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<document id="99E18050A2AACF639BF2FC184598D1FD" ID-DOI="10.5281/zenodo.193731" ID-GBIF-Dataset="4bf51a1d-077f-46dc-bd30-e582ffbf11ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="193731" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460169130497" checkinUser="plazi" docAuthor="Lohrmann, Volker &amp; Ohl, Michael" docDate="2010" docId="03B3C46B392F1C2EFF6722D3D8220BA4" docLanguage="en" docName="zt02384p043.pdf" docOrigin="Zootaxa 2384" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Liosphex tupi Lohrmann, sp. nov." docType="treatment" docVersion="10" lastPageNumber="34" masterDocId="FF8ABC13390F1C0FFFF02244D9340A13" masterDocTitle="World revision of the wasp genus Liosphex Townes, 1977 (Hymenoptera: Rhopalosomatidae)" masterLastPageNumber="43" masterPageNumber="1" pageNumber="33" updateTime="1698235918724" updateUser="plazi">
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<mods:title id="EB5A03B0D0A3E8B48F950F2B319BF933">World revision of the wasp genus Liosphex Townes, 1977 (Hymenoptera: Rhopalosomatidae)</mods:title>
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<mods:namePart id="37666EAB253D6C136181F5BE35B314E7">Lohrmann, Volker</mods:namePart>
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<mods:namePart id="3C8A0CC2B09912927E51DC5F72E510FD">Ohl, Michael</mods:namePart>
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<mods:title id="0659D926CA55512042110D3C79107B88">Zootaxa</mods:title>
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<mods:date id="E5FC1C2B1AF7D281AEDE50DF88D89F03">2010</mods:date>
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<mods:identifier id="B7CD198892A39FEE7A240BEFF7B08C01" type="DOI">10.5281/zenodo.193731</mods:identifier>
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<treatment id="03B3C46B392F1C2EFF6722D3D8220BA4" ID-DOI="http://doi.org/10.5281/zenodo.6209725" ID-GBIF-Taxon="124956311" ID-Zenodo-Dep="6209725" LSID="urn:lsid:zoobank.org:act:08804554-D044-45CA-9126-0FBB07C7029B" httpUri="http://treatment.plazi.org/id/03B3C46B392F1C2EFF6722D3D8220BA4" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="C30026F6392F1C2FFF6722D3D8190AC7" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BA5757D392F1C2FFF6722D3DB1B0AA2" blockId="32.[151,559,151,212]" box="[151,559,151,177]" pageId="32" pageNumber="33">
<heading id="D0EDC211392F1C2FFF6722D3DB1B0AA2" bold="true" box="[151,559,151,177]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="B96EA96F392F1C2FFF6722D3DB1B0AA2" bold="true" box="[151,559,151,177]" pageId="32" pageNumber="33">
<taxonomicName id="4C1A0EFE392F1C2FFF6722D3D8FE0AA2" ID-CoL="3V83G" authority="Lohrmann" authorityName="Lohrmann" box="[151,458,151,177]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="tupi" status="sp. nov.">
<emphasis id="B96EA96F392F1C2FFF6722D3D8080AA2" bold="true" box="[151,316,151,177]" italics="true" pageId="32" pageNumber="33">Liosphex tupi</emphasis>
Lohrmann
</taxonomicName>
<taxonomicNameLabel id="A25D1414392F1C2FFE2222D3DB1B0AA2" box="[466,559,151,177]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA5757D392F1C2FFF6722FED8190AC7" blockId="32.[151,559,151,212]" box="[151,301,186,212]" pageId="32" pageNumber="33">
(
<figureCitation id="132169F8392F1C2FFF6F22FED8120AC7" box="[159,294,186,212]" captionStart="FIGURES 62 64" captionStartId="33.[151,269,1766,1790]" captionTargetBox="[207,1379,495,1739]" captionTargetId="figure@33.[207,1379,495,1740]" captionTargetPageId="33" captionText="FIGURES 62 64. Liosphex tupi sp. nov. (holotype) Ƥ. 62: head in dorsal view; 63: habitus in lateral view; 64: habitus in dorsal view." httpUri="https://zenodo.org/record/193747/files/figure.png" pageId="32" pageNumber="33">Figs 6264</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30026F6392F1C2FFF672341DA760B9C" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BA5757D392F1C2FFF672341D8A80B57" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
<emphasis id="B96EA96F392F1C2FFF672341D8250B0C" bold="true" box="[151,273,261,287]" pageId="32" pageNumber="33">
<typeStatus id="54A1CBDF392F1C2FFF672341D8380B0C" box="[151,268,261,287]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female.
