158 lines
19 KiB
XML
158 lines
19 KiB
XML
<document id="1E649BB3E1658E30322F8384F2595D8B" ID-DOI="10.5281/zenodo.5352440" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352440" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630208260396" checkinUser="felipe" docAuthor="Alvarez, Héctor Jaime Gasca, da, Claudio Ruy Vasconcelos, Fonseca & Ratcliffe, Brett C." docDate="2008" docId="03FC1A09FFC5F77DFF024C610D47F928" docLanguage="en" docName="0061.pdf" docOrigin="Insecta Mundi 2008 (61)" docStyle="DocumentStyle:A87B5EC635B7D5A430FC1D805C706BD1.3:InsectaMundi.2007-2008.journal_article" docStyleId="A87B5EC635B7D5A430FC1D805C706BD1" docStyleName="InsectaMundi.2007-2008.journal_article" docStyleVersion="3" docTitle="Heterogomphus ulysses Burmeister 1847" docType="treatment" docVersion="3" lastPageNumber="25" masterDocId="FFC56271FFDFF767FFC24F620949FFF3" masterDocTitle="Synopsis of the Oryctini (Coleoptera: Scarabaeidae: Dynastinae) from the Brazilian Amazon" masterLastPageNumber="62" masterPageNumber="1" pageNumber="25" updateTime="1699185735107" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="20608A07EB3F147EC52ED458FC55900D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8AB2AC9897E2FC46684BD21ECEAD7874">
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<mods:title id="DB9B2282426825C0B18ECADC00B63A09">Synopsis of the Oryctini (Coleoptera: Scarabaeidae: Dynastinae) from the Brazilian Amazon</mods:title>
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</mods:titleInfo>
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<mods:name id="3FE4B9EFF723A1E9EF1266E2DB88C585" type="personal">
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<mods:role id="65BC40BE7B8EF17C9C3F599C3D662AF4">
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<mods:roleTerm id="BCA97954F9A528F5339E96F636AF490F">Author</mods:roleTerm>
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<mods:namePart id="45E16A7D9FBB76EDFE5401737806A954">Alvarez, Héctor Jaime Gasca</mods:namePart>
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<mods:affiliation id="4DDDEDA8DD15303ECBA5353C34A1228F">Instituto Nacional de Pesquisas da Amazônia-INPA Coordenação de Pesquisas em Entomologia Av. André Araújo, 2936 - Petrópolis CEP 69011 - 970 Manaus, Amazonas, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="0519D2D4E2C8B3C4AC1E8CC4CED3ADAC" type="personal">
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<mods:role id="F335ADF3B7BA2DB3E533BBC28059B301">
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<mods:roleTerm id="ACFC65B07DBE80BE537465AC32FEF85F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0B02E7C67D1661A75AB9E4EBE982429D">da, Claudio Ruy Vasconcelos</mods:namePart>
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<mods:affiliation id="5BD6FB5E2AF5FFDC54C8106F6BFEAF79">Instituto Nacional de Pesquisas da Coordenação de Pesquisas em Av. André Araújo, 2936 - Petrópolis CEP 69011 - 970 Manaus, Amazonas</mods:affiliation>
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</mods:name>
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<mods:name id="B2070629A6B46C3F28712D65661AC6EE" type="personal">
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<mods:role id="7B67BD1918DB4D445415632EC8CE4563">
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<mods:roleTerm id="04612606F63255FC4015F7DF0C92AA59">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="67380393687863BE1EA153E1E425A972">Fonseca</mods:namePart>
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<mods:affiliation id="AC8AC11B05791C76F7E74B840B5BB5FD">Amazônia-INPA Entomologia Petrópolis, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="8F7F388E8C9D65E1C2907591209A0360" type="personal">
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<mods:role id="6D6338719A2A143092136636E2EA7F03">
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<mods:roleTerm id="65A284741ED2BCD9F54E7DD7C1C97AA7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="94622753F147B09699609B96672EF842">Ratcliffe, Brett C.</mods:namePart>
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<mods:affiliation id="9F3D50EEFEF1BDD425CB6BA7C75DCAC0">University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0514, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="80C4A7CD78018319103939C74D7E0958">text</mods:typeOfResource>
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<mods:titleInfo id="ACE1F1AFA6C78279AF046CF94BF64C19">
