treatments-xml/data/03/EB/5A/03EB5A00FFEF7D7DAC0DD0C6FC889A6D.xml

<document id="83B5BA474B90436AE90937B80A7A5637" ID-CLB-Dataset="21966" ID-DOI="10.5852/ejt.2016.195" ID-GBIF-Dataset="39071d6f-d900-4c81-ace1-dd03875018c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071140602" checkinUser="valdenar" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2016" docId="03EB5A00FFEF7D7DAC0DD0C6FC889A6D" docLanguage="en" docName="ejt-2016-195.pdf.imf" docOrigin="European Journal of Taxonomy 195" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tylopus acuminatus Likhitrakarn &amp; Golovatch &amp; Panha 2016, sp. nov." docType="treatment" docUuid="C0DB482F-EE58-4385-9733-7D2E724DEFED" docUuidSource="ZooBank" docVersion="8" lastPageNumber="22" masterDocId="FFD22278FFFF7D68AE62D451FFD19824" masterDocTitle="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina" masterLastPageNumber="47" masterPageNumber="1" pageNumber="17" updateTime="1698840626989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4CE1137AAAE7D5459C623F44BA87FDCC">The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina</mods:title>
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<mods:namePart id="B2CB7E70D56A00A6A6308AB03E1E476C">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="0E5063E2D548ED7114EE4A0FAFF58861">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand. &amp; Email: kongerrrr @ hotmail. com &amp; urn: lsid: zoobank. org: author: 442 E 2 E 6 E- 1 CB 3 - 47 B 2 - 9896 - 1 A 922 CE 509 E 1</mods:affiliation>
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<mods:namePart id="D9ACD96D221F47FBB817E37CAF1EDF03">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="4C8480FBCAA55086BB4D0341878D4072">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia. &amp; Corresponding author: sgolovatch @ yandex. ru &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:namePart id="87403F19461537D8FD36C9A96B37A44C">Panha, Somsak</mods:namePart>
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<mods:affiliation id="AD79FF9DF79E79064F360AFD551C2CD6">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Corresponding author: somsak. pan @ chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:date id="08D687477CC80CA0113F4C43F2AEC472">2016</mods:date>
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<paragraph id="8BFDEB16FFEF7D78AC0DD0C6FC149C95" blockId="16.[365,1222,1175,1272]" box="[623,965,1175,1202]" pageId="16" pageNumber="17">
<heading id="D0B55C7AFFEF7D78AC0DD0C6FC149C95" box="[623,965,1175,1202]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="4C429095FFEF7D78AC0DD0C6FCB29C95" ID-CoL="7CZMC" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[623,867,1175,1202]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="B9363704FFEF7D78AC0DD0C6FCB29C95" bold="true" box="[623,867,1175,1202]" italics="true" pageId="16" pageNumber="17">Tylopus acuminatus</emphasis>
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<taxonomicNameLabel id="A2058A7FFFEF7D78AD08D0C6FC149C95" box="[874,965,1175,1201]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act: 
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<subSubSection id="C358B89DFFEF7D78ACA0D08CFCA09CD3" box="[706,881,1245,1272]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BFDEB16FFEF7D78ACA0D08CFCA09CD3" blockId="16.[365,1222,1175,1272]" box="[706,881,1245,1272]" pageId="16" pageNumber="17">
<figureCitation id="1379F793FFEF7D78ACA0D08CFC999CDC" box="[706,840,1245,1272]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="17.[189,232,1892,1918]" captionStartId-1="18.[189,232,1928,1954]" captionStartId-2="19.[187,230,1928,1954]" captionStartId-3="20.[189,232,1856,1882]" captionTargetBox-0="[189,1398,597,1846]" captionTargetBox-1="[308,1241,1126,1855]" captionTargetBox-2="[189,1398,680,1886]" captionTargetBox-3="[189,1398,605,1811]" captionTargetId-0="figure@17.[189,1398,597,1846]" captionTargetId-1="figure@18.[302,1247,1119,1870]" captionTargetId-2="figure@19.[189,1398,680,1886]" captionTargetId-3="figure@20.[189,1398,605,1811]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." captionText-1="Fig. 13. Tylopus acuminatus sp. nov., holotype, Ƌ, right gonopod. A–B. Lateral and mesal views, respectively. C. Leg 13. Scale bar: A–B. 0.5 mm; C = no scale bar. Abbreviations are explained in the text." captionText-2="Fig. 14. Tylopus acuminatus sp. nov., holotype, Ƌ, right gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, sublateral, submesal, subcaudal and suboral views, respectively. Scale bars = 0.2 mm." captionText-3="Fig. 15. Tylopus acuminatus sp. nov., paratype, Ƌ, from Phufa (A–B), paratype, Ƌ, from Phuthalang forest protected area (C–D), paratype, Ƌ, from forest near road (E), paratype, Ƌ, from Ban Na Thong (F), paratype, Ƌ, from Phagneung Phoukulom waterfall (G–H). A–H. Distal part, sublateral, submesal, dorsal, subdorsal, dorsal, subdorsal, ventral and dorsal views, respectively. Scale bars = 0.2 mm. Red colour shows process h. Green colour shows spine z." figureDoi-0="http://doi.org/10.5281/zenodo.3837898" figureDoi-1="http://doi.org/10.5281/zenodo.3837900" figureDoi-2="http://doi.org/10.5281/zenodo.3837902" figureDoi-3="http://doi.org/10.5281/zenodo.3837904" httpUri-0="https://zenodo.org/record/3837898/files/figure.png" httpUri-1="https://zenodo.org/record/3837900/files/figure.png" httpUri-2="https://zenodo.org/record/3837902/files/figure.png" httpUri-3="https://zenodo.org/record/3837904/files/figure.png" pageId="16" pageNumber="17">Figs 12–15</figureCitation>
, 
<figureCitation id="1379F793FFEF7D78AD36D08CFCA09CD3" box="[852,881,1245,1271]" captionStart="Fig" captionStartId="38.[189,232,1821,1847]" captionTargetBox="[266,1310,1040,1771]" captionTargetId="figure@38.[266,1310,1040,1771]" captionTargetPageId="38" captionText="Fig. 26. Distribution of the new eight species of Tylopus. Filled triangle. T. punctus sp. nov. Filled circle. T. acuminatus sp. nov. Crossed circle. T. acuminatus sp. nov., T. dorsalis sp. nov. Crossed square. T. acuminatus sp. nov. T. retusus sp. nov. Open circle. T. retusus sp. nov. Open triangle. T. baenzigeri Golovatch &amp; Enghoff, 1993. Filled square. T. hongkhraiensis sp. nov. Open diamond. T. thunghaihin sp. nov. Inverted filled triangle. T. flavolineatus sp. nov. Open square. T. moniliformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3837927" httpUri="https://zenodo.org/record/3837927/files/figure.png" pageId="16" pageNumber="17">26</figureCitation>