<collectingCountry id="F30D35ED392F1C2FFE8F2341D8DD0B0C" box="[383,489,261,287]" name="Brazil" pageId="32" pageNumber="33">BRAZIL</collectingCountry>
: Rio de Janeiro,
<date id="FFA453BD392F1C2FFD442341DA620B0C" box="[692,854,261,287]" pageId="32" pageNumber="33" value="1939-01">January 1939</date>
, YelFevServ MESBrazil R. C. Shannon (
<collectionCode id="ED0BEDB8392F1C2FFACE2341DCAC0B0C" LSID="urn:lsid:biocol.org:col:34871" box="[1342,1432,261,287]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="32" pageNumber="33">USNM</collectionCode>
,
<typeStatus id="54A1CBDF392F1C2FFF67236ED9C80B57" box="[151,252,298,324]" pageId="32" pageNumber="33" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C1A0EFE392F1C2FFED3236ED8BF0B50" box="[291,395,298,323]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="varius">
<emphasis id="B96EA96F392F1C2FFED3236ED8BF0B50" box="[291,395,298,323]" italics="true" pageId="32" pageNumber="33">L. varius</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA5757D392F1C2FFF34230BDA760B9C" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
<emphasis id="B96EA96F392F1C2FFF34230BD80A0B7A" bold="true" box="[196,318,335,361]" pageId="32" pageNumber="33">
<typeStatus id="54A1CBDF392F1C2FFF34230BD80D0B7A" box="[196,313,335,361]" pageId="32" pageNumber="33" type="paratype">Paratype</typeStatus>
.
</emphasis>
<collectingCountry id="F30D35ED392F1C2FFEB62314D8850B79" box="[326,433,336,362]" name="Brazil" pageId="32" pageNumber="33">BRAZIL</collectingCountry>
: Paraná, Piraquara, Mananciais de Serra,
<geoCoordinate id="EE2E13BA392F1C2FFC562314DD3C0B79" box="[934,1032,336,362]" direction="south" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-25.483334">25°29S</geoCoordinate>
<geoCoordinate id="EE2E13BA392F1C2FFBE02314DD480B79" box="[1040,1148,336,362]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="-48.983334">48°59W</geoCoordinate>
,
<quantity id="4CE2D898392F1C2FFB7A2314DDEB0B79" box="[1162,1247,336,362]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="32" pageNumber="33" unit="m" value="1050.0">1050m</quantity>
,
<date id="FFA453BD392F1C2FFB1C2314D8720B9C" pageId="32" pageNumber="33" value="2006-01-22" valueMax="2006-02-04" valueMin="2006-01-22">22 January 4 February 2006</date>
, A. Aguíar, am. Malaise trap (1 3,
<collectionCode id="ED0BEDB8392F1C2FFD142331DA020B9C" box="[740,822,373,399]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="32" pageNumber="33">DZUP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30026F6392F1C2FFF3423DED83608A9" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BA5757D392F1C2FFF3423DEDAC10837" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
<emphasis id="B96EA96F392F1C2FFF3423DED8770BA7" bold="true" box="[196,323,410,436]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
The female of
<taxonomicName id="4C1A0EFE392F1C2FFE0923DEDB730BA0" box="[505,583,410,435]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="tupi">