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<mods:title id="00F1C34E387905B02394BBDB82D722F9">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="E68BFD0DF384EB2BAED45AB80F4A516E">
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<mods:date id="C5153481477F54D87C126D77D765C4B3">2008</mods:date>
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<mods:detail id="A3841153100F49E2DD6AEE018C110E8D" type="pubDate">
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<mods:number id="58F9A53229CEE7E80FFB6976A176C52D">2008-12-05</mods:number>
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</mods:detail>
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<mods:detail id="9F0869C5E9D58141F78F0D13707A539D" type="volume">
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<mods:number id="998E81FEED98AFFE6B60C54E8572BCB8">2008</mods:number>
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</mods:detail>
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<mods:detail id="2D6A589E1FB0DBB386B9320DBB826CB8" type="issue">
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<mods:number id="AC8179BBDDCEAE4831AAFAFB873FD7D0">61</mods:number>
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</mods:detail>
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<mods:extent id="F2C9919AE2F67A5BBF119F4FBE934E59" unit="page">
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<mods:start id="0D5ECF31F8AAA3E60EA134A33EFC219B">1</mods:start>
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<mods:end id="E9D2851A78BEB2391ABEA8980E9CEE25">62</mods:end>
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<mods:classification id="5937C7D858E7D07F3B440174F7B8A345">journal article</mods:classification>
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<mods:identifier id="E79BF4B7D0512861AF94C9ED299EA91C" type="DOI">10.5281/zenodo.5352440</mods:identifier>
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<mods:identifier id="A5EE8095A6B6B06E33824CD21D125946" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="09922A6DE3DBBE677C078548D5ABD831" type="Zenodo-Dep">5352440</mods:identifier>
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<treatment id="03FC1A09FFC5F77DFF024C610D47F928" LSID="urn:lsid:plazi:treatment:03FC1A09FFC5F77DFF024C610D47F928" httpUri="http://treatment.plazi.org/id/03FC1A09FFC5F77DFF024C610D47F928" lastPageNumber="25" pageId="26" pageNumber="25">
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<subSubSection id="C34FF894FFC5F77DFF024C610B9FFCE8" box="[192,726,771,795]" pageId="26" pageNumber="25" type="nomenclature">
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<paragraph id="8BEAAB1FFFC5F77DFF024C610B9FFCE8" blockId="26.[192,726,771,795]" box="[192,726,771,795]" pageId="26" pageNumber="25">
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<heading id="D0A21C73FFC5F77DFF024C610B9FFCE8" bold="true" box="[192,726,771,795]" fontSize="10" level="2" pageId="26" pageNumber="25" reason="4">
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<taxonomicName id="4C55D09CFFC5F77DFF024C610B9FFCE8" authority="Burmeister 1847" authorityName="Burmeister" authorityYear="1847" box="[192,726,771,795]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="ulysses">
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<emphasis id="B921770DFFC5F77DFF024C610B9FFCE8" bold="true" box="[192,726,771,795]" pageId="26" pageNumber="25">
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<emphasis id="B921770DFFC5F77DFF024C6108BCFCE8" bold="true" box="[192,501,771,795]" italics="true" pageId="26" pageNumber="25">Heterogomphus ulysses</emphasis>
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<bibRefCitation id="EFC4D6EEFFC5F77DFE384C610B9FFCE8" author="Burmeister, H. C. C." box="[506,726,771,795]" pageId="26" pageNumber="25" refId="ref23787" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. (Coleoptera, Lamellicornia Xylophila et Pectinicornia) Vol. 5. T. C. F. Enslin; Berlin, Germany. 584 p." type="book" year="1847">Burmeister 1847</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FF894FFC5F77DFF024C410801FCC8" box="[192,328,803,827]" pageId="26" pageNumber="25" type="description">
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<paragraph id="8BEAAB1FFFC5F77DFF024C410801FCC8" blockId="26.[192,328,803,827]" box="[192,328,803,827]" pageId="26" pageNumber="25">
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(
