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<document id="BDB28E9B57ACDF6C2101AC656F2D3540" ID-CLB-Dataset="32363" ID-DOI="10.11646/zootaxa.4306.4.7" ID-GBIF-Dataset="bfbbc6d9-281a-4202-8af1-af25d360a72d" ID-ISSN="1175-5326" ID-Zenodo-Dep="845193" ID-ZooBank="0E2DE634-1F32-465A-A8B1-DD2A6EB3F94D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503039325345" checkinUser="plazi" docAuthor="Mcknight, Tristan A. &amp; Cannings, Robert A." docDate="2017" docId="03AC4D43FFB2FFA364C9DAC1E9D7F82D" docLanguage="en" docName="zootaxa.4306.4.7.pdf" docOrigin="Zootaxa 4306 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stackelberginia cerberus Mcknight &amp; Cannings 2017, sp. nov." docType="treatment" docUuid="026A5107-BB60-49B3-B17C-80C1385D6FE9" docUuidSource="ZooBank" docVersion="8" lastPageNumber="574" masterDocId="FF95353BFFB6FFA4645ED90FE87BFFA5" masterDocTitle="Description and phylogenetic classification of Stackelberginia cerberus sp. nov. (Diptera: Asilidae), comprising the first record of this genus from the Nearctic" masterLastPageNumber="579" masterPageNumber="567" pageNumber="571" updateTime="1698466912992" updateUser="ExternalLinkService">
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<mods:title id="1AC56D2D49134E33968BE8C7EEF9263E">Description and phylogenetic classification of Stackelberginia cerberus sp. nov. (Diptera: Asilidae), comprising the first record of this genus from the Nearctic</mods:title>
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<mods:namePart id="7272075E2FCF8A632CA5D6BBD785CFE5">Mcknight, Tristan A.</mods:namePart>
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<paragraph id="8BBAFC55FFB2FFA064C9DAC1EAD4FC4D" blockId="4.[151,687,974,1000]" box="[151,687,974,1000]" pageId="4" pageNumber="571">
<heading id="D0F24B39FFB2FFA064C9DAC1EAD4FC4D" bold="true" box="[151,687,974,1000]" fontSize="11" level="1" pageId="4" pageNumber="571" reason="1">
<emphasis id="B9712047FFB2FFA064C9DAC1EAD4FC4D" bold="true" box="[151,687,974,1000]" pageId="4" pageNumber="571">
<taxonomicName id="4C0587D6FFB2FFA064C9DAC1E9BAFC4D" ID-CoL="9HKHN" authority="Mcknight &amp; Cannings, 2017" authorityName="Mcknight &amp; Cannings" authorityYear="2017" box="[151,449,974,1000]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="cerberus" status="sp. nov.">
<emphasis id="B9712047FFB2FFA064C9DAC1E9BAFC4D" bold="true" box="[151,449,974,1000]" italics="true" pageId="4" pageNumber="571">Stackelberginia cerberus</emphasis>
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<taxonomicNameLabel id="A2429D3CFFB2FFA06597DAC1EA5DFC4D" box="[457,550,974,1000]" pageId="4" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
McKnight
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<subSubSection id="C31FAFDEFFB2FFA064C9DD1BEB8FFB19" pageId="4" pageNumber="571" type="diagnosis">
<paragraph id="8BBAFC55FFB2FFA064C9DD1BEB8FFB19" blockId="4.[151,1437,1044,2005]" pageId="4" pageNumber="571">
<emphasis id="B9712047FFB2FFA064C9DD1BE96AFB88" bold="true" box="[151,273,1044,1069]" pageId="4" pageNumber="571">Diagnosis.</emphasis>
A small grey species similar to
<taxonomicName id="4C0587D6FFB2FFA06620DD1AEB78FB89" authorityName="Loew" authorityYear="1847" box="[638,771,1045,1068]" class="Insecta" family="Asilidae" genus="Lasiopogon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="571" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB2FFA06620DD1AEB78FB89" box="[638,771,1045,1068]" italics="true" pageId="4" pageNumber="571">Lasiopogon</emphasis>
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but with facial swelling mostly flattened; mystax confined to lower facial swelling; bristles of ocellar triangle, thorax, and legs extremely long and strong; occipital setae strong and straight; tarsal claws unusually large; metacoxa with anterior peg blunt; cuticle dark brown or black except for ferruginous tibiae and tarsi, mostly covered in grey tomentum; genitalia ferruginous, hypopygium rotated 120° and epandrium completely divided medially, narrow (
<figureCitation id="133EE0D0FFB2FFA067DFDDABEBB3FB19" box="[897,968,1188,1213]" captionStart="FIGURE 1" captionStartId="5.[151,250,1501,1523]" captionTargetBox="[227,1436,663,1476]" captionTargetId="figure@5.[151,1436,663,1476]" captionTargetPageId="5" captionText="FIGURE 1. Stackelberginia cerberus sp. nov. female habitus and head detail. The male differs only in the terminalia, which are presented in Fig 2." httpUri="https://zenodo.org/record/845195/files/figure.png" pageId="4" pageNumber="571">Figs 1</figureCitation>
,
<figureCitation id="133EE0D0FFB2FFA06788DDABEB9DFB19" box="[982,998,1188,1212]" captionStart="FIGURE 2" captionStartId="6.[151,250,952,974]" captionTargetBox="[151,1436,265,931]" captionTargetId="figure@6.[151,1436,265,931]" captionTargetPageId="6" captionText="FIGURE 2. Stackelberginia cerberus sp. nov. genitalia detail: A) epandrium, dorsal; B) epandrium lateral; C) gonostylus, apical; D) subepandrial sclerite, ventral; E) gonocoxite + hypandrium complex, ventral; F) gonocoxite + hypandrium complex, lateral; G) female ovipositor, lateral; H) phallus, lateral. Scale bars are 0.2 mm." httpUri="https://zenodo.org/record/845197/files/figure.png" pageId="4" pageNumber="571">2</figureCitation>
).
</paragraph>
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<emphasis id="B9712047FFB2FFA06499DDC8E922FB45" bold="true" box="[199,345,1223,1248]" pageId="4" pageNumber="571">Description.</emphasis>
<emphasis id="B9712047FFB2FFA06501DDC8E99DFB45" box="[351,486,1223,1248]" italics="true" pageId="4" pageNumber="571">Body length</emphasis>
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;
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<quantity id="4CFD51B0FFB2FFA066EDDDC8EB31FB7A" box="[691,842,1223,1248]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.15" metricValueMin="0.85" pageId="4" pageNumber="571" unit="mm" value="10.0" valueMax="11.5" valueMin="8.5">8.511.5 mm</quantity>
.
</paragraph>
<paragraph id="8BBAFC55FFB2FFA06499DDE3ECBFF9A5" blockId="4.[151,1437,1044,2005]" pageId="4" pageNumber="571">
<emphasis id="B9712047FFB2FFA06499DDE3E97FFAA0" box="[199,260,1260,1285]" italics="true" pageId="4" pageNumber="571">Head</emphasis>
. Cuticle black; tomentum of face silver, vertex and occiput grey. Facial swelling greatly reduced, leaving only a slightly swollen angle above oral margin but otherwise not extending beyond eyes in lateral view. All setae on head white except ocellar and orbital bristles, which are typically chestnut brown. Mystax setae long, loosely clumped over oral margin with a small gap dorsomedially; frons with a clump of short, thick setae midway between antennae and ocellar triangle; one long, strong orbital seta anteriorly inclined, occasionally with another short fine seta; ocellar triangle with two long, strong setae and several short fine hairs anteriorly inclined along posterior edge, longest bristle to
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; occipital bristles erect, relatively short and straight, in lateral view extending ventrally to midpoint of eye. Palps with fine white hairs along length; robust setae on postgena.
