at light on the bridge above the slowly flowing stony river, M. Fikáček &amp; J. Hájek lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FB7408DF4A54FE57" box="[1229,1317,404,428]" pageId="12" pageNumber="414" type="male">2 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FA8908DF4AEDFE57" box="[1328,1436,404,428]" pageId="12" pageNumber="414" type="female">3 females</specimenCount>
(SYSU): Jinggangshan, Xiangzhou,
<date id="EA481055FFE0FFE7FD8608FC4DC3FE34" box="[575,690,439,463]" pageId="12" pageNumber="414" value="2011-07-02">2.vii.2011</date>
, Weicai Xie, Jinwei Li &amp; Lijun Yang lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FB0508FC4A6BFE34" box="[1212,1306,439,463]" pageId="12" pageNumber="414" type="male">2 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FA9308FC4AEDFE34" box="[1322,1436,439,463]" pageId="12" pageNumber="414" type="female">2 females</specimenCount>
(SYSU): Jinggangshan Mts., Baiyinhu,
<quantity id="590E9B70FFE0FFE7FDE008974DD2FE0F" box="[601,675,476,500]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="414" unit="m" value="800.0">800 m</quantity>
,
<date id="EA481055FFE0FFE7FD0908974C59FE0F" box="[688,808,476,500]" pageId="12" pageNumber="414" value="2011-04-27">27.iv.2011</date>
, Fenglong Jia &amp; Shuang Zhao lgt.
<emphasis id="AC82EA87FFE0FFE7FB0C08904A6EFE0F" bold="true" box="[1205,1311,475,500]" pageId="12" pageNumber="414">Shaanxi:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FA9E08974AF0FE0F" box="[1319,1409,476,500]" pageId="12" pageNumber="414" type="male">2 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FA3708974F9FFDEC" pageId="12" pageNumber="414" type="female">4 females</specimenCount>
(SYSU): Zhashui,
<date id="EA481055FFE0FFE7FE7F08B44D38FDEC" box="[454,585,511,535]" pageId="12" pageNumber="414" value="2001-07-31">31.vii.2001</date>
, Ling Zhao lgt.
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFE0FFE7FF700B684E5FFBF4" pageId="12" pageNumber="414" type="discussion">
<paragraph id="9E493695FFE0FFE7FF700B684EF0FC84" blockId="12.[151,1436,151,895]" pageId="12" pageNumber="414">
<emphasis id="AC82EA87FFE0FFE7FF700B684E20FDC7" bold="true" box="[201,337,547,572]" pageId="12" pageNumber="414">Comments.</emphasis>
Widely distributed species reaching from Russian Far East through whole
<collectingCountry id="E6E17605FFE0FFE7FB780B6F4A75FDC7" box="[1217,1284,548,572]" name="China" pageId="12" pageNumber="414">China</collectingCountry>
to Southeast Asia (
<collectingCountry id="E6E17605FFE0FFE7FF650B0C4E35FDA4" box="[220,324,583,607]" name="Thailand" pageId="12" pageNumber="414">Thailand</collectingCountry>
,
<collectingCountry id="E6E17605FFE0FFE7FEEA0B0C4EC6FDA4" box="[339,439,583,607]" name="Vietnam" pageId="12" pageNumber="414">Vietnam</collectingCountry>
). New for Guangdong, Guangxi, Hainan, Hubei, Hunan, Jiangxi provinces.
<taxonomicName id="59F64D16FFE0FFE7FA900B034F8FFD7F" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="AC82EA87FFE0FFE7FA900B034F8FFD7F" italics="true" pageId="12" pageNumber="414">Laccobius formosus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="59F64D16FFE0FFE7FE380B264DEAFD7F" authority="Nakane, 1966" authorityName="Nakane" authorityYear="1966" box="[385,667,620,644]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="AC82EA87FFE0FFE7FE380B264E82FD7F" box="[385,499,620,644]" italics="true" pageId="12" pageNumber="414">L. fragilis</emphasis>
Nakane, 1966
</taxonomicName>
by the morphology of the aedeagus (the latter occurring in
<collectingCountry id="E6E17605FFE0FFE7FAEC0B264AE9FD7F" box="[1365,1432,621,644]" name="Japan" pageId="12" pageNumber="414">Japan</collectingCountry>
,
<collectingCountry id="E6E17605FFE0FFE7FF2E0BDB4FADFD5C" box="[151,220,656,679]" name="South Korea" pageId="12" pageNumber="414">Korea</collectingCountry>
and northern
<collectingCountry id="E6E17605FFE0FFE7FECE0BC44ECEFD5C" box="[375,447,655,679]" name="China" pageId="12" pageNumber="414">China</collectingCountry>
). However,
<taxonomicName id="59F64D16FFE0FFE7FDFD0BDA4DB8FD53" box="[580,713,656,680]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="AC82EA87FFE0FFE7FDFD0BDA4DB8FD53" box="[580,713,656,680]" italics="true" pageId="12" pageNumber="414">L. formosus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="59F64D16FFE0FFE7FCE50BDA4CBBFD53" box="[860,970,656,680]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="AC82EA87FFE0FFE7FCE50BDA4CBBFD53" box="[860,970,656,680]" italics="true" pageId="12" pageNumber="414">L. fragilis</emphasis>
</taxonomicName>
by the following characters: (1) head and pronotum bears very weak shagreen (in contrast, a distinct shagreen consisting of micropunctures is developed on the head and pronotum of
<taxonomicName id="59F64D16FFE0FFE7FE7B0B924D40FD0B" box="[450,561,728,752]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="AC82EA87FFE0FFE7FE7B0B924D40FD0B" box="[450,561,728,752]" italics="true" pageId="12" pageNumber="414">L. fragilis</emphasis>