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<paragraph id="8BFDEB16FFEF7D78AEDFD14EFEE79D1D" blockId="16.[189,310,1311,1337]" box="[189,310,1311,1337]" pageId="16" pageNumber="17">
<heading id="D0B55C7AFFEF7D78AEDFD14EFEE79D1D" bold="true" box="[189,310,1311,1337]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B9363704FFEF7D78AEDFD14EFEE79D1D" bold="true" box="[189,310,1311,1337]" pageId="16" pageNumber="17">Diagnosis</emphasis>
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<paragraph id="8BFDEB16FFEF7D78AEDFD11FFED69DF5" blockId="16.[189,1398,1357,1489]" pageId="16" pageNumber="17">
Very similar to 
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<emphasis id="B9363704FFEF7D78AFE1D11FFE239D43" box="[387,498,1357,1383]" italics="true" pageId="16" pageNumber="17">T. jeekeli</emphasis>
<bibRefCitation id="EFD396E7FFEF7D78AF9CD11CFC899D43" author="Golovatch S. I. &amp; Enghoff H." box="[510,856,1357,1383]" pageId="16" pageNumber="17" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C429095FFEF7D78ADF9D11FFDDE9DAF" authority="Likhitrakarn" authorityName="Likhitrakarn" authorityYear="2010" class="Diplopoda" family="Paradoxosomatidae" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="parajeekeli">
<emphasis id="B9363704FFEF7D78ADF9D11FFB939D43" box="[923,1090,1357,1383]" italics="true" pageId="16" pageNumber="17">T. parajeekeli</emphasis>
<bibRefCitation id="EFD396E7FFEF7D78AA2CD11CFDDE9DAF" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Prateepasen R. &amp; Panha S." pageId="16" pageNumber="17" pagination="23 - 68" refId="ref20037" refString="Likhitrakarn N., Golovatch S. I., Prateepasen R. &amp; Panha S. 2010. Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23 - 68. http: // dx. doi. org / 10.3897 / zookeys. 72.744" type="journal article" year="2010">Likhitrakarn, Golovatch, Prateepasen &amp; Panha, 2010</bibRefCitation>
</taxonomicName>
, especially in its gonopod conformation, but differs in process 
<emphasis id="B9363704FFEF7D78AB7BD120FAF89DAF" bold="true" box="[1305,1321,1393,1419]" pageId="16" pageNumber="17">h</emphasis>
being higher and more strongly flattened, pointed, while the sternal lobe between Ƌ coxae 4 linguiform and larger.
</paragraph>
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<paragraph id="8BFDEB16FFEF7D78AEDFD1A8FE959E37" blockId="16.[189,324,1529,1555]" box="[189,324,1529,1555]" pageId="16" pageNumber="17">
<heading id="D0B55C7AFFEF7D78AEDFD1A8FE959E37" bold="true" box="[189,324,1529,1555]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B9363704FFEF7D78AEDFD1A8FE959E37" bold="true" box="[189,324,1529,1555]" pageId="16" pageNumber="17">Etymology</emphasis>
</heading>
</paragraph>
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To emphasize the acuminate processes 
<emphasis id="B9363704FFEF7D78ACEBD276FD489E65" bold="true" box="[649,665,1575,1601]" pageId="16" pageNumber="17">h</emphasis>
on the gonopod; adjective.
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<paragraph id="8BFDEB16FFEF7D78AEDFD239FE7B9EA6" blockId="16.[189,426,1640,1666]" box="[189,426,1640,1666]" pageId="16" pageNumber="17">
<heading id="D0B55C7AFFEF7D78AEDFD239FE7B9EA6" bold="true" box="[189,426,1640,1666]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B9363704FFEF7D78AEDFD239FE7B9EA6" bold="true" box="[189,426,1640,1666]" pageId="16" pageNumber="17">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B2AE14BFFEF7D78AE88D2C6FCD19ED3" ID-GBIF-Occurrence="2621711308" collectingDate="2014-10-15" collectionCode="R, CUMZ" collectorName="C. Sutcharit &amp; Srisonchai" country="Laos" elevation="820" latitude="21.530556" location="Boun Neua" longLatPrecision="21" longitude="101.862495" municipality="Ban Sin Chai" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Phongsali" typeStatus="holotype">
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<typeStatus id="54F955B4FFEF7D78AE88D2C6FE8D9E95" box="[234,348,1687,1713]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
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<collectingCountry id="F355AB86FFEF7D78AEDFD2EBFEDF9EF0" box="[189,270,1722,1748]" name="Laos" pageId="16" pageNumber="17">LAOS</collectingCountry>
: Ƌ, 
<collectingMunicipality id="6B99716CFFEF7D78AF23D2EAFE359EF0" box="[321,484,1722,1749]" pageId="16" pageNumber="17">Ban Sin Chai</collectingMunicipality>
, 
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, 
<collectingRegion id="498625F4FFEF7D78ACEAD2EBFD2E9EF0" box="[648,767,1722,1748]" country="Laos" name="Phongsali" pageId="16" pageNumber="17">Phongsali</collectingRegion>
, 
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, 
<geoCoordinate id="EE768DD1FFEF7D78ADD2D2EBFB809EF1" box="[944,1105,1722,1749]" degrees="101" direction="east" minutes="51" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="45" value="101.862495">101°51'45&quot; E</geoCoordinate>
, ca 
<elevation id="006F0C25FFEF7D78AAE0D2EBFADE9EF0" box="[1154,1295,1722,1749]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="16" pageNumber="17" unit="m" value="820.0">
<quantity id="4CBA46F3FFEF7D78AAE0D2EBFB1F9EF1" box="[1154,1230,1722,1749]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="16" pageNumber="17" unit="m" value="820.0">820 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFFCCDD6FFEF7D78AB7CD2EBFF2A9ED3" pageId="16" pageNumber="17" value="2014-10-15">
<collectingDate id="EFB8343EFFEF7D78AB7CD2EBFF2A9ED3" pageId="16" pageNumber="17" value="2014-10-15">15 Oct. 2014</collectingDate>
</date>
, leg. 
<collectorName id="26B78EC0FFEF7D78AF5BD28CFE149ED3" box="[313,453,1757,1784]" pageId="16" pageNumber="17">C. Sutcharit</collectorName>
&amp; 
<collectionCode id="ED5373D3FFEF7D78AF88D28FFE2F9EDC" box="[490,510,1758,1784]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="17">R</collectionCode>
. 
