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<document id="17841314A299CA79D775A2112589CAB8" ID-CLB-Dataset="3612" ID-DOI="10.11646/zootaxa.5061.1.6" ID-GBIF-Dataset="0e4f0459-0892-43b2-b01d-50f384c6c945" ID-ISSN="1175-5326" ID-Zenodo-Dep="5679421" ID-ZooBank="AED1C537-0DEF-4356-BDE9-E590A9EBCDD3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635923379677" checkinUser="plazi" docAuthor="Ngo, Quoc Phu &amp; Nguyen, Minh Ty" docDate="2021" docId="03A8552C1B428446FF32FB99FB9336EC" docLanguage="en" docName="zootaxa.5061.1.6.pdf" docOrigin="Zootaxa 5061 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophiogomphus (Ophionurus) phantoani Ngo &amp; Nguyen 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="139" masterDocId="FF912D541B408443FFA5FFA7FFE83422" masterDocTitle="The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae)" masterLastPageNumber="144" masterPageNumber="134" pageNumber="136" updateTime="1699467663801" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="450AFEC1BF6207581A25D6FAB17685FF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B43298B98B2365FD675552BAB4D863CE">
<mods:title id="62291C59E2A2703C6A941A9D86ED4427">The genus Ophiogomphus Lieftinck, 1964 in Vietnam, with descriptions of the female of O. minimus Karube, 2014 and O. phantoani sp. nov. (Odonata: Gomphidae)</mods:title>
</mods:titleInfo>
<mods:name id="CDFF4CF66CEA4329B57E72B8C07F0071" type="personal">
<mods:role id="CFC15C0627335D8A38E8F2B01F2C01A2">
<mods:roleTerm id="621EEB2C96D50447CA54E012D58E402F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D48B34E31C0B343449C26CC15E3D71C3">Ngo, Quoc Phu</mods:namePart>
<mods:affiliation id="DA3727EFA891DBF49E3F0D25F3C7E01C">Center for Entomology &amp; Parasitology Research, College of Medicine &amp; Pharmacy, Duy Tan University, Da Nang 550000, Vietnam. Institue of Applied Technology, Thu Dau Mot University, No. 06 Tran Van On, Phu Hoa Ward, Thu Dau Mot City, Binh Duong province, Vietnam. https: // orcid. org / 0000 - 0002 - 2023 - 1813</mods:affiliation>
</mods:name>
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<mods:roleTerm id="D8822F46F426EA5C228E5D6DDD8AE98A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47CD23DF078BE1E4EF4FFFC6CEEFBD7D">Nguyen, Minh Ty</mods:namePart>
<mods:nameIdentifier id="24C52A38C1795CB4EBC7205669D719CC" type="ORCID">0000-0002-2023-1813</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="EE04265563B6B921C6DE19CC2D289FAA">text</mods:typeOfResource>
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<mods:titleInfo id="863090A1DAA16D9F8B43F0EC9469297B">
<mods:title id="4A96894856D3C2BC19C0C25B016B2DA9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FEFFB39B8CCA5DF8CB41FE90700F66B2">
<mods:date id="7071455FE4E47EDB082C0B544D63BAF5">2021</mods:date>
<mods:detail id="4126896CF2B7FF51785BB5C81E0222AA" type="pubDate">
<mods:number id="44E2B522DF5EF7637C15C5BC24CCFB28">2021-11-03</mods:number>
</mods:detail>
<mods:detail id="DDEC9D5084EB7394816C344213A56A8F" type="volume">
<mods:number id="74B0296DF9C9293CAE13B7E86020CBE0">5061</mods:number>
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<mods:number id="AC7339CF75916F5AF22DE89F3829F5F5">1</mods:number>
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<mods:classification id="8871F907B823FBB07569E5008CB5DB25">journal article</mods:classification>
<mods:identifier id="962A6F7BF6FE7238500D4AAF890846D3" type="CLB-Dataset">3612</mods:identifier>
<mods:identifier id="8B94A55A1E18C080BAC70CEE1CDC5225" type="DOI">10.11646/zootaxa.5061.1.6</mods:identifier>
<mods:identifier id="DAAD20C3DB4A60EABAD97C178CFA2390" type="GBIF-Dataset">0e4f0459-0892-43b2-b01d-50f384c6c945</mods:identifier>
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<treatment id="03A8552C1B428446FF32FB99FB9336EC" ID-DOI="http://doi.org/10.5281/zenodo.5699201" ID-GBIF-Taxon="190304677" ID-Zenodo-Dep="5699201" LSID="urn:lsid:plazi:treatment:03A8552C1B428446FF32FB99FB9336EC" httpUri="http://treatment.plazi.org/id/03A8552C1B428446FF32FB99FB9336EC" lastPageId="5" lastPageNumber="139" pageId="2" pageNumber="136">
<subSubSection id="C31BB7B11B428441FF32FB99FD0F307A" box="[151,743,1086,1113]" pageId="2" pageNumber="136" type="nomenclature">
<paragraph id="8BBEE43A1B428441FF32FB99FD0F307A" blockId="2.[151,743,1086,1149]" box="[151,743,1086,1113]" pageId="2" pageNumber="136">
<heading id="D0F653561B428441FF32FB99FD0F307A" bold="true" box="[151,743,1086,1113]" fontSize="11" level="1" pageId="2" pageNumber="136" reason="1">
<emphasis id="B97538281B428441FF32FB99FD0F307A" bold="true" box="[151,743,1086,1113]" pageId="2" pageNumber="136">
<taxonomicName id="4C019FB91B428441FF32FB99FD6A307B" ID-CoL="9X5GW" authority="Ngo &amp; Nguyen, 2021" authorityName="Ngo &amp; Nguyen" authorityYear="2021" box="[151,642,1086,1113]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="136" phylum="Arthropoda" rank="species" species="phantoani" status="sp. nov." subGenus="Ophionurus">
<emphasis id="B97538281B428441FF32FB99FD6A307B" bold="true" box="[151,642,1086,1113]" italics="true" pageId="2" pageNumber="136">Ophiogomphus (Ophionurus) phantoani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24685531B428441FD2CFB99FD0F307A" box="[649,743,1086,1112]" pageId="2" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31BB7B11B428441FF32FBC5FD80305E" box="[151,616,1122,1149]" pageId="2" pageNumber="136" type="description">
<paragraph id="8BBEE43A1B428441FF32FBC5FD80305E" blockId="2.[151,743,1086,1149]" box="[151,616,1122,1149]" pageId="2" pageNumber="136">
(
<figureCitation id="133AF8BF1B428441FF3AFBC5FEF9305E" box="[159,273,1122,1148]" captionStart-0="FIGURES 4–5" captionStart-1="FIGURES 6–9" captionStart-2="FIGURES 10–13" captionStartId-0="3.[151,264,1974,1998]" captionStartId-1="4.[151,264,1976,2000]" captionStartId-2="5.[151,264,1645,1669]" captionTargetBox-0="[160,1409,198,1929]" captionTargetBox-1="[151,1436,1065,1953]" captionTargetBox-2="[157,1427,747,1611]" captionTargetId-0="figure-18@3.[155,1433,181,1950]" captionTargetId-1="figure-364@4.[151,1436,1065,1953]" captionTargetId-2="figure-265@5.[151,1436,741,1622]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." captionText-1="FIGURES 6–9. Head of Ophiogomphus phantoani sp. nov. in frontal and dorsal view. (6, 8), Holotype ♂ &amp; (7, 9), Paratype ♀." captionText-2="FIGURES 10–13. Appendages Ophiogomphus spp. in lateral and dorsal view. (10, 11), O. phantoani sp. nov., holotype ♂; (12, 13), O. sinicus, Cao Bang Province." figureDoi-0="http://doi.org/10.5281/zenodo.5679425" figureDoi-1="http://doi.org/10.5281/zenodo.5679427" figureDoi-2="http://doi.org/10.5281/zenodo.5679429" httpUri-0="https://zenodo.org/record/5679425/files/figure.png" httpUri-1="https://zenodo.org/record/5679427/files/figure.png" httpUri-2="https://zenodo.org/record/5679429/files/figure.png" pageId="2" pageNumber="136">Figs 4–11</figureCitation>
,
<figureCitation id="133AF8BF1B428441FEB8FBC5FED3305E" box="[285,315,1122,1148]" captionStart="FIGURES 14" captionStartId="6.[151,264,1141,1165]" captionTargetBox="[159,1428,188,1106]" captionTargetId="figure-17@6.[151,1436,181,1117]" captionTargetPageId="6" captionText="FIGURES 14–15. Genital structures of Ophiogomphus phantoani sp. nov. holotype ♂; (14), vesica spermalis, lateral view; (15), apical segment of vesica spermalis, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5679431" httpUri="https://zenodo.org/record/5679431/files/figure.png" pageId="2" pageNumber="136">14</figureCitation>