</taxonomicName>
); (2) primary and secondary rows of elytra are rather similar to each other in size of punctures (they differ in size of punctures in
<taxonomicName id="59F64D16FFE0FFE7FD4B0BB64C13FCEF" box="[754,866,764,788]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="AC82EA87FFE0FFE7FD4B0BB64C13FCEF" box="[754,866,764,788]" italics="true" pageId="12" pageNumber="414">L. fragilis</emphasis>
</taxonomicName>
); (3) apex of the median lobe is rounded apically (acute in
<taxonomicName id="59F64D16FFE0FFE7FF470A6A4E1DFCC3" box="[254,364,800,824]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="AC82EA87FFE0FFE7FF470A6A4E1DFCC3" box="[254,364,800,824]" italics="true" pageId="12" pageNumber="414">L. fragilis</emphasis>
</taxonomicName>
); and (4) parameres partly sclerotized and hence regularly expanded at apex (apex of parameres membranous and hence irregularly expanded in
<taxonomicName id="59F64D16FFE0FFE7FD0D0A0E4C53FCA7" box="[692,802,836,860]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="AC82EA87FFE0FFE7FD0D0A0E4C53FCA7" box="[692,802,836,860]" italics="true" pageId="12" pageNumber="414">L. fragilis</emphasis>
</taxonomicName>
). The aedeagus of a Chinese specimen of
<taxonomicName id="59F64D16FFE0FFE7FB440A0E4AF3FCA7" box="[1277,1410,836,860]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="AC82EA87FFE0FFE7FB440A0E4AF3FCA7" box="[1277,1410,836,860]" italics="true" pageId="12" pageNumber="414">L. formosus</emphasis>
</taxonomicName>
is illustrated in
<figureCitation id="06CD2A10FFE0FFE7FE930A2C4E0CFC84" box="[298,381,871,895]" captionStart="FIGURES 17 19" captionStartId="8.[151,264,1725,1748]" captionTargetBox="[153,1433,527,1673]" captionTargetId="figure@8.[151,1436,527,1706]" captionTargetPageId="8" captionText="FIGURES 17 19. Aedeagi, dorsal views. 17: Laccobius hingstoni. 18: L. inopinus. 19: L. formosus." httpUri="https://zenodo.org/record/219566/files/figure.png" pageId="12" pageNumber="414">Fig. 19</figureCitation>
.
</paragraph>
<paragraph id="9E493695FFE0FFE7FF2E0A984E5FFBF4" blockId="12.[151,759,979,1039]" pageId="12" pageNumber="414">
<taxonomicName id="59F64D16FFE0FFE7FF2E0A984D86FC17" authority="Gentili, 1984" authorityName="Gentili" authorityYear="1984" box="[151,759,979,1004]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="hammondi" subGenus="Microlaccobius">
<emphasis id="AC82EA87FFE0FFE7FF2E0A984D2DFC17" bold="true" box="[151,604,979,1004]" pageId="12" pageNumber="414">
<emphasis id="AC82EA87FFE0FFE7FF2E0A984E7FFC17" bold="true" box="[151,270,979,1004]" italics="true" pageId="12" pageNumber="414">Laccobius</emphasis>
(
<emphasis id="AC82EA87FFE0FFE7FEA40A984EA1FC17" bold="true" box="[285,464,979,1004]" italics="true" pageId="12" pageNumber="414">Microlaccobius</emphasis>
)
<emphasis id="AC82EA87FFE0FFE7FE590A984D2DFC17" bold="true" box="[480,604,979,1004]" italics="true" pageId="12" pageNumber="414">hammondi</emphasis>
</emphasis>
Gentili, 1984
</taxonomicName>
(
<figureCitation id="06CD2A10FFE0FFE7FF260ABC4E54FBF4" box="[159,293,1015,1039]" captionStart="FIGURES 20 26" captionStartId="13.[151,264,1914,1937]" captionTargetBox="[155,1411,299,1872]" captionTargetId="figure@13.[151,1436,297,1873]" captionTargetPageId="13" captionText="FIGURES 20 26. Laccobius hammondi (20 25) and L. oscillans (26). 20 21: general habitus (20: dorsal view, 21: lateral view). 22 24: aedegus of L. hammondi (22: dorsal view, 23: lateral view, 24 ventral view). 25 26: aedeagus with open parameres showing the shape of the median lobe (25: L. hammondi, 26: L. oscillans from Japan, Sapporo env., Kimotetsu)." httpUri="https://zenodo.org/record/219567/files/figure.png" pageId="12" pageNumber="414">Figs. 2025</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFE0FFE7FF2E0D744DEBF86F" pageId="12" pageNumber="414" type="materials_examined">
<paragraph id="9E493695FFE0FFE7FF2E0D744DEBF86F" blockId="12.[151,1437,1087,2013]" pageId="12" pageNumber="414">
<emphasis id="AC82EA87FFE0FFE7FF2E0D744E94FBA3" bold="true" box="[151,485,1087,1112]" pageId="12" pageNumber="414">Material examined. Beijing:</emphasis>
Shisanling,
<date id="EA481055FFE0FFE7FDC90D744D8BFBAC" box="[624,762,1087,1111]" pageId="12" pageNumber="414" value="1962-08-10">10.viii.1962</date>
, Zhelong Pu lgt., with a label “
<taxonomicName id="59F64D16FFE0FFE7FBEC0D0B4AFCFBAC" authority="Peiping" authorityName="Peiping" box="[1109,1421,1087,1112]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="AC82EA87FFE0FFE7FBEC0D0B4A55FBA3" box="[1109,1316,1088,1112]" italics="true" pageId="12" pageNumber="414">Laccobius gracilis</emphasis>
, Peiping
</taxonomicName>
”;
<specimenCount id="88F0FD1CFFE0FFE7FF2E0D2F4F97FB87" box="[151,230,1124,1148]" pageId="12" pageNumber="414" type="male">1 male</specimenCount>
(IRSNB): Peping [= Beijing], 33 / Collector C.F. Wu // A. dOrchymont det. Lacc.
