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<document id="F331FD55AF0768CE12BAF15868148ED1" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E3008C3EA2EDFF57FA75F80C" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum Verhoeff 1924" docType="key" docVersion="9" lastPageNumber="211" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="211" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:title id="E97BDC4759AAF57D3189FB0315B81F2D">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<treatment id="03EB5506E3008C3EA2EDFF57FA75F80C" ID-DOI="http://doi.org/10.5281/zenodo.5934520" ID-GBIF-Taxon="154474559" ID-Zenodo-Dep="5934520" LSID="urn:lsid:plazi:treatment:03EB5506E3008C3EA2EDFF57FA75F80C" httpUri="http://treatment.plazi.org/id/03EB5506E3008C3EA2EDFF57FA75F80C" lastPageNumber="211" pageId="10" pageNumber="211">
<subSubSection id="C358B79BE3008C3EA2EDFF57FC35FF4E" box="[151,988,151,178]" pageId="10" pageNumber="211" type="nomenclature">
<paragraph id="8BFDE410E3008C3EA2EDFF57FC35FF4E" blockId="10.[151,988,151,178]" box="[151,988,151,178]" pageId="10" pageNumber="211">
<heading id="D0B5537CE3008C3EA2EDFF57FC35FF4E" bold="true" box="[151,988,151,178]" fontSize="11" level="1" pageId="10" pageNumber="211" reason="1">
<emphasis id="B9363802E3008C3EA2EDFF57FC35FF4E" bold="true" box="[151,988,151,178]" pageId="10" pageNumber="211">
Key to species of
<taxonomicName id="4C429F93E3008C3EA317FF58FDF4FF4D" authorityName="Verhoeff" authorityYear="1924" box="[365,541,151,177]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B9363802E3008C3EA317FF58FDF4FF4D" bold="true" box="[365,541,151,177]" italics="true" pageId="10" pageNumber="211">Sphaerobelum</emphasis>
</taxonomicName>
(based mainly on male characters):
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C358B79BE3008C3EA2EDFF16FA75F80C" pageId="10" pageNumber="211" type="key">
<keyStep id="30B6FCB5E3008C3EA2EDFF16FA72FED4" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFF16FA72FEF7" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFF16FA72FEF7" pageId="10" pageNumber="211">
1. Femur of legs at mesal margin extended, with teeth (
<figureCitation id="1379F895E3008C3EA0A3FF16FCDBFF12" box="[729,818,217,238]" captionStart="FIGURE 13" captionStartId="19.[151,250,1951,1973]" captionTargetBox="[191,1401,195,1934]" captionTargetId="figure@19.[191,1401,193,1934]" captionTargetPageId="19" captionText="FIGURE 13. Sphaerobelum denticulatum n. sp., holotype (MHNG); A: 9th left leg; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left posterior telopod, anterior view; E: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601100/files/figure.png" pageId="10" pageNumber="211">Fig. 13A</figureCitation>
). Apical margin of prefemur modified, extended into a process..................................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFEDCFA72FED4" blockId="10.[151,1436,217,2032]" box="[151,1435,275,296]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFEDCFA72FED4" box="[151,1435,275,296]" pageId="10" pageNumber="211">
- Femur of legs not extended, well-rounded (
<figureCitation id="1379F895E3008C3EA0FAFEDCFD3EFED4" box="[640,727,275,296]" captionStart="FIGURE 17" captionStartId="24.[151,250,1981,2003]" captionTargetBox="[151,1436,238,1935]" captionTargetId="figure@24.[151,1436,233,1935]" captionTargetPageId="24" captionText="FIGURE 17. Sphaerobelum lachneeis n. sp., female paratype (MHNG); A: coxa and prefemur of second leg with vulva; B: 9th left leg. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601109/files/figure.png" pageId="10" pageNumber="211">Fig. 17B</figureCitation>
). Prefemur unmodified........................................ 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFEE0FA75FE9D" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFEE0FA72FEB8" blockId="10.[151,1436,217,2032]" box="[151,1435,303,324]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFEE0FA72FEB8" box="[151,1435,303,324]" pageId="10" pageNumber="211">
2. Teeth on femur only at apex: 46 (
<figureCitation id="1379F895E3008C3EA051FEE0FD90FEB8" box="[555,633,303,324]" captionStart="FIGURE 8" captionStartId="12.[151,250,1985,2007]" captionTargetBox="[151,1435,812,1976]" captionTargetId="figure@12.[151,1435,812,1976]" captionTargetPageId="12" captionText="FIGURE 8. Sphaerobelum bolavensis n. sp., female paratype (ZFMK MYR8100); A: 9th left leg; B: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur. Not to scale." httpUri="https://zenodo.org/record/2601088/files/figure.png" pageId="10" pageNumber="211">Fig. 8A</figureCitation>
). Anterior telopod consisting of 3 podomeres............................. 3
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFE83FA75FE9D" blockId="10.[151,1436,217,2032]" box="[151,1436,332,353]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFE83FA75FE9D" box="[151,1436,332,353]" pageId="10" pageNumber="211">- Teeth on whole mesal margin of femur:&gt;10. Anterior telopod always consisting of 4 telopoditomeres................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFEA6FA75FE2A" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFEA6FA75FE67" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFEA6FA75FE67" pageId="10" pageNumber="211">
3. Female vulval operculum strongly projecting in two processes. Posterior telopod with telopoditomere 4 tip curved towards immovable finger like a small hook.....................................................