<collectorName id="26B78EC0FFEF7D78AC6FD28CFD589ED3" box="[525,649,1757,1783]" pageId="16" pageNumber="17">Srisonchai</collectorName>
(
<collectionCode id="ED5373D3FFEF7D78ACF8D28CFD259ED3" box="[666,756,1757,1783]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="16" pageNumber="17">CUMZ</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BFDEB16FFEF7D78AE88D34EFEB69F1D" blockId="16.[189,1399,1823,2026]" box="[234,359,1823,1849]" pageId="16" pageNumber="17">
<emphasis id="B9363704FFEF7D78AE88D34EFEB69F1D" bold="true" box="[234,359,1823,1849]" pageId="16" pageNumber="17">
<typeStatus id="54F955B4FFEF7D78AE88D34EFEB69F1D" box="[234,359,1823,1849]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFDEB16FFEF7D79AEDFD313FD0A994A" blockId="16.[189,1399,1823,2026]" lastBlockId="17.[189,1398,269,366]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<collectingCountry id="F355AB86FFEF7D78AEDFD313FEDF9F78" box="[189,270,1858,1884]" name="Laos" pageId="16" pageNumber="17">LAOS</collectingCountry>
: 1 Ƌ, 
<specimenCount id="9D44209FFFEF7D78AF3ED313FE5B9F79" box="[348,394,1858,1885]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, Phuthalang forest protected area, Boun Neua, 
<collectingRegion id="498625F4FFEF7D78ADAAD313FBEE9F78" box="[968,1087,1858,1884]" country="Laos" name="Phongsali" pageId="16" pageNumber="17">Phongsali</collectingRegion>
, 
<geoCoordinate id="EE768DD1FFEF7D78AA2FD313FB359F79" box="[1101,1252,1858,1885]" degrees="21" direction="north" minutes="37" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="30" value="21.625">21°37'30&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE768DD1FFEF7D78AA91D313FF009FA4" degrees="101" direction="east" minutes="55" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="55" value="101.93194">101°55'55&quot; E</geoCoordinate>
, ca 
<quantity id="4CBA46F3FFEF7D78AF66D334FE8E9FA4" box="[260,351,1893,1920]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="16" pageNumber="17" unit="m" value="1125.0">1125 m</quantity>
a.s.l., 
<date id="FFFCCDD6FFEF7D78AFD2D334FD9F9F5B" box="[432,590,1893,1919]" pageId="16" pageNumber="17" value="2014-10-16">16 Oct. 2014</date>
(
<collectionCode id="ED5373D3FFEF7D78AC02D334FD6A9F5B" box="[608,699,1893,1919]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="16" pageNumber="17">CUMZ</collectionCode>
); 1 Ƌ, 
<specimenCount id="9D44209FFFEF7D78AD70D334FC869FA4" box="[786,855,1893,1920]" pageId="16" pageNumber="17" type="female">2 ♀♀</specimenCount>
, Phufa, Boun Neua, 
<collectingRegion id="498625F4FFEF7D78AA34D334FB1C9F5B" box="[1110,1229,1893,1919]" country="Laos" name="Phongsali" pageId="16" pageNumber="17">Phongsali</collectingRegion>
, 
<geoCoordinate id="EE768DD1FFEF7D78AAB9D334FAA39FA4" box="[1243,1394,1893,1920]" degrees="21" direction="north" minutes="41" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="05" value="21.684721">21°41'05&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE768DD1FFEF7D78AEDFD3D9FE8D9F87" box="[189,348,1928,1955]" degrees="102" direction="east" minutes="06" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="35" value="102.10972">102°06'35&quot; E</geoCoordinate>
, ca 
<quantity id="4CBA46F3FFEF7D78AFE9D3D8FE349F87" box="[395,485,1929,1955]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="16" pageNumber="17" unit="m" value="1470.0">1470 m</quantity>
a.s.l, 
<date id="FFFCCDD6FFEF7D78AC49D3D9FD159F87" box="[555,708,1928,1955]" pageId="16" pageNumber="17" value="2014-10-16">
16 Oct. 
<date id="FFFCCDD6FFEF7D78ACEBD3D8FD159F87" box="[649,708,1929,1955]" pageId="16" pageNumber="17" value="2014" year="2014">2014</date>
</date>
(
<collectionCode id="ED5373D3FFEF7D78ACB6D3D8FCFE9F87" box="[724,815,1929,1955]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="16" pageNumber="17">CUMZ</collectionCode>
); 
<specimenCount id="9D44209FFFEF7D78AD26D3D8FCA09F87" box="[836,881,1929,1955]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, near Ban Borkhun, Boun Neua, 
<collectingRegion id="498625F4FFEF7D78AA99D3D9FAA39F86" box="[1275,1394,1928,1954]" country="Laos" name="Phongsali" pageId="16" pageNumber="17">Phongsali</collectingRegion>
, 
<geoCoordinate id="EE768DD1FFEF7D78AEDFD3FAFE819FE2" box="[189,336,1963,1990]" degrees="21" direction="north" minutes="27" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="57" value="21.465834">21°27'57&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE768DD1FFEF7D78AF39D3FDFE289FE2" box="[347,505,1964,1990]" degrees="101" direction="east" minutes="47" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="20" value="101.78889">101°47'20&quot; E</geoCoordinate>
, ca 
<quantity id="4CBA46F3FFEF7D78AC46D3FDFDBF9FE2" box="[548,622,1964,1990]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="16" pageNumber="17" unit="m" value="880.0">880 m</quantity>
a.s.l., 
<date id="FFFCCDD6FFEF7D78ACDAD3FDFC9F9FE2" box="[696,846,1964,1990]" pageId="16" pageNumber="17" value="2014-10-18">