<taxonomicName id="59F64D16FFE0FFE7FB7B0D2F4AE8FB87" box="[1218,1433,1124,1148]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="variety" species="gracilis" variety="sculpt">gracilis var. sculpt</taxonomicName>
. Mots..
<emphasis id="AC82EA87FFE0FFE7FF5F0DCC4E0DFB5B" bold="true" box="[230,380,1159,1184]" pageId="12" pageNumber="414">Guangdong:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FE3D0DCC4EA3FB64" box="[388,466,1159,1183]" pageId="12" pageNumber="414" type="male">1 male</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FE660DCC4D30FB64" box="[479,577,1159,1183]" pageId="12" pageNumber="414" type="female">1 female</specimenCount>
(SYSU): Danxiashan Mt.,
<date id="EA481055FFE0FFE7FCCF0DCC4C9EFB64" box="[886,1007,1159,1183]" pageId="12" pageNumber="414" value="2008-04-20">20.iv.2008</date>
, Fenglong Jia lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FB710DCC4A53FB64" box="[1224,1314,1159,1183]" pageId="12" pageNumber="414" type="male">3 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FA960DCC4AEDFB64" box="[1327,1436,1159,1183]" pageId="12" pageNumber="414" type="female">6 females</specimenCount>
(SYSU): Lianzhou (Lianxian), Dadongshan,
<date id="EA481055FFE0FFE7FD260DE74C6BFB3F" box="[671,794,1196,1220]" pageId="12" pageNumber="414" value="2008-09-27">27.ix.2008</date>
, Yun Wang lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FC5C0DE74B33FB3F" box="[997,1090,1196,1220]" pageId="12" pageNumber="414" type="male">2 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FBEB0DE74BB3FB3F" box="[1106,1218,1196,1220]" pageId="12" pageNumber="414" type="female">2 females</specimenCount>
(SYSU): Nanling, Dadongshan,
<date id="EA481055FFE0FFE7FE830D844EC5FB1C" box="[314,436,1231,1255]" pageId="12" pageNumber="414" value="2009-06-22">22.vi.2009</date>
, Fenglong Jia lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FD140D844C72FB1C" box="[685,771,1231,1255]" pageId="12" pageNumber="414" type="male">1 male</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FCAE0D844CF0FB1C" box="[791,897,1231,1255]" pageId="12" pageNumber="414" type="female">1 female</specimenCount>
(SYSU): Lianxian (Lianzhou), Dadongshan,
<date id="EA481055FFE0FFE7FF2E0DBF4E72FAF7" box="[151,259,1268,1292]" pageId="12" pageNumber="414" value="1992-09-07">7.ix.1992</date>
, Fenglong Jia &amp; Ru Dai lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FDC90DBF4DBFFAF7" box="[624,718,1268,1292]" pageId="12" pageNumber="414" type="male">5 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FD670DBF4C3EFAF7" box="[734,847,1268,1292]" pageId="12" pageNumber="414" type="female">3 females</specimenCount>
(SYSU): Guangzhou, Baiyunshan,
<date id="EA481055FFE0FFE7FB4C0DBF4AE9FAF7" box="[1269,1432,1268,1292]" pageId="12" pageNumber="414" value="1958-04-18" valueMax="1958-04-20" valueMin="1958-04-18">1820.iv.1958</date>
, Zhelong Pu lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FEE60C5C4ECDFAD4" box="[351,444,1303,1327]" pageId="12" pageNumber="414" type="male">2 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FE720C5C4D4AFAD4" box="[459,571,1303,1327]" pageId="12" pageNumber="414" type="female">2 females</specimenCount>
(SYSU): Xingning, Luofu, Huangmaozhang,
<date id="EA481055FFE0FFE7FBEC0C5C4BB8FAD4" box="[1109,1225,1303,1327]" pageId="12" pageNumber="414" value="2004-07-02">2.vii.2004</date>
, Fenglong Jia lgt.
