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<document id="4F9175259F84E6978E9031B39795B0DA" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFD21F40FF50FA17FBFC89AF" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax fagildei Borhidi &amp; Muniz 1985" docType="treatment" docVersion="2" lastPageNumber="48" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="46" updateTime="1695980191673" updateUser="ExternalLinkService">
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<mods:title id="82A549CDC8A2CAEF89C3BF377367F0BD">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<mods:namePart id="78A7A15B16CC876B678EEE085BBD4C6B">Henderson, Andrew</mods:namePart>
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<mods:title id="3E77ED9410E95295F83CC95BA09D3055">Phytotaxa</mods:title>
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<mods:date id="35FA606AE14A3578C2C793E6536FD775">2023</mods:date>
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<mods:number id="B95484AC8D78B994B1F6B648E2B2B13F">2023-09-19</mods:number>
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<mods:number id="CA036349423025F73BACC9AC31DFB2B4">614</mods:number>
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<mods:classification id="5DA6697F6C40B65C66B284D5B1DF4E42">journal article</mods:classification>
<mods:identifier id="50E99AE4811321A2009814A20D0CDCE5" type="DOI">10.11646/phytotaxa.614.1.1</mods:identifier>
<mods:identifier id="70CC4AB2CD6B8938AB99D867741D6A84" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="D8013D6991B120D8A9E982A185D4510C" type="Zenodo-Dep">8389307</mods:identifier>
</mods:mods>
<treatment id="03B387DAFFD21F40FF50FA17FBFC89AF" ID-DOI="http://doi.org/10.5281/zenodo.8389293" ID-Zenodo-Dep="8389293" LSID="urn:lsid:plazi:treatment:03B387DAFFD21F40FF50FA17FBFC89AF" httpUri="http://treatment.plazi.org/id/03B387DAFFD21F40FF50FA17FBFC89AF" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46">
<subSubSection id="C3006547FFD21F42FF50FA17FC078DF7" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8BA536CCFFD21F42FF50FA17FC078DF7" blockId="45.[136,1451,1454,1517]" pageId="45" pageNumber="46">
<emphasis id="B96EEADEFFD21F42FF50FA17FCDE8DD3" bold="true" box="[136,807,1454,1481]" pageId="45" pageNumber="46">
1.13.
<taxonomicName id="4C1A4D4FFFD21F42FF1FFA17FCD88DD3" ID-CoL="WLYD" authority="Borhidi &amp; Muniz (1985: 227)" authorityName="Borhidi &amp; Muniz" authorityPageNumber="227" authorityYear="1985" box="[199,801,1454,1481]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="fagildei">
<emphasis id="B96EEADEFFD21F42FF1FFA17FE388DD2" bold="true" box="[199,449,1454,1481]" italics="true" pageId="45" pageNumber="46">Coccothrinax fagildei</emphasis>
<bibRefCitation id="EF8B4B3DFFD21F42FE10FA17FCD88DD3" author="Borhidi, A. &amp; Muniz, O." box="[456,801,1454,1480]" pageId="45" pageNumber="46" pagination="225 - 230" refId="ref61125" refString="Borhidi, A. &amp; Muniz, O. (1985) Adiciones al catalogo de las palmas de Cuba. Acta Botanica Hungarica 31: 225 - 230." type="journal article" year="1985">Borhidi &amp; Muñiz (1985: 227)</bibRefCitation>
</taxonomicName>
.
</emphasis>
<materialsCitation id="3B723C91FFD21F42FCF6FA17FEB08DF6" collectingDate="1984-04-25" collectionCode="O" country="Cuba" location="Rio Verraco" municipality="Type" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Santiago de Cuba">
<collectingMunicipality id="6BC1ACB6FFD21F42FCF6FA17FC908DD3" box="[814,873,1454,1480]" pageId="45" pageNumber="46">Type</collectingMunicipality>
:—
<collectingCountry id="F30D765CFFD21F42FC52FA17FC238DD3" box="[906,986,1454,1480]" name="Cuba" pageId="45" pageNumber="46">CUBA</collectingCountry>
.