, 15, 17, 19, 24, 25, 32, 33)
</paragraph>
</subSubSection>
<subSubSection id="C31BB7B11B428441FF32FB02FA74316D" pageId="2" pageNumber="136" type="materials_examined">
<paragraph id="8BBEE43A1B428441FF32FB02FC3D30C1" blockId="2.[151,1437,1189,2043]" pageId="2" pageNumber="136">
<materialsCitation id="3B69EE671B428441FF32FB02FC3D30C1" ID-GBIF-Occurrence="3400671311" collectingDate="2020-06-28" collectorName="Q. P. Ngo" country="Vietnam" county="Nam Giang District" elevation="1105" latitude="15.508" location="Song Thanh National Park" longLatPrecision="7" longitude="107.472" municipality="Dak Re Commune" pageId="2" pageNumber="136" specimenCount="1" specimenCount-male="1" stateProvince="Quang Nam" typeStatus="holotype">
<emphasis id="B97538281B428441FF32FB02FEEC309D" bold="true" box="[151,260,1189,1215]" pageId="2" pageNumber="136">
<typeStatus id="54BA5A981B428441FF32FB02FEEC309D" box="[151,260,1189,1215]" pageId="2" pageNumber="136" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D072FB31B428441FEB7FB02FEC1309D" box="[274,297,1189,1215]" pageId="2" pageNumber="136" type="male">♂</specimenCount>
,
<location id="8EDEB2E11B428441FE91FB02FDB3309D" LSID="urn:lsid:plazi:treatment:03A8552C1B428446FF32FB99FB9336EC:8EDEB2E11B428441FE91FB02FDB3309D" box="[308,603,1189,1215]" country="Vietnam" county="Nam Giang District" latitude="15.508" longLatPrecision="7" longitude="107.472" municipality="Dak Re Commune" name="Song Thanh National Park" pageId="2" pageNumber="136" stateProvince="Quang Nam">Song Thanh National Park</location>
(
<geoCoordinate id="EE3582FD1B428441FDCCFB02FD0B309D" box="[617,739,1189,1216]" degrees="15.5080" direction="north" orientation="latitude" pageId="2" pageNumber="136" precision="5" value="15.508">15.5080 N</geoCoordinate>
,
<geoCoordinate id="EE3582FD1B428441FD4BFB02FC98309D" box="[750,880,1189,1215]" degrees="107.4720" direction="east" orientation="longitude" pageId="2" pageNumber="136" precision="5" value="107.472">107.4720 E</geoCoordinate>
, altitude
<quantity id="4CF949DF1B428441FC70FB02FBC4309D" box="[981,1068,1189,1216]" metricMagnitude="3" metricUnit="m" metricValue="1.105" pageId="2" pageNumber="136" unit="m" value="1105.0">
<elevation id="002C03091B428441FC70FB02FBC4309D" box="[981,1068,1189,1216]" metricMagnitude="3" metricUnit="m" metricValue="1.105" pageId="2" pageNumber="136" unit="m" value="1105.0">1105 m</elevation>
</quantity>
),
<collectingMunicipality id="6BDA7E401B428441FB9AFB02FAF9309D" box="[1087,1297,1189,1215]" pageId="2" pageNumber="136">Dak Re Commune</collectingMunicipality>
,
<collectingCounty id="62DF9CB61B428441FABEFB02FF0630C1" pageId="2" pageNumber="136">Nam Giang District</collectingCounty>
,
<collectingRegion id="49C52AD81B428441FF5CFB6EFE0630C1" box="[249,494,1225,1251]" country="Vietnam" name="Quang Nam" pageId="2" pageNumber="136">Quang Nam Province</collectingRegion>
, central
<collectingCountry id="F316A4AA1B428441FDE9FB6EFD4630C1" box="[588,686,1225,1251]" name="Vietnam" pageId="2" pageNumber="136">Vietnam</collectingCountry>
,
<date id="FFBFC2FA1B428441FD1CFB6EFCDC30C1" box="[697,820,1225,1251]" pageId="2" pageNumber="136" value="2020-06-28">
<collectingDate id="EFFB3B121B428441FD1CFB6EFCDC30C1" box="[697,820,1225,1251]" pageId="2" pageNumber="136" value="2020-06-28">28.vi.2020</collectingDate>
</date>
,
<collectorName id="26F481EC1B428441FC9AFB6EFC4D30C1" box="[831,933,1225,1251]" pageId="2" pageNumber="136">Q.P. Ngo</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BBEE43A1B428441FF62FB4AFA74316D" blockId="2.[151,1437,1189,2043]" pageId="2" pageNumber="136">
<materialsCitation id="3B69EE671B428441FF62FB4AFB2F3125" ID-GBIF-Occurrence="3400671308" box="[199,1223,1261,1288]" collectingDate="2020-06-28" collectionCode="ZCDTU" collectorName="Q. P. Ngo" country="Vietnam" county="Nam Giang District" elevation="1105" latitude="15.508" location="Song Thanh National Park" longLatPrecision="7" longitude="107.472" municipality="Dak Re Commune" pageId="2" pageNumber="136" specimenCount="1" specimenCount-female="1" stateProvince="Quang Nam" typeStatus="paratype">
<emphasis id="B97538281B428441FF62FB4AFED73125" bold="true" box="[199,319,1261,1287]" pageId="2" pageNumber="136">
<typeStatus id="54BA5A981B428441FF62FB4AFED73125" box="[199,319,1261,1287]" pageId="2" pageNumber="136" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D072FB31B428441FEF5FB4AFE953125" box="[336,381,1261,1287]" pageId="2" pageNumber="136" type="female">1 ♀</specimenCount>
(in copula with
<typeStatus id="54BA5A981B428441FD9AFB4AFD493125" box="[575,673,1261,1287]" pageId="2" pageNumber="136" type="holotype">holotype</typeStatus>
male), same data as holotype, both in
<collectionCode id="ED107CFF1B428441FBC3FB4AFB2F3125" box="[1126,1223,1261,1287]" pageId="2" pageNumber="136">ZCDTU</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B69EE671B428441FB70FB4AFA71316D" ID-GBIF-Occurrence="3400671309" collectingDate="2018-07-12" collectorName="Alisa J. &amp; Thomas W. Donnelly " country="Vietnam" county="Phuoc Son District" elevation="725" latitude="15.3113" location="Song Thanh National Park" longLatPrecision="7" longitude="107.9097" municipality="Phuoc Kim Commune" pageId="2" pageNumber="136" specimenCount="1" specimenCount-female="1" stateProvince="Quang Nam" typeStatus="paratype">
<specimenCount id="9D072FB31B428441FB70FB4AFAE93125" box="[1237,1281,1261,1287]" pageId="2" pageNumber="136" type="female">1 ♀</specimenCount>
,
<location id="8EDEB2E11B428441FAB7FB4AFEDB3109" LSID="urn:lsid:plazi:treatment:03A8552C1B428446FF32FB99FB9336EC:8EDEB2E11B428441FAB7FB4AFEDB3109" country="Vietnam" county="Phuoc Son District" latitude="15.3113" longLatPrecision="7" longitude="107.9097" municipality="Phuoc Kim Commune" name="Song Thanh National Park" pageId="2" pageNumber="136" stateProvince="Quang Nam">Song Thanh National Park</location>
(
<geoCoordinate id="EE3582FD1B428441FEE1FAB6FE283109" box="[324,448,1297,1324]" degrees="15.3113" direction="north" orientation="latitude" pageId="2" pageNumber="136" precision="5" value="15.3113">15.3113 N</geoCoordinate>
,
<geoCoordinate id="EE3582FD1B428441FE68FAB6FDB93109" box="[461,593,1297,1323]" degrees="107.9097" direction="east" orientation="longitude" pageId="2" pageNumber="136" precision="5" value="107.9097">107.9097 E</geoCoordinate>
, altitude
<quantity id="4CF949DF1B428441FD19FAB6FCE03109" box="[700,776,1297,1324]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="2" pageNumber="136" unit="m" value="725.0">
<elevation id="002C03091B428441FD19FAB6FCE03109" box="[700,776,1297,1324]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="2" pageNumber="136" unit="m" value="725.0">725 m</elevation>
</quantity>
),
<collectingMunicipality id="6BDA7E401B428441FCBBFAB6FBF73109" box="[798,1055,1297,1323]" pageId="2" pageNumber="136">Phuoc Kim Commune</collectingMunicipality>
,
<collectingCounty id="62DF9CB61B428441FB89FAB6FAEE3109" box="[1068,1286,1297,1323]" pageId="2" pageNumber="136">Phuoc Son District</collectingCounty>
