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<document id="9562A1A22EFEFB0A47992079B3C707E4" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E3018C3AA2EDFF58FE9BFE2C" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum bolavensis Wesener 2019, new species" docType="treatment" docVersion="9" lastPageNumber="215" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="212" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:title id="DC0D7FA4234F4DC346EB8BB405B8C9E9">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:namePart id="D6A3B403C679C1FB614CAECB438D201A">Wesener, Thomas</mods:namePart>
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<mods:title id="B5930E1A14792D62D28C09F87C070751">Zootaxa</mods:title>
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<mods:date id="62634519A506FCD641614F92C1452712">2019</mods:date>
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<treatment id="03EB5506E3018C3AA2EDFF58FE9BFE2C" ID-DOI="http://doi.org/10.5281/zenodo.5934522" ID-GBIF-Taxon="154474558" ID-Zenodo-Dep="5934522" LSID="urn:lsid:plazi:treatment:03EB5506E3018C3AA2EDFF58FE9BFE2C" httpUri="http://treatment.plazi.org/id/03EB5506E3018C3AA2EDFF58FE9BFE2C" lastPageId="14" lastPageNumber="215" pageId="11" pageNumber="212">
<subSubSection id="C358B79BE3018C3FA2EDFF58FD8AFF4E" box="[151,611,151,178]" pageId="11" pageNumber="212" type="nomenclature">
<paragraph id="8BFDE410E3018C3FA2EDFF58FD8AFF4E" blockId="11.[151,611,151,211]" box="[151,611,151,178]" pageId="11" pageNumber="212">
<heading id="D0B5537CE3018C3FA2EDFF58FD8AFF4E" bold="true" box="[151,611,151,178]" fontSize="11" level="1" pageId="11" pageNumber="212" reason="1">
<emphasis id="B9363802E3018C3FA2EDFF58FD8AFF4E" bold="true" box="[151,611,151,178]" pageId="11" pageNumber="212">
<taxonomicName id="4C429F93E3018C3FA2EDFF58FE22FF4D" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,459,151,177]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3018C3FA2EDFF58FE22FF4D" bold="true" box="[151,459,151,177]" italics="true" pageId="11" pageNumber="212">Sphaerobelum bolavensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058579E3018C3FA3A9FF57FD8AFF4E" box="[467,611,152,178]" pageId="11" pageNumber="212" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE3018C3FA2EDFF75FE44FF2F" box="[151,429,186,211]" pageId="11" pageNumber="212" type="description">
<paragraph id="8BFDE410E3018C3FA2EDFF75FE44FF2F" blockId="11.[151,611,151,211]" box="[151,429,186,211]" pageId="11" pageNumber="212">
<figureCitation id="1379F895E3018C3FA2EDFF75FEFFFF2F" box="[151,278,186,211]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="11" pageNumber="212">Figures 6A</figureCitation>
,
<figureCitation id="1379F895E3018C3FA358FF74FEDBFF2E" box="[290,306,187,210]" captionStart="FIGURE 7" captionStartId="12.[151,250,712,734]" captionTargetBox="[150,1436,193,690]" captionTargetId="figure@12.[150,1437,193,691]" captionTargetPageId="12" captionText="FIGURE 7. Sphaerobelum bolavensis n. sp., holotype (ZMUC), left antenna, SEM, A: lateral view; B: apical disc with sensory cones. Abbreviations: d = apical disc; sb = sensilla basiconica; sc = sensory cones; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601086/files/figure.png" pageId="11" pageNumber="212">7</figureCitation>
,
<figureCitation id="1379F895E3018C3FA344FF74FEA4FF2F" box="[318,333,187,211]" captionStart="FIGURE 8" captionStartId="12.[151,250,1985,2007]" captionTargetBox="[151,1435,812,1976]" captionTargetId="figure@12.[151,1435,812,1976]" captionTargetPageId="12" captionText="FIGURE 8. Sphaerobelum bolavensis n. sp., female paratype (ZFMK MYR8100); A: 9th left leg; B: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur. Not to scale." httpUri="https://zenodo.org/record/2601088/files/figure.png" pageId="11" pageNumber="212">8</figureCitation>
,
<figureCitation id="1379F895E3018C3FA320FF74FE96FF2F" box="[346,383,187,211]" captionStart="FIGURE 9" captionStartId="13.[151,250,1423,1445]" captionTargetBox="[152,1434,1021,1400]" captionTargetId="figure@13.[152,1435,1021,1402]" captionTargetPageId="13" captionText="FIGURE 9. Endoterga of the body ring 9 of different Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum spinatum n. sp.; D: Sphaerobelum splendidum n. sp. Abbreviations: cp = cuticular patterns; iA = inner area." httpUri="https://zenodo.org/record/2601090/files/figure.png" pageId="11" pageNumber="212">9A</figureCitation>
,
<figureCitation id="1379F895E3018C3FA3F0FF74FE40FF2F" box="[394,425,187,211]" captionStart="FIGURE 10" captionStartId="16.[151,250,1946,1968]" captionTargetBox="[177,1416,193,1934]" captionTargetId="figure@16.[177,1416,193,1934]" captionTargetPageId="16" captionText="FIGURE 10. Sphaerobelum bolavensis n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601094/files/figure.png" pageId="11" pageNumber="212">10</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE3018C3FA2EDFECDFEB1FE33" pageId="11" pageNumber="212" type="materials_examined">
<paragraph id="8BFDE410E3018C3FA2EDFECDFC35FE9F" blockId="11.[151,1436,258,931]" pageId="11" pageNumber="212">
<emphasis id="B9363802E3018C3FA2EDFECDFE6BFEE7" bold="true" box="[151,386,258,283]" pageId="11" pageNumber="212">Material examined:</emphasis>
Type specimens. 1 M
<typeStatus id="54F95AB2E3018C3FA004FECDFD09FEE7" box="[638,736,258,283]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
(ZMUC00040256) from
<collectingCountry id="F355A480E3018C3FA185FECCFBD1FEE6" box="[1023,1080,259,282]" name="Laos" pageId="11" pageNumber="212">Laos</collectingCountry>
,
<collectingRegion id="49862AF2E3018C3FA639FECDFAD8FEE7" box="[1091,1329,258,283]" country="Laos" name="Champasak" pageId="11" pageNumber="212">Champasak Province</collectingRegion>
, Bolaven Plateau, Ban Houayteuay,
<quantity id="4CBA49F5E3018C3FA3B2FEE8FDC8FEC2" box="[456,545,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="212" unit="m" value="1200.0">1200 m</quantity>
(
<geoCoordinate id="EE7682D7E3018C3FA049FEE8FD28FEC3" box="[563,705,294,319]" direction="north" orientation="latitude" pageId="11" pageNumber="212" precision="1" value="15.077583">N 15°4.655'</geoCoordinate>
,
<geoCoordinate id="EE7682D7E3018C3FA0B5FEE8FC9DFEC3" box="[719,884,294,319]" direction="east" orientation="longitude" pageId="11" pageNumber="212" precision="1" value="106.2808">E 106°16.848'</geoCoordinate>
) disturbed montane forest, leg.