<emphasis id="B96EA96F392F1C2FFE0923DEDB730BA0" box="[505,583,410,435]" italics="true" pageId="32" pageNumber="33">L. tupi</emphasis>
</taxonomicName>
is unique among all female
<taxonomicName id="4C1A0EFE392F1C2FFC6623DEDD340BA0" box="[918,1024,410,435]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B96EA96F392F1C2FFC6623DEDD340BA0" box="[918,1024,410,435]" italics="true" pageId="32" pageNumber="33">Liosphex</emphasis>
</taxonomicName>
in showing the combination of the following characters: Distal margin of the mid- and hindtibia with a group of stout and thick bristles (as in
<figureCitation id="132169F8392F1C2FFA9D2384D98D0BEC" captionStart="FIGURES 14 25" captionStartId="9.[151,272,1579,1603]" captionTargetBox="[206,1380,194,1558]" captionTargetId="figure@9.[206,1381,194,1558]" captionTargetPageId="9" captionText="FIGURES 14 25. Liosphex trichopleurum Ƥ. 14: forebasitarsus. Liosphex atratus sp. nov. Ƥ. 15: forebasitarsus; 21: apex of hindtibia in lateral view; 24: tergite III. Liosphex bribri sp. nov. Ƥ. 16: head in frontal view; 19: pronotum in dorsal view. Liosphex guarani sp. nov. Ƥ. 17: head in frontal view; 23: apex of hindtibia in lateral view. Liosphex longicornis sp. nov. Ƥ. 18: head in frontal view. Liosphex maleku sp. nov. Ƥ. 20: pronotum in dorsal view. Liosphex boreus sp. nov. Ƥ. 22: apex of hindtibia in lateral view; 25: tergite III. Abbreviations: bt: basitarsus, ht: hindtibia, its: inseration of tibialspur, Lac: Length of antennal cleaner, Lbt: Length of forebasitarsomere." httpUri="https://zenodo.org/record/193734/files/figure.png" pageId="32" pageNumber="33">Fig. 23</figureCitation>
); subantennal black marking not restricted to subantennal sclerite, reaching laterally down to cover the dorsal margin of clypeus (as in
<figureCitation id="132169F8392F1C2FFDF7204EDB690837" box="[519,605,522,548]" captionStart="FIGURES 14 25" captionStartId="9.[151,272,1579,1603]" captionTargetBox="[206,1380,194,1558]" captionTargetId="figure@9.[206,1381,194,1558]" captionTargetPageId="9" captionText="FIGURES 14 25. Liosphex trichopleurum Ƥ. 14: forebasitarsus. Liosphex atratus sp. nov. Ƥ. 15: forebasitarsus; 21: apex of hindtibia in lateral view; 24: tergite III. Liosphex bribri sp. nov. Ƥ. 16: head in frontal view; 19: pronotum in dorsal view. Liosphex guarani sp. nov. Ƥ. 17: head in frontal view; 23: apex of hindtibia in lateral view. Liosphex longicornis sp. nov. Ƥ. 18: head in frontal view. Liosphex maleku sp. nov. Ƥ. 20: pronotum in dorsal view. Liosphex boreus sp. nov. Ƥ. 22: apex of hindtibia in lateral view; 25: tergite III. Abbreviations: bt: basitarsus, ht: hindtibia, its: inseration of tibialspur, Lac: Length of antennal cleaner, Lbt: Length of forebasitarsomere." httpUri="https://zenodo.org/record/193734/files/figure.png" pageId="32" pageNumber="33">Fig. 17</figureCitation>
); and brownish mid- and hindlegs.