<collectingRegion id="49DEF82EFFD21F42FC3CFA17FB558DD3" box="[996,1196,1454,1480]" country="Cuba" name="Santiago de Cuba" pageId="45" pageNumber="46">Santiago de Cuba</collectingRegion>
, oeste del
<location id="8EC56017FFD21F42FAC7FA17FA5E8DD2" LSID="urn:lsid:plazi:treatment:03B387DAFFD21F40FF50FA17FBFC89AF:8EC56017FFD21F42FAC7FA17FA5E8DD2" box="[1311,1447,1454,1481]" country="Cuba" municipality="Type" name="Rio Verraco" pageId="45" pageNumber="46" stateProvince="Santiago de Cuba">Río Verraco</location>
,
<date id="FFA4100CFFD21F42FF50FA6BFEDE8DF7" box="[136,295,1490,1517]" pageId="45" pageNumber="46" value="1984-04-25">
<collectingDate id="EFE0E9E4FFD21F42FF50FA6BFEDE8DF7" box="[136,295,1490,1517]" pageId="45" pageNumber="46" value="1984-04-25">25 April 1984</collectingDate>
</date>
,
<collectionCode id="ED0BAE09FFD21F42FEEAFA6AFEB08DF6" box="[306,329,1491,1517]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="45" pageNumber="46" type="Herbarium">
<emphasis id="B96EEADEFFD21F42FEEAFA6AFEB08DF6" box="[306,329,1491,1517]" italics="true" pageId="45" pageNumber="46">O</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="B96EEADEFFD21F42FE91FA6AFDAC8DF7" box="[329,597,1490,1517]" italics="true" pageId="45" pageNumber="46">
.
<materialsCitation id="3B723C91FFD21F42FE8CFA6AFE418DF7" box="[340,440,1491,1516]" collectionCode="J" country="Cuba" location="Muniz" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Santiago de Cuba">
<location id="8EC56017FFD21F42FE8CFA6AFE648DF7" LSID="urn:lsid:plazi:treatment:03B387DAFFD21F40FF50FA17FBFC89AF:8EC56017FFD21F42FE8CFA6AFE648DF7" box="[340,413,1491,1516]" country="Cuba" name="Muniz" pageId="45" pageNumber="46" stateProvince="Santiago de Cuba">Muñiz</location>
,
<collectionCode id="ED0BAE09FFD21F42FE70FA6AFE418DF7" box="[424,440,1491,1516]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="45" pageNumber="46" type="Herbarium">J</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B723C91FFD21F42FE1AFA6BFDA88DF7" box="[450,593,1490,1517]" collectionCode="B" country="Cuba" location="Fagilde" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Santiago de Cuba">
<location id="8EC56017FFD21F42FE1AFA6BFDE08DF7" LSID="urn:lsid:plazi:treatment:03B387DAFFD21F40FF50FA17FBFC89AF:8EC56017FFD21F42FE1AFA6BFDE08DF7" box="[450,537,1490,1516]" country="Cuba" name="Fagilde" pageId="45" pageNumber="46" stateProvince="Santiago de Cuba">Fagilde</location>
&amp;
<collectionCode id="ED0BAE09FFD21F42FDE5FA6AFDA88DF7" box="[573,593,1491,1516]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="45" pageNumber="46" type="Herbarium">B</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation id="3B723C91FFD21F42FD84FA6BFC078DF7" box="[604,1022,1490,1516]" collectionCode="HAC" country="Cuba" location="Gonzalez" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Santiago de Cuba" typeStatus="holotype">
<emphasis id="B96EEADEFFD21F42FD84FA6BFCE88DF7" box="[604,785,1490,1516]" italics="true" pageId="45" pageNumber="46">
<location id="8EC56017FFD21F42FD84FA6BFD3D8DF7" LSID="urn:lsid:plazi:treatment:03B387DAFFD21F40FF50FA17FBFC89AF:8EC56017FFD21F42FD84FA6BFD3D8DF7" box="[604,708,1490,1516]" country="Cuba" name="Gonzalez" pageId="45" pageNumber="46" stateProvince="Santiago de Cuba">González</location>
15021
</emphasis>
(
<typeStatus id="54A1886EFFD21F42FCF8FA6BFC7D8DF7" box="[800,900,1490,1516]" pageId="45" pageNumber="46">holotype</typeStatus>
<collectionCode id="ED0BAE09FFD21F42FC53FA6BFC3F8DF7" box="[907,966,1490,1516]" country="Cuba" httpUri="http://grbio.org/cool/cenz-te88" name="Instituto de Ecología y Sistemática" pageId="45" pageNumber="46" type="Herbarium">HAC</collectionCode>
<emphasis id="B96EEADEFFD21F42FC15FA6AFC178DF7" box="[973,1006,1491,1516]" italics="true" pageId="45" pageNumber="46">n.v</emphasis>
.)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFD21F42FF50F9A3FCCB8037" pageId="45" pageNumber="46" type="description">
<paragraph id="8BA536CCFFD21F42FF50F9A3FCCB8037" blockId="45.[136,1452,1562,2093]" pageId="45" pageNumber="46">
<emphasis id="B96EEADEFFD21F42FF50F9A3FF298E2F" bold="true" box="[136,208,1562,1588]" pageId="45" pageNumber="46">Stems</emphasis>
length not recorded,
<quantity id="4CE29B29FFD21F42FE65F9A3FDF08E2E" box="[445,521,1562,1589]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="45" pageNumber="46" unit="cm" value="5.3">5.3 cm</quantity>
diameter, clustered.