<date id="FFFCC2D0E3018C3FA69CFEE8FA9BFEC3" box="[1254,1394,295,319]" pageId="11" pageNumber="212" value="2007-12-10">10.XII.2007</date>
, by hand, S. Tarasov. 1 F
<typeStatus id="54F95AB2E3018C3FA3F3FE84FE00FE9F" box="[393,489,331,355]" pageId="11" pageNumber="212" type="paratype">paratype</typeStatus>
(ZFMK MYR8100), same data as
<typeStatus id="54F95AB2E3018C3FA108FE85FC31FE9F" box="[882,984,330,355]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BFDE410E3018C3FA2BDFEA1FEB1FE33" blockId="11.[151,1436,258,931]" pageId="11" pageNumber="212">
<emphasis id="B9363802E3018C3FA2BDFEA1FE42FE7B" bold="true" box="[199,427,366,391]" pageId="11" pageNumber="212">Non-type material:</emphasis>
1 F (MHNG LT-10/24) from
<collectingCountry id="F355A480E3018C3FA08CFEA0FCC6FE7A" box="[758,815,367,390]" name="Laos" pageId="11" pageNumber="212">Laos</collectingCountry>
,
<collectingRegion id="49862AF2E3018C3FA141FEA1FBC3FE7B" box="[827,1066,366,391]" country="Laos" name="Champasak" pageId="11" pageNumber="212">Champasak Province</collectingRegion>
, Bolaven Plateau,
<quantity id="4CBA49F5E3018C3FA680FEA0FADAFE7B" box="[1274,1331,366,391]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="212" unit="km" value="3.0">3 km</quantity>
S of Ban Nong Luang, Tad Kameud,
<quantity id="4CBA49F5E3018C3FA3ABFE5CFDCEFE56" box="[465,551,403,427]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="11" pageNumber="212" unit="m" value="1270.0">1270 m</quantity>
(
<geoCoordinate id="EE7682D7E3018C3FA04CFE5CFD2FFE57" box="[566,710,402,427]" direction="north" orientation="latitude" pageId="11" pageNumber="212" precision="15" value="15.073055">N 15°04'23&quot;</geoCoordinate>
,
<geoCoordinate id="EE7682D7E3018C3FA0ABFE5DFC82FE57" box="[721,875,402,427]" direction="east" orientation="longitude" pageId="11" pageNumber="212" precision="15" value="106.21">106°12'36&quot; E</geoCoordinate>
), undisturbed evergreen hill forest,
<date id="FFFCC2D0E3018C3FA686FE5CFA8EFE57" box="[1276,1383,403,427]" pageId="11" pageNumber="212" value="2010-10-01">1.X.2010</date>
, leg. P. Schwendinger.
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE3018C3FA2BDFE15FB7DFD36" pageId="11" pageNumber="212" type="diagnosis">
<paragraph id="8BFDE410E3018C3FA2BDFE15FB7DFD36" blockId="11.[151,1436,258,931]" pageId="11" pageNumber="212">
<emphasis id="B9363802E3018C3FA2BDFE15FED3FE0F" bold="true" box="[199,314,474,499]" pageId="11" pageNumber="212">Diagnosis</emphasis>
:
<taxonomicName id="4C429F93E3018C3FA330FE14FE35FE0F" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[330,476,475,499]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3018C3FA330FE14FE35FE0F" box="[330,476,475,499]" italics="true" pageId="11" pageNumber="212">S. bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA39EFE15FDCDFE0F" bold="true" box="[484,548,474,499]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA39EFE15FDCDFE0F" box="[484,548,474,499]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
belongs to a group of
<taxonomicName id="4C429F93E3018C3FA15FFE14FC2FFE0F" authorityName="Verhoeff" authorityYear="1924" box="[805,966,475,499]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B9363802E3018C3FA15FFE14FC2FFE0F" box="[805,966,475,499]" italics="true" pageId="11" pageNumber="212">Sphaerobelum</emphasis>
</taxonomicName>
species in which the mesal margin of the femur is extended and apically tipped with several teeth (
<figureCitation id="1379F895E3018C3FA167FE31FC91FDEB" box="[797,888,510,535]" captionStart="FIGURE 8" captionStartId="12.[151,250,1985,2007]" captionTargetBox="[151,1435,812,1976]" captionTargetId="figure@12.[151,1435,812,1976]" captionTargetPageId="12" captionText="FIGURE 8. Sphaerobelum bolavensis n. sp., female paratype (ZFMK MYR8100); A: 9th left leg; B: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur. Not to scale." httpUri="https://zenodo.org/record/2601088/files/figure.png" pageId="11" pageNumber="212">Fig. 8A</figureCitation>
).
<taxonomicName id="4C429F93E3018C3FA1F6FE31FB42FDEA" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[908,1195,510,534]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3018C3FA1F6FE31FB42FDEA" box="[908,1195,510,534]" italics="true" pageId="11" pageNumber="212">Sphaerobelum bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA6C8FE31FB1BFDEB" bold="true" box="[1202,1266,510,535]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA6C8FE31FB1BFDEB" box="[1202,1266,510,535]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C429F93E3018C3FA750FE30FA72FDEA" authorityName="Attems" authorityYear="1938" box="[1322,1435,510,534]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="B9363802E3018C3FA750FE30FA72FDEA" box="[1322,1435,510,534]" italics="true" pageId="11" pageNumber="212">S. bicorne</emphasis>
</taxonomicName>
from
<collectingCountry id="F355A480E3018C3FA2AEFDEDFEDAFDC7" box="[212,307,546,571]" name="Vietnam" pageId="11" pageNumber="212">Vietnam</collectingCountry>
are the only species with an extension only in the apical part of the femur.