</paragraph>
<paragraph id="8BA5757D392F1C2FFF342074D83608A9" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
The male of
<taxonomicName id="4C1A0EFE392F1C2FFEAB2074D89D085A" box="[347,425,560,585]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="tupi">
<emphasis id="B96EA96F392F1C2FFEAB2074D89D085A" box="[347,425,560,585]" italics="true" pageId="32" pageNumber="33">L. tupi</emphasis>
</taxonomicName>
is unique among all male
<taxonomicName id="4C1A0EFE392F1C2FFD142074DA7A085A" box="[740,846,560,585]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B96EA96F392F1C2FFD142074DA7A085A" box="[740,846,560,585]" italics="true" pageId="32" pageNumber="33">Liosphex</emphasis>
</taxonomicName>
by showing the following character combination: Subantennal black marking not restricted to subantennal sclerite, reaching laterally down to cover the dorsal margin of clypeus (as in
<figureCitation id="132169F8392F1C2FFE49203EDB3B0887" box="[441,527,634,660]" captionStart="FIGURES 14 25" captionStartId="9.[151,272,1579,1603]" captionTargetBox="[206,1380,194,1558]" captionTargetId="figure@9.[206,1381,194,1558]" captionTargetPageId="9" captionText="FIGURES 14 25. Liosphex trichopleurum Ƥ. 14: forebasitarsus. Liosphex atratus sp. nov. Ƥ. 15: forebasitarsus; 21: apex of hindtibia in lateral view; 24: tergite III. Liosphex bribri sp. nov. Ƥ. 16: head in frontal view; 19: pronotum in dorsal view. Liosphex guarani sp. nov. Ƥ. 17: head in frontal view; 23: apex of hindtibia in lateral view. Liosphex longicornis sp. nov. Ƥ. 18: head in frontal view. Liosphex maleku sp. nov. Ƥ. 20: pronotum in dorsal view. Liosphex boreus sp. nov. Ƥ. 22: apex of hindtibia in lateral view; 25: tergite III. Abbreviations: bt: basitarsus, ht: hindtibia, its: inseration of tibialspur, Lac: Length of antennal cleaner, Lbt: Length of forebasitarsomere." httpUri="https://zenodo.org/record/193734/files/figure.png" pageId="32" pageNumber="33">Fig. 17</figureCitation>
); anterior margin of pronotum notched (as in
<figureCitation id="132169F8392F1C2FFBD7203EDD4F0887" box="[1063,1147,634,660]" captionStart="FIGURES 14 25" captionStartId="9.[151,272,1579,1603]" captionTargetBox="[206,1380,194,1558]" captionTargetId="figure@9.[206,1381,194,1558]" captionTargetPageId="9" captionText="FIGURES 14 25. Liosphex trichopleurum Ƥ. 14: forebasitarsus. Liosphex atratus sp. nov. Ƥ. 15: forebasitarsus; 21: apex of hindtibia in lateral view; 24: tergite III. Liosphex bribri sp. nov. Ƥ. 16: head in frontal view; 19: pronotum in dorsal view. Liosphex guarani sp. nov. Ƥ. 17: head in frontal view; 23: apex of hindtibia in lateral view. Liosphex longicornis sp. nov. Ƥ. 18: head in frontal view. Liosphex maleku sp. nov. Ƥ. 20: pronotum in dorsal view. Liosphex boreus sp. nov. Ƥ. 22: apex of hindtibia in lateral view; 25: tergite III. Abbreviations: bt: basitarsus, ht: hindtibia, its: inseration of tibialspur, Lac: Length of antennal cleaner, Lbt: Length of forebasitarsomere." httpUri="https://zenodo.org/record/193734/files/figure.png" pageId="32" pageNumber="33">Fig. 19</figureCitation>
) and brownish mid- and hindlegs.
</paragraph>
</subSubSection>
<subSubSection id="C30026F6392F1C2EFF342081D8220BA4" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="8BA5757D392F1C2FFF342081DA41095C" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
<emphasis id="B96EA96F392F1C2FFF342081DB2008CC" bold="true" box="[196,532,709,735]" pageId="32" pageNumber="33">Description of the female.</emphasis>
Head (
<figureCitation id="132169F8392F1C2FFD9E2081DBF808CC" box="[622,716,709,735]" captionStart="FIGURES 62 64" captionStartId="33.[151,269,1766,1790]" captionTargetBox="[207,1379,495,1739]" captionTargetId="figure@33.[207,1379,495,1740]" captionTargetPageId="33" captionText="FIGURES 62 64. Liosphex tupi sp. nov. (holotype) Ƥ. 62: head in dorsal view; 63: habitus in lateral view; 64: habitus in dorsal view." httpUri="https://zenodo.org/record/193747/files/figure.png" pageId="32" pageNumber="33">Fig. 62</figureCitation>
): Maximum width in frontal view
<quantity id="4CE2D898392F1C2FFB852081DDE308CC" box="[1141,1239,709,735]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="32" pageNumber="33" unit="mm" value="1.7">1.7 mm</quantity>
. UID 1.4× LID. Flagellomeres short (flagellomere II 2.9×, flagellomere VII 1.9× as long as wide). Apical bristles present on flagellomeres IV, longer bristle on flagellomere I 0.6× its length. Lateral ocellus distad from eye by 2.4× its longitudinal diameter. OOD 1.1×, IOD 1.1×, MOD 1.0× LOD.