<emphasis id="B96EEADEFFD21F42FD29F9A3FCBA8E2F" bold="true" box="[753,835,1562,1588]" pageId="45" pageNumber="46">Leaves</emphasis>
more or less deciduous or only leaf bases persisting on stem; leaf sheath fibers 0.9(0.8–1.0) mm diameter, stout, woody, loosely woven, ± joined or briefly free at the apices; petioles 8.5(6.9–10.0) mm diameter just below the apex; palmans 6.0 cm long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades wedge-shaped; segments 20(18– 22) per leaf, the middle ones 29.5(29.0–30.0) cm long and 2.1(1.9–2.3) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments tapering from base to apex, often folded, stiff and leathery, with or without scarcely developed shoulders, the apices sharply pointed and briefly splitting; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially, densely indumentose abaxially, with irregularly shaped, persistent, interlocking, fimbriate hairs, each one with a rounded, raised, light green to greenish-brown or reddish-brown center, without transverse veinlets.
<emphasis id="B96EEADEFFD21F42FBCCF8E7FB448F63" bold="true" box="[1044,1213,1886,1912]" pageId="45" pageNumber="46">Inflorescences</emphasis>
curving, arching, or pendulous amongst the leaves, with few partial inflorescences; rachis bracts somewhat flattened, loosely sheathing, usually tomentose with a dense tuft of erect hairs at the apex; partial inflorescences 3; proximalmost rachillae straight,
<quantity id="4CE29B29FFD21F42FF50F873FF2A8FFE" box="[136,211,1994,2021]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="45" pageNumber="46" unit="cm" value="4.7">4.7 cm</quantity>
long and
<quantity id="4CE29B29FFD21F42FE98F873FE6F8FFE" box="[320,406,1994,2021]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="45" pageNumber="46" unit="mm" value="0.9">0.9 mm</quantity>
diameter in fruit; rachillae glabrous at or near anthesis; stamens 9; fruit pedicels
<quantity id="4CE29B29FFD21F42FACFF873FA958FFE" box="[1303,1388,1994,2021]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="45" pageNumber="46" unit="mm" value="3.3">3.3 mm</quantity>
long;
<emphasis id="B96EEADEFFD21F42FF50F857FF308013" bold="true" box="[136,201,2030,2056]" pageId="45" pageNumber="46">fruits</emphasis>
9.0 mm long and 9.0 mm diameter, color not recorded; fruit surfaces with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
</paragraph>
</subSubSection>
<caption id="DF656644FFD11F41FF50F8C4FAFF8F88" ID-DOI="http://doi.org/10.5281/zenodo.8389377" ID-Zenodo-Dep="8389377" box="[136,1286,1917,1939]" httpUri="https://zenodo.org/record/8389377/files/figure.png" pageId="46" pageNumber="47" startId="46.[136,214,1917,1938]" targetBox="[151,1433,194,1891]" targetPageId="46" targetType="plate">
<paragraph id="8BA536CCFFD11F41FF50F8C4FAFF8F88" blockId="46.[136,1286,1917,1939]" box="[136,1286,1917,1939]" pageId="46" pageNumber="47">
<emphasis id="B96EEADEFFD11F41FF50F8C4FF0A8F89" bold="true" box="[136,243,1917,1938]" pageId="46" pageNumber="47">PLATE 11</emphasis>
.
<taxonomicName id="4C1A4D4FFFD11F41FF27F8C4FE2D8F89" box="[255,468,1917,1938]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="ekmanii">
<emphasis id="B96EEADEFFD11F41FF27F8C4FE2D8F89" box="[255,468,1917,1938]" italics="true" pageId="46" pageNumber="47">Coccothrinax ekmanii</emphasis>
</taxonomicName>
, Dominican Republic, with wedge-shaped leaf blades. (Image by Eladio Fernández).