18 Oct. 
<date id="FFFCCDD6FFEF7D78AD71D3FDFC9F9FE2" box="[787,846,1964,1990]" pageId="16" pageNumber="17" value="2014" year="2014">2014</date>
</date>
(
<collectionCode id="ED5373D3FFEF7D78AD3ED3FDFC669FE2" box="[860,951,1964,1990]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="16" pageNumber="17">CUMZ</collectionCode>
); 
<specimenCount id="9D44209FFFEF7D78ADA9D3FDFC249FE2" box="[971,1013,1964,1990]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, Khoua, 
<collectingRegion id="498625F4FFEF7D78AA3DD3FAFB079FE1" box="[1119,1238,1963,1989]" country="Laos" name="Phongsali" pageId="16" pageNumber="17">Phongsali</collectingRegion>
, 
<geoCoordinate id="EE768DD1FFEF7D78AA82D3FAFAA39FE2" box="[1248,1394,1963,1990]" degrees="21" direction="north" minutes="11" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="55" value="21.19861">21°11'55&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE768DD1FFEF7D78AEDFD39EFE8A9FCD" box="[189,347,1999,2025]" degrees="102" direction="east" minutes="06" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="40" value="102.11111">102°06'40&quot; E</geoCoordinate>
, ca 
<quantity id="4CBA46F3FFEF7D78AFEAD39EFE039FCD" box="[392,466,1999,2025]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="16" pageNumber="17" unit="m" value="840.0">840 m</quantity>
a.s.l., 
<date id="FFFCCDD6FFEF7D78AC7FD39EFD659FCD" box="[541,692,1999,2026]" pageId="16" pageNumber="17" value="2014-10-15">
15 Oct. 
<date id="FFFCCDD6FFEF7D78AC1BD39EFD659FCD" box="[633,692,1999,2025]" pageId="16" pageNumber="17" value="2014" year="2014">2014</date>
</date>
(
<collectionCode id="ED5373D3FFEF7D78ACA1D39EFCCF9FCD" box="[707,798,1999,2025]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="16" pageNumber="17">CUMZ</collectionCode>
); 1 Ƌ, 
<specimenCount id="9D44209FFFEF7D78AD09D39EFC469FCD" box="[875,919,1999,2025]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5373D3FFEF7D78ADC4D39EFBDD9FCD" box="[934,1036,1999,2025]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="17" type="Museum">NHMW</collectionCode>
), 1 Ƌ (
<collectionCode id="ED5373D3FFEF7D78AA3BD39EFB679FCD" box="[1113,1206,1999,2025]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="16" pageNumber="17">ZMUM</collectionCode>
ρ3059), Ban Na Thong, Namo, 
<collectingRegion id="498625F4FFEE7D79AF13D55CFE249903" box="[369,501,269,295]" country="Laos" name="Oudomxai" pageId="17" pageNumber="18">Oudomxay</collectingRegion>
, 
<geoCoordinate id="EE768DD1FFEE7D79AC63D55CFD47990C" box="[513,662,269,296]" degrees="20" direction="north" minutes="52" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="25" value="20.873611">20°52'25&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE768DD1FFEE7D79ACC1D55CFC92990C" box="[675,835,269,296]" degrees="101" direction="east" minutes="47" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="00" value="101.78333">101°47'00&quot; E</geoCoordinate>
, ca 
<quantity id="4CBA46F3FFEE7D79AD11D55CFC6F990C" box="[883,958,269,296]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="17" pageNumber="18" unit="m" value="650.0">650 m</quantity>
a.s.l., 
<date id="FFFCCDD6FFEE7D79AA6FD55CFB7B9903" box="[1037,1194,269,295]" pageId="17" pageNumber="18" value="2014-10-14">
14 Oct. 
<date id="FFFCCDD6FFEE7D79AA0ED55CFB7B9903" box="[1132,1194,269,295]" pageId="17" pageNumber="18" value="2014" year="2014">2014</date>
</date>
; 1 Ƌ, Phagneung Phoukulom Waterfall, Sing, 
<collectingRegion id="498625F4FFEE7D79AC6FD560FD11996E" box="[525,704,304,331]" country="Laos" name="Louangnamtha" pageId="17" pageNumber="18">Luang Namtha</collectingRegion>
, 
<geoCoordinate id="EE768DD1FFEE7D79ACAED561FCB0996F" box="[716,865,304,331]" degrees="21" direction="north" minutes="07" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="15" value="21.120834">21°07'15&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE768DD1FFEE7D79AD0FD560FBDC996F" box="[877,1037,305,331]" degrees="101" direction="east" minutes="14" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="40" value="101.24444">101°14'40&quot; E</geoCoordinate>
, ca 
<quantity id="4CBA46F3FFEE7D79AA5FD560FB59996F" box="[1085,1160,305,331]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="18" unit="m" value="900.0">900 m</quantity>
a.s.l., 
<date id="FFFCCDD6FFEE7D79AAB7D560FAA3996F" box="[1237,1394,305,331]" pageId="17" pageNumber="18" value="2014-10-13">
13 Oct. 
<date id="FFFCCDD6FFEE7D79AB56D560FAA3996F" box="[1332,1394,305,331]" pageId="17" pageNumber="18" value="2014" year="2014">2014</date>
</date>
, all leg. 
<collectionCode id="ED5373D3FFEE7D79AF76D505FEF9994A" box="[276,296,340,366]" country="Denmark" httpUri="http://grbio.org/cool/na0h-6vfh" name="University of Copenhagen" pageId="17" pageNumber="18" type="Herbarium">C</collectionCode>
. Sutcharit &amp; 
<collectionCode id="ED5373D3FFEE7D79AFA4D505FE0B994A" box="[454,474,340,366]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="18">R</collectionCode>
. Srisonchai (
<collectionCode id="ED5373D3FFEE7D79AC11D505FD1B994A" box="[627,714,340,366]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="17" pageNumber="18">ZMUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFEE7D7DAEDFD5C4FAC49991" lastPageId="21" lastPageNumber="22" pageId="17" pageNumber="18" type="description">
<paragraph id="8BFDEB16FFEE7D79AEDFD5C4FE9F998B" blockId="17.[189,334,405,431]" box="[189,334,405,431]" pageId="17" pageNumber="18">