</paragraph>
<paragraph id="8BA5757D392F1C2FFF34211ED9F10989" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
Mesosoma: Length of mesosoma:
<quantity id="4CE2D898392F1C2FFDAF211EDB820967" box="[607,694,858,884]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="32" pageNumber="33" unit="mm" value="3.3">3.3mm</quantity>
. Anterior margin of pronotum with a median notch (as in
<figureCitation id="132169F8392F1C2FFA9D211ED9830989" captionStart="FIGURES 14 25" captionStartId="9.[151,272,1579,1603]" captionTargetBox="[206,1380,194,1558]" captionTargetId="figure@9.[206,1381,194,1558]" captionTargetPageId="9" captionText="FIGURES 14 25. Liosphex trichopleurum Ƥ. 14: forebasitarsus. Liosphex atratus sp. nov. Ƥ. 15: forebasitarsus; 21: apex of hindtibia in lateral view; 24: tergite III. Liosphex bribri sp. nov. Ƥ. 16: head in frontal view; 19: pronotum in dorsal view. Liosphex guarani sp. nov. Ƥ. 17: head in frontal view; 23: apex of hindtibia in lateral view. Liosphex longicornis sp. nov. Ƥ. 18: head in frontal view. Liosphex maleku sp. nov. Ƥ. 20: pronotum in dorsal view. Liosphex boreus sp. nov. Ƥ. 22: apex of hindtibia in lateral view; 25: tergite III. Abbreviations: bt: basitarsus, ht: hindtibia, its: inseration of tibialspur, Lac: Length of antennal cleaner, Lbt: Length of forebasitarsomere." httpUri="https://zenodo.org/record/193734/files/figure.png" pageId="32" pageNumber="33">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8BA5757D392F1C2FFF3421E1DD0F0E19" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
Wings: Maximum length of forewing:
<quantity id="4CE2D898392F1C2FFD5621E1DBCB09AC" box="[678,767,933,959]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="32" pageNumber="33" unit="mm" value="7.5">7.5mm</quantity>
, 2.25× as long as mesosoma. Forewing 1cu-a slightly curved, basal section vertical, distad M by at least 2.2× its length. M-cu slightly curved. 1Rs 2.25× as long as high, its basal angle clearly smaller than 90° (
<figureCitation id="132169F8392F1C2FFD5E21B4DBC20E19" box="[686,758,1008,1034]" captionStart="FIGURES 8 13" captionStartId="8.[151,276,1841,1865]" captionTargetBox="[206,1380,381,1820]" captionTargetId="figure@8.[206,1381,381,1820]" captionTargetPageId="8" captionText="FIGURES 8 13. Liosphex trichopleurum Ƥ. 8: forewing cell 1 Rs; 13: mesosoma in lateral view (showing the distribution of setae). Liosphex atratus sp. nov. Ƥ. 9: forewing cell 1 Rs; 11 &amp; 12: mesosoma in lateral view; general morphology (11) and showing the distribution of setae (12). Liosphex varius Ƥ. 10: forewing cell 1 Rs. Abbreviations: al: anal lobe, bh: basal hamuli, ep: endophragmal pit, es: epistrnal scrobe, hpl: humural plate, jl: jugal lobe, msc: mesoscutum, mn: metanotum, msl: mesosternal lobe, msm: mesepimeron, mss: mesepisternum, mtm: metepimeron, mts: metepisternum, pn: pronotum, prl: pronotal lobe, ps: parapsidal sulcus, sp: spiracle, st: streptaulus, tg: tegula." httpUri="https://zenodo.org/record/193733/files/figure.png" pageId="32" pageNumber="33">Fig. 9</figureCitation>
). Rs of hindwing reclivous.