</paragraph>
<paragraph id="8BBAFC55FFB2FFA06499DF02EC5AF9C9" blockId="4.[151,1437,1044,2005]" pageId="4" pageNumber="571">
HW ♂
<quantity id="4CFD51B0FFB2FFA06546DF03E9CCF981" box="[280,439,1548,1572]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" metricValueMax="1.88" metricValueMin="1.48" pageId="4" pageNumber="571" unit="mm" value="1.68" valueMax="1.88" valueMin="1.48">1.481.88mm</quantity>
; ♀
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. FW ♂
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; ♀
<quantity id="4CFD51B0FFB2FFA067C2DF03EC40F980" box="[924,1083,1548,1573]" metricMagnitude="-4" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.5" metricValueMin="3.2" pageId="4" pageNumber="571" unit="mm" value="0.33499999999999996" valueMax="0.35" valueMin="0.32">0.320.35mm</quantity>
. VW ♂
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; ♀
<quantity id="4CFD51B0FFB2FFA06103DF03E88CF9ED" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" metricValueMax="7.7" metricValueMin="6.7" pageId="4" pageNumber="571" unit="mm" value="0.72" valueMax="0.77" valueMin="0.67">0.67 0.77mm</quantity>
. GL ♂
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; ♀
<quantity id="4CFD51B0FFB2FFA06653DF20EAD7F9ED" box="[525,684,1583,1608]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.3" metricValueMin="2.5" pageId="4" pageNumber="571" unit="mm" value="0.29" valueMax="0.33" valueMin="0.25">0.250.33mm</quantity>
. VW/HW = ♂ 0.390.45; ♀ 0.41. FW/VW = ♂ 0.420.46; ♀ 0.46 0.48. VD/VW = ♂ 0.100.14; ♀ 0.120.13. GH/GL = ♂ 0.430.52 ♀ 0.420.48.
</paragraph>
<paragraph id="8BBAFC55FFB2FFA06499DF77EAF4F97D" blockId="4.[151,1437,1044,2005]" pageId="4" pageNumber="571">
<emphasis id="B9712047FFB2FFA06499DF77E94AF92A" box="[199,305,1656,1679]" italics="true" pageId="4" pageNumber="571">Antennae</emphasis>
. Dark brown, base of pedicel pale brown. Robust white setae on scape and pedicel. Postpedicel rectangular with a slight bulge at midlength; stylus long, with pronounced apical spine. WPP/LPP =
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0.22;
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0.200.21. LAS/LPP =
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0.50;
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0.670.70.
</paragraph>
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<emphasis id="B9712047FFB2FFA06499DFEBE963F958" box="[199,280,1764,1789]" italics="true" pageId="4" pageNumber="571">Thorax</emphasis>
. Thoracic cuticle dark brown, covered in thick grey tomentum. Prothorax with hairs white; postpronotal lobes chestnut brown with lateral angle reddish, hairs white; postalar lobes reddish. Scutum tomentum grey, without stripes or spots; short hairs sparse, white around perimeter and brown/black over dorsum. Dorsocentral bristles brown, prominent (longest to
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), 1 anterior and 2 posterior to transverse suture. Lateral bristles of the scutum brown, very long (longest to
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): one postalar, one supra-alar, and two notopleural (one each of the presutural and posthumeral subgroups of
<bibRefCitation id="EF9481A4FFB2FFA0674BDE98EBB3F815" author="Cannings" box="[789,968,1943,1968]" pageId="4" pageNumber="571" refString="Cannings, R. A. (2002) The systematics of Lasiopogon (Diptera: Asilidae). Royal British Columbia Museum, Victoria, 354 pp." type="book" year="2002">Cannings 2002</bibRefCitation>
). Scutellum covered in grey tomentum, without impressed rim, bare on disc, 5-6 long white bristles apically with a few short fine hairs just inside.
</paragraph>
<paragraph id="8BBAFC55FFB3FFA16499D998EB44FEB8" blockId="5.[151,1437,151,608]" pageId="5" pageNumber="572">
Pleural tomentum grey, all hairs and bristles white. Katatergal setae 67 with a few finer hairs mixed dorsally; katepisternal setae fine, short; anepisternum with 2 long, prominent setae (longest
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) at posterior edge and a patch of fine hairs along dorsal margin; anepimeron without setae. Antepronotum with 812 fine setae; proepisternum with 46 strong long setae and 02 fine hairs.
</paragraph>
<paragraph id="8BBAFC55FFB3FFA16499D826EC3DFDC5" blockId="5.[151,1437,151,608]" pageId="5" pageNumber="572">
<emphasis id="B9712047FFB3FFA16499D826E887FEE5" box="[199,252,297,320]" italics="true" pageId="5" pageNumber="572">Legs</emphasis>
. Cuticle black except reddish tibia, trochanters, extreme medial and apical tips of femorae, and tarsal bases; tomentum grey, very thin below femora. Anterior face of metacoxa bear pronounced peg with apex rounded, not tapered; 2 fine setae projecting ventrally at base of peg. All hairs and bristles of legs white except on tarsi, where mostly brown. Coxae with dense, long, strong bristles facing anteriorly (on procoxa) or laterally (on meso- and metacoxa); 40+ on procoxa, 312 on mesocoxa, 4 on metacoxa. Femora with fine, short procumbent hairs and strong erect bristles; 56 ventral bristles longer than adjacent hairs on pro- and mesofemur, metafemur with only fine hairs just longer than surrounding setae. Apical and dorsolateral bristles strong, numerous: profemur with 35, mesofemur with 24, metafemur with 57. Bristles on tibiae and tarsi strong, long; protibia with longest bristle 3.5x longer than tibial width. Tarsal claws large, with reddish base and black apices.
</paragraph>
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<emphasis id="B9712047FFB3FFA164C9DCD2E96FFA57" bold="true" box="[151,276,1501,1523]" pageId="5" pageNumber="572">FIGURE 1.</emphasis>
<taxonomicName id="4C0587D6FFB3FFA16542DCD2EA62FA56" authority="Mcknight &amp; Cannings, 2017" authorityName="Mcknight &amp; Cannings" authorityYear="2017" box="[284,537,1501,1523]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="cerberus" status="sp. nov.">
<emphasis id="B9712047FFB3FFA16542DCD2EA62FA56" box="[284,537,1501,1523]" italics="true" pageId="5" pageNumber="572">Stackelberginia cerberus</emphasis>
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<emphasis id="B9712047FFB3FFA1667EDCD1EA09FA56" bold="true" box="[544,626,1502,1523]" pageId="5" pageNumber="572">sp. nov.</emphasis>
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female habitus and head detail. The male differs only in the terminalia, which are presented in Fig 2.
</paragraph>
</caption>
<paragraph id="8BBAFC55FFB3FFA16499DF4FE9DAF967" blockId="5.[151,1437,1598,2019]" pageId="5" pageNumber="572">
<emphasis id="B9712047FFB3FFA16499DF4FE971F9F2" box="[199,266,1600,1623]" italics="true" pageId="5" pageNumber="572">Wings</emphasis>
. Veins yellow to dark brown, darker apically; membrane hyaline, slightly brown in oblique view. DCI = 0.520.58; cell m3 broadly open; cell cu
<emphasis id="B9712047FFB3FFA16605DF6BEA12F9DE" box="[603,617,1636,1659]" italics="true" pageId="5" pageNumber="572">p</emphasis>
(sensu
<bibRefCitation id="EF9481A4FFB3FFA166E0DF6DEB0AF9DE" author="McAlpine" box="[702,881,1634,1659]" pageId="5" pageNumber="572" refString="McAlpine, J. F. (1981) 2. Morphology and terminology-adults. In: McAlpine, J. F., Peterson, B. V., Shewell, G. E., Teskey, H. J., Vockeroth, J. R. &amp; Wood, D. M. (Coords.), Manual of Nearctic Diptera. Uol. 1. Agriculture Canada, Ottawa, pp. 9 - 63." type="journal article" year="1981">McAlpine 1981</bibRefCitation>
; cu
<emphasis id="B9712047FFB3FFA167C9DF6BEBDEF9DE" box="[919,933,1636,1659]" italics="true" pageId="5" pageNumber="572">a</emphasis>
sensu
<bibRefCitation id="EF9481A4FFB3FFA167ACDF6CEC80F9DE" author="Wootton" box="[1010,1275,1635,1659]" pageId="5" pageNumber="572" refString="Wootton, R. J. &amp; Ennos, A. R. (1989) The implications of function on the origin and homologies of the dipterous wing. Systematic Entomology, 14, 507 - 520." type="journal article" year="1989">Wootton &amp; Ennos 1989</bibRefCitation>
) just closed at wing margin, varying from slightly stalked to slightly open in different individuals. Halter cream to light brown; knob without dark spot.
</paragraph>
<paragraph id="8BBAFC55FFB3FFA16499DFC1EC9CF8D2" blockId="5.[151,1437,1598,2019]" pageId="5" pageNumber="572">
<emphasis id="B9712047FFB3FFA16499DFC1E94AF942" box="[199,305,1742,1767]" italics="true" pageId="5" pageNumber="572">Abdomen</emphasis>
. Tergite base color dark brown, light brown apically; covered in pale grey tomentum without other patterning. Tergite 1 with 68 strong bristles basolaterally, other tergites with strong short lateral setae, longest apically. Lateral setae white, dorsal setulae white laterally, black near midline. Sternite tomentum grey; setae white; sternite 1 with clump of 58 prominent erect setae. Tergite 8 red, visible between tergite 7 and terminalia. Terminalia red, with long hairs (white in male, black in female), in male rotated ~120° to one side.