<taxonomicName id="4C429F93E3018C3FA604FDECFA75FDC7" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1150,1436,547,571]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3018C3FA604FDECFA75FDC7" box="[1150,1436,547,571]" italics="true" pageId="11" pageNumber="212">Sphaerobelum bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA2EDFD89FF30FDA3" bold="true" box="[151,217,582,607]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA2EDFD89FF30FDA3" box="[151,217,582,607]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
possesses a female vulval operculum only projecting in a single mesal process (
<figureCitation id="1379F895E3018C3FA6F3FD89FB0DFDA3" box="[1161,1252,582,607]" captionStart="FIGURE 8" captionStartId="12.[151,250,1985,2007]" captionTargetBox="[151,1435,812,1976]" captionTargetId="figure@12.[151,1435,812,1976]" captionTargetPageId="12" captionText="FIGURE 8. Sphaerobelum bolavensis n. sp., female paratype (ZFMK MYR8100); A: 9th left leg; B: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur. Not to scale." httpUri="https://zenodo.org/record/2601088/files/figure.png" pageId="11" pageNumber="212">Fig. 8B</figureCitation>
); in contrast,
<taxonomicName id="4C429F93E3018C3FA7FDFD88FF05FD7F" authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="B9363802E3018C3FA7FDFD88FF05FD7F" italics="true" pageId="11" pageNumber="212">S. bicorne</emphasis>
</taxonomicName>
has two processes.
<taxonomicName id="4C429F93E3018C3FA3B2FDA4FD0FFD7F" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[456,742,619,643]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3018C3FA3B2FDA4FD0FFD7F" box="[456,742,619,643]" italics="true" pageId="11" pageNumber="212">Sphaerobelum bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA096FDA5FCC2FD7F" bold="true" box="[748,811,618,643]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA096FDA5FCC2FD7F" box="[748,811,618,643]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has a telopoditomere 4 of the posterior telopod straight, apically without a recessed hook (
<figureCitation id="1379F895E3018C3FA062FD41FD48FD5B" box="[536,673,654,679]" captionStart="FIGURE 10" captionStartId="16.[151,250,1946,1968]" captionTargetBox="[177,1416,193,1934]" captionTargetId="figure@16.[177,1416,193,1934]" captionTargetPageId="16" captionText="FIGURE 10. Sphaerobelum bolavensis n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601094/files/figure.png" pageId="11" pageNumber="212">Figs 10E, F</figureCitation>
); in contrast
<taxonomicName id="4C429F93E3018C3FA14EFD40FC4EFD5A" authorityName="Attems" authorityYear="1938" box="[820,935,654,678]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="B9363802E3018C3FA14EFD40FC4EFD5A" box="[820,935,654,678]" italics="true" pageId="11" pageNumber="212">S. bicorne</emphasis>
</taxonomicName>
possesses a hook.
<taxonomicName id="4C429F93E3018C3FA607FD41FA75FD5A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1149,1436,654,678]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3018C3FA607FD41FA75FD5A" box="[1149,1436,654,678]" italics="true" pageId="11" pageNumber="212">Sphaerobelum bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA2EDFD7DFF3FFD37" bold="true" box="[151,214,690,715]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA2EDFD7DFF3FFD37" box="[151,214,690,715]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has a short locking carina on the anal shield; whereas the carina is long in
<taxonomicName id="4C429F93E3018C3FA666FD7CFB64FD37" authorityName="Attems" authorityYear="1938" box="[1052,1165,691,715]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="B9363802E3018C3FA666FD7CFB64FD37" box="[1052,1165,691,715]" italics="true" pageId="11" pageNumber="212">S. bicorne</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE3018C3AA2BDFD19FEA9FEC3" lastPageId="14" lastPageNumber="215" pageId="11" pageNumber="212" type="description">
<paragraph id="8BFDE410E3018C3FA2BDFD19FCB6FD13" blockId="11.[151,1436,258,931]" box="[199,863,726,751]" pageId="11" pageNumber="212">
<emphasis id="B9363802E3018C3FA2BDFD19FEB0FD13" bold="true" box="[199,345,726,751]" pageId="11" pageNumber="212">Description.</emphasis>
Based on
<typeStatus id="54F95AB2E3018C3FA3B4FD19FDD9FD13" box="[462,560,726,751]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
male and
<typeStatus id="54F95AB2E3018C3FA0DEFD18FCEDFD13" box="[676,772,727,751]" pageId="11" pageNumber="212" type="paratype">paratype</typeStatus>
female.
</paragraph>
<paragraph id="8BFDE410E3018C3FA2BDFD35FB74FC5F" blockId="11.[151,1436,258,931]" pageId="11" pageNumber="212">
<emphasis id="B9363802E3018C3FA2BDFD35FE9FFCEF" bold="true" box="[199,374,762,787]" pageId="11" pageNumber="212">Measurements</emphasis>
:
<typeStatus id="54F95AB2E3018C3FA3FFFD35FE04FCEF" box="[389,493,762,787]" pageId="11" pageNumber="212" type="holotype">Holotype</typeStatus>
male. Body length ca
<quantity id="4CBA49F5E3018C3FA097FD34FCBCFCEE" box="[749,853,763,787]" metricMagnitude="-2" metricUnit="m" metricValue="3.29" pageId="11" pageNumber="212" unit="mm" value="32.9">32.9 mm</quantity>
. Width, of thoracic shield =
<quantity id="4CBA49F5E3018C3FA6ECFD34FB14FCEE" box="[1174,1277,763,787]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="11" pageNumber="212" unit="mm" value="14.2">14.2 mm</quantity>
, of tergite 8 =
<quantity id="4CBA49F5E3018C3FA2EDFCD0FF12FCCA" box="[151,251,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="11" pageNumber="212" unit="mm" value="14.9">14.9 mm</quantity>
(= broadest). Height, of thoracic shield =
<quantity id="4CBA49F5E3018C3FA0A9FCD0FCC5FCCA" box="[723,812,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="11" pageNumber="212" unit="mm" value="7.1">7.1 mm</quantity>
, of tergite 7 =
<quantity id="4CBA49F5E3018C3FA1AAFCD0FBCFFCCA" box="[976,1062,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="212" unit="mm" value="8.2">8.2 mm</quantity>
(= highest). Female: body length = ca
<quantity id="4CBA49F5E3018C3FA2B4FC8CFEDFFCA6" box="[206,310,835,859]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="11" pageNumber="212" unit="mm" value="28.8">28.8 mm</quantity>
. Width, of thoracic shield =
<quantity id="4CBA49F5E3018C3FA00CFC8DFD34FCA6" box="[630,733,834,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="11" pageNumber="212" unit="mm" value="12.6">12.6 mm</quantity>
, of tergite 8 =
<quantity id="4CBA49F5E3018C3FA1F9FC8DFC0EFCA6" box="[899,999,834,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="11" pageNumber="212" unit="mm" value="13.6">13.6 mm</quantity>
(=broadest). Height, of thoracic shield =
<quantity id="4CBA49F5E3018C3FA2CAFCA9FEE2FC82" box="[176,267,870,895]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="11" pageNumber="212" unit="mm" value="7.6">7.6 mm</quantity>
, of tergite 7,
<quantity id="4CBA49F5E3018C3FA3D3FCA8FDE8FC82" box="[425,513,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="11" pageNumber="212" unit="mm" value="8.3">8.3 mm</quantity>
(= highest).