</paragraph>
<paragraph id="8BA5757D392F1C2FFF342651D8DD0E69" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
Legs: Forebasitarsomere short, length of antennal cleaner 0.4× its length. Mid- and hindfemur with a few bristles spread over its distal dorsal area. Mid- and hindtibia with a group of stout and thick bristles at the apical margin (as in
<figureCitation id="132169F8392F1C2FFE752624D8E80E69" box="[389,476,1120,1146]" captionStart="FIGURES 14 25" captionStartId="9.[151,272,1579,1603]" captionTargetBox="[206,1380,194,1558]" captionTargetId="figure@9.[206,1381,194,1558]" captionTargetPageId="9" captionText="FIGURES 14 25. Liosphex trichopleurum Ƥ. 14: forebasitarsus. Liosphex atratus sp. nov. Ƥ. 15: forebasitarsus; 21: apex of hindtibia in lateral view; 24: tergite III. Liosphex bribri sp. nov. Ƥ. 16: head in frontal view; 19: pronotum in dorsal view. Liosphex guarani sp. nov. Ƥ. 17: head in frontal view; 23: apex of hindtibia in lateral view. Liosphex longicornis sp. nov. Ƥ. 18: head in frontal view. Liosphex maleku sp. nov. Ƥ. 20: pronotum in dorsal view. Liosphex boreus sp. nov. Ƥ. 22: apex of hindtibia in lateral view; 25: tergite III. Abbreviations: bt: basitarsus, ht: hindtibia, its: inseration of tibialspur, Lac: Length of antennal cleaner, Lbt: Length of forebasitarsomere." httpUri="https://zenodo.org/record/193734/files/figure.png" pageId="32" pageNumber="33">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8BA5757D392F1C2FFF3426C1DB840E8C" blockId="32.[151,1437,261,2004]" box="[196,688,1157,1183]" pageId="32" pageNumber="33">Metasoma: Tergite I 1.9× as long as wide.</paragraph>
<paragraph id="8BA5757D392F1C2FFF3426EEDD4D0FB7" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
Color (as in
<figureCitation id="132169F8392F1C2FFEAE26EED8D90ED7" box="[350,493,1194,1220]" captionStart="FIGURES 62 64" captionStartId="33.[151,269,1766,1790]" captionTargetBox="[207,1379,495,1739]" captionTargetId="figure@33.[207,1379,495,1740]" captionTargetPageId="33" captionText="FIGURES 62 64. Liosphex tupi sp. nov. (holotype) Ƥ. 62: head in dorsal view; 63: habitus in lateral view; 64: habitus in dorsal view." httpUri="https://zenodo.org/record/193747/files/figure.png" pageId="32" pageNumber="33">Figs 6364</figureCitation>
): Basically dark brown to black. Subantennal black marking not restricted to subantennal sclerite, reaching laterally down to cover the dorsal margin of clypeus (as in
<figureCitation id="132169F8392F1C2FFB562694DDC80EF9" box="[1190,1276,1232,1258]" captionStart="FIGURES 14 25" captionStartId="9.[151,272,1579,1603]" captionTargetBox="[206,1380,194,1558]" captionTargetId="figure@9.[206,1381,194,1558]" captionTargetPageId="9" captionText="FIGURES 14 25. Liosphex trichopleurum Ƥ. 14: forebasitarsus. Liosphex atratus sp. nov. Ƥ. 15: forebasitarsus; 21: apex of hindtibia in lateral view; 24: tergite III. Liosphex bribri sp. nov. Ƥ. 16: head in frontal view; 19: pronotum in dorsal view. Liosphex guarani sp. nov. Ƥ. 17: head in frontal view; 23: apex of hindtibia in lateral view. Liosphex longicornis sp. nov. Ƥ. 18: head in frontal view. Liosphex maleku sp. nov. Ƥ. 20: pronotum in dorsal view. Liosphex boreus sp. nov. Ƥ. 22: apex of hindtibia in lateral view; 25: tergite III. Abbreviations: bt: basitarsus, ht: hindtibia, its: inseration of tibialspur, Lac: Length of antennal cleaner, Lbt: Length of forebasitarsomere." httpUri="https://zenodo.org/record/193734/files/figure.png" pageId="32" pageNumber="33">Fig. 17</figureCitation>
). Ventral side of antennae, area within eye notch, clypeus (except its medioventral part), labrum, gena, posterior margin of pronotum, tegula, posterior section of propodeum, forefemur (except two longitudinal stripes on its inner and outer side), foretibia, basal quarter/third of mid- and hindtibia, tibial spurs, posterior section of metasomal segment I, first quarter/third of metasomal segment II and anterior half section of metasomal segment III stramineous. Wings brownish, more intensively on distal section. Wing veins brown.