<emphasis id="AC82EA87FFE0FFE7FF2E0C704E76FAAF" bold="true" box="[151,263,1339,1364]" pageId="12" pageNumber="414">Guangxi:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FEA90C774E1AFAAF" box="[272,363,1340,1364]" pageId="12" pageNumber="414" type="male">5 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FEC10C774E97FAAF" box="[376,486,1340,1364]" pageId="12" pageNumber="414" type="female">5 females</specimenCount>
(SYSU): Hechi,
<date id="EA481055FFE0FFE7FD140C774C68FAAF" box="[685,793,1340,1364]" pageId="12" pageNumber="414" value="1941-11-04">4.xi.1941</date>
, Zhelong Pu lgt.
<emphasis id="AC82EA87FFE0FFE7FC580C704B20FAAF" bold="true" box="[993,1105,1339,1364]" pageId="12" pageNumber="414">Guizhou:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FBE20C774BC7FAAF" box="[1115,1206,1340,1364]" pageId="12" pageNumber="414" type="male">3 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FB7A0C774A40FAAF" box="[1219,1329,1340,1364]" pageId="12" pageNumber="414" type="female">4 females</specimenCount>
(SYSU): Rongxian County, Pingyang, Xiaodanjiang River,
<quantity id="590E9B70FFE0FFE7FD610C144C53FA83" box="[728,802,1375,1400]" metricMagnitude="2" metricUnit="m" metricValue="6.85" pageId="12" pageNumber="414" unit="m" value="685.0">685 m</quantity>
a.s.l.,
<date id="EA481055FFE0FFE7FCC80C144C9AFA8C" box="[881,1003,1375,1399]" pageId="12" pageNumber="414" value="2005-09-15">15.ix.2005</date>
, Shuang Zhao lgt.
<emphasis id="AC82EA87FFE0FFE7FB700C2B4A54FA83" bold="true" box="[1225,1317,1376,1400]" pageId="12" pageNumber="414">Hunan:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FA960C144AF1FA8C" box="[1327,1408,1375,1399]" pageId="12" pageNumber="414" type="male">1 male</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FA370C144F9FFA67" pageId="12" pageNumber="414" type="female">3 females</specimenCount>
(NHMW): ca.
<quantity id="590E9B70FFE0FFE7FE200CCF4E91FA67" box="[409,480,1412,1436]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="414" unit="km" value="25.0">25 km</quantity>
N Pingjang City, 113°38ʹ0 5ʺE 28°50ʹ10ʺN, ca.
<quantity id="590E9B70FFE0FFE7FC400CCF4B31FA67" box="[1017,1088,1412,1436]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="414" unit="m" value="200.0">200 m</quantity>
a.s.l.,
<date id="EA481055FFE0FFE7FB300CCF4A74FA67" box="[1161,1285,1412,1436]" pageId="12" pageNumber="414" value="2003-03-20">20.iii.2003</date>
, Schönmann, Komarek &amp; Wang lgt. (CWBS 500);
<specimenCount id="88F0FD1CFFE0FFE7FD850CEC4DE7FA44" box="[572,662,1447,1471]" pageId="12" pageNumber="414" type="male">2 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FD1B0CEC4C7EFA44" box="[674,783,1447,1471]" pageId="12" pageNumber="414" type="female">2 females</specimenCount>
(SYSU): Nanyue (Hengshan Mt.),
<date id="EA481055FFE0FFE7FB180CEC4A7DFA44" box="[1185,1292,1447,1471]" pageId="12" pageNumber="414" value="1941-09-02">2.ix.1941</date>
, Zhelong Pu lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FF730C874E5BFA1F" box="[202,298,1484,1508]" pageId="12" pageNumber="414" type="female">1 female</specimenCount>
(SYSU): Huaihua, Yushuwan,
<date id="EA481055FFE0FFE7FD3E0C874C73FA1F" box="[647,770,1484,1508]" pageId="12" pageNumber="414" value="1965-06-17">17.vi.1965</date>
, Zhenyao Chen lgt.