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<figureCitation id="136EB79AFFC5F77DFF0B4C410809FCC8" box="[201,320,803,827]" captionStart-0="Figure 39-44" captionStart-1="Figure 45" captionStartId-0="27.[186,270,1836,1858]" captionStartId-1="28.[186,270,1094,1116]" captionTargetBox-0="[278,1349,310,1745]" captionTargetBox-1="[201,1429,196,1066]" captionTargetId-0="figure-53@27.[214,1417,191,1794]" captionTargetId-1="figure-207@28.[191,1446,186,1074]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionText-0="Figure 39-44. Heterogomphus ulysses. 39) Habitus. 40) Male head and pronotum (lateral view). 41) Male head (dorsal view). 42) Metatibia (scale line: 5 mm). 43) Parameres frontal view. 44) Parameres lateral view (scale line: 1 mm)." captionText-1="Figure 45. Distribution of Heterogomphus ulysses in Brazilian Amazonia." figureDoi-0="http://doi.org/10.5281/zenodo.5352474" figureDoi-1="http://doi.org/10.5281/zenodo.5352476" httpUri-0="https://zenodo.org/record/5352474/files/figure.png" httpUri-1="https://zenodo.org/record/5352476/files/figure.png" pageId="26" pageNumber="25">Fig. 39-45</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FF894FFC5F77DFF024C010BAEFC88" box="[192,743,867,891]" pageId="26" pageNumber="25" type="reference_group">
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<paragraph id="8BEAAB1FFFC5F77DFF024C010BAEFC88" blockId="26.[192,743,867,891]" box="[192,743,867,891]" pageId="26" pageNumber="25">
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<treatmentCitationGroup id="AB458C31FFC5F77DFF024C010BAEFC88" box="[192,743,867,891]" pageId="26" pageNumber="25">
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<treatmentCitation id="0AF48D0EFFC5F77DFF024C010BAEFC88" author="Burmeister, H. C. C." box="[192,743,867,891]" page="232" pageId="26" pageNumber="25" year="1847">
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<taxonomicName id="4C55D09CFFC5F77DFF024C010BAEFC88" authority="Burmeister 1847: 232" authorityName="Burmeister" authorityPageNumber="232" authorityYear="1847" box="[192,743,867,891]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="ulysses">
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<emphasis id="B921770DFFC5F77DFF024C010892FC88" box="[192,475,867,891]" italics="true" pageId="26" pageNumber="25">Heterogomphus ulysses</emphasis>
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<bibRefCitation id="EFC4D6EEFFC5F77DFE234C010BAEFC88" author="Burmeister, H. C. C." box="[481,743,867,891]" pageId="26" pageNumber="25" refId="ref23787" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. (Coleoptera, Lamellicornia Xylophila et Pectinicornia) Vol. 5. T. C. F. Enslin; Berlin, Germany. 584 p." type="book" year="1847">Burmeister 1847: 232</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FF894FFC5F77DFF024CC10A0EF9A8" pageId="26" pageNumber="25" type="description">
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<paragraph id="8BEAAB1FFFC5F77DFF024CC10AD3FC28" blockId="26.[192,1441,931,1627]" pageId="26" pageNumber="25">
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<emphasis id="B921770DFFC5F77DFF024CC108D0FC48" bold="true" box="[192,409,931,955]" pageId="26" pageNumber="25">DESCRIPTION.</emphasis>
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Length:
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<quantity id="4CAD06FAFFC5F77DFDC14CC10BE1FC48" box="[515,680,931,955]" metricMagnitude="-2" metricUnit="m" metricValue="4.985" metricValueMax="5.25" metricValueMin="4.72" pageId="26" pageNumber="25" unit="mm" value="49.85" valueMax="52.5" valueMin="47.2">47.2-52.5 mm</quantity>
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(males);
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<quantity id="4CAD06FAFFC5F77DFCD64CC10AF0FC48" box="[788,953,931,955]" metricMagnitude="-2" metricUnit="m" metricValue="4.79" metricValueMax="4.91" metricValueMin="4.67" pageId="26" pageNumber="25" unit="mm" value="47.9" valueMax="49.1" valueMin="46.7">46.7-49.1 mm</quantity>
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(females). Width:
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<quantity id="4CAD06FAFFC5F77DFB564CC10C70FC48" box="[1172,1337,931,955]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.73" metricValueMin="2.27" pageId="26" pageNumber="25" unit="mm" value="25.0" valueMax="27.3" valueMin="22.7">22.7-27.3 mm</quantity>
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(males); 21.4-24,5 mm (females). Color: Dark reddish brown to black.