<heading id="D0B55C7AFFEE7D79AEDFD5C4FE9F998B" bold="true" box="[189,334,405,431]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B9363704FFEE7D79AEDFD5C4FE9F998B" bold="true" box="[189,334,405,431]" pageId="17" pageNumber="18">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFEE7D79AEDFD595FAA79A00" blockId="17.[189,1398,451,549]" pageId="17" pageNumber="18">
MEASUREMENTS AND COLOUR. Length 24.5–31.6 (Ƌ) or 
<quantity id="4CBA46F3FFEE7D79AD23D592FC3F99FA" box="[833,1006,451,478]" metricMagnitude="-2" metricUnit="m" metricValue="2.79" metricValueMax="3.15" metricValueMin="2.43" pageId="17" pageNumber="18" unit="mm" value="27.9" valueMax="31.5" valueMin="24.3">24.3–31.5 mm</quantity>
(
<specimenCount id="9D44209FFFEE7D79AA61D595FBC799FA" box="[1027,1046,452,478]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
), width of midbody pro- and metazonae 1.98–2.6 and 
<quantity id="4CBA46F3FFEE7D79AFBDD5B6FDAB9A25" box="[479,634,487,514]" metricMagnitude="-3" metricUnit="m" metricValue="3.275" metricValueMax="3.6" metricValueMin="2.95" pageId="17" pageNumber="18" unit="mm" value="3.275" valueMax="3.6" valueMin="2.95">2.95–3.6 mm</quantity>
(Ƌ) or 2.7–2.9 and 
<quantity id="4CBA46F3FFEE7D79AD3DD5B6FC3D9A25" box="[863,1004,487,513]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="4.2" metricValueMin="3.1" pageId="17" pageNumber="18" unit="mm" value="3.65" valueMax="4.2" valueMin="3.1">3.1–4.2 mm</quantity>
(
<specimenCount id="9D44209FFFEE7D79AD9CD5B6FBC09A25" box="[1022,1041,487,513]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
), respectively. Live coloration dark brown (
<figureCitation id="1379F793FFEE7D79AF3AD65BFE169A00" box="[344,455,522,548]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="17" pageNumber="18">Fig. 12A</figureCitation>
); antennae and epiproct light brown; legs yellow-brown. Coloration of alcohol
</paragraph>
<caption id="DF3DBB9EFFEE7D79AEDFD335FC6E9FCC" ID-DOI="http://doi.org/10.5281/zenodo.3837898" ID-Zenodo-Dep="3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1892,1918]" targetBox="[189,1398,597,1846]" targetPageId="17">
<paragraph id="8BFDEB16FFEE7D79AEDFD335FC6E9FCC" blockId="17.[189,1399,1892,2025]" pageId="17" pageNumber="18">
<emphasis id="B9363704FFEE7D79AEDFD335FECF9F5A" bold="true" box="[189,286,1892,1918]" pageId="17" pageNumber="18">Fig. 12.</emphasis>
<taxonomicName id="4C429095FFEE7D79AF4BD334FDC69F5A" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[297,535,1893,1919]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="B9363704FFEE7D79AF4BD334FDC69F5A" box="[297,535,1893,1919]" italics="true" pageId="17" pageNumber="18">Tylopus acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFEE7D79AC41D334FD549F5B" box="[547,645,1893,1919]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ. 
<emphasis id="B9363704FFEE7D79AD5AD335FC9C9F5A" bold="true" box="[824,845,1892,1918]" pageId="17" pageNumber="18">A</emphasis>
. Habitus, live coloration. 
<emphasis id="B9363704FFEE7D79AAF3D335FB199F5A" bold="true" box="[1169,1224,1892,1918]" pageId="17" pageNumber="18">B–C</emphasis>
. Anterior part of body, dorsal and lateral views, respectively. 
<emphasis id="B9363704FFEE7D79AC9ED3D9FCE29F86" bold="true" box="[764,819,1928,1954]" pageId="17" pageNumber="18">D–E</emphasis>
. Segments 10 and 11, dorsal and lateral views, respectively. 
<emphasis id="B9363704FFEE7D79AF38D3FAFE409FE1" bold="true" box="[346,401,1963,1989]" pageId="17" pageNumber="18">F–H</emphasis>
. Posterior part of body, dorsal, ventral and lateral views, respectively. 
<emphasis id="B9363704FFEE7D79AA8BD3FAFAC39FE1" bold="true" box="[1257,1298,1963,1989]" pageId="17" pageNumber="18">I–J</emphasis>
. Sternal cones between coxae 4, caudal and sublateral views, respectively.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFED7D7AAEDFD55AFB45996D" blockId="18.[189,1399,267,329]" pageId="18" pageNumber="19">
material after one year of preservation faded to brown to yellow-brown; head, antennae, epiproct light brown to yellow-brown, legs and venter yellow-brown to light yellow (
<figureCitation id="1379F793FFED7D7AAA63D57FFB85996D" box="[1025,1108,302,329]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
B–J).
</paragraph>
<paragraph id="8BFDEB16FFED7D7AAEDFD521FB379A1E" blockId="18.[189,1399,368,570]" pageId="18" pageNumber="19">
HEAD. Clypeolabral region densely, vertex sparsely, setose; epicranial suture distinct. Antennae (
<figureCitation id="1379F793FFED7D7AAEA4D5C2FEE09989" box="[198,305,403,429]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="18" pageNumber="19">Fig. 12A</figureCitation>
) short, surpassing body segment 2 (Ƌ, 
<specimenCount id="9D44209FFFED7D7AAC9DD5C5FCC2998A" box="[767,787,404,430]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
) when stretched dorsally. In width, head &lt;segment 3 &lt;collum &lt;segment 4 &lt;2 &lt;5–16 (Ƌ), or head &lt;segment 3 &lt;4 &lt;collum &lt;segment 2 &lt;5–16 (
<specimenCount id="9D44209FFFED7D7AAB33D5E7FAB499F4" box="[1361,1381,438,464]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
); thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4+4 anterior, 1+1 intermediate and 2+2 posterior paraterga declined ventrad, with a small lateral incision at about midway; caudal corner very broadly rounded, not surpassing rear tergal margin (
<figureCitation id="1379F793FFED7D7AAA0FD671FB089A1E" box="[1133,1241,544,570]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="18" pageNumber="19">Fig. 12C</figureCitation>
).