,
<collectingRegion id="49C52AD81B428441FAB7FAB6FF15316D" country="Vietnam" name="Quang Nam" pageId="2" pageNumber="136">Quang Nam Province</collectingRegion>
, central
<collectingCountry id="F316A4AA1B428441FEFEFA92FE55316D" box="[347,445,1333,1359]" name="Vietnam" pageId="2" pageNumber="136">Vietnam</collectingCountry>
,
<date id="FFBFC2FA1B428441FE6DFA92FDA2316D" box="[456,586,1333,1359]" pageId="2" pageNumber="136" value="2018-07-12">
<collectingDate id="EFFB3B121B428441FE6DFA92FDA2316D" box="[456,586,1333,1359]" pageId="2" pageNumber="136" value="2018-07-12">12.vii.2018</collectingDate>
</date>
,
<collectorName id="26F481EC1B428441FDF1FA92FD40316D" box="[596,680,1333,1359]" pageId="2" pageNumber="136">Alisa J.</collectorName>
&amp;
<collectorName id="26F481EC1B428441FD6EFA92FC52316D" box="[715,954,1333,1359]" pageId="2" pageNumber="136">Thomas W. Donnelly</collectorName>
leg.,
<collectorName id="26F481EC1B428441FC5DFA92FABA316D" box="[1016,1362,1333,1359]" pageId="2" pageNumber="136">Rosser W. Garrison Collection</collectorName>
, USA
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31BB7B11B428441FF62FAFEFB9C31B5" pageId="2" pageNumber="136" type="etymology">
<paragraph id="8BBEE43A1B428441FF62FAFEFB9C31B5" blockId="2.[151,1437,1189,2043]" pageId="2" pageNumber="136">
<emphasis id="B97538281B428441FF62FAFEFEA63151" bold="true" box="[199,334,1369,1395]" pageId="2" pageNumber="136">Etymology.</emphasis>
The new species is named in honor of Dr. Phan Quoc Toan, a Vietnamese odonatologist, for his great contribution to the knowledge of Southeast Asian
<taxonomicName id="4C019FB91B428441FCA2FADAFC8E31B5" authorityName="Fabricius" authorityYear="1793" box="[775,870,1405,1431]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Odonata" pageId="2" pageNumber="136" phylum="Tracheophyta" rank="order">Odonata</taxonomicName>
(noun in genitive case).
</paragraph>
</subSubSection>
<subSubSection id="C31BB7B11B428447FF62FA06FBAB3787" lastPageId="4" lastPageNumber="138" pageId="2" pageNumber="136" type="description">
<paragraph id="8BBEE43A1B428441FF62FA06FDF73199" blockId="2.[151,1437,1189,2043]" box="[199,543,1441,1467]" pageId="2" pageNumber="136">
<emphasis id="B97538281B428441FF62FA06FDF73199" bold="true" box="[199,543,1441,1467]" pageId="2" pageNumber="136">
Description of
<typeStatus id="54BA5A981B428441FED2FA06FE363199" box="[375,478,1441,1467]" pageId="2" pageNumber="136" type="holotype">holotype</typeStatus>
male
</emphasis>
</paragraph>
<paragraph id="8BBEE43A1B428441FF62FA62FC043269" blockId="2.[151,1437,1189,2043]" pageId="2" pageNumber="136">
<emphasis id="B97538281B428441FF62FA62FEEC31FD" box="[199,260,1477,1503]" italics="true" pageId="2" pageNumber="136">Head</emphasis>
(
<figureCitation id="133AF8BF1B428441FEB7FA62FEB331FD" box="[274,347,1477,1503]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1974,1998]" captionTargetBox="[160,1409,198,1929]" captionTargetId="figure-18@3.[155,1433,181,1950]" captionTargetPageId="3" captionText="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679425" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="2" pageNumber="136">Figs 4</figureCitation>
,
<figureCitation id="133AF8BF1B428441FEC3FA62FE7A31FD" box="[358,402,1477,1503]" captionStart="FIGURES 6–9" captionStartId="4.[151,264,1976,2000]" captionTargetBox="[151,1436,1065,1953]" captionTargetId="figure-364@4.[151,1436,1065,1953]" captionTargetPageId="4" captionText="FIGURES 6–9. Head of Ophiogomphus phantoani sp. nov. in frontal and dorsal view. (6, 8), Holotype ♂ &amp; (7, 9), Paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.5679427" httpUri="https://zenodo.org/record/5679427/files/figure.png" pageId="2" pageNumber="136">6, 8</figureCitation>
). Labium black, labrum black with a pair of greenish yellow markings at dorso-lateralyl; anteclypeus greenish yellow; postclypeus black with a small yellow spot at dorso-laterally; frons broadly marginated with yellow fascia lying interrupted medially with black; antennae black, yellowish basally; vertex black with two rounded prominences near lateral ocelli. Occiput black, prominent centrally.
</paragraph>
<paragraph id="8BBEE43A1B428441FF62F9F2FD3F32DD" blockId="2.[151,1437,1189,2043]" pageId="2" pageNumber="136">
<emphasis id="B97538281B428441FF62F9F2FEF0324D" box="[199,280,1621,1647]" italics="true" pageId="2" pageNumber="136">Thorax</emphasis>
(
<figureCitation id="133AF8BF1B428441FE82F9F2FE84324D" box="[295,364,1621,1647]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1974,1998]" captionTargetBox="[160,1409,198,1929]" captionTargetId="figure-18@3.[155,1433,181,1950]" captionTargetPageId="3" captionText="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679425" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="2" pageNumber="136">Fig. 4</figureCitation>
). Prothorax entirely black. Synthorax mostly black with greenish yellow markings; pale antehumeral stripe and not connected with collar stripe; humeral stripe absent; dorsal carina and post-ventral part of mesinfraepisternum with yellow markings; median part of mesepimeron with pale stripe about half as wide as combined black antehumeral and humeral stripes; a small mid-dorsal yellow stripe on metepisternum; metepimeron largely greenish yellow, anterior and ventral borders black.
</paragraph>
<paragraph id="8BBEE43A1B428441FF62F8ADFC383365" blockId="2.[151,1437,1189,2043]" pageId="2" pageNumber="136">
<emphasis id="B97538281B428441FF62F8ADFF143301" box="[199,252,1802,1827]" italics="true" pageId="2" pageNumber="136">Legs</emphasis>
(
<figureCitation id="133AF8BF1B428441FEABF8AEFEBD3301" box="[270,341,1801,1827]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1974,1998]" captionTargetBox="[160,1409,198,1929]" captionTargetId="figure-18@3.[155,1433,181,1950]" captionTargetPageId="3" captionText="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679425" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="2" pageNumber="136">Fig. 4</figureCitation>
). Legs stoutly built, entirely black. Femur and tibia thick and robust with closely set, gradually lengthening spurs apically. Coxa of front legs black with greenish yellow.
</paragraph>
<paragraph id="8BBEE43A1B428441FF62F8F5FB1233AD" blockId="2.[151,1437,1189,2043]" pageId="2" pageNumber="136">
<emphasis id="B97538281B428441FF62F8F5FEE33349" box="[199,267,1874,1899]" italics="true" pageId="2" pageNumber="136">Wings</emphasis>
(
<figureCitation id="133AF8BF1B428441FEBFF8F6FEB63349" box="[282,350,1873,1899]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1974,1998]" captionTargetBox="[160,1409,198,1929]" captionTargetId="figure-18@3.[155,1433,181,1950]" captionTargetPageId="3" captionText="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679425" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="2" pageNumber="136">Fig. 4</figureCitation>
). Wings hyaline, with brown patches at wing base; anal loop 2-celled; nodal index: Ax/Px = 17/18 (FW) and 12/19 (HW); Pt dark brown,
<quantity id="4CF949DF1B428441FDEBF8D2FD2833AD" box="[590,704,1909,1936]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="2" pageNumber="136" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
length in both wings, covering about 5 or 6-celles.