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</paragraph>
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<paragraph id="8BEAAB1FFFC5F77DFF324C810859FA28" blockId="26.[192,1441,931,1627]" pageId="26" pageNumber="25">
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<emphasis id="B921770DFFC5F77DFF324C810801FC08" bold="true" box="[240,328,995,1019]" pageId="26" pageNumber="25">Males.</emphasis>
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<emphasis id="B921770DFFC5F77DFE8C4C8108D9FC08" box="[334,400,995,1019]" italics="true" pageId="26" pageNumber="25">Head</emphasis>
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: Frons with long, recurved horn, sides of horn subparallel, apex rounded and truncate (
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<figureCitation id="136EB79AFFC5F77DFF0A4B610852FBE8" box="[200,283,1027,1051]" captionStart="Figure 39-44" captionStartId="27.[186,270,1836,1858]" captionTargetBox="[278,1349,310,1745]" captionTargetId="figure-53@27.[214,1417,191,1794]" captionTargetPageId="27" captionText="Figure 39-44. Heterogomphus ulysses. 39) Habitus. 40) Male head and pronotum (lateral view). 41) Male head (dorsal view). 42) Metatibia (scale line: 5 mm). 43) Parameres frontal view. 44) Parameres lateral view (scale line: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5352474" httpUri="https://zenodo.org/record/5352474/files/figure.png" pageId="26" pageNumber="25">Fig 40</figureCitation>
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). Eye canthus with rounded apex, surface rugose, anterior margin weakly crenulate. Clypeus reflexed, truncate, apex broad with notch at center (
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<figureCitation id="136EB79AFFC5F77DFCE74B410A31FBC8" box="[805,888,1059,1083]" captionStart="Figure 39-44" captionStartId="27.[186,270,1836,1858]" captionTargetBox="[278,1349,310,1745]" captionTargetId="figure-53@27.[214,1417,191,1794]" captionTargetPageId="27" captionText="Figure 39-44. Heterogomphus ulysses. 39) Habitus. 40) Male head and pronotum (lateral view). 41) Male head (dorsal view). 42) Metatibia (scale line: 5 mm). 43) Parameres frontal view. 44) Parameres lateral view (scale line: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5352474" httpUri="https://zenodo.org/record/5352474/files/figure.png" pageId="26" pageNumber="25">Fig. 41</figureCitation>
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), surface smooth to punctate. Antenna with 10 segments, club subequal in length to segments 2-7. Mandibles with a single, large tooth with rounded apex.
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<emphasis id="B921770DFFC5F77DFEC64B060833FB88" box="[260,378,1124,1147]" italics="true" pageId="26" pageNumber="25">Pronotum</emphasis>
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: Majors with broad, high prominence, posterior surface convexly rounded; apex cariniform, with emargination at center. Sides with broad, tuberculiform process (
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<figureCitation id="136EB79AFFC5F77DFBC44BE10D15FB68" box="[1030,1116,1155,1179]" captionStart="Figure 39-44" captionStartId="27.[186,270,1836,1858]" captionTargetBox="[278,1349,310,1745]" captionTargetId="figure-53@27.[214,1417,191,1794]" captionTargetPageId="27" captionText="Figure 39-44. Heterogomphus ulysses. 39) Habitus. 40) Male head and pronotum (lateral view). 41) Male head (dorsal view). 42) Metatibia (scale line: 5 mm). 43) Parameres frontal view. 44) Parameres lateral view (scale line: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5352474" httpUri="https://zenodo.org/record/5352474/files/figure.png" pageId="26" pageNumber="25">Fig. 40</figureCitation>
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). Base with marginal bead. Surface of fovea with small, sparse, setigerous punctures; setae dense, moderately long, reddish brown. Lateral margins rugose. Minors with frontal horn and pronotal prominence variably reduced.
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<emphasis id="B921770DFFC5F77DFA884BA10CD0FB28" box="[1354,1433,1219,1243]" italics="true" pageId="26" pageNumber="25">Elytra</emphasis>
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: Impressed sutural stria present. Surface occasionally with weak furrows in the central area. Deep punctures present near humerus. Sides with irregular rows of ocellate and deep punctures behind humerus. Apex with small setae, reddish brown setae.
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<taxonomicName id="4C55D09CFFC5F77DFD104A410AF8FAC8" authority=": Surface" authorityName="Surface" box="[722,945,1315,1339]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="26" pageNumber="25" phylum="Chordata" rank="genus">
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<emphasis id="B921770DFFC5F77DFD104A410A01FAC8" box="[722,840,1315,1339]" italics="true" pageId="26" pageNumber="25">Pygidium</emphasis>
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: Surface
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</taxonomicName>
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rugopunctate, base with transverse band of long, dense, reddish brown setae. Apical half smooth, densely setigerous in small specimens.