</paragraph>
<paragraph id="8BFDEB16FFED7D7CAEDFD633FDF6996C" blockId="18.[189,1399,609,1058]" lastBlockId="20.[189,1399,267,328]" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
BODY. Tegument smooth and shining, prozonae finely shagreened, metaterga smooth and leathery (
<figureCitation id="1379F793FFED7D7AAEA4D6D4FEA69ABB" box="[198,375,645,671]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="18" pageNumber="19">Fig. 12B, D, F</figureCitation>
); surface below paraterga microgranulate (
<figureCitation id="1379F793FFED7D7AAD10D6D4FBF49ABB" box="[882,1061,645,671]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="18" pageNumber="19">Fig. 12C, E, H</figureCitation>
). Postcollum metaterga with an anterior transverse row of 2+2, mostly abraded setae; posterior row barely traceable as 4+4 insertion points, these being better visible on segments 17–19 (
<figureCitation id="1379F793FFED7D7AAD55D69AFC719AC1" box="[823,928,715,741]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="18" pageNumber="19">Fig. 12F</figureCitation>
). Tergal setae simple and slender, about 2/3 as long as metaterga. Axial line traceable on prozonae and anterior halves of metazonae. Paraterga well-developed (
<figureCitation id="1379F793FFED7D7AAFE3D740FE029B08" box="[385,467,785,812]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
B–F, H), especially so in Ƌ, set rather high (at upper 1/3 of body height), slightly upturned, but lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded, drawn behind tergal margin, increasingly protruding and pointed starting with segment 16 (
<figureCitation id="1379F793FFED7D7AAC35D72AFD309BB2" box="[599,737,891,918]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="18" pageNumber="19">Fig. 12F, H</figureCitation>
). Paraterga 2 with two evident incisions at lateral edge in anterior 1/3. Paraterga 3 and 4 each with two similar incisions at lateral edge, one in anterior 1/3, the other about the middle. Following paraterga each with one evident lateral incision in anterior 1/3 and, in pore-bearing segments, a very small denticle in the middle (
<figureCitation id="1379F793FFED7D7AADCED7B4FBB19BDB" box="[940,1120,997,1023]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="18" pageNumber="19">Fig. 12B, D, F</figureCitation>
). Calluses on paraterga delimited by a sulcus both dorsally and ventrally. Ozopores evident, lateral, lying in an ovoid groove at about 1/3 of metatergal length in front of posterior edge of metaterga. Transverse sulcus evident on metaterga 5–17, incomplete on metaterga 18, narrow, line-shaped, rather deep, ribbed at bottom, not reaching bases of paraterga (
<figureCitation id="1379F793FFEC7D7BAC70D502FD159949" box="[530,708,339,365]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="19" pageNumber="20">Fig. 12B, D, F</figureCitation>
). Stricture between pro- and metazonae evident, wide and deep, ribbed at bottom down to base of paraterga (
<figureCitation id="1379F793FFEC7D7BAD66D527FC6099B4" box="[772,945,374,400]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="19" pageNumber="20">
Fig. 12B, D 
<emphasis id="B9363704FFEC7D7BADECD527FC4C99B4" bold="true" box="[910,925,374,400]" pageId="19" pageNumber="20">–</emphasis>
F
</figureCitation>
). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2 and 3 (Ƌ, 
<specimenCount id="9D44209FFFEC7D7BAD48D5CBFCEF9990" box="[810,830,410,436]" pageId="19" pageNumber="20" type="female">♀</specimenCount>
), increasingly reduced and remaining a strong, sharp, caudal tooth on segments 4–8, retained as a small sharp tooth on segments 9–16, a minute tooth on segment 17 (Ƌ), or an evident, sharp, caudal tooth on segments 4–15, then remaining a small rounded tooth on segment 16, thereafter missing (
<specimenCount id="9D44209FFFEC7D7BACC7D652FD699A39" box="[677,696,515,541]" pageId="19" pageNumber="20" type="female">♀</specimenCount>
) (
<figureCitation id="1379F793FFEC7D7BACB6D652FC569A39" box="[724,903,515,541]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="19" pageNumber="20">Fig. 12C, E, H</figureCitation>
). Epiproct (
<figureCitation id="1379F793FFEC7D7BAA70D652FBB79A39" box="[1042,1126,515,541]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="19" pageNumber="20">Fig. 12</figureCitation>
F–H) conical, flattened dorsoventrally, with two evident, rounded, apical papillae; tip subtruncate; lateral pre-apical papillae evident, lying close to tip. Hypoproct (
<figureCitation id="1379F793FFEB7D7CACF0D55AFCD19901" box="[658,768,267,293]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="20" pageNumber="21">Fig. 12G</figureCitation>
) roundly subtrapeziform, setigerous knobs at caudal edge small and well-separated.
</paragraph>
<caption id="DF3DBB9EFFED7D7AAEDFD3D9FF209FCD" ID-DOI="http://doi.org/10.5281/zenodo.3837900" ID-Zenodo-Dep="3837900" httpUri="https://zenodo.org/record/3837900/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1928,1954]" targetBox="[308,1241,1126,1855]" targetPageId="18">
<paragraph id="8BFDEB16FFED7D7AAEDFD3D9FF209FCD" blockId="18.[189,1399,1928,2025]" pageId="18" pageNumber="19">
<emphasis id="B9363704FFED7D7AAEDFD3D9FECF9F86" bold="true" box="[189,286,1928,1954]" pageId="18" pageNumber="19">Fig. 13.</emphasis>
<taxonomicName id="4C429095FFED7D7AAF48D3D9FDC99F86" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[298,536,1928,1954]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="B9363704FFED7D7AAF48D3D9FDC99F86" box="[298,536,1928,1954]" italics="true" pageId="18" pageNumber="19">Tylopus acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFED7D7AAC47D3D8FD569F87" box="[549,647,1929,1955]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ, right gonopod. 
<emphasis id="B9363704FFED7D7AAD9BD3D9FBE19F86" bold="true" box="[1017,1072,1928,1954]" pageId="18" pageNumber="19">A–B</emphasis>
. Lateral and mesal views, respectively. 
<emphasis id="B9363704FFED7D7AAF3AD3FAFEBC9FE1" bold="true" box="[344,365,1963,1989]" pageId="18" pageNumber="19">C</emphasis>
. Leg 13. Scale bar: A–B. 0.5 mm; C = no scale bar. Abbreviations are explained in the text.
</paragraph>
</caption>
<caption id="DF3DBB9EFFEC7D7BAED9D3D9FEBC9FCD" ID-DOI="http://doi.org/10.5281/zenodo.3837902" ID-Zenodo-Dep="3837902" httpUri="https://zenodo.org/record/3837902/files/figure.png" pageId="19" pageNumber="20" startId="19.[187,230,1928,1954]" targetBox="[189,1398,680,1886]" targetPageId="19">
<paragraph id="8BFDEB16FFEC7D7BAED9D3D9FEBC9FCD" blockId="19.[187,1399,1928,2025]" pageId="19" pageNumber="20">
<emphasis id="B9363704FFEC7D7BAED9D3D9FECC9F86" bold="true" box="[187,285,1928,1954]" pageId="19" pageNumber="20">Fig. 14.</emphasis>
<taxonomicName id="4C429095FFEC7D7BAF48D3D9FDC99F86" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[298,536,1928,1954]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="B9363704FFEC7D7BAF48D3D9FDC99F86" box="[298,536,1928,1954]" italics="true" pageId="19" pageNumber="20">Tylopus acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFEC7D7BAC47D3D8FD599F87" box="[549,648,1929,1955]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ, right gonopod. 
<emphasis id="B9363704FFEC7D7BAD9FD3D9FBE59F86" bold="true" box="[1021,1076,1928,1954]" pageId="19" pageNumber="20">A–B</emphasis>
. Mesal and lateral views, respectively. 