</paragraph>
</caption>
<subSubSection id="C3006547FFD01F40FF65FF27FBFC89AF" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8BA536CCFFD01F40FF65FF27FDA188C7" blockId="47.[136,1452,158,437]" pageId="47" pageNumber="48">
<emphasis id="B96EEADEFFD01F40FF65FF27FDF588A3" bold="true" box="[189,524,158,184]" pageId="47" pageNumber="48">Distribution and habitat:—</emphasis>
<collectingCountry id="F30D765CFFD01F40FDD4FF27FDBE88A3" box="[524,583,158,184]" name="Cuba" pageId="47" pageNumber="48">Cuba</collectingCountry>
(
<collectingRegion id="49DEF82EFFD01F40FD81FF27FCD188A3" box="[601,808,158,184]" country="Cuba" name="Santiago de Cuba" pageId="47" pageNumber="48">Santiago de Cuba</collectingRegion>
) (
<figureCitation id="13212A49FFD01F40FC9AFF27FC6F88A3" box="[834,918,158,184]" captionStart="FIGURE 12" captionStartId="47.[136,229,1777,1799]" captionTargetBox="[155,1429,469,1747]" captionTargetId="figure-147@47.[150,1434,464,1752]" captionTargetPageId="47" captionText="FIGURE 12. Distribution of Coccothrinax fagildei, C. fragrans, C. garciana, and C. gonaivensis." figureDoi="http://doi.org/10.5281/zenodo.8389379" httpUri="https://zenodo.org/record/8389379/files/figure.png" pageId="47" pageNumber="48">Fig. 12</figureCitation>
) in scrubby woodland on marine terraces and dog’s tooth limestone at
<quantity id="4CE29B29FFD01F40FE43FF7BFE1B88C6" box="[411,482,194,221]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="100.0">100 m</quantity>
elevation.
</paragraph>
<paragraph id="8BA536CCFFD01F40FF65FF5FFBFC89AF" blockId="47.[136,1452,158,437]" pageId="47" pageNumber="48">
<emphasis id="B96EEADEFFD01F40FF65FF5FFE56891B" bold="true" box="[189,431,230,256]" pageId="47" pageNumber="48">Taxonomic notes:—</emphasis>
As a preliminary species,
<taxonomicName id="4C1A4D4FFFD01F40FD06FF5FFC2F891B" authorityName="Borhidi &amp; Muniz" authorityYear="1985" box="[734,982,230,256]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="fagildei">
<emphasis id="B96EEADEFFD01F40FD06FF5FFC2F891B" box="[734,982,230,256]" italics="true" pageId="47" pageNumber="48">Coccothrinax fagildei</emphasis>
</taxonomicName>
has a unique combination of qualitative character states and is recognized as a phylogenetic species. It is notable for its clustered stems and wedge-shaped leaves.
<bibRefCitation id="EF8B4B3DFFD01F40FF06FE96FE8B8953" author="Moya, C." box="[222,370,302,329]" pageId="47" pageNumber="48" pagination="1 - 27" refId="ref63288" refString="Moya, C. (2021) An annotated checklist of Cuban palms 6. Coccothrinax, Pt. 2. 1972 - 1995. Nomenclature, typification, and distribution. PalmArbor 3: 1 - 27." type="journal article" year="2021">Moya (2021)</bibRefCitation>
reported that the
<typeStatus id="54A1886EFFD01F40FDE6FE96FD978952" box="[574,622,303,329]" pageId="47" pageNumber="48">type</typeStatus>
of
<taxonomicName id="4C1A4D4FFFD01F40FD4FFE96FCF68953" authorityName="Borhidi &amp; Muniz" authorityYear="1985" box="[663,783,302,329]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="fagildei">
<emphasis id="B96EEADEFFD01F40FD4FFE96FCF68953" box="[663,783,302,329]" italics="true" pageId="47" pageNumber="48">C. fagildei</emphasis>
</taxonomicName>
is not present at HAC. The only other specimen (
<emphasis id="B96EEADEFFD01F40FA94FE97FF4B8977" italics="true" pageId="47" pageNumber="48">Salzman 505</emphasis>
) known is from the
<typeStatus id="54A1886EFFD01F40FE4BFEEAFE3A8976" box="[403,451,339,365]" pageId="47" pageNumber="48">type</typeStatus>
locality. The abaxial surface of the segments of this specimen is unusual
<emphasis id="B96EEADEFFD01F40FB32FEEBFAFF8977" bold="true" box="[1258,1286,338,364]" pageId="47" pageNumber="48">—</emphasis>
it could almost be scored as
<taxonomicName id="4C1A4D4FFFD01F40FECFFECEFE5A898B" box="[279,419,375,401]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="argentata">
<emphasis id="B96EEADEFFD01F40FECFFECEFE5A898B" box="[279,419,375,401]" italics="true" pageId="47" pageNumber="48">C. argentata</emphasis>
</taxonomicName>
-like. Fruits have not been seen (fruit size is scored from the protologue) but are described in the protologue as covered with “puntos prominulos dense dispuestos ruguloso”.
</paragraph>
</subSubSection>
</treatment>
</document>
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