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<emphasis id="B921770DFFC5F77DFAA24A260CD1FAA8" box="[1376,1432,1348,1371]" italics="true" pageId="26" pageNumber="25">Legs</emphasis>
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: Protibia quadridentate. Apex of posterior tibia crenulate, with a row of thick spinules (
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<figureCitation id="136EB79AFFC5F77DFB374A010C06FA88" box="[1269,1359,1379,1403]" captionStart="Figure 39-44" captionStartId="27.[186,270,1836,1858]" captionTargetBox="[278,1349,310,1745]" captionTargetId="figure-53@27.[214,1417,191,1794]" captionTargetPageId="27" captionText="Figure 39-44. Heterogomphus ulysses. 39) Habitus. 40) Male head and pronotum (lateral view). 41) Male head (dorsal view). 42) Metatibia (scale line: 5 mm). 43) Parameres frontal view. 44) Parameres lateral view (scale line: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5352474" httpUri="https://zenodo.org/record/5352474/files/figure.png" pageId="26" pageNumber="25">Fig. 42</figureCitation>
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). First tarsomere of posterior tarsus long, quadrangular.
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<emphasis id="B921770DFFC5F77DFCD64AE60A2AFA68" box="[788,867,1412,1435]" italics="true" pageId="26" pageNumber="25">Venter</emphasis>
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: Prosternal process long, apex rounded, densely setose; setae long, reddish brown.
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<emphasis id="B921770DFFC5F77DFD9C4AC60BABFA48" box="[606,738,1444,1467]" italics="true" pageId="26" pageNumber="25">Parameres</emphasis>
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: Elongated, oval, strongly contracted forward apices (
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<figureCitation id="136EB79AFFC5F77DFAB24AC1084DFA28" captionStart="Figure 39-44" captionStartId="27.[186,270,1836,1858]" captionTargetBox="[278,1349,310,1745]" captionTargetId="figure-53@27.[214,1417,191,1794]" captionTargetPageId="27" captionText="Figure 39-44. Heterogomphus ulysses. 39) Habitus. 40) Male head and pronotum (lateral view). 41) Male head (dorsal view). 42) Metatibia (scale line: 5 mm). 43) Parameres frontal view. 44) Parameres lateral view (scale line: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.5352474" httpUri="https://zenodo.org/record/5352474/files/figure.png" pageId="26" pageNumber="25">Fig. 43-44</figureCitation>
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).
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</paragraph>
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<paragraph id="8BEAAB1FFFC5F77DFF324A810A0EF9A8" blockId="26.[192,1441,931,1627]" pageId="26" pageNumber="25">
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<emphasis id="B921770DFFC5F77DFF324A810821FA08" bold="true" box="[240,360,1507,1531]" pageId="26" pageNumber="25">Females.</emphasis>
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As males except in the following respects:
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<emphasis id="B921770DFFC5F77DFC994A810AEAFA08" box="[859,931,1507,1531]" italics="true" pageId="26" pageNumber="25">Head:</emphasis>
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Frons with small, conical tubercle. Clypeal surface strongly punctate. Mandibles smaller in size.
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<emphasis id="B921770DFFC5F77DFCFC49660AF6F9E8" box="[830,959,1540,1563]" italics="true" pageId="26" pageNumber="25">Pronotum:</emphasis>
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Surface of anterior half strongly rugose, posterior half smooth, with sparse punctures.
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<emphasis id="B921770DFFC5F77DFD2049410A16F9C8" box="[738,863,1571,1595]" italics="true" pageId="26" pageNumber="25">
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<taxonomicName id="4C55D09CFFC5F77DFD2049410A11F9C8" box="[738,856,1571,1595]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="26" pageNumber="25" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
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:
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</emphasis>
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Apex with small, reddish brown setae. In lateral view, basal half convex, anterior half weakly concave.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FF894FFC5F77DFF0249E10D47F928" pageId="26" pageNumber="25" type="diagnosis">
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|
<paragraph id="8BEAAB1FFFC5F77DFF0249E10D47F928" blockId="26.[192,1441,1667,1755]" pageId="26" pageNumber="25">
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|
<emphasis id="B921770DFFC5F77DFF0249E10826F968" bold="true" box="[192,367,1667,1691]" pageId="26" pageNumber="25">DIAGNOSIS.</emphasis>
|
|
<taxonomicName id="4C55D09CFFC5F77DFEB749E10BC7F968" authorityName="Burmeister" authorityYear="1847" box="[373,654,1667,1691]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="ulysses">
|
|
<emphasis id="B921770DFFC5F77DFEB749E10BC7F968" box="[373,654,1667,1691]" italics="true" pageId="26" pageNumber="25">Heterogomphus ulysses</emphasis>
|
|
</taxonomicName>
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|
may be distinguished by the following combination of characters: apex of metatibia with a row of spinules, sides of pronotum with a small tuberculiform process, pronotal fovea deep and densely setigerous, frontal horn long with bilobed apex.
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</paragraph>
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|
</subSubSection>
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</treatment>
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</document> |