<emphasis id="B9363802E3018C3FA0E2FCA9FCF0FC83" bold="true" box="[664,793,870,895]" pageId="11" pageNumber="212">Coloration</emphasis>
: in preserved specimens dark green, tergites with black posterior margin. Head and collum chestnut brown. Antennae orange, legs brown (
<figureCitation id="1379F895E3018C3FA64CFC45FB79FC5F" box="[1078,1168,906,931]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="11" pageNumber="212">Fig. 6A</figureCitation>
).
</paragraph>
<caption id="DF3DB498E3018C3FA2EDF8B3FDA5F811" httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="11" pageNumber="212" startId="11.[151,250,1916,1938]" targetBox="[150,1423,987,1897]" targetPageId="11">
<paragraph id="8BFDE410E3018C3FA2EDF8B3FDA5F811" blockId="11.[151,1436,1916,2029]" pageId="11" pageNumber="212">
<emphasis id="B9363802E3018C3FA2EDF8B3FEF1F86E" bold="true" box="[151,280,1916,1938]" pageId="11" pageNumber="212">FIGURE 6.</emphasis>
Habitus multi-layer stacked photographs of
<taxonomicName id="4C429F93E3018C3FA085F8B3FC79F86E" authorityName="Verhoeff" authorityYear="1924" box="[767,912,1916,1938]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B9363802E3018C3FA085F8B3FC79F86E" box="[767,912,1916,1938]" italics="true" pageId="11" pageNumber="212">Sphaerobelum</emphasis>
</taxonomicName>
species.
<emphasis id="B9363802E3018C3FA182F8B3FBE3F86E" bold="true" box="[1016,1034,1916,1938]" pageId="11" pageNumber="212">A</emphasis>
:
<taxonomicName id="4C429F93E3018C3FA667F8B3FACFF86E" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1053,1318,1916,1938]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3018C3FA667F8B3FACFF86E" box="[1053,1318,1916,1938]" italics="true" pageId="11" pageNumber="212">Sphaerobelum bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA748F8B3FA98F86E" bold="true" box="[1330,1393,1916,1938]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA748F8B3FA98F86E" box="[1330,1393,1916,1938]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3018C3FA7FEF8B3FA7CF86E" bold="true" box="[1412,1429,1916,1938]" pageId="11" pageNumber="212">B</emphasis>
:
<taxonomicName id="4C429F93E3018C3FA2EDF855FE62F84C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,395,1946,1968]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
<emphasis id="B9363802E3018C3FA2EDF855FE62F84C" box="[151,395,1946,1968]" italics="true" pageId="11" pageNumber="212">Sphaerobelum phouloei</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA3EEF855FE39F84C" bold="true" box="[404,464,1946,1968]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA3EEF855FE39F84C" box="[404,464,1946,1968]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3018C3FA39AF855FE12F84C" bold="true" box="[480,507,1946,1968]" pageId="11" pageNumber="212">C:</emphasis>
<taxonomicName id="4C429F93E3018C3FA07EF855FCCAF84C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[516,803,1946,1968]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
<emphasis id="B9363802E3018C3FA07EF855FCCAF84C" box="[516,803,1946,1968]" italics="true" pageId="11" pageNumber="212">Sphaerobelum denticulatum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA156F855FC80F84C" bold="true" box="[812,873,1946,1968]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA156F855FC80F84C" box="[812,873,1946,1968]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3018C3FA103F855FC62F84C" bold="true" box="[889,907,1946,1968]" pageId="11" pageNumber="212">D</emphasis>
:
<taxonomicName id="4C429F93E3018C3FA1E6F855FB7AF84C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[924,1171,1946,1968]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
<emphasis id="B9363802E3018C3FA1E6F855FB7AF84C" box="[924,1171,1946,1968]" italics="true" pageId="11" pageNumber="212">Sphaerobelum spinatum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA6E7F855FB30F84C" bold="true" box="[1181,1241,1946,1968]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA6E7F855FB30F84C" box="[1181,1241,1946,1968]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3018C3FA693F855FB13F84C" bold="true" box="[1257,1274,1946,1968]" pageId="11" pageNumber="212">E</emphasis>
:
<taxonomicName id="4C429F93E3018C3FA771F855FF1EF833" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="lachneeis" status="sp. nov.">
<emphasis id="B9363802E3018C3FA771F855FF1EF833" italics="true" pageId="11" pageNumber="212">Sphaerobelum lachneeis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA287F876FEDFF833" bold="true" box="[253,310,1977,1999]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA287F876FEDFF833" box="[253,310,1977,1999]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3018C3FA33EF876FEBAF833" bold="true" box="[324,339,1977,1999]" pageId="11" pageNumber="212">F</emphasis>
:
<taxonomicName id="4C429F93E3018C3FA31AF876FD9DF833" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[352,628,1977,1999]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="peterjaegeri" status="sp. nov.">
<emphasis id="B9363802E3018C3FA31AF876FD9DF833" box="[352,628,1977,1999]" italics="true" pageId="11" pageNumber="212">Sphaerobelum peterjaegeri</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA000F876FD5AF833" bold="true" box="[634,691,1977,1999]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA000F876FD5AF833" box="[634,691,1977,1999]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3018C3FA0BBF876FD3CF833" bold="true" box="[705,725,1977,1999]" pageId="11" pageNumber="212">G</emphasis>
:
<taxonomicName id="4C429F93E3018C3FA098F876FC2AF833" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[738,963,1977,1999]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
<emphasis id="B9363802E3018C3FA098F876FC2AF833" box="[738,963,1977,1999]" italics="true" pageId="11" pageNumber="212">Sphaerobelum nigrum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA1B0F876FBEAF833" bold="true" box="[970,1027,1977,1999]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA1B0F876FBEAF833" box="[970,1027,1977,1999]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3018C3FA66AF876FBCDF833" bold="true" box="[1040,1060,1977,1999]" pageId="11" pageNumber="212">H</emphasis>
:
<taxonomicName id="4C429F93E3018C3FA64BF876FAD7F833" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1073,1342,1977,1999]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
<emphasis id="B9363802E3018C3FA64BF876FAD7F833" box="[1073,1342,1977,1999]" italics="true" pageId="11" pageNumber="212">Sphaerobelum splendidum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA73EF876FA94F833" bold="true" box="[1348,1405,1977,1999]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA73EF876FA94F833" box="[1348,1405,1977,1999]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3018C3FA7F1F876FA7CF833" bold="true" box="[1419,1429,1977,1999]" pageId="11" pageNumber="212">I</emphasis>
:
<taxonomicName id="4C429F93E3018C3FA2EDF818FE6EF811" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,391,2007,2029]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
<emphasis id="B9363802E3018C3FA2EDF818FE6EF811" box="[151,391,2007,2029]" italics="true" pageId="11" pageNumber="212">Sphaerobelum laoticum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3018C3FA3F7F818FE2FF811" bold="true" box="[397,454,2007,2029]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A2058579E3018C3FA3F7F818FE2FF811" box="[397,454,2007,2029]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Not to scale.