</paragraph>
<paragraph id="8BBEE43A1B428441FF62F83EFAC933D9" blockId="2.[151,1437,1189,2043]" pageId="2" pageNumber="136">
<emphasis id="B97538281B428441FF62F83EFED93391" box="[199,305,1945,1971]" italics="true" pageId="2" pageNumber="136">Abdomen</emphasis>
(
<figureCitation id="133AF8BF1B428441FEE7F83EFE603391" box="[322,392,1945,1971]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1974,1998]" captionTargetBox="[160,1409,198,1929]" captionTargetId="figure-18@3.[155,1433,181,1950]" captionTargetPageId="3" captionText="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679425" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="2" pageNumber="136">Fig. 4</figureCitation>
). Black with greenish yellow markings; S1 with a medio-ventral spot connecting with postventral spot; S2 with two large lateral spots, one on auricle, the other along posterior carina; S3–6 mostly black each with a narrow basal ring; S7 with a basal spot covering about half of the segment; S8–10 entirely black.
</paragraph>
<caption id="DF7EB4B21B438440FF32F811FAF433EC" ID-DOI="http://doi.org/10.5281/zenodo.5679425" ID-Zenodo-Dep="5679425" box="[151,1308,1974,1999]" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="3" pageNumber="137" startId="3.[151,264,1974,1998]" targetBox="[160,1409,198,1929]" targetPageId="3">
<paragraph id="8BBEE43A1B438440FF32F811FAF433EC" blockId="3.[151,1308,1974,1999]" box="[151,1308,1974,1999]" pageId="3" pageNumber="137">
<emphasis id="B97538281B438440FF32F811FEDC33EC" bold="true" box="[151,308,1974,1998]" pageId="3" pageNumber="137">FIGURES 4–5</emphasis>
. Habitus of
<taxonomicName id="4C019FB91B438440FE14F811FD5233EC" authority="Ngo &amp; Nguyen, 2021" authorityName="Ngo &amp; Nguyen" authorityYear="2021" box="[433,698,1974,1998]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="137" phylum="Arthropoda" rank="species" species="phantoani" status="sp. nov.">
<emphasis id="B97538281B438440FE14F811FD5233EC" box="[433,698,1974,1998]" italics="true" pageId="3" pageNumber="137">Ophiogomphus phantoani</emphasis>
</taxonomicName>
<emphasis id="B97538281B438440FD65F811FCF833EC" bold="true" box="[704,784,1974,1998]" pageId="3" pageNumber="137">
<taxonomicNameLabel id="A24685531B438440FD65F811FCF833EC" box="[704,784,1974,1998]" pageId="3" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm.
</paragraph>
</caption>
<paragraph id="8BBEE43A1B448447FF62FF30FC5034DB" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF62FF30FE623493" box="[199,394,151,177]" italics="true" pageId="4" pageNumber="138">Anal appendages</emphasis>
(
<figureCitation id="133AF8BF1B448447FE3CFF30FDF13493" box="[409,537,151,177]" captionStart="FIGURES 10–13" captionStartId="5.[151,264,1645,1669]" captionTargetBox="[157,1427,747,1611]" captionTargetId="figure-265@5.[151,1436,741,1622]" captionTargetPageId="5" captionText="FIGURES 10–13. Appendages Ophiogomphus spp. in lateral and dorsal view. (10, 11), O. phantoani sp. nov., holotype ♂; (12, 13), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679429" httpUri="https://zenodo.org/record/5679429/files/figure.png" pageId="4" pageNumber="138">Figs 10, 11</figureCitation>
). Entirely black; cercus broad basally, narrower apically, gently curving ventroposteriorly, apex hooked with a dorsal small tooth near the base of cercus. Epiproct curved posterodorsally, subequal in length to cerci with a small dorsal tooth approximately at basal third.
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FEA4FB183563" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF62FEA4FE4E353F" box="[199,422,259,285]" italics="true" pageId="4" pageNumber="138">Accessory genitalia</emphasis>
(
<figureCitation id="133AF8BF1B448447FE1DFEA4FDE7353F" box="[440,527,259,285]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="4" pageNumber="138">Fig. 24</figureCitation>
). Anterior hamulus broad medially, abruptly attenuate with apical half strongly curved posteriorly. Posterior hamulus broad basally, narrower and slightly curved cephalad at apex.
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FEECFB0735AB" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF62FEECFE6B3547" box="[199,387,331,357]" italics="true" pageId="4" pageNumber="138">Vesica spermalis</emphasis>
(
<figureCitation id="133AF8BF1B448447FE37FEECFE023547" box="[402,490,331,357]" captionStart="FIGURES 14" captionStartId="6.[151,264,1141,1165]" captionTargetBox="[159,1428,188,1106]" captionTargetId="figure-17@6.[151,1436,181,1117]" captionTargetPageId="6" captionText="FIGURES 14–15. Genital structures of Ophiogomphus phantoani sp. nov. holotype ♂; (14), vesica spermalis, lateral view; (15), apical segment of vesica spermalis, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5679431" httpUri="https://zenodo.org/record/5679431/files/figure.png" pageId="4" pageNumber="138">Figs 14</figureCitation>
, 15, 25). Prepuce protruded ventroposteriorly, its stem slender; posterior lobe of vesica spermalis slender and short. Cornua of glans with two long, acute V-shaped flagellae (
<figureCitation id="133AF8BF1B448447FB2BFEC8FB0A35AB" box="[1166,1250,366,393]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="4" pageNumber="138">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FE34FC36358F" blockId="4.[151,1437,151,1041]" box="[199,990,403,430]" pageId="4" pageNumber="138">Measurements (in mm). Abdomen (including appendages) 40; HW 33.</paragraph>
<paragraph id="8BBEE43A1B448447FF62FE11FDA535F2" blockId="4.[151,1437,151,1041]" box="[199,589,438,465]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF62FE11FDA535F2" bold="true" box="[199,589,438,465]" pageId="4" pageNumber="138">
Description of a
<typeStatus id="54BA5A981B448447FE29FE10FE1F35F3" box="[396,503,439,465]" pageId="4" pageNumber="138" type="paratype">paratype</typeStatus>
female
</emphasis>
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FE7CFB8435D7" blockId="4.[151,1437,151,1041]" box="[199,1132,475,501]" pageId="4" pageNumber="138">
Colour pattern similar to that of
<typeStatus id="54BA5A981B448447FD97FE7CFD7C35D7" box="[562,660,475,501]" pageId="4" pageNumber="138" type="holotype">holotype</typeStatus>
male apart from the following characters:
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FE58FF013643" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF62FE58FEEC363B" box="[199,260,511,537]" italics="true" pageId="4" pageNumber="138">Head</emphasis>
(
<figureCitation id="133AF8BF1B448447FEB0FE58FE88363B" box="[277,352,510,537]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1974,1998]" captionTargetBox="[160,1409,198,1929]" captionTargetId="figure-18@3.[155,1433,181,1950]" captionTargetPageId="3" captionText="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679425" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="4" pageNumber="138">Figs 5</figureCitation>
,
<figureCitation id="133AF8BF1B448447FECBFE58FE74363B" box="[366,412,511,537]" captionStart="FIGURES 6–9" captionStartId="4.[151,264,1976,2000]" captionTargetBox="[151,1436,1065,1953]" captionTargetId="figure-364@4.[151,1436,1065,1953]" captionTargetPageId="4" captionText="FIGURES 6–9. Head of Ophiogomphus phantoani sp. nov. in frontal and dorsal view. (6, 8), Holotype ♂ &amp; (7, 9), Paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.5679427" httpUri="https://zenodo.org/record/5679427/files/figure.png" pageId="4" pageNumber="138">7, 9</figureCitation>
,
<figureCitation id="133AF8BF1B448447FE0FFE58FE20363B" box="[426,456,511,537]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1269,1293]" captionTargetBox="[151,1436,181,1245]" captionTargetId="figure-18@7.[151,1436,181,1245]" captionTargetPageId="7" captionText="FIGURES 16–19. Posterior head and vulva lamina of Ophiogomphus spp., ♀. (16, 18), O. minimus &amp; (17, 19), O. phantoani sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5679433" httpUri="https://zenodo.org/record/5679433/files/figure.png" pageId="4" pageNumber="138">17</figureCitation>
,