</paragraph>
<paragraph id="8BBAFC55FFB3FFA26499DE8DEB92FF71" blockId="5.[151,1437,1598,2019]" lastBlockId="6.[151,1437,151,213]" lastPageId="6" lastPageNumber="573" pageId="5" pageNumber="572">
<emphasis id="B9712047FFB3FFA16499DE8DE914F83E" box="[199,367,1922,1947]" italics="true" pageId="5" pageNumber="572">Male genitalia</emphasis>
. Epandrium and gonocoxite-hypandrial complex reddish-orange with darker apices, without tomentum, clothed in long white setae. Epandrium completely divided medially, in lateral view each half with width about 48% the length, widest at apex, ventral and dorsal margins mostly parallel, slumping ventrally, apex truncate with rounded corners. In dorsal view, medial margins of epandrial halves very shallowly concave, almost straight. Epandrial apodeme shallow; basal sclerite absent (
<bibRefCitation id="EF9481A4FFB0FFA26773D9B3EBA6FF71" author="Cannings" box="[813,989,188,213]" pageId="6" pageNumber="573" refString="Cannings, R. A. (2002) The systematics of Lasiopogon (Diptera: Asilidae). Royal British Columbia Museum, Victoria, 354 pp." type="book" year="2002">Cannings 2002</bibRefCitation>
).
</paragraph>
<caption id="DF7AACDDFFB0FFA264C9DAB7EBC3FBAE" httpUri="https://zenodo.org/record/845197/files/figure.png" pageId="6" pageNumber="573" targetBox="[151,1436,265,931]" targetPageId="6">
<paragraph id="8BBAFC55FFB0FFA264C9DAB7EBC3FBAE" blockId="6.[151,1436,952,1035]" pageId="6" pageNumber="573">
<emphasis id="B9712047FFB0FFA264C9DAB7E96DFC68" bold="true" box="[151,278,952,974]" pageId="6" pageNumber="573">FIGURE 2.</emphasis>
<taxonomicName id="4C0587D6FFB0FFA26541DAB7EA64FC6B" authority="Mcknight &amp; Cannings, 2017" authorityName="Mcknight &amp; Cannings" authorityYear="2017" box="[287,543,952,974]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="cerberus" status="sp. nov.">
<emphasis id="B9712047FFB0FFA26541DAB7EA64FC6B" box="[287,543,952,974]" italics="true" pageId="6" pageNumber="573">Stackelberginia cerberus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2429D3CFFB0FFA26676DAB7EA00FC68" box="[552,635,952,973]" pageId="6" pageNumber="573" rank="species">
<emphasis id="B9712047FFB0FFA26676DAB7EA00FC68" bold="true" box="[552,635,952,973]" pageId="6" pageNumber="573">sp. nov.</emphasis>
</taxonomicNameLabel>
genitalia detail: A) epandrium, dorsal; B) epandrium lateral; C) gonostylus, apical; D) subepandrial sclerite, ventral; E) gonocoxite + hypandrium complex, ventral; F) gonocoxite + hypandrium complex, lateral; G) female ovipositor, lateral; H) phallus, lateral. Scale bars are 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BBAFC55FFB0FFA26499DD37E994FAA0" blockId="6.[151,1436,1080,2005]" pageId="6" pageNumber="573">
Gonostylus elongate, slender, curved dorsally like a hooked finger, with another point dorsolaterally at the base. Long hairs over basal part of gonostylus. Gonocoxite-hypandrial complex in ventral view with width about 86% the length, partially fused laterally, medial transverse slit semicircular with distally pointed apex. Medial setal brush on gonocoxite-hypandrial complex white, setae long but relatively sparse compared to
<taxonomicName id="4C0587D6FFB0FFA2614FDDAAEDEEFB19" authorityName="Loew" authorityYear="1847" box="[1297,1429,1189,1212]" class="Insecta" family="Asilidae" genus="Lasiopogon" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB0FFA2614FDDAAEDEEFB19" box="[1297,1429,1189,1212]" italics="true" pageId="6" pageNumber="573">Lasiopogon</emphasis>
</taxonomicName>
. Gonocoxal apodemes long, in lateral view exposed length about 70% the basal width of the hypandrium; apodeme with sclerotized web ventrally.
</paragraph>
<paragraph id="8BBAFC55FFB0FFA26499DC1FEABDFA30" blockId="6.[151,1436,1080,2005]" pageId="6" pageNumber="573">Phallus paramere sheath long and slender, curved ventrally like a goose neck. Dorsal carina a low ridge no wider than the paramere sheath itself that follows the curvature of the paramere sheath and terminates before the apex without a notch. Subepandrial sclerite V-shaped; triangular medial unsclerotized portion in basal 65%; spines slender, attenuate, sparsely scattered over surface.</paragraph>
<paragraph id="8BBAFC55FFB0FFA26499DCAFEAFAF9A4" blockId="6.[151,1436,1080,2005]" pageId="6" pageNumber="573">
<emphasis id="B9712047FFB0FFA26499DCAFE9F0FA1C" box="[199,395,1440,1465]" italics="true" pageId="6" pageNumber="573">Female genitalia</emphasis>
. (Undissected) Hairs on segment 8 dark brown/black, erect, abundant. Tergite 8 black, sternite 8 red/yellow, hypogynial valves dark brown, with many dark hairs; lateral lobe with dark setae; acanthophorite spines black, 3 on each half.
</paragraph>
</subSubSection>
<subSubSection id="C31FAFDEFFB0FFA36499DF03EAB2FF52" lastPageId="7" lastPageNumber="574" pageId="6" pageNumber="573" type="materials_examined">
<paragraph id="8BBAFC55FFB0FFA26499DF03EBF1F910" blockId="6.[151,1436,1080,2005]" pageId="6" pageNumber="573">
<emphasis id="B9712047FFB0FFA26499DF03E902F980" bold="true" box="[199,377,1548,1573]" pageId="6" pageNumber="573">Type Material.</emphasis>
HOLOTYPE. ♂ labelled: &quot;[rectangular white label] NEV: Nye Co: NTS [Nuclear Test Site]/ Rock Valley/
<date id="FFBBDA95FFB0FFA26569DF3FE9D9F9EC" box="[311,418,1584,1609]" pageId="6" pageNumber="573" value="1976-09-12">IX-12-76</date>
, can tr./E.L. Sleeper et al.&quot;; “[rectangular white label]?/ det. EMFisher”; “[rectangular white label] FISH”; “[rectangular white label] mirus”. A holotype label &quot;HOLOTYPE/
<taxonomicName id="4C0587D6FFB0FFA26026DF5BED5EF9C8" authorityName="Lehr" authorityYear="1964" box="[1144,1317,1620,1645]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="573" phylum="Arthropoda" rank="genus">Stackelberginia</taxonomicName>
/
<taxonomicName id="4C0587D6FFB0FFA2616BDF5BEDE8F9C8" authorityName="Mcknight &amp; Cannings" authorityYear="2017" box="[1333,1427,1620,1645]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="cerberus">cerberus</taxonomicName>
/ des. T.A. McKnight 2017 [red, black-bordered label]&quot; has been attached to this specimen. USNM type #USNMENT01295437. Both antennae are missing the postpedicel.
</paragraph>
<paragraph id="8BBAFC55FFB0FFA26499DFCFECDFF815" blockId="6.[151,1436,1080,2005]" pageId="6" pageNumber="573">
<materialsCitation id="3B6DF608FFB0FFA26499DFCFECDFF815" ID-GBIF-Occurrence="1586208346" collectingDate="1975-10-03" collectingDateMax="2013-10-06" collectingDateMin="1975-10-03" collectionCode="DNA, NTS, RBCM, FISH" collectorName="T. A. McKnight &amp; E. L. Sleeper &amp; Hwy" country="United States of America" county="Clark Co" elevation="1251" latitude="36.63359" location="Nye Co" longLatPrecision="1" longitude="-116.31018" municipality="Wheeler" pageId="6" pageNumber="573" specimenCode="ENT017-001946, ENT017- 001945" specimenCount="7" specimenCount-female="6" specimenCount-male="1" stateProvince="Nevada" typeStatus="paratype">
<typeStatus id="54BE42F7FFB0FFA26499DFCFE919F97D" box="[199,354,1728,1752]" pageId="6" pageNumber="573" type="paratype">PARATYPES</typeStatus>
(8 designated, including 3 with genomic
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extracts).