<emphasis id="AC82EA87FFE0FFE7FC5B0C804B33FA1F" bold="true" box="[994,1090,1483,1508]" pageId="12" pageNumber="414">Jiangxi:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FBF00C874BDEFA1F" box="[1097,1199,1484,1508]" pageId="12" pageNumber="414" type="male">65 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FB030C874A42FA1F" box="[1210,1331,1484,1508]" pageId="12" pageNumber="414" type="female">55 females</specimenCount>
(SYSU): Shangrao, Sanqingshan,
<date id="EA481055FFE0FFE7FE780CA44D15F9FC" box="[449,612,1519,1543]" pageId="12" pageNumber="414" value="2007-04-15" valueMax="2007-04-20" valueMin="2007-04-15">1520.iv.2007</date>
, Fenglong Jia lgt.; 20 spec. (SYSU, MSNV), Jinggangshan Mts., Xiangzhou,
<date id="EA481055FFE0FFE7FE9B0F5F4EE8F9D7" box="[290,409,1556,1580]" pageId="12" pageNumber="414" value="2011-04-26">26.iv.2011</date>
, Shuang Zhao &amp; Fenglong Jia lgt.; 12 spec. (NMPC, MSNV, SYSU): Jinggangshan Mts., Xiangzhou (forested valley south of the village), 26°35.5ʹN 114°16.0ʹE,
<quantity id="590E9B70FFE0FFE7FC7B0F7C4B78F9B4" box="[962,1033,1591,1615]" metricMagnitude="2" metricUnit="m" metricValue="3.7399999999999998" pageId="12" pageNumber="414" unit="m" value="374.0">374 m</quantity>
a.s.l., under stones and among sand at the bank of a stony river below the bridge,
<date id="EA481055FFE0FFE7FD1D0F174C6AF98F" box="[676,795,1628,1652]" pageId="12" pageNumber="414" value="2011-04-26">26.iv.2011</date>
, Fikáček &amp; Hájek lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FB910F174BFAF98F" box="[1064,1163,1628,1652]" pageId="12" pageNumber="414" type="female">1 female</specimenCount>
(NMPC): same locality and date, rice field and exposed stream directly in the village; 5 spec. (NMPC): same locality and date, at light on the bridge above the slowly running stony river, 17 spec. (NMPC, MSNV): Jinggangshan Mts., Baiyinhu env., 26°36.8ʹN 114°11.1ʹE,
<quantity id="590E9B70FFE0FFE7FE190F8C4E9BF924" box="[416,490,1735,1759]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="414" unit="m" value="800.0">800 m</quantity>
a.s.l., drying-up stream in the stony bed, night collecting in isolated pools in the stream, on the wet rocks with algae,
<date id="EA481055FFE0FFE7FDFF0FA74D96F8FF" box="[582,743,1772,1796]" pageId="12" pageNumber="414" value="2011-04-23" valueMax="2011-04-29" valueMin="2011-04-23">2329.iv.2011</date>
, Fikáček, Hájek &amp; Kubeček lgt.
<emphasis id="AC82EA87FFE0FFE7FBDE0FA04BA0F8FF" bold="true" box="[1127,1233,1771,1796]" pageId="12" pageNumber="414">Shaanxi:</emphasis>
41 spec. (SYSU, MSNV): Xian, Dayu,
<date id="EA481055FFE0FFE7FE260E444D7EF8DC" box="[415,527,1807,1831]" pageId="12" pageNumber="414" value="2011-05-12">12.v.2011</date>
, Fenglong Jia lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FD570E444C3BF8DC" box="[750,842,1807,1831]" pageId="12" pageNumber="414" type="male">3 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FCEE0E5B4CB4F8DC" box="[855,965,1807,1831]" pageId="12" pageNumber="414" type="female">7 females</specimenCount>
(NMPC, MSNV): Dayu village, Chanhe river ca.
<quantity id="590E9B70FFE0FFE7FEB90E7F4E3BF8B7" box="[256,330,1844,1868]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="414" unit="km" value="30.0">30 km</quantity>
SSE of Xian, 34°01.0ʹN 109°06.9ʹE,
<quantity id="590E9B70FFE0FFE7FCBB0E7F4C3DF8B7" box="[770,844,1844,1868]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="12" pageNumber="414" unit="m" value="720.0">720 m</quantity>
a.s.l.,
<date id="EA481055FFE0FFE7FC220E7F4B7BF8B7" box="[923,1034,1844,1868]" pageId="12" pageNumber="414" value="2011-05-11">11.v.2011</date>
, M. Balke &amp; J. Hájek lgt.;
<specimenCount id="88F0FD1CFFE0FFE7FAF40E7F4AEAF8B7" box="[1357,1435,1844,1868]" pageId="12" pageNumber="414" type="male">1 male</specimenCount>
(SYSU): Changan, Nanwutai,
<date id="EA481055FFE0FFE7FDA70E1C4DD6F894" box="[542,679,1879,1903]" pageId="12" pageNumber="414" value="1984-08-23">23.viii.1984</date>
, Zhihe Huang lgt.
<emphasis id="AC82EA87FFE0FFE7FC170E1C4B69F88B" bold="true" box="[942,1048,1879,1904]" pageId="12" pageNumber="414">Sichuan:</emphasis>
<specimenCount id="88F0FD1CFFE0FFE7FB940E1C4BE2F894" box="[1069,1171,1879,1903]" pageId="12" pageNumber="414" type="male">3 males</specimenCount>
,
<specimenCount id="88F0FD1CFFE0FFE7FB150E1C4A54F894" box="[1196,1317,1879,1903]" pageId="12" pageNumber="414" type="female">2 females</specimenCount>
(SYSU): Qingchengshan,
<date id="EA481055FFE0FFE7FEEA0E374EBEF86F" box="[339,463,1916,1940]" pageId="12" pageNumber="414" value="1982-08-08">8.viii.1982</date>
, Zhihe Huang lgt.
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFE0FFE5FF700ED44C60FDA4" lastPageId="14" lastPageNumber="416" pageId="12" pageNumber="414" type="discussion">
<paragraph id="9E493695FFE0FFE6FF700ED44B1DFF2F" blockId="12.[151,1437,1087,2013]" lastBlockId="13.[151,1436,151,212]" lastPageId="13" lastPageNumber="415" pageId="12" pageNumber="414">
<emphasis id="AC82EA87FFE0FFE7FF700ED44E20F843" bold="true" box="[201,337,1951,1976]" pageId="12" pageNumber="414">Comments.</emphasis>
New for Beijing, Guangxi, and Jiangxi.