</paragraph>
</caption>
<caption id="DF3DB498E3068C38A2EDFD07FA87FD01" httpUri="https://zenodo.org/record/2601086/files/figure.png" pageId="12" pageNumber="213" startId="12.[151,250,712,734]" targetBox="[150,1436,193,690]" targetPageId="12">
<paragraph id="8BFDE410E3068C38A2EDFD07FA87FD01" blockId="12.[151,1435,712,765]" pageId="12" pageNumber="213">
<emphasis id="B9363802E3068C38A2EDFD07FEFBFD22" bold="true" box="[151,274,712,734]" pageId="12" pageNumber="213">FIGURE 7.</emphasis>
<taxonomicName id="4C429F93E3068C38A362FD07FDF3FD22" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[280,538,712,734]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3068C38A362FD07FDF3FD22" box="[280,538,712,734]" italics="true" pageId="12" pageNumber="213">Sphaerobelum bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3068C38A065FD07FDB1FD22" bold="true" box="[543,600,712,734]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="A2058579E3068C38A065FD07FDB1FD22" box="[543,600,712,734]" pageId="12" pageNumber="213" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (ZMUC), left antenna, SEM,
<emphasis id="B9363802E3068C38A191FD07FC14FD22" bold="true" box="[1003,1021,712,734]" pageId="12" pageNumber="213">A</emphasis>
: lateral view;
<emphasis id="B9363802E3068C38A6F7FD07FB74FD22" bold="true" box="[1165,1181,712,734]" pageId="12" pageNumber="213">B</emphasis>
: apical disc with sensory cones. Abbreviations:
<emphasis id="B9363802E3068C38A300FD28FE61FD01" bold="true" box="[378,392,743,765]" pageId="12" pageNumber="213">d</emphasis>
= apical disc;
<emphasis id="B9363802E3068C38A066FD28FDDDFD01" bold="true" box="[540,564,743,765]" pageId="12" pageNumber="213">sb</emphasis>
= sensilla basiconica;
<emphasis id="B9363802E3068C38A163FD28FCC7FD01" bold="true" box="[793,814,743,765]" pageId="12" pageNumber="213">sc</emphasis>
= sensory cones; numbers refer to antennomere number.
</paragraph>
</caption>
<caption id="DF3DB498E3068C38A2EDF80EFB29F80A" httpUri="https://zenodo.org/record/2601088/files/figure.png" pageId="12" pageNumber="213" startId="12.[151,250,1985,2007]" targetBox="[151,1435,812,1976]" targetPageId="12">
<paragraph id="8BFDE410E3068C38A2EDF80EFB29F80A" blockId="12.[151,1435,1985,2038]" pageId="12" pageNumber="213">
<emphasis id="B9363802E3068C38A2EDF80EFEE6F82B" bold="true" box="[151,271,1985,2007]" pageId="12" pageNumber="213">FIGURE 8</emphasis>
.
<taxonomicName id="4C429F93E3068C38A367F80EFDC8F82B" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[285,545,1985,2007]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3068C38A367F80EFDC8F82B" box="[285,545,1985,2007]" italics="true" pageId="12" pageNumber="213">Sphaerobelum bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3068C38A053F80EFD8DF82B" bold="true" box="[553,612,1985,2007]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="A2058579E3068C38A053F80EFD8DF82B" box="[553,612,1985,2007]" pageId="12" pageNumber="213" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female paratype (ZFMK MYR8100);
<emphasis id="B9363802E3068C38A180F80EFBE5F82B" bold="true" box="[1018,1036,1985,2007]" pageId="12" pageNumber="213">A</emphasis>
: 9th left leg;
<emphasis id="B9363802E3068C38A6E7F80EFB47F82B" bold="true" box="[1181,1198,1985,2007]" pageId="12" pageNumber="213">B</emphasis>
: coxa and prefemur of second leg with vulva. Abbreviations:
<emphasis id="B9363802E3068C38A064F82FFDD4F80A" bold="true" box="[542,573,2016,2038]" pageId="12" pageNumber="213">Cx</emphasis>
= coxa;
<emphasis id="B9363802E3068C38A0EEF82FFD41F80A" bold="true" box="[660,680,2016,2038]" pageId="12" pageNumber="213">O</emphasis>
= operculum of vulva;
<emphasis id="B9363802E3068C38A1EDF82FFC54F80A" bold="true" box="[919,957,2016,2038]" pageId="12" pageNumber="213">Pre</emphasis>
= prefemur. Not to scale.