<emphasis id="B9712047FFB0FFA267A4DFCFECC8F97C" bold="true" box="[1018,1203,1728,1753]" pageId="6" pageNumber="573">
<collectingCountry id="F312BCC5FFB0FFA267A4DFCFEC31F97C" box="[1018,1098,1728,1753]" name="United States of America" pageId="6" pageNumber="573">U.S.A.</collectingCountry>
:
<collectingRegion id="49C132B7FFB0FFA26007DFCFECC8F97C" box="[1113,1203,1728,1753]" country="United States of America" name="Nevada" pageId="6" pageNumber="573">Nevada</collectingRegion>
</emphasis>
,
<collectingCounty id="62DB84D9FFB0FFA2609ADFCFED4BF97D" box="[1220,1328,1728,1753]" pageId="6" pageNumber="573">Clark Co</collectingCounty>
:
<collectingMunicipality id="6BDE662FFFB0FFA26163DFCFEDE7F97C" box="[1341,1436,1728,1753]" pageId="6" pageNumber="573">Wheeler</collectingMunicipality>
wash
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<quantity id="4CFD51B0FFB0FFA26482DFEBE969F958" box="[220,274,1764,1789]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="6" pageNumber="573" unit="mi" value="6.0">6 mi</quantity>
NE of Pahrump
</locationDeviation>
,
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<geoCoordinate id="EE319A92FFB0FFA266D6DFEAEB49F958" box="[648,818,1764,1789]" direction="west" orientation="longitude" pageId="6" pageNumber="573" precision="1" value="-115.89428">W 115.89428°</geoCoordinate>
, elev:
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<quantity id="4CFD51B0FFB0FFA267DDDFEBEBA4F959" box="[899,991,1764,1789]" metricMagnitude="3" metricUnit="m" metricValue="1.251" pageId="6" pageNumber="573" unit="m" value="1251.0">1251 m</quantity>
</elevation>
,
<collectingDate id="EFFF237DFFB0FFA267B1DFEBEC2FF958" box="[1007,1108,1764,1789]" pageId="6" pageNumber="573" value="2013-10-05">
<date id="FFBBDA95FFB0FFA267B1DFEBEC2FF958" box="[1007,1108,1764,1789]" pageId="6" pageNumber="573" value="2013-10-05">5.x.2013</date>
</collectingDate>
,
<collectorName id="26F09983FFB0FFA2603DDFEAED68F958" box="[1123,1299,1764,1789]" pageId="6" pageNumber="573">T.A. McKnight</collectorName>
(
<specimenCount id="9D0337DCFFB0FFA26174DFEAED30F958" box="[1322,1355,1765,1789]" pageId="6" pageNumber="573" type="female">1♀</specimenCount>
[
<collectionCode id="ED146490FFB0FFA26102DFEAEDE7F958" box="[1372,1436,1765,1789]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="6" pageNumber="573">DNA</collectionCode>
voucher] UMMZ-TAM-509-1,
<specimenCount id="9D0337DCFFB0FFA2665DDE07EA5DF885" box="[515,550,1800,1824]" pageId="6" pageNumber="573" type="female">1♀</specimenCount>
[
<collectionCode id="ED146490FFB0FFA26664DE06EA01F884" box="[570,634,1801,1825]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="6" pageNumber="573">DNA</collectionCode>
voucher] UMMZ-TAM-509-2;
<specimenCount id="9D0337DCFFB0FFA267AADE07EC6CF885" box="[1012,1047,1800,1824]" pageId="6" pageNumber="573" type="female">1♀</specimenCount>
[EtOH] UMMZ-TAM-509-3; 1? [damaged by dermestids] UMMZ-TAM-509-4);
<location id="8EDAAA8EFFB0FFA2669EDE22EB66F8E0" LSID="urn:lsid:plazi:treatment:03AC4D43FFB2FFA364C9DAC1E9D7F82D:8EDAAA8EFFB0FFA2669EDE22EB66F8E0" box="[704,797,1837,1861]" country="United States of America" county="Clark Co" latitude="36.63359" longLatPrecision="1" longitude="-116.31018" municipality="Wheeler" name="Nye Co" pageId="6" pageNumber="573" stateProvince="Nevada">Nye Co</location>
:
<collectionCode id="ED146490FFB0FFA26777DE22EB25F8E0" box="[809,862,1837,1861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14925" name="Nevada Operations Office, U.S. Department of Energy" pageId="6" pageNumber="573">NTS</collectionCode>
[
<location id="8EDAAA8EFFB0FFA26730DE23EC43F8E0" LSID="urn:lsid:plazi:treatment:03AC4D43FFB2FFA364C9DAC1E9D7F82D:8EDAAA8EFFB0FFA26730DE23EC43F8E0" box="[878,1080,1836,1861]" country="United States of America" county="Clark Co" latitude="36.63359" longLatPrecision="1" longitude="-116.31018" municipality="Wheeler" name="Nuclear Test Site" pageId="6" pageNumber="573" stateProvince="Nevada">Nuclear Test Site</location>
]
<location id="8EDAAA8EFFB0FFA26016DE23ECA8F8E0" LSID="urn:lsid:plazi:treatment:03AC4D43FFB2FFA364C9DAC1E9D7F82D:8EDAAA8EFFB0FFA26016DE23ECA8F8E0" box="[1096,1235,1836,1861]" country="United States of America" county="Clark Co" latitude="36.63359" longLatPrecision="1" longitude="-116.31018" municipality="Wheeler" name="Rock Valley" pageId="6" pageNumber="573" stateProvince="Nevada">Rock Valley</location>
can tr.,
<collectingDate id="EFFF237DFFB0FFA2616DDE23EDECF8E0" box="[1331,1431,1836,1861]" pageId="6" pageNumber="573" value="1975-10-03">
<date id="FFBBDA95FFB0FFA2616DDE23EDECF8E0" box="[1331,1431,1836,1861]" pageId="6" pageNumber="573" value="1975-10-03">3.x.1975</date>
</collectingDate>
,
<collectorName id="26F09983FFB0FFA264C9DE5EE959F8CC" box="[151,290,1872,1897]" pageId="6" pageNumber="573">E.L. Sleeper</collectorName>
(
<specimenCount id="9D0337DCFFB0FFA26568DE5FE92DF8CD" box="[310,342,1872,1896]" pageId="6" pageNumber="573" type="female">1♀</specimenCount>
[dissected]
<collectionCode id="ED146490FFB0FFA26580DE5FEA54F8CD" box="[478,559,1872,1896]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="6" pageNumber="573">RBCM</collectionCode>
<specimenCode id="DBA3542EFFB0FFA26669DE5FEA82F8CC" box="[567,761,1872,1897]" collectionCode="ENT" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="6" pageNumber="573">ENT017-001946</specimenCode>
),
<collectingDate id="EFFF237DFFB0FFA26752DE5FEBFCF8CC" box="[780,903,1872,1897]" pageId="6" pageNumber="573" value="1976-09-12">
<date id="FFBBDA95FFB0FFA26752DE5FEBFCF8CC" box="[780,903,1872,1897]" pageId="6" pageNumber="573" value="1976-09-12">12.ix.1976</date>
</collectingDate>
,
<collectorName id="26F09983FFB0FFA267CCDE5EEC65F8CC" box="[914,1054,1872,1897]" pageId="6" pageNumber="573">E.L. Sleeper</collectorName>
(
<specimenCount id="9D0337DCFFB0FFA26071DE5FEC29F8CD" box="[1071,1106,1872,1896]" pageId="6" pageNumber="573" type="male">1♂</specimenCount>
[dissected]
<collectionCode id="ED146490FFB0FFA26084DE5FED50F8CD" box="[1242,1323,1872,1896]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="6" pageNumber="573">RBCM</collectionCode>
<specimenCode id="DBA3542EFFB0FFA2616CDE5FE88BF828" collectionCode="ENT" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="6" pageNumber="573">ENT017- 001945</specimenCode>
);
<location id="8EDAAA8EFFB0FFA2655ADE7BE9F4F828" LSID="urn:lsid:plazi:treatment:03AC4D43FFB2FFA364C9DAC1E9D7F82D:8EDAAA8EFFB0FFA2655ADE7BE9F4F828" box="[260,399,1908,1933]" country="United States of America" county="Clark Co" latitude="36.63359" longLatPrecision="1" longitude="-116.31018" municipality="Wheeler" name="Rock Valley" pageId="6" pageNumber="573" stateProvince="Nevada">Rock Valley</location>
wash jct
<collectorName id="26F09983FFB0FFA265A1DE7AEA4EF828" box="[511,565,1909,1933]" pageId="6" pageNumber="573">Hwy</collectorName>
95, 5 mi E of
<location id="8EDAAA8EFFB0FFA266BEDE7AEBD8F828" LSID="urn:lsid:plazi:treatment:03AC4D43FFB2FFA364C9DAC1E9D7F82D:8EDAAA8EFFB0FFA266BEDE7AEBD8F828" box="[736,931,1908,1933]" country="United States of America" county="Clark Co" latitude="36.63359" longLatPrecision="1" longitude="-116.31018" municipality="Wheeler" name="Amargosa Valley" pageId="6" pageNumber="573" stateProvince="Nevada">Amargosa Valley</location>