<taxonomicName id="59F64D16FFE0FFE7FCAF0EEB4B2AF843" authority="Gentili" authorityName="Gentili" box="[790,1115,1952,1976]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="AC82EA87FFE0FFE7FCAF0EEB4B77F843" box="[790,1030,1952,1976]" italics="true" pageId="12" pageNumber="414">Laccobius hammondi</emphasis>
Gentili
</taxonomicName>
is a very common species in
<collectingCountry id="E6E17605FFE0FFE7FF2E0E8F4FABF827" box="[151,218,1988,2012]" name="China" pageId="12" pageNumber="414">China</collectingCountry>
and is very close to
<taxonomicName id="59F64D16FFE0FFE7FE6C0E8E4DB9F827" authority="Sharp. It" authorityName="Sharp. It" box="[469,712,1988,2012]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="414" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="AC82EA87FFE0FFE7FE6C0E8E4D2BF827" box="[469,602,1988,2012]" italics="true" pageId="12" pageNumber="414">L. oscillans</emphasis>
Sharp. It
</taxonomicName>
may be separated from the latter by the aedeagus, having the median lobe more dilated at the apical third than
<taxonomicName id="59F64D16FFE1FFE6FD5D09D24C1BFF4B" box="[740,874,152,176]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="415" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="AC82EA87FFE1FFE6FD5D09D24C1BFF4B" box="[740,874,152,176]" italics="true" pageId="13" pageNumber="415">L. oscillans</emphasis>
</taxonomicName>
(compare
<figureCitation id="06CD2A10FFE1FFE6FC5309DC4BD9FF54" box="[1002,1192,151,175]" captionStart="FIGURES 20 26" captionStartId="13.[151,264,1914,1937]" captionTargetBox="[155,1411,299,1872]" captionTargetId="figure@13.[151,1436,297,1873]" captionTargetPageId="13" captionText="FIGURES 20 26. Laccobius hammondi (20 25) and L. oscillans (26). 20 21: general habitus (20: dorsal view, 21: lateral view). 22 24: aedegus of L. hammondi (22: dorsal view, 23: lateral view, 24 ventral view). 25 26: aedeagus with open parameres showing the shape of the median lobe (25: L. hammondi, 26: L. oscillans from Japan, Sapporo env., Kimotetsu)." httpUri="https://zenodo.org/record/219567/files/figure.png" pageId="13" pageNumber="415">Figs. 25 and 26</figureCitation>
). Another distinctive character seems to be the pronotal surface, which is strongly shagreened in
<taxonomicName id="59F64D16FFE1FFE6FC5D09F64B14FF2F" box="[996,1125,188,212]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="415" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="AC82EA87FFE1FFE6FC5D09F64B14FF2F" box="[996,1125,188,212]" italics="true" pageId="13" pageNumber="415">L. oscillans</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="CA89661DFFE1FFE6FF2E0E314A1FF835" httpUri="https://zenodo.org/record/219567/files/figure.png" pageId="13" pageNumber="415" targetBox="[155,1411,299,1872]" targetPageId="13">
<paragraph id="9E493695FFE1FFE6FF2E0E314A1FF835" blockId="13.[151,1436,1914,1998]" pageId="13" pageNumber="415">
<emphasis id="AC82EA87FFE1FFE6FF2E0E314E26F86B" bold="true" box="[151,343,1914,1937]" pageId="13" pageNumber="415">FIGURES 2026.</emphasis>
<taxonomicName id="59F64D16FFE1FFE6FED90E304D4CF86A" box="[352,573,1915,1937]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="415" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="AC82EA87FFE1FFE6FED90E304D4CF86A" box="[352,573,1915,1937]" italics="true" pageId="13" pageNumber="415">Laccobius hammondi</emphasis>
</taxonomicName>
(2025) and
<taxonomicName id="59F64D16FFE1FFE6FD760E304C4FF86A" box="[719,830,1915,1937]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="415" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="AC82EA87FFE1FFE6FD760E304C4FF86A" box="[719,830,1915,1937]" italics="true" pageId="13" pageNumber="415">L.oscillans</emphasis>
</taxonomicName>
(26). 2021: general habitus (20: dorsal view, 21: lateral view). 2224: aedegus of
<taxonomicName id="59F64D16FFE1FFE6FE090ED04D4BF854" box="[432,570,1945,1968]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="415" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="AC82EA87FFE1FFE6FE090ED04D4BF854" box="[432,570,1945,1968]" italics="true" pageId="13" pageNumber="415">L. hammondi</emphasis>
</taxonomicName>
(22: dorsal view, 23: lateral view, 24 ventral view). 2526: aedeagus with open parameres showing the shape of the median lobe (25:
<taxonomicName id="59F64D16FFE1FFE6FD000EF24C4EF835" box="[697,831,1976,1998]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="415" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="AC82EA87FFE1FFE6FD000EF24C4EF835" box="[697,831,1976,1998]" italics="true" pageId="13" pageNumber="415">L. hammondi</emphasis>
</taxonomicName>
, 26:
<taxonomicName id="59F64D16FFE1FFE6FCCB0EF34C90F835" box="[882,993,1976,1998]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="415" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="AC82EA87FFE1FFE6FCCB0EF34C90F835" box="[882,993,1976,1998]" italics="true" pageId="13" pageNumber="415">L.oscillans</emphasis>
</taxonomicName>
from Japan, Sapporo env., Kimotetsu).