</paragraph>
</caption>
<paragraph id="8BFDE410E3078C39A2BDFF58FAEFFE74" blockId="13.[151,1437,151,968]" pageId="13" pageNumber="214">
<emphasis id="B9363802E3078C39A2BDFF58FEEEFF4C" bold="true" box="[199,263,151,176]" pageId="13" pageNumber="214">Head</emphasis>
: Eyes with&gt;70 ocelli. Aberrant ocellus located inside antennal groove. Antennae short, with rounded joints, extending posteriorly to leg-pair 3. First antennomere with cuticular scales (
<figureCitation id="1379F895E3078C39A6E0FF73FB14FF28" box="[1178,1277,188,213]" captionStart="FIGURE 7" captionStartId="12.[151,250,712,734]" captionTargetBox="[150,1436,193,690]" captionTargetId="figure@12.[150,1437,193,691]" captionTargetPageId="12" captionText="FIGURE 7. Sphaerobelum bolavensis n. sp., holotype (ZMUC), left antenna, SEM, A: lateral view; B: apical disc with sensory cones. Abbreviations: d = apical disc; sb = sensilla basiconica; sc = sensory cones; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601086/files/figure.png" pageId="13" pageNumber="214">Fig. 7A</figureCitation>
). Lengths of antennomeres: 1&gt;2=3=4=5&lt;&lt;6 (
<figureCitation id="1379F895E3078C39A070FF10FD81FF04" box="[522,616,223,248]" captionStart="FIGURE 7" captionStartId="12.[151,250,712,734]" captionTargetBox="[150,1436,193,690]" captionTargetId="figure@12.[150,1437,193,691]" captionTargetPageId="12" captionText="FIGURE 7. Sphaerobelum bolavensis n. sp., holotype (ZMUC), left antenna, SEM, A: lateral view; B: apical disc with sensory cones. Abbreviations: d = apical disc; sb = sensilla basiconica; sc = sensory cones; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601086/files/figure.png" pageId="13" pageNumber="214">Fig. 7A</figureCitation>
). All antennomeres densely pubescent, sensilla basiconica surrounding apical disc. Shape of antennae sexually dimorphic, cylindrical in female; thickened, apically widened and slightly flattened in male (
<figureCitation id="1379F895E3078C39A319FEE8FE52FEBC" box="[355,443,295,320]" captionStart="FIGURE 7" captionStartId="12.[151,250,712,734]" captionTargetBox="[150,1436,193,690]" captionTargetId="figure@12.[150,1437,193,691]" captionTargetPageId="12" captionText="FIGURE 7. Sphaerobelum bolavensis n. sp., holotype (ZMUC), left antenna, SEM, A: lateral view; B: apical disc with sensory cones. Abbreviations: d = apical disc; sb = sensilla basiconica; sc = sensory cones; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601086/files/figure.png" pageId="13" pageNumber="214">Fig. 7B</figureCitation>
). Apical disc with ca 47/52 apical cones (male) (
<figureCitation id="1379F895E3078C39A1A4FEE8FBDDFEBC" box="[990,1076,295,320]" captionStart="FIGURE 7" captionStartId="12.[151,250,712,734]" captionTargetBox="[150,1436,193,690]" captionTargetId="figure@12.[150,1437,193,691]" captionTargetPageId="12" captionText="FIGURE 7. Sphaerobelum bolavensis n. sp., holotype (ZMUC), left antenna, SEM, A: lateral view; B: apical disc with sensory cones. Abbreviations: d = apical disc; sb = sensilla basiconica; sc = sensory cones; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601086/files/figure.png" pageId="13" pageNumber="214">Fig. 7B</figureCitation>
) or 30/32 (female). Apical cone shape typical of Diplopoda (
<figureCitation id="1379F895E3078C39A399FE83FDA9FE98" box="[483,576,332,357]" captionStart="FIGURE 7" captionStartId="12.[151,250,712,734]" captionTargetBox="[150,1436,193,690]" captionTargetId="figure@12.[150,1437,193,691]" captionTargetPageId="12" captionText="FIGURE 7. Sphaerobelum bolavensis n. sp., holotype (ZMUC), left antenna, SEM, A: lateral view; B: apical disc with sensory cones. Abbreviations: d = apical disc; sb = sensilla basiconica; sc = sensory cones; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601086/files/figure.png" pageId="13" pageNumber="214">Fig. 7B</figureCitation>
). Organ of Tömösváry located inside antennal groove.
<emphasis id="B9363802E3078C39A6BFFE83FA7DFE99" bold="true" box="[1221,1428,332,357]" pageId="13" pageNumber="214">Gnathochilarium</emphasis>
: structure typical of the order. Palpi with sensory cones arranged in clusters.
<emphasis id="B9363802E3078C39A192FEA0FB8FFE74" bold="true" box="[1000,1126,367,392]" pageId="13" pageNumber="214">Mandibles</emphasis>
not dissected.
</paragraph>
<paragraph id="8BFDE410E3078C39A2BDFE5BFDD0FC34" blockId="13.[151,1437,151,968]" pageId="13" pageNumber="214">
<emphasis id="B9363802E3078C39A2BDFE5BFE61FE51" bold="true" box="[199,392,404,429]" pageId="13" pageNumber="214">Stigmatic plates</emphasis>
: first stigmatic plate widely rounded, apex well-rounded, weakly curved towards coxa 1.
<emphasis id="B9363802E3078C39A2EDFE78FEA9FE2C" bold="true" box="[151,320,439,464]" pageId="13" pageNumber="214">Laterotergites</emphasis>
: laterotergite 1 strongly projecting into a sharp tip. Laterotergite 2 with a broad, stout, much shorter projection.
<emphasis id="B9363802E3078C39A361FE13FE9DFE09" bold="true" box="[283,372,476,501]" pageId="13" pageNumber="214">Collum</emphasis>
with glabrous surface, margins with few isolated setae.
<emphasis id="B9363802E3078C39A675FE13FB25FE09" bold="true" box="[1039,1228,476,501]" pageId="13" pageNumber="214">Thoracic shield</emphasis>
: surface glabrous unlike tergites, setae only in grooves. Shallow grooves beset with numerous long setae, sloped towards groove with 9 continuous anterior and posterior keels.
<emphasis id="B9363802E3078C39A008FDEBFD38FDC1" bold="true" box="[626,721,548,573]" pageId="13" pageNumber="214">Tergites</emphasis>
: surface densely setose with short setae. Tips of paratergites of midbody tergites projecting posteriorly.
<emphasis id="B9363802E3078C39A02DFD88FD07FD9C" bold="true" box="[599,750,583,608]" pageId="13" pageNumber="214">Endotergum</emphasis>
: inner section lacking any spines or setae. Middle area with a single row of large, sparse, elliptical, cuticular impressions. Distance between impressions twice as wide as their diameter. Apically, 2 dense rows of long marginal bristles, their tips slightly protruding beyond tergal margin (
<figureCitation id="1379F895E3078C39A715FD40FF55FD30" captionStart="FIGURE 9" captionStartId="13.[151,250,1423,1445]" captionTargetBox="[152,1434,1021,1400]" captionTargetId="figure@13.[152,1435,1021,1402]" captionTargetPageId="13" captionText="FIGURE 9. Endoterga of the body ring 9 of different Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum spinatum n. sp.; D: Sphaerobelum splendidum n. sp. Abbreviations: cp = cuticular patterns; iA = inner area." httpUri="https://zenodo.org/record/2601090/files/figure.png" pageId="13" pageNumber="214">Fig. 9A</figureCitation>
). Bristles not smooth, but with numerous small spicules.
<emphasis id="B9363802E3078C39A127FD7BFC0EFD31" bold="true" box="[861,999,692,717]" pageId="13" pageNumber="214">Anal shield</emphasis>
: large, sexually dimorphic: in female weakly bell-shaped, in male strongly bell-shaped. Surface in both sexes completely covered by tiny setae. Underside with a single, short, black, locking carina, the latter located close to last laterotergite.