city, N of hwy,
<geoCoordinate id="EE319A92FFB0FFA26000DE7AEC94F828" box="[1118,1263,1908,1933]" direction="north" orientation="latitude" pageId="6" pageNumber="573" precision="1" value="36.63359">N 36.63359°</geoCoordinate>
<geoCoordinate id="EE319A92FFB0FFA260A6DE7AEDE7F828" box="[1272,1436,1908,1933]" direction="west" orientation="longitude" pageId="6" pageNumber="573" precision="1" value="-116.31018">W 116.31018°</geoCoordinate>
elev:
<elevation id="00281B66FFB0FFA2648ADE97E966F815" box="[212,285,1944,1968]" metricMagnitude="2" metricUnit="m" metricValue="8.78" pageId="6" pageNumber="573" unit="m" value="878.0">
<quantity id="4CFD51B0FFB0FFA2648ADE97E966F815" box="[212,285,1944,1968]" metricMagnitude="2" metricUnit="m" metricValue="8.78" pageId="6" pageNumber="573" unit="m" value="878.0">878 m</quantity>
</elevation>
,
<collectingDate id="EFFF237DFFB0FFA26577DE97E9F6F814" box="[297,397,1944,1969]" pageId="6" pageNumber="573" value="2013-10-06">
<date id="FFBBDA95FFB0FFA26577DE97E9F6F814" box="[297,397,1944,1969]" pageId="6" pageNumber="573" value="2013-10-06">6.x.2013</date>
</collectingDate>
,
<collectorName id="26F09983FFB0FFA265C7DE96EA3FF814" box="[409,580,1944,1969]" pageId="6" pageNumber="573">T.A. McKnight</collectorName>
(
<specimenCount id="9D0337DCFFB0FFA26609DE97EA03F815" box="[599,632,1944,1968]" pageId="6" pageNumber="573" type="female">1♀</specimenCount>
[
<collectionCode id="ED146490FFB0FFA266DBDE96EABEF814" box="[645,709,1945,1969]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="6" pageNumber="573">DNA</collectionCode>
voucher] UMMZ-TAM-512-1;
<specimenCount id="9D0337DCFFB0FFA26072DE97EC34F815" box="[1068,1103,1944,1968]" pageId="6" pageNumber="573" type="female">1♀</specimenCount>
<collectionCode id="ED146490FFB0FFA26008DE97ECE3F815" box="[1110,1176,1944,1968]" pageId="6" pageNumber="573">FISH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BBAFC55FFB0FFA26499DEB2ED74F870" blockId="6.[151,1436,1080,2005]" box="[199,1295,1980,2005]" pageId="6" pageNumber="573">
<materialsCitation id="3B6DF608FFB0FFA26499DEB2ED74F870" ID-GBIF-Occurrence="1586208344" box="[199,1295,1980,2005]" country="United States of America" county="Nye Co." latitude="36.694" location="Rock Valley" longLatPrecision="70" longitude="-116.179" municipality="Nuclear Test Site" pageId="6" pageNumber="573" specimenCount="1" stateProvince="Nevada" typeStatus="holotype">
<emphasis id="B9712047FFB0FFA26499DEB2E927F870" box="[199,348,1980,2005]" italics="true" pageId="6" pageNumber="573">
<typeStatus id="54BE42F7FFB0FFA26499DEB2E880F871" box="[199,251,1981,2004]" pageId="6" pageNumber="573">Type</typeStatus>
Locality
</emphasis>
.
<collectingCountry id="F312BCC5FFB0FFA26534DEB2E9C1F870" box="[362,442,1981,2005]" name="United States of America" pageId="6" pageNumber="573">U.S.A.</collectingCountry>
;
<collectingRegion id="49C132B7FFB0FFA26598DEB3EA65F870" box="[454,542,1980,2005]" country="United States of America" name="Nevada" pageId="6" pageNumber="573">Nevada</collectingRegion>
,
<collectingCounty id="62DB84D9FFB0FFA26677DEB2EAF2F870" box="[553,649,1981,2005]" pageId="6" pageNumber="573">Nye Co.</collectingCounty>
,
<collectingMunicipality id="6BDE662FFFB0FFA266CBDEB3EB23F870" box="[661,856,1980,2005]" pageId="6" pageNumber="573">Nuclear Test Site</collectingMunicipality>
,
<location id="8EDAAA8EFFB0FFA2673DDEB3EB96F870" LSID="urn:lsid:plazi:treatment:03AC4D43FFB2FFA364C9DAC1E9D7F82D:8EDAAA8EFFB0FFA2673DDEB3EB96F870" box="[867,1005,1980,2005]" country="United States of America" county="Nye Co." latitude="36.694" longLatPrecision="70" longitude="-116.179" municipality="Nuclear Test Site" name="Rock Valley" pageId="6" pageNumber="573" stateProvince="Nevada">Rock Valley</location>
[
<geoCoordinate id="EE319A92FFB0FFA267A5DEB2EC0AF870" box="[1019,1137,1980,2005]" direction="north" orientation="latitude" pageId="6" pageNumber="573" precision="55" value="36.694">N 36.694°</geoCoordinate>
<geoCoordinate id="EE319A92FFB0FFA26026DEB2ED7FF870" box="[1144,1284,1980,2005]" direction="west" orientation="longitude" pageId="6" pageNumber="573" precision="55" value="-116.179">W 116.179°</geoCoordinate>
].
</materialsCitation>
</paragraph>
<paragraph id="8BBAFC55FFB1FFA36499D998EAB2FF52" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="574">
<emphasis id="B9712047FFB1FFA36499D998E9E1FF15" bold="true" box="[199,410,151,176]" pageId="7" pageNumber="574">Taxonomic Notes.</emphasis>
This species was included in
<bibRefCitation id="EF9481A4FFB1FFA366B8D998EBDAFF15" author="Cannings" box="[742,929,151,176]" pageId="7" pageNumber="574" refString="Cannings, R. A. (2002) The systematics of Lasiopogon (Diptera: Asilidae). Royal British Columbia Museum, Victoria, 354 pp." type="book" year="2002">Cannings (2002)</bibRefCitation>
as “L. unc-
<specimenCount id="9D0337DCFFB1FFA3607DD997EC2CFF15" box="[1059,1111,151,176]" pageId="7" pageNumber="574" type="generic">
7
<emphasis id="B9712047FFB1FFA36068D998EC2CFF15" bold="true" box="[1078,1111,151,176]" pageId="7" pageNumber="574">sp.</emphasis>
</specimenCount>
nov.” and had been assigned a manuscript name of “mirus” because it was thought to be a miraculously unusual
<taxonomicName id="4C0587D6FFB1FFA36010D9B2ECA9FF71" authorityName="Loew" authorityYear="1847" box="[1102,1234,189,212]" class="Insecta" family="Asilidae" genus="Lasiopogon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA36010D9B2ECA9FF71" box="[1102,1234,189,212]" italics="true" pageId="7" pageNumber="574">Lasiopogon</emphasis>
</taxonomicName>
. This epithet was changed after reclassification outside
<taxonomicName id="4C0587D6FFB1FFA36660D9EEEAB9FF5D" authorityName="Loew" authorityYear="1847" box="[574,706,225,248]" class="Insecta" family="Asilidae" genus="Lasiopogon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA36660D9EEEAB9FF5D" box="[574,706,225,248]" italics="true" pageId="7" pageNumber="574">Lasiopogon</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31FAFDEFFB1FFA36499D80BEBF7FE2D" pageId="7" pageNumber="574" type="etymology">
<paragraph id="8BBAFC55FFB1FFA36499D80BEBF7FE2D" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="574">
<emphasis id="B9712047FFB1FFA36499D80BE935FEB8" bold="true" box="[199,334,260,285]" pageId="7" pageNumber="574">Etymology.</emphasis>
Latin
<emphasis id="B9712047FFB1FFA365C8D80BE985FEB8" box="[406,510,260,285]" italics="true" pageId="7" pageNumber="574">Cerberus</emphasis>
from the Greek Κέρβερος, used as a noun in apposition, named in similarity to the hound guarding the gates of the underworld in Greek mythology. The
<typeStatus id="54BE42F7FFB1FFA367F1D827EBA4FEE5" box="[943,991,296,320]" pageId="7" pageNumber="574">type</typeStatus>
locality for this predator is just outside the boundaries of both Death Valley National Park (the lowest elevation in North America) and the Yucca Mountain nuclear waste facility—sites with a suitably Hadean flair.