</paragraph>
<paragraph id="8BA5757D392F1C2FFF3427F4DB2F0C29" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">Pilosity: Body, including legs, largely covered with dense setae. Setae lacking on following structures: occiput, ventrolateral side of pronotum, ventrolateral side of scutellum, median area of mesepisternum, ventral part of metepimeron, anteroventral of metepisternum, median part of lateral side of propodeum and posteroventral half of hindfemur.</paragraph>
<paragraph id="8BA5757D392F1C2FFF342401DC360C4C" blockId="32.[151,1437,261,2004]" box="[196,1282,1605,1631]" pageId="32" pageNumber="33">
<emphasis id="B96EA96F392F1C2FFF342401D8D90C4C" bold="true" box="[196,493,1605,1631]" pageId="32" pageNumber="33">Description of the male.</emphasis>
Head, mesosoma and metasoma as in female except the following:
</paragraph>
<paragraph id="8BA5757D392F1C2FFF34242ED9DC0CDC" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
Head: Maximum width in frontal view
<quantity id="4CE2D898392F1C2FFD50242EDBCB0C97" box="[672,767,1642,1668]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="32" pageNumber="33" unit="mm" value="1.4">1.4 mm</quantity>
. UID 1.5× LID. Flagellomere II 2.3×, flagellomere VII 1.7× as long as wide. Apical bristles present on flagellomeres IVI, longer bristle on flagellomere I 0.7× its length.
</paragraph>
<paragraph id="8BA5757D392F1C2FFF34249EDB8D0CE7" blockId="32.[151,1437,261,2004]" box="[196,697,1754,1780]" pageId="32" pageNumber="33">
Mesosoma: Length of mesosoma:
<quantity id="4CE2D898392F1C2FFDA7249EDB800CE7" box="[599,692,1754,1780]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="32" pageNumber="33" unit="mm" value="2.4">2.4 mm</quantity>
.
</paragraph>
<paragraph id="8BA5757D392F1C2FFF342544DA240D2C" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">
Wings: Maximum length of forewing:
<quantity id="4CE2D898392F1C2FFD7E2544DBD80D09" box="[654,748,1792,1818]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="32" pageNumber="33" unit="mm" value="5.9">5.9 mm</quantity>
, 2.45× as long as mesosoma. Forewing 1cu-a distad M by at least 1.8× its length. 1Rs cell 2.15× as long as high.
</paragraph>
<paragraph id="8BA5757D392F1C2FFF34250ED8710D99" blockId="32.[151,1437,261,2004]" pageId="32" pageNumber="33">Legs: Mid- and hindtibia without a group of stout and thick bristles at its apical margin. Tarsomeres IIIV with plantulae.</paragraph>
<paragraph id="8BA5757D392F1C2FFF3425D1DB840DBC" blockId="32.[151,1437,261,2004]" box="[196,688,1941,1967]" pageId="32" pageNumber="33">Metasoma: Tergite I 1.7× as long as wide.</paragraph>
<paragraph id="8BA5757D392F1C2FFF3425FEDB080DC7" blockId="32.[151,1437,261,2004]" box="[196,572,1978,2004]" pageId="32" pageNumber="33">
<emphasis id="B96EA96F392F1C2FFF3425FED86F0DC7" bold="true" box="[196,347,1978,2004]" pageId="32" pageNumber="33">Life history.</emphasis>
Nothing is known.