<emphasis id="B9363802E3078C39A69CFD33FAF7FCE9" bold="true" box="[1254,1310,764,789]" pageId="13" pageNumber="214">Legs</emphasis>
: leg-pair 1 with 2 ventral spines, leg-pair 2 with 4, leg-pair 3 with 5 or 6. First two leg-pairs without an apical spine. Leg-pairs 421 with 79 ventral spines and one dorso-apical spine. In leg-pair 9, femur 1.3 times, tarsus 4 times longer than wide (
<figureCitation id="1379F895E3078C39A2A7FCA8FED0FC7C" box="[221,313,871,896]" captionStart="FIGURE 8" captionStartId="12.[151,250,1985,2007]" captionTargetBox="[151,1435,812,1976]" captionTargetId="figure@12.[151,1435,812,1976]" captionTargetPageId="12" captionText="FIGURE 8. Sphaerobelum bolavensis n. sp., female paratype (ZFMK MYR8100); A: 9th left leg; B: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur. Not to scale." httpUri="https://zenodo.org/record/2601088/files/figure.png" pageId="13" pageNumber="214">Fig. 8A</figureCitation>
). All podomeres densely setose. Coxa with a large and well-rounded process. Coxal process absent on leg-pair 1 and 2. Prefemur apico-mesally with a weak projection. Femur in apical part extended mesally into a dentate margin tipped with 46 teeth.
</paragraph>
<caption id="DF3DB498E3078C39A2EDFA40FE1AFA1E" httpUri="https://zenodo.org/record/2601090/files/figure.png" pageId="13" pageNumber="214" startId="13.[151,250,1423,1445]" targetBox="[152,1434,1021,1400]" targetPageId="13">
<paragraph id="8BFDE410E3078C39A2EDFA40FE1AFA1E" blockId="13.[151,1436,1423,1506]" pageId="13" pageNumber="214">
<emphasis id="B9363802E3078C39A2EDFA40FEF0FA59" bold="true" box="[151,281,1423,1445]" pageId="13" pageNumber="214">FIGURE 9.</emphasis>
Endoterga of the body ring 9 of different
<taxonomicName id="4C429F93E3078C39A081FA40FC65FA59" authorityName="Verhoeff" authorityYear="1924" box="[763,908,1423,1445]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B9363802E3078C39A081FA40FC65FA59" box="[763,908,1423,1445]" italics="true" pageId="13" pageNumber="214">Sphaerobelum</emphasis>
</taxonomicName>
species.
<emphasis id="B9363802E3078C39A18FFA40FBEEFA59" bold="true" box="[1013,1031,1423,1445]" pageId="13" pageNumber="214">A</emphasis>
:
<taxonomicName id="4C429F93E3078C39A661FA40FACDFA59" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1051,1316,1423,1445]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3078C39A661FA40FACDFA59" box="[1051,1316,1423,1445]" italics="true" pageId="13" pageNumber="214">Sphaerobelum bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3078C39A74BFA40FA99FA59" bold="true" box="[1329,1392,1423,1445]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="A2058579E3078C39A74BFA40FA99FA59" box="[1329,1392,1423,1445]" pageId="13" pageNumber="214" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3078C39A7FEFA40FA7CFA59" bold="true" box="[1412,1429,1423,1445]" pageId="13" pageNumber="214">B</emphasis>
:
<taxonomicName id="4C429F93E3078C39A2EDFA61FE63FA38" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,394,1454,1476]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
<emphasis id="B9363802E3078C39A2EDFA61FE63FA38" box="[151,394,1454,1476]" italics="true" pageId="13" pageNumber="214">Sphaerobelum phouloei</emphasis>
</taxonomicName>
<emphasis id="B9363802E3078C39A3E9FA61FE27FA38" bold="true" box="[403,462,1454,1476]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="A2058579E3078C39A3E9FA61FE27FA38" box="[403,462,1454,1476]" pageId="13" pageNumber="214" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3078C39A3A4FA61FE10FA38" bold="true" box="[478,505,1454,1476]" pageId="13" pageNumber="214">C:</emphasis>
<taxonomicName id="4C429F93E3078C39A078FA61FD10FA38" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[514,761,1454,1476]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
<emphasis id="B9363802E3078C39A078FA61FD10FA38" box="[514,761,1454,1476]" italics="true" pageId="13" pageNumber="214">Sphaerobelum spinatum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3078C39A178FA61FCD4FA38" bold="true" box="[770,829,1454,1476]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="A2058579E3078C39A178FA61FCD4FA38" box="[770,829,1454,1476]" pageId="13" pageNumber="214" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9363802E3078C39A137FA61FCB6FA38" bold="true" box="[845,863,1454,1476]" pageId="13" pageNumber="214">D</emphasis>
:
<taxonomicName id="4C429F93E3078C39A115FA61FB96FA38" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[879,1151,1454,1476]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
<emphasis id="B9363802E3078C39A115FA61FB96FA38" box="[879,1151,1454,1476]" italics="true" pageId="13" pageNumber="214">Sphaerobelum splendidum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3078C39A6FDFA61FB2AFA38" bold="true" box="[1159,1219,1454,1476]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="A2058579E3078C39A6FDFA61FB2AFA38" box="[1159,1219,1454,1476]" pageId="13" pageNumber="214" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Abbreviations:
<emphasis id="B9363802E3078C39A711FA61FA6CFA38" bold="true" box="[1387,1413,1454,1476]" pageId="13" pageNumber="214">cp</emphasis>
= cuticular patterns;
<emphasis id="B9363802E3078C39A329FA03FE85FA1E" bold="true" box="[339,364,1484,1506]" pageId="13" pageNumber="214">iA</emphasis>
= inner area.
</paragraph>
</caption>
<paragraph id="8BFDE410E3078C39A2BDF9C0FC66F98C" blockId="13.[151,1437,1551,2008]" pageId="13" pageNumber="214">
<emphasis id="B9363802E3078C39A2BDF9C0FE1FF9D4" bold="true" box="[199,502,1551,1576]" pageId="13" pageNumber="214">Female sexual characters</emphasis>
: vulva large, covering 2/3 of coxa, extending mesally to anterior third of prefemur (
<figureCitation id="1379F895E3078C39A2E5F9FCFF10F9B0" box="[159,249,1587,1612]" captionStart="FIGURE 8" captionStartId="12.[151,250,1985,2007]" captionTargetBox="[151,1435,812,1976]" captionTargetId="figure@12.[151,1435,812,1976]" captionTargetPageId="12" captionText="FIGURE 8. Sphaerobelum bolavensis n. sp., female paratype (ZFMK MYR8100); A: 9th left leg; B: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur. Not to scale." httpUri="https://zenodo.org/record/2601088/files/figure.png" pageId="13" pageNumber="214">Fig. 8B</figureCitation>
). Operculum rounded, very slightly invaginated medially, mesal margin strongly projecting into a wellrounded lobe twice as long as operculum.