</paragraph>
</subSubSection>
<subSubSection id="C31FAFDEFFB1FFA36499D89BE997FB5A" pageId="7" pageNumber="574" type="distribution">
<paragraph id="8BBAFC55FFB1FFA36499D89BE95BFDBD" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="574">
<emphasis id="B9712047FFB1FFA36499D89BE91BFE08" bold="true" box="[199,352,404,429]" pageId="7" pageNumber="574">Distribution.</emphasis>
Nearctic:
<collectingCountry id="F312BCC5FFB1FFA36585D89BEA6DFE09" box="[475,534,404,428]" name="United States of America" pageId="7" pageNumber="574">USA</collectingCountry>
; known only from two sites in the Amargosa desert of southwestern
<collectingRegion id="49C132B7FFB1FFA3611ED89BEDE2FE08" box="[1344,1433,404,429]" country="United States of America" name="Nevada" pageId="7" pageNumber="574">Nevada</collectingRegion>
. Previously described species of
<taxonomicName id="4C0587D6FFB1FFA3665ED8B8EAD5FE75" authorityName="Lehr" authorityYear="1964" box="[512,686,439,464]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA3665ED8B8EAD5FE75" box="[512,686,439,464]" italics="true" pageId="7" pageNumber="574">Stackelberginia</emphasis>
</taxonomicName>
are known from deserts in
<collectingCountry id="F312BCC5FFB1FFA367BAD8B8EC1CFE75" box="[996,1127,439,464]" name="Kazakhstan" pageId="7" pageNumber="574">Kazakhstan</collectingCountry>
and
<collectingCountry id="F312BCC5FFB1FFA360C0D8B8ED42FE75" box="[1182,1337,439,464]" name="Turkmenistan" pageId="7" pageNumber="574">Turkmenistan</collectingCountry>
(
<figureCitation id="133EE0D0FFB1FFA36116D8B8EDF7FE75" box="[1352,1420,439,464]" captionStart="FIGURE 3" captionStartId="7.[151,250,1085,1107]" captionText="FIGURE 3. Stackelberginia world distribution." pageId="7" pageNumber="574">Fig. 3</figureCitation>
); we have summarized localities from the literature (
<bibRefCitation id="EF9481A4FFB1FFA36754D8D3EBF0FE50" author="Lehr" box="[778,907,476,501]" pageId="7" pageNumber="574" refString="Lehr, P. A. (1964) [New genera and new species of the robber-flies (Diptera, Asilidae) in the fauna of the USSR]. Entomologicheskoe Obozrenie, 18 (4), 914 - 934. [in Russian]" type="journal article" year="1964">Lehr 1964</bibRefCitation>
,
<bibRefCitation id="EF9481A4FFB1FFA367C0D8D3EBACFE51" author="Lehr" box="[926,983,476,500]" pageId="7" pageNumber="574" refString="Lehr, P. A. (1980) [Eurybiotic mode of life as the initial stage of evolution of a taxon, by the example of robber flies (Diptera, Asilidae)]. In: [Parallelism and Regularity in the Evolution of Insects] Far East Division of the Academy of Sciences of the USSR, Vladivostok, pp. 4 - 69. [in Russian]" type="book chapter" year="1980">1980</bibRefCitation>
) below with estimated georeferenced coordinates:
</paragraph>
<caption id="DF7AACDDFFB1FFA364C9DD32EA02FBF6" box="[151,633,1085,1107]" pageId="7" pageNumber="574">
<paragraph id="8BBAFC55FFB1FFA364C9DD32EA02FBF6" blockId="7.[151,633,1085,1107]" box="[151,633,1085,1107]" pageId="7" pageNumber="574">
<emphasis id="B9712047FFB1FFA364C9DD32E969FBF6" bold="true" box="[151,274,1085,1107]" pageId="7" pageNumber="574">FIGURE 3.</emphasis>
<taxonomicName id="4C0587D6FFB1FFA36547DD32E9CDFBF6" authorityName="Lehr" authorityYear="1964" box="[281,438,1085,1107]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA36547DD32E9CDFBF6" box="[281,438,1085,1107]" italics="true" pageId="7" pageNumber="574">Stackelberginia</emphasis>
</taxonomicName>
world distribution.
</paragraph>
</caption>
<paragraph id="8BBAFC55FFB1FFA36499DD73E997FB5A" blockId="7.[151,1437,1147,1928]" pageId="7" pageNumber="574">
<materialsCitation id="3B6DF608FFB1FFA36499DD73E997FB5A" ID-GBIF-Occurrence="1586208345" collectingDate="1952-09-07" collectingDateMax="1960-09-07" collectingDateMin="1952-09-07" collectorName="P. A. Lehr" country="Kazakhstan" county="River Ila" latitude="43.696" location="Baskuduk district" longLatPrecision="68" longitude="51.227" municipality="Ayakkalkan" pageId="7" pageNumber="574" specimenCount="1" specimenCount-male="1" stateProvince="Almaty Oblast">
<taxonomicName id="4C0587D6FFB1FFA36499DD73E94CFB31" authorityName="Lehr" authorityYear="1964" box="[199,311,1147,1172]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B9712047FFB1FFA36499DD73E94CFB31" box="[199,311,1147,1172]" italics="true" pageId="7" pageNumber="574">S. gracilis</emphasis>
</taxonomicName>
:
<collectingCountry id="F312BCC5FFB1FFA36518DD74E9B6FB31" box="[326,461,1147,1172]" name="Kazakhstan" pageId="7" pageNumber="574">Kazakhstan</collectingCountry>
:
<collectingRegion id="49C132B7FFB1FFA36589DD74EA04FB31" box="[471,639,1147,1172]" country="Kazakhstan" name="Almaty Oblysy" pageId="7" pageNumber="574">Almaty Oblast</collectingRegion>
:
<collectingCounty id="62DB84D9FFB1FFA366D7DD74EA97FB31" box="[649,748,1147,1172]" pageId="7" pageNumber="574">River Ila</collectingCounty>
at
<collectingMunicipality id="6BDE662FFFB1FFA36753DD74EBE9FB31" box="[781,914,1147,1172]" pageId="7" pageNumber="574">Ayakkalkan</collectingMunicipality>
resort, sandy crest among gravel, claypan, and marshy hollows on left bank [
<geoCoordinate id="EE319A92FFB1FFA365BBDDAFEA20FB1D" box="[485,603,1183,1208]" direction="north" orientation="latitude" pageId="7" pageNumber="574" precision="55" value="43.861">N 43.861°</geoCoordinate>
<geoCoordinate id="EE319A92FFB1FFA3663CDDAFEAA3FB12" box="[610,728,1183,1207]" direction="east" orientation="longitude" pageId="7" pageNumber="574" precision="55" value="78.424">E 78.424°</geoCoordinate>
],
<collectingDate id="EFFF237DFFB1FFA366B4DD90EBFAFB1D" box="[746,897,1183,1208]" pageId="7" pageNumber="574" value="1952-09-07" valueMax="1952-10-09" valueMin="1952-09-07">
<date id="FFBBDA95FFB1FFA366B4DD90EBFAFB1D" box="[746,897,1183,1208]" pageId="7" pageNumber="574" value="1952-09-07" valueMax="1952-10-09" valueMin="1952-09-07">7.ix-9.x.1952</date>
</collectingDate>
(number of specimens not noted); Mangishlyak Peninsula:
<location id="8EDAAA8EFFB1FFA3654CDDCCE9A1FB79" LSID="urn:lsid:plazi:treatment:03AC4D43FFB2FFA364C9DAC1E9D7F82D:8EDAAA8EFFB1FFA3654CDDCCE9A1FB79" box="[274,474,1219,1244]" country="Kazakhstan" county="River Ila" latitude="43.696" longLatPrecision="68" longitude="51.227" municipality="Ayakkalkan" name="Baskuduk district" pageId="7" pageNumber="574" stateProvince="Almaty Oblast">Baskuduk district</location>
, salt flats and claypan amidst marshy hollows and dune chains [
<geoCoordinate id="EE319A92FFB1FFA360FEDDCBED6EFB79" box="[1184,1301,1219,1244]" direction="north" orientation="latitude" pageId="7" pageNumber="574" precision="55" value="43.696">N 43.696°</geoCoordinate>
<geoCoordinate id="EE319A92FFB1FFA36142DDCBEDEAFB79" box="[1308,1425,1219,1244]" direction="east" orientation="longitude" pageId="7" pageNumber="574" precision="55" value="51.227">E 51.227°</geoCoordinate>
],
<collectingDate id="EFFF237DFFB1FFA364C9DDE8E978FAA5" box="[151,259,1255,1280]" pageId="7" pageNumber="574" value="1960-09-07">
<date id="FFBBDA95FFB1FFA364C9DDE8E978FAA5" box="[151,259,1255,1280]" pageId="7" pageNumber="574" value="1960-09-07">7.ix.1960</date>
</collectingDate>
, col.