<figureCitation id="133AF8BF1B448447FE73FE58FE1E363B" box="[470,502,511,537]" captionStart="FIGURES 28–35" captionStartId="8.[151,264,1738,1762]" captionTargetBox="[152,1436,1138,1714]" captionTargetId="figure-206@8.[152,1436,1138,1714]" captionTargetPageId="8" captionText="FIGURES 28–35. Female occiput [28, 30, 32, 34] &amp; abdominal end [29, 31, 33, 35] in ventral view of Ophiogomphus spp. (28, 29), O. longihamulus (rearranged from Karube 2014: Fig. 16 j, k); (30, 31), O. minimus, Nam Nung Nature Reserve; (32, 33), O. phantoani sp. nov.; (34, 35), O. sinicus (rearranged from Chao 1990: Fig. 10 &amp; Zhang 2019: p. 424)." figureDoi="http://doi.org/10.5281/zenodo.5679437" httpUri="https://zenodo.org/record/5679437/files/figure.png" pageId="4" pageNumber="138">32</figureCitation>
). The greenish yellow markings on dorsal frons much narrower (
<figureCitation id="133AF8BF1B448447FB44FE58FACF363B" box="[1249,1319,511,537]" captionStart="FIGURES 6–9" captionStartId="4.[151,264,1976,2000]" captionTargetBox="[151,1436,1065,1953]" captionTargetId="figure-364@4.[151,1436,1065,1953]" captionTargetPageId="4" captionText="FIGURES 6–9. Head of Ophiogomphus phantoani sp. nov. in frontal and dorsal view. (6, 8), Holotype ♂ &amp; (7, 9), Paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.5679427" httpUri="https://zenodo.org/record/5679427/files/figure.png" pageId="4" pageNumber="138">Fig. 9</figureCitation>
), (
<figureCitation id="133AF8BF1B448447FAE2FE58FA64363B" box="[1351,1420,511,537]" captionStart="FIGURES 6–9" captionStartId="4.[151,264,1976,2000]" captionTargetBox="[151,1436,1065,1953]" captionTargetId="figure-364@4.[151,1436,1065,1953]" captionTargetPageId="4" captionText="FIGURES 6–9. Head of Ophiogomphus phantoani sp. nov. in frontal and dorsal view. (6, 8), Holotype ♂ &amp; (7, 9), Paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.5679427" httpUri="https://zenodo.org/record/5679427/files/figure.png" pageId="4" pageNumber="138">Fig. 8</figureCitation>
); occiput flattened, a projection medially on occipital crest as shown in
<figureCitation id="133AF8BF1B448447FC17FD84FBE5361F" box="[946,1037,547,573]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1269,1293]" captionTargetBox="[151,1436,181,1245]" captionTargetId="figure-18@7.[151,1436,181,1245]" captionTargetPageId="7" captionText="FIGURES 16–19. Posterior head and vulva lamina of Ophiogomphus spp., ♀. (16, 18), O. minimus &amp; (17, 19), O. phantoani sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5679433" httpUri="https://zenodo.org/record/5679433/files/figure.png" pageId="4" pageNumber="138">Figs. 17</figureCitation>
and
<figureCitation id="133AF8BF1B448447FBE3FD84FB8C361F" box="[1094,1124,547,573]" captionStart="FIGURES 28–35" captionStartId="8.[151,264,1738,1762]" captionTargetBox="[152,1436,1138,1714]" captionTargetId="figure-206@8.[152,1436,1138,1714]" captionTargetPageId="8" captionText="FIGURES 28–35. Female occiput [28, 30, 32, 34] &amp; abdominal end [29, 31, 33, 35] in ventral view of Ophiogomphus spp. (28, 29), O. longihamulus (rearranged from Karube 2014: Fig. 16 j, k); (30, 31), O. minimus, Nam Nung Nature Reserve; (32, 33), O. phantoani sp. nov.; (34, 35), O. sinicus (rearranged from Chao 1990: Fig. 10 &amp; Zhang 2019: p. 424)." figureDoi="http://doi.org/10.5281/zenodo.5679437" httpUri="https://zenodo.org/record/5679437/files/figure.png" pageId="4" pageNumber="138">32</figureCitation>
, its dorsal surface minutely serrate.
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FDCCFD2B368B" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="138">
Thorax (
<figureCitation id="133AF8BF1B448447FE8FFDCCFE9836A7" box="[298,368,618,645]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1974,1998]" captionTargetBox="[160,1409,198,1929]" captionTargetId="figure-18@3.[155,1433,181,1950]" captionTargetPageId="3" captionText="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679425" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="4" pageNumber="138">Fig. 5</figureCitation>
). Prothorax entirely black. Synthorax robust, yellow making same as in
<typeStatus id="54BA5A981B448447FB17FDCCFAFC36A7" box="[1202,1300,619,645]" pageId="4" pageNumber="138" type="holotype">holotype</typeStatus>
but lacking small mid-dorsal yellow patch on metepisternum.
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FD14FBF236D3" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF62FD14FEE336EE" box="[199,267,691,716]" italics="true" pageId="4" pageNumber="138">Wings</emphasis>
(
<figureCitation id="133AF8BF1B448447FEBFFD14FEB536EF" box="[282,349,690,717]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1974,1998]" captionTargetBox="[160,1409,198,1929]" captionTargetId="figure-18@3.[155,1433,181,1950]" captionTargetPageId="3" captionText="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679425" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="4" pageNumber="138">Fig. 5</figureCitation>
). Wings hyaline, with dark brown patches at the base; nodal index:Ax/Px = 17/21 (FW) and 12/21 (HW); Pt dark brown,
<quantity id="4CF949DF1B448447FE31FD71FDEE36D3" box="[404,518,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="4" pageNumber="138" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
length in both wings, covering about 5–7 cells.
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FD5CFBAF371B" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF62FD5CFED93737" box="[199,305,763,789]" italics="true" pageId="4" pageNumber="138">Abdomen</emphasis>
(
<figureCitation id="133AF8BF1B448447FEE7FD5CFE603737" box="[322,392,762,789]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1974,1998]" captionTargetBox="[160,1409,198,1929]" captionTargetId="figure-18@3.[155,1433,181,1950]" captionTargetPageId="3" captionText="FIGURES 4–5. Habitus of Ophiogomphus phantoani sp. nov. (4), holotype ♂; (5), paratype ♀. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679425" httpUri="https://zenodo.org/record/5679425/files/figure.png" pageId="4" pageNumber="138">Fig. 5</figureCitation>
). S1 with lateral triangular patch; lateral S2 laterally with a large greenish yellow marking; S3–6 each with a pale basal ring; anterior fourth of S7 yellow; S8–10 entirely black.
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FCE4FC7F37A3" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF62FCE4FE61377F" box="[199,393,835,861]" italics="true" pageId="4" pageNumber="138">Anal appendages</emphasis>
(
<figureCitation id="133AF8BF1B448447FE32FCE4FE05377F" box="[407,493,835,861]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1269,1293]" captionTargetBox="[151,1436,181,1245]" captionTargetId="figure-18@7.[151,1436,181,1245]" captionTargetPageId="7" captionText="FIGURES 16–19. Posterior head and vulva lamina of Ophiogomphus spp., ♀. (16, 18), O. minimus &amp; (17, 19), O. phantoani sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5679433" httpUri="https://zenodo.org/record/5679433/files/figure.png" pageId="4" pageNumber="138">Figs 19</figureCitation>
,
<figureCitation id="133AF8BF1B448447FE5CFCE4FDF0377F" box="[505,536,835,861]" captionStart="FIGURES 28–35" captionStartId="8.[151,264,1738,1762]" captionTargetBox="[152,1436,1138,1714]" captionTargetId="figure-206@8.[152,1436,1138,1714]" captionTargetPageId="8" captionText="FIGURES 28–35. Female occiput [28, 30, 32, 34] &amp; abdominal end [29, 31, 33, 35] in ventral view of Ophiogomphus spp. (28, 29), O. longihamulus (rearranged from Karube 2014: Fig. 16 j, k); (30, 31), O. minimus, Nam Nung Nature Reserve; (32, 33), O. phantoani sp. nov.; (34, 35), O. sinicus (rearranged from Chao 1990: Fig. 10 &amp; Zhang 2019: p. 424)." figureDoi="http://doi.org/10.5281/zenodo.5679437" httpUri="https://zenodo.org/record/5679437/files/figure.png" pageId="4" pageNumber="138">33</figureCitation>
). Cerci as the length of S10, dark yellowish, slender, parallel, pointed at tip.
<taxonomicName id="4C019FB91B448447FACEFCE4FEE637A3" pageId="4" pageNumber="138" rank="variety" variety="lamina">Vulvar lamina</taxonomicName>
triangular-shaped, incised apically but not forming a gap.