<emphasis id="B9363802E3078C39A015F998FCFAF98C" bold="true" box="[623,787,1623,1648]" pageId="13" pageNumber="214">Subanal plate</emphasis>
: damaged.
</paragraph>
<paragraph id="8BFDE410E3078C3AA2BDF9B4FEA9FEC3" blockId="13.[151,1437,1551,2008]" lastBlockId="14.[151,1437,151,464]" lastPageId="14" lastPageNumber="215" pageId="13" pageNumber="214">
<emphasis id="B9363802E3078C39A2BDF9B4FE08F968" bold="true" box="[199,481,1659,1684]" pageId="13" pageNumber="214">Male sexual characters</emphasis>
: gonopore covered with a single, undivided, circular, sclerotized plate (
<figureCitation id="1379F895E3078C39A75EF9B4FA66F968" box="[1316,1423,1659,1684]" captionStart="FIGURE 10" captionStartId="16.[151,250,1946,1968]" captionTargetBox="[177,1416,193,1934]" captionTargetId="figure@16.[177,1416,193,1934]" captionTargetPageId="16" captionText="FIGURE 10. Sphaerobelum bolavensis n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601094/files/figure.png" pageId="13" pageNumber="214">Fig. 10A</figureCitation>
).
<emphasis id="B9363802E3078C39A2EDF950FE82F944" bold="true" box="[151,363,1695,1720]" pageId="13" pageNumber="214">Anterior telopods</emphasis>
(
<figureCitation id="1379F895E3078C39A300F950FE26F944" box="[378,463,1695,1720]" captionStart="FIGURE 10" captionStartId="16.[151,250,1946,1968]" captionTargetBox="[177,1416,193,1934]" captionTargetId="figure@16.[177,1416,193,1934]" captionTargetPageId="16" captionText="FIGURE 10. Sphaerobelum bolavensis n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601094/files/figure.png" pageId="13" pageNumber="214">Figs 10</figureCitation>
BD): consisting of only 3 telopoditomeres above syncoxite. Telopoditomeres 3 and 4 apparently fused. Telopoditomere 1 rectangular, as long as wide. Telopoditomere 2 large, half as long as telopoditomere 3. Process of telopoditomere 2 located posteriorly, visible in anterior view; process slender, projecting to half length of telopoditomere 3. Process of telopoditomere 2 conspicuously 'hooked' apically with a sharp tip. Telopoditomere 3 massive, cylindrical, straight, apically slightly tapering; posterior side with a black sclerotized spot and a small, triangular spine. Telopoditomere
<quantity id="4CBA49F5E3078C39A137F89BFC9EF890" box="[845,887,1875,1900]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="214" unit="in" value="1.0">1 in</quantity>
apical view covered with long setae. In posterior view all telopoditomeres almost glabrous.
<emphasis id="B9363802E3078C39A00BF8B8FCA3F86C" bold="true" box="[625,842,1911,1936]" pageId="13" pageNumber="214">Posterior telopods</emphasis>
(
<figureCitation id="1379F895E3078C39A122F8B8FC37F86C" box="[856,990,1911,1936]" captionStart="FIGURE 10" captionStartId="16.[151,250,1946,1968]" captionTargetBox="[177,1416,193,1934]" captionTargetId="figure@16.[177,1416,193,1934]" captionTargetPageId="16" captionText="FIGURE 10. Sphaerobelum bolavensis n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601094/files/figure.png" pageId="13" pageNumber="214">Figs 10E, F</figureCitation>
): telopoditomere 1 short, half as long as wide. Immovable finger (process of telopoditomere 2) shorter than movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a characteristic distally swollen apex; well rounded, apex therefore twice as wide as base. Telopoditomere 3 rectangular, well rounded. Telopoditomere 4 as long as but slightly slenderer than telopoditomere 3. Telopoditomere 4 2.7 times longer than wide; apically weakly tapering, very slightly curved towards immovable finger. Telopoditomere 1 at both sides covered by setae, remaining telopoditomeres in posterior view almost glabrous, in anterior view with few isolated setae except for immovable finger which is more densely setose.
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE3048C3AA2BDFE83FEC9FE74" pageId="14" pageNumber="215" type="discussion">
<paragraph id="8BFDE410E3048C3AA2BDFE83FEC9FE74" blockId="14.[151,1437,151,464]" pageId="14" pageNumber="215">
<emphasis id="B9363802E3048C3AA2BDFE83FEDDFE99" bold="true" box="[199,308,332,357]" pageId="14" pageNumber="215">Remarks</emphasis>
: This species occurs in syntopy with a female of an undetermined species,
<taxonomicName id="4C429F93E3048C3AA6E0FE83FAC6FE99" authorityName="Gray" authorityYear="1843" box="[1178,1327,332,357]" class="Diplopoda" family="Zephroniidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="215" phylum="Arthropoda" rank="family">Zephroniidae</taxonomicName>
sp. L04b (see below).
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE3048C3AA2BDFE5BFE9BFE2C" pageId="14" pageNumber="215" type="etymology">
<paragraph id="8BFDE410E3048C3AA2BDFE5BFE9BFE2C" blockId="14.[151,1437,151,464]" pageId="14" pageNumber="215">
<emphasis id="B9363802E3048C3AA2BDFE5BFE77FE51" bold="true" box="[199,414,404,429]" pageId="14" pageNumber="215">Derivatio nominis</emphasis>
: bolavensis, noun in apposition, from the Bolaven plateau in
<collectingCountry id="F355A480E3048C3AA621FE5AFB78FE50" box="[1115,1169,405,428]" name="Laos" pageId="14" pageNumber="215">Laos</collectingCountry>
where this species was discovered (
<figureCitation id="1379F895E3048C3AA365FE78FE8DFE2C" box="[287,356,439,464]" captionStart="FIGURE 5" captionStartId="9.[151,250,2005,2027]" captionTargetBox="[185,1415,664,1984]" captionTargetId="figure@9.[181,1415,663,1984]" captionTargetPageId="9" captionText="FIGURE 5. Distribution map of Sphaerobelum species, including the unnamed females." httpUri="https://zenodo.org/record/2601082/files/figure.png" pageId="14" pageNumber="215">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>