</paragraph>
</caption>
<paragraph id="9E493695FFE2FFE5FF7009DC4B44FE9F" blockId="14.[151,1436,151,501]" pageId="14" pageNumber="416">
The examined male specimen from Beijing has the aedeagus identical to the specimens from Shaanxi and Jiangxi. Reexamination of the specimens from the Jilin province published as
<taxonomicName id="59F64D16FFE2FFE5FBBD09F64BF4FF2F" box="[1028,1157,188,212]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="AC82EA87FFE2FFE5FBBD09F64BF4FF2F" box="[1028,1157,188,212]" italics="true" pageId="14" pageNumber="416">L. oscillans</emphasis>
</taxonomicName>
by Gentili (1995) shows that the specimens are actually more similar to
<taxonomicName id="59F64D16FFE2FFE5FD1A09AA4C47FF03" box="[675,822,224,248]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="AC82EA87FFE2FFE5FD1A09AA4C47FF03" box="[675,822,224,248]" italics="true" pageId="14" pageNumber="416">L. hammondi</emphasis>
</taxonomicName>
than to
<taxonomicName id="59F64D16FFE2FFE5FC2909AA4B61FF03" box="[912,1040,224,248]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="AC82EA87FFE2FFE5FC2909AA4B61FF03" box="[912,1040,224,248]" italics="true" pageId="14" pageNumber="416">L. oscillans</emphasis>
</taxonomicName>
in the above characters. These data suggest that the previous records of
<taxonomicName id="59F64D16FFE2FFE5FD89084E4DC2FEE7" box="[560,691,260,284]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="AC82EA87FFE2FFE5FD89084E4DC2FEE7" box="[560,691,260,284]" italics="true" pageId="14" pageNumber="416">L. oscillans</emphasis>
</taxonomicName>
from Beijing, northern
<collectingCountry id="E6E17605FFE2FFE5FC78084F4B75FEE7" box="[961,1028,260,284]" name="China" pageId="14" pageNumber="416">China</collectingCountry>
and possibly even Russian Far East may refer to
<emphasis id="AC82EA87FFE2FFE5FE9408624EBBFEBB" box="[301,458,296,320]" italics="true" pageId="14" pageNumber="416">
<taxonomicName id="59F64D16FFE2FFE5FE9408624EB7FEBB" box="[301,454,296,320]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="hammondi">L. hammondi</taxonomicName>
.
</emphasis>
A revision of the specimens from continental Asia as well as from southern
<collectingCountry id="E6E17605FFE2FFE5FAF908634AF1FEC4" box="[1344,1408,296,319]" name="Japan" pageId="14" pageNumber="416">Japan</collectingCountry>
is necessary to understand the real distribution of both
<taxonomicName id="59F64D16FFE2FFE5FD5A08064C15FE9F" box="[739,868,332,356]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="AC82EA87FFE2FFE5FD5A08064C15FE9F" box="[739,868,332,356]" italics="true" pageId="14" pageNumber="416">L. oscillans</emphasis>
</taxonomicName>
and
<taxonomicName id="59F64D16FFE2FFE5FC2308064B5FFE9F" box="[922,1070,332,356]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="AC82EA87FFE2FFE5FC2308064B5FFE9F" box="[922,1070,332,356]" italics="true" pageId="14" pageNumber="416">L. hammondi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9E493695FFE2FFE5FF7008244C34FE0F" blockId="14.[151,1436,151,501]" pageId="14" pageNumber="416">
The aedeagus of
<taxonomicName id="59F64D16FFE2FFE5FE3C083B4E89FE73" box="[389,504,368,392]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="AC82EA87FFE2FFE5FE3C083B4E89FE73" box="[389,504,368,392]" italics="true" pageId="14" pageNumber="416">Laccobius</emphasis>
</taxonomicName>
(
<emphasis id="AC82EA87FFE2FFE5FDB1083A4D56FE73" box="[520,551,369,392]" italics="true" pageId="14" pageNumber="416">M.</emphasis>
)
<taxonomicName id="59F64D16FFE2FFE5FD8F083A4C0CFE7C" authority="Matsui, 1993" authorityName="Matsui" authorityYear="1993" box="[566,893,367,392]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="yonanguniensis">
<emphasis id="AC82EA87FFE2FFE5FD8F083A4D95FE73" box="[566,740,369,392]" italics="true" pageId="14" pageNumber="416">yonanguniensis</emphasis>
Matsui, 1993
</taxonomicName>
from the Ryukyu Islands seems to be extremelly similar to that of
<taxonomicName id="59F64D16FFE2FFE5FEDA08DE4E8BFE57" box="[355,506,404,428]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="AC82EA87FFE2FFE5FEDA08DE4E8BFE57" box="[355,506,404,428]" italics="true" pageId="14" pageNumber="416">L. hammondi</emphasis>
</taxonomicName>
based on the drawing published by Matsui (1993).