<emphasis id="B9363704FFEC7D7BAF3AD3FAFE5F9FE1" bold="true" box="[344,398,1963,1989]" pageId="19" pageNumber="20">C–F</emphasis>
. Distal part, sublateral, submesal, subcaudal and suboral views, respectively. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFEB7D7CAEDFD521FCB89A33" blockId="20.[189,1399,368,535]" pageId="20" pageNumber="21">
STERNA. Moderately setose and densely microtuberculate; cross-impressing shallow; an entire, linguiform, sternal lobe between Ƌ coxae 4 (
<figureCitation id="1379F793FFEB7D7CACBAD5C2FC84998A" box="[728,853,403,430]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[189,1398,597,1846]" captionTargetId="figure@17.[189,1398,597,1846]" captionTargetPageId="17" captionText="Fig. 12. Tylopus acuminatus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837898" httpUri="https://zenodo.org/record/3837898/files/figure.png" pageId="20" pageNumber="21">
Fig. 
<date id="FFFCCDD6FFEB7D7CAD73D5C2FCE99989" box="[785,824,403,429]" pageId="20" pageNumber="21" value="2012-01">12I</date>
<emphasis id="B9363704FFEB7D7CAD5AD5C2FC969989" bold="true" box="[824,839,403,429]" pageId="20" pageNumber="21">–</emphasis>
J
</figureCitation>
). A paramedian pair of conspicuous ridges in front of gonopod aperture. Legs rather long and slender, midbody ones ca 1.2–1.3 (Ƌ) or 1.0–1.1 (
<specimenCount id="9D44209FFFEB7D7CAB3AD5E7FABA99F4" box="[1368,1387,438,464]" pageId="20" pageNumber="21" type="female">♀</specimenCount>
) as long as body height; Ƌ legs with prefemora distinctly swollen laterally, acropodites with particularly dense setae ventrally (
<figureCitation id="1379F793FFEB7D7CAFA3D5ACFDFB9A33" box="[449,554,509,535]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[308,1241,1126,1855]" captionTargetId="figure@18.[302,1247,1119,1870]" captionTargetPageId="18" captionText="Fig. 13. Tylopus acuminatus sp. nov., holotype, Ƌ, right gonopod. A–B. Lateral and mesal views, respectively. C. Leg 13. Scale bar: A–B. 0.5 mm; C = no scale bar. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837900" httpUri="https://zenodo.org/record/3837900/files/figure.png" pageId="20" pageNumber="21">Fig. 13C</figureCitation>
), but tarsal brushes absent.
</paragraph>
<caption id="DF3DBB9EFFEB7D7CAEDFD311FCF89FCC" ID-DOI="http://doi.org/10.5281/zenodo.3837904" ID-Zenodo-Dep="3837904" httpUri="https://zenodo.org/record/3837904/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1856,1882]" targetBox="[189,1398,605,1811]" targetPageId="20">
<paragraph id="8BFDEB16FFEB7D7CAEDFD311FCF89FCC" blockId="20.[189,1399,1856,2024]" pageId="20" pageNumber="21">
<emphasis id="B9363704FFEB7D7CAEDFD311FECD9F7F" bold="true" box="[189,284,1856,1883]" pageId="20" pageNumber="21">Fig. 15.</emphasis>
<taxonomicName id="4C429095FFEB7D7CAF47D310FDC09F7F" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[293,529,1857,1883]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis id="B9363704FFEB7D7CAF47D310FDC09F7F" box="[293,529,1857,1883]" italics="true" pageId="20" pageNumber="21">Tylopus acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFEB7D7CAC79D310FDAB9F7F" box="[539,634,1857,1883]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, Ƌ, from Phufa (
<emphasis id="B9363704FFEB7D7CADA2D311FC269F7E" bold="true" box="[960,1015,1856,1882]" pageId="20" pageNumber="21">A–B</emphasis>
), paratype, Ƌ, from Phuthalang forest protected area (
<emphasis id="B9363704FFEB7D7CAFA3D335FE2B9F5A" bold="true" box="[449,506,1892,1918]" pageId="20" pageNumber="21">C–D</emphasis>
), paratype, Ƌ, from forest near road (
<emphasis id="B9363704FFEB7D7CADDCD335FC039F5A" bold="true" box="[958,978,1892,1918]" pageId="20" pageNumber="21">E</emphasis>
), paratype, Ƌ, from Ban Na Thong (
<emphasis id="B9363704FFEB7D7CAEA5D3D6FF089F85" bold="true" box="[199,217,1927,1953]" pageId="20" pageNumber="21">F</emphasis>
), paratype, Ƌ, from Phagneung Phoukulom waterfall (
<emphasis id="B9363704FFEB7D7CAD38D3D6FC479F85" bold="true" box="[858,918,1927,1953]" pageId="20" pageNumber="21">G–H</emphasis>
). 