</paragraph>
<paragraph id="8BBEE43A1B448447FF62FC2CFBAB3787" blockId="4.[151,1437,151,1041]" box="[199,1091,907,934]" pageId="4" pageNumber="138">
Measurements (in mm). Abdomen (including appendages)
<quantity id="4CF949DF1B448447FCFEFC2CFC453787" box="[859,941,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="138" unit="mm" value="30.0">30 mm</quantity>
. HW
<quantity id="4CF949DF1B448447FC48FC2CFBD73787" box="[1005,1087,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="138" unit="mm" value="48.0">48 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C31BB7B11B448446FF62FC09FF3E368A" lastPageId="5" lastPageNumber="139" pageId="4" pageNumber="138" type="diagnosis">
<paragraph id="8BBEE43A1B448447FF62FC09FB2B3033" blockId="4.[151,1437,151,1041]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF62FC09FE3937EA" bold="true" box="[199,465,942,968]" pageId="4" pageNumber="138">Differential diagnosis.</emphasis>
Among the known species of the genus
<emphasis id="B97538281B448447FC0BFC08FB8837EB" box="[942,1120,943,969]" italics="true" pageId="4" pageNumber="138">
<taxonomicName id="4C019FB91B448447FC0BFC08FBB437EB" authorityName="Selys" authorityYear="1854" box="[942,1116,943,969]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="138" phylum="Arthropoda" rank="genus">Ophiogomphus</taxonomicName>
,
</emphasis>
the new species shares the following characters with
<emphasis id="B97538281B448447FE18FC74FCEF37CE" box="[445,775,979,1005]" italics="true" pageId="4" pageNumber="138">
<taxonomicName id="4C019FB91B448447FE18FC74FD9237CF" authorityName="Karube" authorityYear="2014" box="[445,634,979,1005]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="138" phylum="Arthropoda" rank="species" species="longihamulus">O. longihamulus</taxonomicName>
,
<taxonomicName id="4C019FB91B448447FD20FC74FCEF37CE" authorityName="Karube" authorityYear="2014" box="[645,775,979,1005]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="138" phylum="Arthropoda" rank="species" species="minimus">O. minimus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C019FB91B448447FC98FC74FC4637CE" baseAuthorityName="Chao" baseAuthorityYear="1954" box="[829,942,979,1005]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="138" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="B97538281B448447FC98FC74FC4637CE" box="[829,942,979,1005]" italics="true" pageId="4" pageNumber="138">O. sinicus</emphasis>
</taxonomicName>
, i.e. large pincer-like appendages with cerci curving posteroventrally and epiproct curving posterodorsally, with apex notched (
<figureCitation id="133AF8BF1B448447FB91FC50FB5E3033" box="[1076,1206,1015,1041]" captionStart="FIGURES 10–13" captionStartId="5.[151,264,1645,1669]" captionTargetBox="[157,1427,747,1611]" captionTargetId="figure-265@5.[151,1436,741,1622]" captionTargetPageId="5" captionText="FIGURES 10–13. Appendages Ophiogomphus spp. in lateral and dorsal view. (10, 11), O. phantoani sp. nov., holotype ♂; (12, 13), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679429" httpUri="https://zenodo.org/record/5679429/files/figure.png" pageId="4" pageNumber="138">Figs 10, 12</figureCitation>
).
</paragraph>
<caption id="DF7EB4B21B448447FF32F81FFF5833D6" ID-DOI="http://doi.org/10.5281/zenodo.5679427" ID-Zenodo-Dep="5679427" httpUri="https://zenodo.org/record/5679427/files/figure.png" pageId="4" pageNumber="138" startId="4.[151,264,1976,2000]" targetBox="[151,1436,1065,1953]" targetPageId="4">
<paragraph id="8BBEE43A1B448447FF32F81FFF5833D6" blockId="4.[151,1437,1976,2036]" pageId="4" pageNumber="138">
<emphasis id="B97538281B448447FF32F81FFEDE33F2" bold="true" box="[151,310,1976,2000]" pageId="4" pageNumber="138">FIGURES 6–9</emphasis>
. Head of
<taxonomicName id="4C019FB91B448447FE38F81EFD4E33F3" authority="Ngo &amp; Nguyen, 2021" authorityName="Ngo &amp; Nguyen" authorityYear="2021" box="[413,678,1977,2001]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="138" phylum="Arthropoda" rank="species" species="phantoani" status="sp. nov.">
<emphasis id="B97538281B448447FE38F81EFD4E33F3" box="[413,678,1977,2001]" italics="true" pageId="4" pageNumber="138">Ophiogomphus phantoani</emphasis>
</taxonomicName>
<emphasis id="B97538281B448447FD08F81EFD1733F3" bold="true" box="[685,767,1977,2001]" pageId="4" pageNumber="138">
<taxonomicNameLabel id="A24685531B448447FD08F81EFD1733F3" box="[685,767,1977,2001]" pageId="4" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in frontal and dorsal view. (6, 8), Holotype ♂ &amp; (7, 9), Paratype ♀.
</paragraph>
</caption>
<paragraph id="8BBEE43A1B458446FF62FF30FC863560" blockId="5.[151,1437,151,718]" pageId="5" pageNumber="139">
<taxonomicName id="4C019FB91B458446FF62FF30FE053493" authorityName="Ngo &amp; Nguyen" authorityYear="2021" box="[199,493,151,177]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="phantoani">
<emphasis id="B97538281B458446FF62FF30FE053493" box="[199,493,151,177]" italics="true" pageId="5" pageNumber="139">Ophiogomphus phantoani</emphasis>
</taxonomicName>
can be distinguished from the other congeners by the following characters: In the male, the cornua of glans has two long, acute widely separated, V-shaped flagella (
<figureCitation id="133AF8BF1B458446FB99FF1CFB6734F7" box="[1084,1167,186,213]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="5" pageNumber="139">Fig. 25</figureCitation>
), whereas they are relatively short, U-shaped flagella in
<taxonomicName id="4C019FB91B458446FDA8FF78FD2F34DB" authorityName="Karube" authorityYear="2014" box="[525,711,223,249]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="longihamulus">
<emphasis id="B97538281B458446FDA8FF78FD2F34DB" box="[525,711,223,249]" italics="true" pageId="5" pageNumber="139">O. longihamulus</emphasis>
</taxonomicName>
(
<figureCitation id="133AF8BF1B458446FD72FF78FCC134DB" box="[727,809,223,249]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="5" pageNumber="139">Fig. 21</figureCitation>
),
<taxonomicName id="4C019FB91B458446FC9BFF78FC5734DA" authorityName="Karube" authorityYear="2014" box="[830,959,223,249]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B97538281B458446FC9BFF78FC5734DA" box="[830,959,223,249]" italics="true" pageId="5" pageNumber="139">O. minimus</emphasis>
</taxonomicName>
(
<figureCitation id="133AF8BF1B458446FC6BFF78FBC834DB" box="[974,1056,223,249]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="5" pageNumber="139">Fig. 23</figureCitation>
) and
<taxonomicName id="4C019FB91B458446FBFBFF78FB2634DA" baseAuthorityName="Chao" baseAuthorityYear="1954" box="[1118,1230,223,249]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="B97538281B458446FBFBFF78FB2634DA" box="[1118,1230,223,249]" italics="true" pageId="5" pageNumber="139">O. sinicus</emphasis>
</taxonomicName>
(
<figureCitation id="133AF8BF1B458446FB78FF78FAD934DB" box="[1245,1329,223,249]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="5" pageNumber="139">Fig. 27</figureCitation>
); anterior hamulus of
<taxonomicName id="4C019FB91B458446FEB9FEA4FE58353F" authorityName="Ngo &amp; Nguyen" authorityYear="2021" box="[284,432,259,285]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="phantoani">
<emphasis id="B97538281B458446FEB9FEA4FE58353F" box="[284,432,259,285]" italics="true" pageId="5" pageNumber="139">O. phantoani</emphasis>
</taxonomicName>
with well-developed developed posterior shoulder (
<figureCitation id="133AF8BF1B458446FC5DFEA4FBA2353F" box="[1016,1098,259,285]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="5" pageNumber="139">Fig. 24</figureCitation>
) shoulder smaller in
<taxonomicName id="4C019FB91B458446FA92FEA4FF1E3563" authorityName="Karube" authorityYear="2014" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="longihamulus">
<emphasis id="B97538281B458446FA92FEA4FF1E3563" italics="true" pageId="5" pageNumber="139">O. longihamulus</emphasis>
</taxonomicName>
(
<figureCitation id="133AF8BF1B458446FEA0FE80FEB13563" box="[261,345,295,321]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="5" pageNumber="139">Fig. 20</figureCitation>
),
<taxonomicName id="4C019FB91B458446FEC8FE80FE063562" authorityName="Karube" authorityYear="2014" box="[365,494,295,321]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B97538281B458446FEC8FE80FE063562" box="[365,494,295,321]" italics="true" pageId="5" pageNumber="139">O. minimus</emphasis>