<collectorName id="26F09983FFB1FFA36561DDE7E9D1FAA5" box="[319,426,1255,1280]" pageId="7" pageNumber="574">P.A. Lehr</collectorName>
(
<specimenCount id="9D0337DCFFB1FFA365E5DDE8E9A4FB5A" box="[443,479,1255,1279]" pageId="7" pageNumber="574" type="male">1♂</specimenCount>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31FAFDEFFB1FFA36499DC03E9D7F82D" pageId="7" pageNumber="574" type="description">
<paragraph id="8BBAFC55FFB1FFA36499DC03E9A2FAE2" blockId="7.[151,1437,1147,1928]" pageId="7" pageNumber="574">
<materialsCitation id="3B6DF608FFB1FFA36499DC03E9A2FAE2" ID-GBIF-Occurrence="1586208347" collectingDate="1959-10-09" collectorName="D. M. Charykulievi" country="Turkmenistan" latitude="37.291" location="Leskhoz" longLatPrecision="70" longitude="62.351" pageId="7" pageNumber="574" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Mary">
<taxonomicName id="4C0587D6FFB1FFA36499DC03E90FFA81" authorityName="Lehr" authorityYear="1964" box="[199,372,1291,1316]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="tsharykulievi">
<emphasis id="B9712047FFB1FFA36499DC03E90FFA81" box="[199,372,1291,1316]" italics="true" pageId="7" pageNumber="574">S. tsharykulievi</emphasis>
</taxonomicName>
:
<collectingCountry id="F312BCC5FFB1FFA365DBDC04EA5FFA81" box="[389,548,1291,1316]" name="Turkmenistan" pageId="7" pageNumber="574">Turkmenistan</collectingCountry>
:
<collectingRegion id="49C132B7FFB1FFA3666EDC03EA0AFA81" box="[560,625,1292,1316]" country="Turkmenistan" name="Mary Welayaty" pageId="7" pageNumber="574">Mary</collectingRegion>
:
<location id="8EDAAA8EFFB1FFA36620DC04EA9BFA81" LSID="urn:lsid:plazi:treatment:03AC4D43FFB2FFA364C9DAC1E9D7F82D:8EDAAA8EFFB1FFA36620DC04EA9BFA81" box="[638,736,1291,1316]" country="Turkmenistan" latitude="37.291" longLatPrecision="70" longitude="62.351" name="Leskhoz" pageId="7" pageNumber="574" stateProvince="Mary">Leskhoz</location>
, desert region, on sand [
<geoCoordinate id="EE319A92FFB1FFA367A8DC03EC15FA86" box="[1014,1134,1291,1315]" direction="north" orientation="latitude" pageId="7" pageNumber="574" precision="55" value="37.291">N 37.291°</geoCoordinate>
<geoCoordinate id="EE319A92FFB1FFA36028DC03EC96FA81" box="[1142,1261,1291,1316]" direction="east" orientation="longitude" pageId="7" pageNumber="574" precision="55" value="62.351">E 62.351°</geoCoordinate>
],
<collectingDate id="EFFF237DFFB1FFA3615FDC04ED1DFA81" box="[1281,1382,1291,1316]" pageId="7" pageNumber="574" value="1959-10-09">
<date id="FFBBDA95FFB1FFA3615FDC04ED1DFA81" box="[1281,1382,1291,1316]" pageId="7" pageNumber="574" value="1959-10-09">9.x.1959</date>
</collectingDate>
, col.
<collectorName id="26F09983FFB1FFA364C9DC3FE910FAED" box="[151,363,1327,1352]" pageId="7" pageNumber="574">D.M. Charykulievi</collectorName>
(
<specimenCount id="9D0337DCFFB1FFA36522DC20E9E4FAE2" box="[380,415,1327,1351]" pageId="7" pageNumber="574" type="male">1♂</specimenCount>
<specimenCount id="9D0337DCFFB1FFA365FBDC20E9B1FAE2" box="[421,458,1327,1351]" pageId="7" pageNumber="574" type="female">2♀</specimenCount>
).
</materialsCitation>
</paragraph>
<paragraph id="8BBAFC55FFB1FFA36499DC5CEC71FA11" blockId="7.[151,1437,1147,1928]" pageId="7" pageNumber="574">
<emphasis id="B9712047FFB1FFA36499DC5CE9F7FAC9" bold="true" box="[199,396,1363,1388]" pageId="7" pageNumber="574">Natural History.</emphasis>
Habitat: sandy dry desert washes, usually found perching on fist-sized rocks in sandy areas near clumps of vegetation (
<figureCitation id="133EE0D0FFB1FFA36594DC78EA74FA2A" box="[458,527,1399,1424]" captionStart="FIGURE 4" captionStartId="8.[151,250,683,705]" captionTargetBox="[169,1417,195,661]" captionTargetId="figure@8.[169,1417,195,661]" captionTargetPageId="8" captionText="FIGURE 4. Habitats where Stackelberginia cerberus sp. nov. were collected in Nevada (see specimen list for more precise locality data): A) Clark Co.: Wheeler wash; B) Nye Co.: Rock Valley wash. Individuals were caught perching on small stones and sand along the washes." httpUri="https://zenodo.org/record/845199/files/figure.png" pageId="7" pageNumber="574">Fig. 4</figureCitation>
). One female was observed preying on a slightly-smaller fly (not collected). The fall flight period (
<date id="FFBBDA95FFB1FFA36500DC94EA02FA11" box="[350,633,1435,1460]" pageId="7" pageNumber="574">12 September6 October</date>
) is distinctive within the subfamily.
</paragraph>
<paragraph id="8BBAFC55FFB1FFA36499DCB0ED16F985" blockId="7.[151,1437,1147,1928]" pageId="7" pageNumber="574">
Other assassin flies were also active at the
<taxonomicName id="4C0587D6FFB1FFA366F2DCB0EB21FA7D" authorityName="Lehr" authorityYear="1964" box="[684,858,1471,1496]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA366F2DCB0EB21FA7D" box="[684,858,1471,1496]" italics="true" pageId="7" pageNumber="574">Stackelberginia</emphasis>
</taxonomicName>
sites at the same time. These were identified using keys in
<bibRefCitation id="EF9481A4FFB1FFA364ACDCECE9FCFA5E" author="Wood" box="[242,391,1507,1532]" pageId="7" pageNumber="574" refString="Wood, G. C. (1981) Family Asilidae. In: McAlpine, J. R., Peterson, B. V., Shell, G. E., Teskey, H. J., Vockeroth, J. R. &amp; Wood, D. M. (Eds.), Manual of Nearctic Diptera. Uol. 1. Agriculture Canada, Hull, pp. 549 - 573." type="journal article" year="1981">Wood (1981)</bibRefCitation>
and
<bibRefCitation id="EF9481A4FFB1FFA3659CDCECEA1BFA59" author="Wilcox" box="[450,608,1507,1532]" pageId="7" pageNumber="574" refString="Wilcox, J. (1961) The genus Hodophylax James (Diptera: Asilidae). Bulletin of the Brooklyn Entomological Society, 61, 112 - 116." type="journal article" year="1961">Wilcox (1961</bibRefCitation>
,
<bibRefCitation id="EF9481A4FFB1FFA36630DCECEAD2FA59" author="Wilcox" box="[622,681,1507,1532]" pageId="7" pageNumber="574" refString="Wilcox, J. (1965) New Cophura, with revised keys to the groups and to the Nearctic species (Diptera: Asilidae). Annals of the Entomological Society of America, 58 (6), 805 - 816. https: // doi. org / 10.1093 / aesa / 58.6.805" type="journal article" year="1965">1965</bibRefCitation>
,
<bibRefCitation id="EF9481A4FFB1FFA366E8DCECEA94FA59" author="Wilcox" box="[694,751,1507,1532]" pageId="7" pageNumber="574" refString="Wilcox, J. (1966) Efferia Coquillett in America north of Mexico (Diptera: Asilidae). Proceedings of the California Academy of Sciences, 34 (2), 85 - 234." type="journal article" year="1966">1966</bibRefCitation>
) and voucher specimens have been deposited at the RBCM and UMMZ:
<taxonomicName id="4C0587D6FFB1FFA36572DF08EAEDF985" authority="Pritchard" authorityName="Pritchard" authorityYear="1938" box="[300,662,1543,1568]" class="Insecta" family="Asilidae" genus="Hodophylax" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="basingeri">
<emphasis id="B9712047FFB1FFA36572DF08EA5DF985" box="[300,550,1543,1568]" italics="true" pageId="7" pageNumber="574">Hodophylax basingeri</emphasis>
Pritchard
</taxonomicName>
,
<taxonomicName id="4C0587D6FFB1FFA366FFDF08EBD4F985" authority="Wilcox" authorityName="Wilcox" authorityYear="1965" box="[673,943,1543,1568]" class="Insecta" family="Asilidae" genus="Cophura" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="B9712047FFB1FFA366FFDF08EB28F985" box="[673,851,1543,1568]" italics="true" pageId="7" pageNumber="574">Cophura fisheri</emphasis>
Wilcox
</taxonomicName>
,
<taxonomicName id="4C0587D6FFB1FFA367E4DF08ED16F985" authority="Wilcox." authorityName="Wilcox." box="[954,1389,1543,1568]" class="Insecta" family="Asilidae" genus="Efferia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="basingeri" subGenus="Aridefferia">
<emphasis id="B9712047FFB1FFA367E4DF08ED76F985" box="[954,1293,1543,1568]" italics="true" pageId="7" pageNumber="574">Efferia (Aridefferia) basingeri</emphasis>
Wilcox.