</paragraph>
<paragraph id="8BA5757D392E1C2EFF3422D3D84D0AEF" blockId="33.[151,1436,151,439]" pageId="33" pageNumber="34">
<emphasis id="B96EA96F392E1C2EFF3422D3D8860AA2" bold="true" box="[196,434,151,177]" pageId="33" pageNumber="34">Etymological note.</emphasis>
<taxonomicName id="4C1A0EFE392E1C2EFE3522DCD8CD0AA2" box="[453,505,152,177]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="40" phylum="Arthropoda" rank="species" species="tupi">
<emphasis id="B96EA96F392E1C2EFE3522DCD8CD0AA2" box="[453,505,152,177]" italics="true" pageId="33" pageNumber="34">Tupí</emphasis>
</taxonomicName>
is the name of an indigenous language which was spoken in a wide area all along the Brazilian Atlantic coast before and during the conquest and colonization by the Portuguese; it is a noun in apposition.
</paragraph>
<paragraph id="8BA5757D392E1C2EFF342343D8220BA4" blockId="33.[151,1436,151,439]" pageId="33" pageNumber="34">
<emphasis id="B96EA96F392E1C2EFF342343D8320B32" bold="true" box="[196,262,263,289]" pageId="33" pageNumber="34">Note.</emphasis>
The
<typeStatus id="54A1CBDF392E1C2EFEB2234CD89C0B31" box="[322,424,264,290]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1A0EFE392E1C2EFE3F234CDB280B32" box="[463,540,264,289]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tupi">
<emphasis id="B96EA96F392E1C2EFE3F234CDB280B32" box="[463,540,264,289]" italics="true" pageId="33" pageNumber="34">L. tupi</emphasis>
</taxonomicName>
was part of the original
<typeStatus id="54A1CBDF392E1C2EFCCA234CDA580B31" box="[826,876,264,290]" pageId="33" pageNumber="34">type</typeStatus>
series of
<taxonomicName id="4C1A0EFE392E1C2EFC2E234CDDC10B31" authority="Townes, 1977" authorityName="Townes" authorityYear="1977" box="[990,1269,264,290]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="varius">
<emphasis id="B96EA96F392E1C2EFC2E234CDD710B32" box="[990,1093,264,289]" italics="true" pageId="33" pageNumber="34">L. varius</emphasis>
<bibRefCitation id="EF8B088C392E1C2EFBBD234CDDC10B31" author="Townes" box="[1101,1269,264,290]" pageId="33" pageNumber="43" refString="Townes, H. (1977) A revision of the Rhopalosomatidae (Hymenoptera). Contributions of the American Entomological Institute, 15 (1), 1 - 34." type="journal article" year="1977">Townes, 1977</bibRefCitation>
</taxonomicName>
. This species shares most of the main diagnostic features with
<taxonomicName id="4C1A0EFE392E1C2EFC482369DD0C0B55" box="[952,1080,301,326]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="B96EA96F392E1C2EFC482369DD0C0B55" box="[952,1080,301,326]" italics="true" pageId="33" pageNumber="34">L. guarani</emphasis>
</taxonomicName>
, but the completely different coloration makes it more likely that this specimen is to be treated as a valid species. The overall coloration is not simply darker than in
<taxonomicName id="4C1A0EFE392E1C2EFE37233CDB770B82" box="[455,579,376,401]" class="Insecta" family="Rhopalosomatidae" genus="Liosphex" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="B96EA96F392E1C2EFE37233CDB770B82" box="[455,579,376,401]" italics="true" pageId="33" pageNumber="34">L. guarani</emphasis>
</taxonomicName>
, it shows furthermore a completely different pattern on the clypeus, legs and wings.
</paragraph>
</subSubSection>
</treatment>
</document>