<emphasis id="B9363704FFEB7D7CADCDD3D6FC3B9F85" bold="true" box="[943,1002,1927,1953]" pageId="20" pageNumber="21">A–H</emphasis>
. Distal part, sublateral, submesal, dorsal, subdorsal, dorsal, subdorsal, ventral and dorsal views, respectively. Scale bars = 0.2 mm. Red colour shows process 
<emphasis id="B9363704FFEB7D7CAFDDD39FFE069FCC" bold="true" box="[447,471,1998,2024]" pageId="20" pageNumber="21">h.</emphasis>
Green colour shows spine 
<emphasis id="B9363704FFEB7D7CAD77D39FFCF39FCC" bold="true" box="[789,802,1998,2024]" pageId="20" pageNumber="21">z</emphasis>
.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFEA7D7DAEDFD55FFAC49991" blockId="21.[189,1399,270,437]" pageId="21" pageNumber="22">
GONOPOD. Rather simple (
<figureCitation id="1379F793FFEA7D7DAC6CD55FFD7C990C" box="[526,685,270,296]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[308,1241,1126,1855]" captionTargetId="figure@18.[302,1247,1119,1870]" captionTargetPageId="18" captionText="Fig. 13. Tylopus acuminatus sp. nov., holotype, Ƌ, right gonopod. A–B. Lateral and mesal views, respectively. C. Leg 13. Scale bar: A–B. 0.5 mm; C = no scale bar. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837900" httpUri="https://zenodo.org/record/3837900/files/figure.png" pageId="21" pageNumber="22">
Figs 13A 
<emphasis id="B9363704FFEA7D7DACEAD55FFD46990C" bold="true" box="[648,663,270,296]" pageId="21" pageNumber="22">–</emphasis>
B
</figureCitation>
, 
<figureCitation id="1379F793FFEA7D7DACA7D55FFCC0990D" box="[709,785,270,297]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="19.[187,230,1928,1954]" captionStartId-1="20.[189,232,1856,1882]" captionTargetBox-0="[189,1398,680,1886]" captionTargetBox-1="[189,1398,605,1811]" captionTargetId-0="figure@19.[189,1398,680,1886]" captionTargetId-1="figure@20.[189,1398,605,1811]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="Fig. 14. Tylopus acuminatus sp. nov., holotype, Ƌ, right gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, sublateral, submesal, subcaudal and suboral views, respectively. Scale bars = 0.2 mm." captionText-1="Fig. 15. Tylopus acuminatus sp. nov., paratype, Ƌ, from Phufa (A–B), paratype, Ƌ, from Phuthalang forest protected area (C–D), paratype, Ƌ, from forest near road (E), paratype, Ƌ, from Ban Na Thong (F), paratype, Ƌ, from Phagneung Phoukulom waterfall (G–H). A–H. Distal part, sublateral, submesal, dorsal, subdorsal, dorsal, subdorsal, ventral and dorsal views, respectively. Scale bars = 0.2 mm. Red colour shows process h. Green colour shows spine z." figureDoi-0="http://doi.org/10.5281/zenodo.3837902" figureDoi-1="http://doi.org/10.5281/zenodo.3837904" httpUri-0="https://zenodo.org/record/3837902/files/figure.png" httpUri-1="https://zenodo.org/record/3837904/files/figure.png" pageId="21" pageNumber="22">
14 
<emphasis id="B9363704FFEA7D7DAC80D55FFD20990C" bold="true" box="[738,753,270,296]" pageId="21" pageNumber="22">–</emphasis>
15
</figureCitation>
); coxa slightly curved caudad, sparsely setose distoventrally. Femorite slightly curved and stout, with an evident mesal groove and a clear distolateral sulcus demarcating a postfemoral part; process 
<emphasis id="B9363704FFEA7D7DAC9AD505FCD9994A" bold="true" box="[760,776,340,366]" pageId="21" pageNumber="22">h</emphasis>
prominent, evidently curved, pointed, with a small, but evident dorsal tooth and sometimes with a small ventral denticle (
<figureCitation id="1379F793FFEA7D7DAA6FD526FB4F99B6" box="[1037,1182,375,402]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[308,1241,1126,1855]" captionTargetId="figure@18.[302,1247,1119,1870]" captionTargetPageId="18" captionText="Fig. 13. Tylopus acuminatus sp. nov., holotype, Ƌ, right gonopod. A–B. Lateral and mesal views, respectively. C. Leg 13. Scale bar: A–B. 0.5 mm; C = no scale bar. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837900" httpUri="https://zenodo.org/record/3837900/files/figure.png" pageId="21" pageNumber="22">
Fig. 13A 
<emphasis id="B9363704FFEA7D7DAA1BD529FB5999B6" bold="true" box="[1145,1160,376,402]" pageId="21" pageNumber="22">–</emphasis>
B
</figureCitation>
); spine 
<emphasis id="B9363704FFEA7D7DAB62D529FADC99B6" bold="true" box="[1280,1293,376,402]" pageId="21" pageNumber="22">z</emphasis>
pointed, sometimes small (
<figureCitation id="1379F793FFEA7D7DAFF3D5CAFE2F9991" box="[401,510,410,437]" captionStart="Fig" captionStartId="20.[189,232,1856,1882]" captionTargetBox="[189,1398,605,1811]" captionTargetId="figure@20.[189,1398,605,1811]" captionTargetPageId="20" captionText="Fig. 15. Tylopus acuminatus sp. nov., paratype, Ƌ, from Phufa (A–B), paratype, Ƌ, from Phuthalang forest protected area (C–D), paratype, Ƌ, from forest near road (E), paratype, Ƌ, from Ban Na Thong (F), paratype, Ƌ, from Phagneung Phoukulom waterfall (G–H). A–H. Distal part, sublateral, submesal, dorsal, subdorsal, dorsal, subdorsal, ventral and dorsal views, respectively. Scale bars = 0.2 mm. Red colour shows process h. Green colour shows spine z." figureDoi="http://doi.org/10.5281/zenodo.3837904" httpUri="https://zenodo.org/record/3837904/files/figure.png" pageId="21" pageNumber="22">Fig. 15G</figureCitation>
); solenophore (
<emphasis id="B9363704FFEA7D7DACD6D5CAFD319991" bold="true" box="[692,736,411,437]" pageId="21" pageNumber="22">sph</emphasis>
) large, expanded, flattened and typically coiled.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFEA7D7DAEDFD58DFC889A6D" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8BFDEB16FFEA7D7DAEDFD58DFEF599D2" blockId="21.[189,292,476,502]" box="[189,292,476,502]" pageId="21" pageNumber="22">
<heading id="D0B55C7AFFEA7D7DAEDFD58DFEF599D2" bold="true" box="[189,292,476,502]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B9363704FFEA7D7DAEDFD58DFEF599D2" bold="true" box="[189,292,476,502]" pageId="21" pageNumber="22">Remark</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFEA7D7DAEDFD65AFC889A6D" blockId="21.[189,1398,523,585]" pageId="21" pageNumber="22">
This species shows slight variations in gonopod structure, process 
<emphasis id="B9363704FFEA7D7DADDCD65AFC1F9A01" bold="true" box="[958,974,523,549]" pageId="21" pageNumber="22">h</emphasis>
often being rather narrow to slightly broader, while spine 
<emphasis id="B9363704FFEA7D7DAFD0D67FFE6E9A6C" bold="true" box="[434,447,558,584]" pageId="21" pageNumber="22">z</emphasis>
small to slightly broader (
<figureCitation id="1379F793FFEA7D7DAC96D67FFC9A9A6D" box="[756,843,558,585]" captionStart="Fig" captionStartId="20.[189,232,1856,1882]" captionTargetBox="[189,1398,605,1811]" captionTargetId="figure@20.[189,1398,605,1811]" captionTargetPageId="20" captionText="Fig. 15. Tylopus acuminatus sp. nov., paratype, Ƌ, from Phufa (A–B), paratype, Ƌ, from Phuthalang forest protected area (C–D), paratype, Ƌ, from forest near road (E), paratype, Ƌ, from Ban Na Thong (F), paratype, Ƌ, from Phagneung Phoukulom waterfall (G–H). A–H. Distal part, sublateral, submesal, dorsal, subdorsal, dorsal, subdorsal, ventral and dorsal views, respectively. Scale bars = 0.2 mm. Red colour shows process h. Green colour shows spine z." figureDoi="http://doi.org/10.5281/zenodo.3837904" httpUri="https://zenodo.org/record/3837904/files/figure.png" pageId="21" pageNumber="22">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>