</taxonomicName>
</paragraph>
<paragraph id="8BBAFC55FFB1FFA36499DF24E9D7F82D" blockId="7.[151,1437,1147,1928]" pageId="7" pageNumber="574">
<emphasis id="B9712047FFB1FFA36499DF24E932F9E1" bold="true" box="[199,329,1579,1604]" pageId="7" pageNumber="574">Phylogeny.</emphasis>
The species tree supports
<taxonomicName id="4C0587D6FFB1FFA36630DF24EB67F9E1" authorityName="Lehr" authorityYear="1964" box="[622,796,1579,1604]" class="Insecta" family="Asilidae" genus="Stackelberginia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA36630DF24EB67F9E1" box="[622,796,1579,1604]" italics="true" pageId="7" pageNumber="574">Stackelberginia</emphasis>
</taxonomicName>
as sister to
<taxonomicName id="4C0587D6FFB1FFA367C0DF23EC59F9E6" authorityName="Loew" authorityYear="1847" box="[926,1058,1580,1603]" class="Insecta" family="Asilidae" genus="Lasiopogon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA367C0DF23EC59F9E6" box="[926,1058,1580,1603]" italics="true" pageId="7" pageNumber="574">Lasiopogon</emphasis>
</taxonomicName>
, and this clade as sister to the rest of the subfamily (
<figureCitation id="133EE0D0FFB1FFA36538DF40E9D5F9CD" box="[358,430,1615,1640]" captionStart="FIGURE 5" captionStartId="8.[151,250,1598,1620]" captionTargetBox="[171,1400,828,1571]" captionTargetId="figure@8.[159,1428,816,1577]" captionTargetPageId="8" captionText="FIGURE 5. Species tree for representatives of Stackelberginia, Lasiopogon, Stichopogon, Townsendia, and Asilidae and Mydidae outgroups, estimated via Bayesian inference in STARBEAST 2 from one mitochondrial locus (COI) and three nuclear protein coding loci (wingless, PEPCK, AATS). Branch labels show support for nodes when posterior probability &lt;1." httpUri="https://zenodo.org/record/845201/files/figure.png" pageId="7" pageNumber="574">Fig. 5</figureCitation>
); these relationships were also recovered in each of the four individual gene trees (not shown). Node posterior probabilities are reasonably high (± 0.92) throughout the tree, except for some uncertainty for phylogenetic relationships within
<taxonomicName id="4C0587D6FFB1FFA3661ADF97EAB3F90A" authorityName="Loew" authorityYear="1847" box="[580,712,1688,1711]" class="Insecta" family="Asilidae" genus="Lasiopogon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA3661ADF97EAB3F90A" box="[580,712,1688,1711]" italics="true" pageId="7" pageNumber="574">Lasiopogon</emphasis>
</taxonomicName>
and at the
<taxonomicName id="4C0587D6FFB1FFA36713DF98EC72F915" box="[845,1033,1687,1712]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="subFamily" subFamily="Stichopogoninae">Stichopogoninae</taxonomicName>
root. Lower support values for the subfamily root appear to stem from a basal polytomy in COI; this locus is known to have trouble resolving deeper level phylogenetic relationships (
<bibRefCitation id="EF9481A4FFB1FFA36677DFD0EB55F952" author="Matsuda" box="[553,814,1759,1784]" pageId="7" pageNumber="574" refString="Matsuda, T., Morishita, M., Hinomoto, N. &amp; Gotoh, T. (2014) Phylogenetic Analysis of the Spider Mite Sub-Family Tetranychinae (Acari: Tetranychidae) Based on the Mitochondrial COI Gene and the 18 S and the 5 ' End of the 28 S rRNA Genes Indicates That Several Genera Are Polyphyletic. PLoS ONE, 9 (10), e 108672. https: // doi. org / 10.1371 / journal. pone. 0108672" type="journal article" year="2014">
Matsuda
<emphasis id="B9712047FFB1FFA366C3DFEFEAA4F95D" box="[669,735,1759,1784]" italics="true" pageId="7" pageNumber="574">et al.</emphasis>
2014
</bibRefCitation>
). Intrageneric relationships within
<taxonomicName id="4C0587D6FFB1FFA36083DFEFED1AF952" authorityName="Loew" authorityYear="1847" box="[1245,1377,1760,1783]" class="Insecta" family="Asilidae" genus="Lasiopogon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA36083DFEFED1AF952" box="[1245,1377,1760,1783]" italics="true" pageId="7" pageNumber="574">Lasiopogon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0587D6FFB1FFA364C9DE0CE959F8B9" authorityName="Loew" authorityYear="1847" box="[151,290,1795,1820]" class="Insecta" family="Asilidae" genus="Stichopogon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA364C9DE0CE959F8B9" box="[151,290,1795,1820]" italics="true" pageId="7" pageNumber="574">Stichopogon</emphasis>
</taxonomicName>
will be covered in future publications with more comprehensive taxon sampling, but the overall genus-level topology matches our prior expectations from morphology—e.g.,
<taxonomicName id="4C0587D6FFB1FFA36002DE28ECA5F8E5" authorityName="Williston" authorityYear="1895" box="[1116,1246,1831,1856]" class="Insecta" family="Asilidae" genus="Townsendia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA36002DE28ECA5F8E5" box="[1116,1246,1831,1856]" italics="true" pageId="7" pageNumber="574">Townsendia</emphasis>
</taxonomicName>
grouping with
<taxonomicName id="4C0587D6FFB1FFA364C9DE44E959F8C1" authorityName="Loew" authorityYear="1847" box="[151,290,1867,1892]" class="Insecta" family="Asilidae" genus="Stichopogon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA364C9DE44E959F8C1" box="[151,290,1867,1892]" italics="true" pageId="7" pageNumber="574">Stichopogon</emphasis>
</taxonomicName>
, and the Asilinae (
<taxonomicName id="4C0587D6FFB1FFA36652DE44EB53F8C6" authority="Macquart" authorityName="Macquart" authorityYear="1838" box="[524,808,1867,1892]" class="Insecta" family="Asilidae" genus="Proctacanthus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA36652DE44EAD5F8C1" box="[524,686,1867,1892]" italics="true" pageId="7" pageNumber="574">Proctacanthus</emphasis>
Macquart
</taxonomicName>
) and
<taxonomicName id="4C0587D6FFB1FFA3672FDE44EB87F8C1" box="[881,1020,1867,1892]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="subFamily" subFamily="Ommatiinae">Ommatiinae</taxonomicName>
(
<taxonomicName id="4C0587D6FFB1FFA3604CDE43ED62F8C1" authority="Wiedemann" authorityName="Wiedemann" authorityYear="1821" box="[1042,1305,1867,1892]" class="Insecta" family="Asilidae" genus="Ommatius" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B9712047FFB1FFA3604CDE43ECFEF8C6" box="[1042,1157,1868,1891]" italics="true" pageId="7" pageNumber="574">Ommatius</emphasis>
Wiedemann
</taxonomicName>
) clustering together in the outgroup.
</paragraph>
</subSubSection>
</treatment>
</document>