<taxonomicName id="59F64D16FFE2FFE5FBED08DF4A0EFE57" box="[1108,1407,404,428]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="yonanguniensis">
<emphasis id="AC82EA87FFE2FFE5FBED08DF4A0EFE57" box="[1108,1407,404,428]" italics="true" pageId="14" pageNumber="416">Laccobius yonanguniensis</emphasis>
</taxonomicName>
is most likely a junior synonym of
<taxonomicName id="59F64D16FFE2FFE5FDBD08F24DE6FE2B" box="[516,663,440,464]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="AC82EA87FFE2FFE5FDBD08F24DE6FE2B" box="[516,663,440,464]" italics="true" pageId="14" pageNumber="416">L. hammondi</emphasis>
</taxonomicName>
but we were not able to examine the
<typeStatus id="414D8837FFE2FFE5FB8308F34B04FE2B" box="[1082,1141,440,464]" pageId="14" pageNumber="416">types</typeStatus>
of the former species. For that reason, we refrain from establishing this synonymy here.
</paragraph>
<paragraph id="9E493695FFE2FFE5FF2E0B0C4C60FDA4" blockId="14.[151,785,583,608]" box="[151,785,583,608]" pageId="14" pageNumber="416">
<taxonomicName id="59F64D16FFE2FFE5FF2E0B0C4C60FDA4" authority="Gentili, 1988" authorityName="Gentili" authorityYear="1988" box="[151,785,583,608]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="416" phylum="Arthropoda" rank="species" species="himalayanus" subGenus="Microlaccobius">
<emphasis id="AC82EA87FFE2FFE5FF2E0B0C4D04FD9B" bold="true" box="[151,629,583,608]" pageId="14" pageNumber="416">
<emphasis id="AC82EA87FFE2FFE5FF2E0B0C4E7CFD9B" bold="true" box="[151,269,583,608]" italics="true" pageId="14" pageNumber="416">Laccobius</emphasis>
(
<emphasis id="AC82EA87FFE2FFE5FEA40B0C4EA1FD9B" bold="true" box="[285,464,583,608]" italics="true" pageId="14" pageNumber="416">Microlaccobius</emphasis>
)
<emphasis id="AC82EA87FFE2FFE5FE590B0C4D04FD9B" bold="true" box="[480,629,583,608]" italics="true" pageId="14" pageNumber="416">himalayanus</emphasis>
</emphasis>
Gentili, 1988
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFE2FFE5FF2E0BC44DDEFD37" pageId="14" pageNumber="416" type="materials_examined">
<paragraph id="9E493695FFE2FFE5FF2E0BC44DDEFD37" blockId="14.[151,1436,654,788]" pageId="14" pageNumber="416">
<emphasis id="AC82EA87FFE2FFE5FF2E0BC44E9EFD53" bold="true" box="[151,495,655,680]" pageId="14" pageNumber="416">Material examined. Yunnan</emphasis>
:
<specimenCount id="88F0FD1CFFE2FFE5FDBF0BC44D26FD5C" box="[518,599,655,679]" pageId="14" pageNumber="416" type="male">1 male</specimenCount>
(ASHC): Xishuang-banna,
<quantity id="590E9B70FFE2FFE5FC1A0BC44C81FD5C" box="[931,1008,655,679]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="416" unit="km" value="20.0">20 km</quantity>
NW Jinghong, Man Dian, 22°07ʹN 100°40ʹE,
<quantity id="590E9B70FFE2FFE5FEB40BFF4E25FD37" box="[269,340,692,716]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="14" pageNumber="416" unit="m" value="730.0">730 m</quantity>
a.s.l.,
<date id="EA481055FFE2FFE5FE240BFF4D7FFD37" box="[413,526,692,716]" pageId="14" pageNumber="416" value="2008-05-26">26.v.2008</date>
, A Weigel lgt.
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFE2FFE5FF7E0B9C4EE2FCEF" pageId="14" pageNumber="416" type="discussion">
<paragraph id="9E493695FFE2FFE5FF7E0B9C4EE2FCEF" blockId="14.[151,1436,654,788]" pageId="14" pageNumber="416">
<emphasis id="AC82EA87FFE2FFE5FF7E0B9C4E3EFD0B" bold="true" box="[199,335,727,752]" pageId="14" pageNumber="416">Comments.</emphasis>
Reaching from
<collectingCountry id="E6E17605FFE2FFE5FDAF0B9C4D28FD14" box="[534,601,727,751]" name="Nepal" pageId="14" pageNumber="416">Nepal</collectingCountry>
and northwestern
<collectingCountry id="E6E17605FFE2FFE5FC830B9C4C05FD14" box="[826,884,727,751]" name="India" pageId="14" pageNumber="416">India</collectingCountry>
(West Bengal, Meghalaya) to SE Asia (
<collectingCountry id="E6E17605FFE2FFE5FAE40B9C4AE6FD14" box="[1373,1431,727,751]" name="Laos" pageId="14" pageNumber="416">Laos</collectingCountry>
,
<collectingCountry id="E6E17605FFE2FFE5FF2E0BB74F89FCEF" box="[151,248,764,788]" name="Vietnam" pageId="14" pageNumber="416">Vietnam</collectingCountry>
) and Yunnan.
</paragraph>
</subSubSection>
</treatment>
</document>