</taxonomicName>
(
<figureCitation id="133AF8BF1B458446FE5BFE80FDA73563" box="[510,591,295,321]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="5" pageNumber="139">Fig. 22</figureCitation>
) and
<taxonomicName id="4C019FB91B458446FD2BFE80FD163562" baseAuthorityName="Chao" baseAuthorityYear="1954" box="[654,766,295,321]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="B97538281B458446FD2BFE80FD163562" box="[654,766,295,321]" italics="true" pageId="5" pageNumber="139">O. sinicus</emphasis>
</taxonomicName>
(
<figureCitation id="133AF8BF1B458446FCA8FE80FC893560" box="[781,865,295,322]" captionStart="FIGURES 20–27" captionStartId="8.[151,264,1017,1041]" captionTargetBox="[152,1436,227,994]" captionTargetId="figure-141@8.[152,1436,227,994]" captionTargetPageId="8" captionText="FIGURES 20–27. Accessory genitalia [20, 22, 24, 26] in lateral view and cornua of glans [21, 23, 25, 27] in ventral view of Ophiogomphus spp. (20–21), O. longihamulus (rearranged from Karube 2014: Fig. 16 e, g); (22, 23), O. minimus (rearranged from Karube 2014: Fig. 18 e, g); (24, 25), O. phantoani sp. nov.; (26, 27), O. sinicus, Cao Bang Province." figureDoi="http://doi.org/10.5281/zenodo.5679435" httpUri="https://zenodo.org/record/5679435/files/figure.png" pageId="5" pageNumber="139">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="8BBEE43A1B458446FF62FEECFCBE35F3" blockId="5.[151,1437,151,718]" pageId="5" pageNumber="139">
In the female, the occiput of
<taxonomicName id="4C019FB91B458446FDA3FEECFD723547" authorityName="Ngo &amp; Nguyen" authorityYear="2021" box="[518,666,331,357]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="phantoani">
<emphasis id="B97538281B458446FDA3FEECFD723547" box="[518,666,331,357]" italics="true" pageId="5" pageNumber="139">O. phantoani</emphasis>
</taxonomicName>
has a projection with two prominent dorsally microserrated branches (
<figureCitation id="133AF8BF1B458446FF05FEC8FF1C35AB" box="[160,244,367,393]" captionStart="FIGURES 28–35" captionStartId="8.[151,264,1738,1762]" captionTargetBox="[152,1436,1138,1714]" captionTargetId="figure-206@8.[152,1436,1138,1714]" captionTargetPageId="8" captionText="FIGURES 28–35. Female occiput [28, 30, 32, 34] &amp; abdominal end [29, 31, 33, 35] in ventral view of Ophiogomphus spp. (28, 29), O. longihamulus (rearranged from Karube 2014: Fig. 16 j, k); (30, 31), O. minimus, Nam Nung Nature Reserve; (32, 33), O. phantoani sp. nov.; (34, 35), O. sinicus (rearranged from Chao 1990: Fig. 10 &amp; Zhang 2019: p. 424)." figureDoi="http://doi.org/10.5281/zenodo.5679437" httpUri="https://zenodo.org/record/5679437/files/figure.png" pageId="5" pageNumber="139">Fig. 32</figureCitation>
), more widely branched than in
<taxonomicName id="4C019FB91B458446FDC4FEC8FCF435AB" authorityName="Karube" authorityYear="2014" box="[609,796,367,393]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="longihamulus">
<emphasis id="B97538281B458446FDC4FEC8FCF435AB" box="[609,796,367,393]" italics="true" pageId="5" pageNumber="139">O. longihamulus</emphasis>
</taxonomicName>
(
<figureCitation id="133AF8BF1B458446FC88FEC8FC6935AB" box="[813,897,367,393]" captionStart="FIGURES 28–35" captionStartId="8.[151,264,1738,1762]" captionTargetBox="[152,1436,1138,1714]" captionTargetId="figure-206@8.[152,1436,1138,1714]" captionTargetPageId="8" captionText="FIGURES 28–35. Female occiput [28, 30, 32, 34] &amp; abdominal end [29, 31, 33, 35] in ventral view of Ophiogomphus spp. (28, 29), O. longihamulus (rearranged from Karube 2014: Fig. 16 j, k); (30, 31), O. minimus, Nam Nung Nature Reserve; (32, 33), O. phantoani sp. nov.; (34, 35), O. sinicus (rearranged from Chao 1990: Fig. 10 &amp; Zhang 2019: p. 424)." figureDoi="http://doi.org/10.5281/zenodo.5679437" httpUri="https://zenodo.org/record/5679437/files/figure.png" pageId="5" pageNumber="139">Fig. 28</figureCitation>
),
<taxonomicName id="4C019FB91B458446FC33FEC8FBFF35AA" authorityName="Karube" authorityYear="2014" box="[918,1047,367,393]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B97538281B458446FC33FEC8FBFF35AA" box="[918,1047,367,393]" italics="true" pageId="5" pageNumber="139">O. minimus</emphasis>
</taxonomicName>
(
<figureCitation id="133AF8BF1B458446FB82FEC8FB9135AB" box="[1063,1145,367,393]" captionStart="FIGURES 28–35" captionStartId="8.[151,264,1738,1762]" captionTargetBox="[152,1436,1138,1714]" captionTargetId="figure-206@8.[152,1436,1138,1714]" captionTargetPageId="8" captionText="FIGURES 28–35. Female occiput [28, 30, 32, 34] &amp; abdominal end [29, 31, 33, 35] in ventral view of Ophiogomphus spp. (28, 29), O. longihamulus (rearranged from Karube 2014: Fig. 16 j, k); (30, 31), O. minimus, Nam Nung Nature Reserve; (32, 33), O. phantoani sp. nov.; (34, 35), O. sinicus (rearranged from Chao 1990: Fig. 10 &amp; Zhang 2019: p. 424)." figureDoi="http://doi.org/10.5281/zenodo.5679437" httpUri="https://zenodo.org/record/5679437/files/figure.png" pageId="5" pageNumber="139">Fig. 30</figureCitation>
) and
<taxonomicName id="4C019FB91B458446FB1CFEC8FAC235AA" baseAuthorityName="Chao" baseAuthorityYear="1954" box="[1209,1322,367,393]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="B97538281B458446FB1CFEC8FAC235AA" box="[1209,1322,367,393]" italics="true" pageId="5" pageNumber="139">O. sinicus</emphasis>
</taxonomicName>
(
<figureCitation id="133AF8BF1B458446FA9FFEC8FA6735AB" box="[1338,1423,367,393]" captionStart="FIGURES 28–35" captionStartId="8.[151,264,1738,1762]" captionTargetBox="[152,1436,1138,1714]" captionTargetId="figure-206@8.[152,1436,1138,1714]" captionTargetPageId="8" captionText="FIGURES 28–35. Female occiput [28, 30, 32, 34] &amp; abdominal end [29, 31, 33, 35] in ventral view of Ophiogomphus spp. (28, 29), O. longihamulus (rearranged from Karube 2014: Fig. 16 j, k); (30, 31), O. minimus, Nam Nung Nature Reserve; (32, 33), O. phantoani sp. nov.; (34, 35), O. sinicus (rearranged from Chao 1990: Fig. 10 &amp; Zhang 2019: p. 424)." figureDoi="http://doi.org/10.5281/zenodo.5679437" httpUri="https://zenodo.org/record/5679437/files/figure.png" pageId="5" pageNumber="139">Fig. 34</figureCitation>
); the vulvar lamina of
<taxonomicName id="4C019FB91B458446FEDAFE34FDFB358F" authorityName="Ngo &amp; Nguyen" authorityYear="2021" box="[383,531,403,429]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="phantoani">
<emphasis id="B97538281B458446FEDAFE34FDFB358F" box="[383,531,403,429]" italics="true" pageId="5" pageNumber="139">O. phantoani</emphasis>
</taxonomicName>
differs from the other species by a pointed tip (
<figureCitation id="133AF8BF1B458446FB87FE34FB9D358F" box="[1058,1141,403,429]" captionStart="FIGURES 28–35" captionStartId="8.[151,264,1738,1762]" captionTargetBox="[152,1436,1138,1714]" captionTargetId="figure-206@8.[152,1436,1138,1714]" captionTargetPageId="8" captionText="FIGURES 28–35. Female occiput [28, 30, 32, 34] &amp; abdominal end [29, 31, 33, 35] in ventral view of Ophiogomphus spp. (28, 29), O. longihamulus (rearranged from Karube 2014: Fig. 16 j, k); (30, 31), O. minimus, Nam Nung Nature Reserve; (32, 33), O. phantoani sp. nov.; (34, 35), O. sinicus (rearranged from Chao 1990: Fig. 10 &amp; Zhang 2019: p. 424)." figureDoi="http://doi.org/10.5281/zenodo.5679437" httpUri="https://zenodo.org/record/5679437/files/figure.png" pageId="5" pageNumber="139">Fig. 33</figureCitation>
), which is incised apically but not widely opened as in the other species (
<figureCitation id="133AF8BF1B458446FD39FE10FCA035F3" box="[668,840,438,465]" captionStart="FIGURES 28–35" captionStartId="8.[151,264,1738,1762]" captionTargetBox="[152,1436,1138,1714]" captionTargetId="figure-206@8.[152,1436,1138,1714]" captionTargetPageId="8" captionText="FIGURES 28–35. Female occiput [28, 30, 32, 34] &amp; abdominal end [29, 31, 33, 35] in ventral view of Ophiogomphus spp. (28, 29), O. longihamulus (rearranged from Karube 2014: Fig. 16 j, k); (30, 31), O. minimus, Nam Nung Nature Reserve; (32, 33), O. phantoani sp. nov.; (34, 35), O. sinicus (rearranged from Chao 1990: Fig. 10 &amp; Zhang 2019: p. 424)." figureDoi="http://doi.org/10.5281/zenodo.5679437" httpUri="https://zenodo.org/record/5679437/files/figure.png" pageId="5" pageNumber="139">Figs 29, 31, 35</figureCitation>
).
</paragraph>
<paragraph id="8BBEE43A1B458446FF62FE7DFF3E368A" blockId="5.[151,1437,151,718]" pageId="5" pageNumber="139">
<emphasis id="B97538281B458446FF62FE7DFE2C35D7" bold="true" box="[199,452,474,501]" pageId="5" pageNumber="139">Habitat and ecology.</emphasis>
<taxonomicName id="4C019FB91B458446FE6AFE7CFD1F35D7" authorityName="Ngo &amp; Nguyen" authorityYear="2021" box="[463,759,475,501]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="phantoani">
<emphasis id="B97538281B458446FE6AFE7CFD1F35D7" box="[463,759,475,501]" italics="true" pageId="5" pageNumber="139">Ophiogomphus phantoani</emphasis>
</taxonomicName>
was found on a small, slowly running, densely vegetated stream (about
<quantity id="4CF949DF1B458446FE93FE58FE95363B" box="[310,381,511,537]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="139" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
wide) within pristine forest. The stream originates from a marsh with muddy bottom. The new species shares the same habitat with
<taxonomicName id="4C019FB91B458446FD93FD84FC2D361F" authority="Phan &amp; To, 2019" authorityName="Phan &amp; To" authorityYear="2019" box="[566,965,547,573]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="schorri">
<emphasis id="B97538281B458446FD93FD84FD12361F" box="[566,762,547,573]" italics="true" pageId="5" pageNumber="139">Coeliccia schorri</emphasis>
Phan &amp; To, 2019
</taxonomicName>
;
<taxonomicName id="4C019FB91B458446FC77FD84FAE2361F" authority="Laidlaw, 1932" authorityName="Laidlaw" authorityYear="1932" box="[978,1290,547,573]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="B97538281B458446FC77FD84FBB6361F" box="[978,1118,547,573]" italics="true" pageId="5" pageNumber="139">C. scutellum</emphasis>
Laidlaw, 1932
</taxonomicName>
;
<taxonomicName id="4C019FB91B458446FAB3FD84FDDD3643" authority="van Tol &amp; Rozeendal, 1995" authorityName="van Tol &amp; Rozeendal" authorityYear="1995" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis id="B97538281B458446FAB3FD84FF063642" italics="true" pageId="5" pageNumber="139">Rhinocypha watsoni</emphasis>
van Tol &amp; Rozeendal, 1995
</taxonomicName>
;
<taxonomicName id="4C019FB91B458446FDE6FDE0FCCE3643" box="[579,806,583,609]" class="Insecta" family="Megapodagrionidae" genus="Burmargiolestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97538281B458446FDE6FDE0FD133643" box="[579,763,583,609]" italics="true" pageId="5" pageNumber="139">Burmargiolestes</emphasis>
sp.
</taxonomicName>
;
<taxonomicName id="4C019FB91B458446FC91FDE0FBA33643" authority="Ris, 1916" authorityName="Ris" authorityYear="1916" box="[820,1099,583,609]" class="Insecta" family="Calopterygidae" genus="Mnais" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="mneme">
<emphasis id="B97538281B458446FC91FDE0FC3A3642" box="[820,978,583,608]" italics="true" pageId="5" pageNumber="139">Mnais mneme</emphasis>
Ris, 1916
</taxonomicName>
;
<taxonomicName id="4C019FB91B458446FBFCFDE0FF3336A7" authority="(Brauer, 1865)" baseAuthorityName="Brauer" baseAuthorityYear="1865" class="Insecta" family="Libellulidae" genus="Orthetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="glaucum">
<emphasis id="B97538281B458446FBFCFDE0FAD03643" box="[1113,1336,583,609]" italics="true" pageId="5" pageNumber="139">Orthetrum glaucum</emphasis>
(Brauer, 1865)
</taxonomicName>
;
<taxonomicName id="4C019FB91B458446FF4CFDCCFDAE36A7" authority="Zhou, 1998" authorityName="Zhou" authorityYear="1998" box="[233,582,619,645]" class="Insecta" family="Cordulegastridae" genus="Anotogaster" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="chaoi">
<emphasis id="B97538281B458446FF4CFDCCFE5136A7" box="[233,441,619,645]" italics="true" pageId="5" pageNumber="139">Anotogaster chaoi</emphasis>
Zhou, 1998
</taxonomicName>
;
<taxonomicName id="4C019FB91B458446FDFCFDCCFBFE36A7" authority="Martin, 1904" authorityName="Martin" authorityYear="1904" box="[601,1046,619,645]" class="Insecta" family="Gomphidae" genus="Lamelligomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="camelus">
<emphasis id="B97538281B458446FDFCFDCCFC9136A7" box="[601,889,619,645]" italics="true" pageId="5" pageNumber="139">Lamelligomphus camelus</emphasis>
Martin, 1904
</taxonomicName>
and
<taxonomicName id="4C019FB91B458446FBF4FDCCFF27368B" authority="Martin, 1904" authorityName="Martin" authorityYear="1904" class="Insecta" family="Gomphidae" genus="Merogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="pavici">
<emphasis id="B97538281B458446FBF4FDCCFAD736A6" box="[1105,1343,619,645]" italics="true" pageId="5" pageNumber="139">Merogomphus pavici</emphasis>
Martin, 1904
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31BB7B11B458446FF62FD15FB9336EC" box="[199,1147,690,718]" pageId="5" pageNumber="139" type="distribution">
<paragraph id="8BBEE43A1B458446FF62FD15FB9336EC" blockId="5.[151,1437,151,718]" box="[199,1147,690,718]" pageId="5" pageNumber="139">
<emphasis id="B97538281B458446FF62FD15FEB136EE" bold="true" box="[199,345,690,716]" pageId="5" pageNumber="139">Distribution</emphasis>
(
<figureCitation id="133AF8BF1B458446FECCFD14FE5236EC" box="[361,442,691,718]" captionStart="FIGURE 36" captionStartId="9.[151,250,1415,1439]" captionTargetBox="[151,1436,181,1391]" captionTargetId="figure-18@9.[151,1436,181,1391]" captionTargetPageId="9" captionText="FIGURE 36. Distribution map of four Ophiogomphus spp. in Vietnam. O. longihamulus (red circle); O. minimus (purple circle); O. phantoani sp. nov. (yellow circle); O. sinicus (green circle). Map background source © GoogleTM Earth Service version 7.3.3.7786." figureDoi="http://doi.org/10.5281/zenodo.5679441" httpUri="https://zenodo.org/record/5679441/files/figure.png" pageId="5" pageNumber="139">Fig. 36</figureCitation>
).
<collectingCountry id="F316A4AA1B458446FE75FD14FDDB36EF" box="[464,563,691,717]" name="Vietnam" pageId="5" pageNumber="139">Vietnam</collectingCountry>
:
<collectingRegion id="49C52AD81B458446FD9BFD14FCD936EF" box="[574,817,691,717]" country="Vietnam" name="Quang Nam" pageId="5" pageNumber="139">Quang Nam Province</collectingRegion>
(Song Thanh National Park).
</paragraph>
</subSubSection>
</treatment>
</document>
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