<taxonomicName id="4C429F93E3008C3EA6C8FE49FA75FE67" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[1202,1436,390,412]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="B9363802E3008C3EA6C8FE49FAFAFE60" box="[1202,1299,390,412]" italics="true" pageId="10" pageNumber="211">S. bicorne</emphasis>
<bibRefCitation id="EFD399E1E3008C3EA761FE49FA75FE67" author="Attems, C." box="[1307,1436,390,411]" pageId="10" pageNumber="211" pagination="187 - 353" refId="ref23863" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle, N. S. 6 (2), 187 - 353." type="journal article" year="1938">Attems, 1938</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFE6CFA75FE2A" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFE6CFA75FE2A" pageId="10" pageNumber="211">
- Female vulva projecting in a mesal process (
<figureCitation id="1379F895E3008C3EA0F6FE6CFD3EFE44" box="[652,727,419,440]" captionStart="FIGURE 8" captionStartId="12.[151,250,1985,2007]" captionTargetBox="[151,1435,812,1976]" captionTargetId="figure@12.[151,1435,812,1976]" captionTargetPageId="12" captionText="FIGURE 8. Sphaerobelum bolavensis n. sp., female paratype (ZFMK MYR8100); A: 9th left leg; B: coxa and prefemur of second leg with vulva. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur. Not to scale." httpUri="https://zenodo.org/record/2601088/files/figure.png" pageId="10" pageNumber="211">Fig. 8B</figureCitation>
). Posterior telopod: telopoditomere 4 straight, not hooked (
<figureCitation id="1379F895E3008C3EA775FE6CFA8AFE44" box="[1295,1379,419,440]" captionStart="FIGURE 10" captionStartId="16.[151,250,1946,1968]" captionTargetBox="[177,1416,193,1934]" captionTargetId="figure@16.[177,1416,193,1934]" captionTargetPageId="16" captionText="FIGURE 10. Sphaerobelum bolavensis n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: left posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601094/files/figure.png" pageId="10" pageNumber="211">Fig. 10E</figureCitation>
).........................................................................................
<taxonomicName id="4C429F93E3008C3EA698FE70FAB6FE29" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1250,1375,447,469]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B9363802E3008C3EA698FE70FAB6FE29" box="[1250,1375,447,469]" italics="true" pageId="10" pageNumber="211">S. bolavensis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3008C3EA71CFE0FFA75FE2A" bold="true" box="[1382,1436,448,470]" pageId="10" pageNumber="211">
<taxonomicNameLabel id="A2058579E3008C3EA71CFE0FFA75FE2A" box="[1382,1436,448,470]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFE12FA72FD9A" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFE12FA72FDD0" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFE12FA72FDD0" pageId="10" pageNumber="211">
4. Posterior telopod: telopoditomere 4 apex weakly curved towards and overlapping immovable finger. Anterior side of immovable finger with sclerotized spots opposite to tip of telopoditomere 4 (
<figureCitation id="1379F895E3008C3EA10DFE35FC27FDF3" box="[887,974,506,527]" captionStart="FIGURE 12" captionStartId="18.[151,250,1942,1964]" captionTargetBox="[151,1436,247,1876]" captionTargetId="figure@18.[151,1436,243,1876]" captionTargetPageId="18" captionText="FIGURE 12. Sphaerobelum phouloei n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601098/files/figure.png" pageId="10" pageNumber="211">Fig. 12E</figureCitation>
). Immovable finger in anterior view with a large sclerotized spine...................................................................................... 5
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFDFBFA72FD9A" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFDFBFA72FD9A" pageId="10" pageNumber="211">
- Posterior telopod: telopoditomere 4 straight, not curved. No sclerotized spots or a spine at immovable finger (
<figureCitation id="1379F895E3008C3EA681FDFBFAB9FDB5" box="[1275,1360,564,585]" captionStart="FIGURE 16" captionStartId="22.[151,250,1927,1949]" captionTargetBox="[151,1435,373,1906]" captionTargetId="figure@22.[151,1435,373,1906]" captionTargetPageId="22" captionText="FIGURE 16. Sphaerobelum spinatum n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Not to scale." httpUri="https://zenodo.org/record/2601106/files/figure.png" pageId="10" pageNumber="211">Fig. 16E</figureCitation>
)......................................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFDA1FA75FD26" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFDA1FA75FD5C" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFDA1FA75FD5C" pageId="10" pageNumber="211">
5. Male antennae with 3035 apical cones. Tergites covered with short setae. Legs orange. Prefemur mesal margin rounded (
<figureCitation id="1379F895E3008C3EA70FFDA1FEE2FD5C" captionStart="FIGURE 11" captionStartId="17.[151,250,1968,1990]" captionTargetBox="[150,1436,250,1894]" captionTargetId="figure@17.[150,1436,250,1894]" captionTargetPageId="17" captionText="FIGURE 11. Sphaerobelum phouloei n. sp., male holotype (ZMUC); A: left first coxa with stigmatic plate; B: 9th left leg. Abbreviations: Cx = coxa; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601096/files/figure.png" pageId="10" pageNumber="211">Fig. 11B</figureCitation>
)...................................................................................
<taxonomicName id="4C429F93E3008C3EA689FD45FAB6FD5C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1267,1375,650,672]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
<emphasis id="B9363802E3008C3EA689FD45FAB6FD5C" box="[1267,1375,650,672]" italics="true" pageId="10" pageNumber="211">S. phouloei</emphasis>
</taxonomicName>
<emphasis id="B9363802E3008C3EA71CFD45FA75FD5C" bold="true" box="[1382,1436,650,672]" pageId="10" pageNumber="211">
<taxonomicNameLabel id="A2058579E3008C3EA71CFD45FA75FD5C" box="[1382,1436,650,672]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFD67FA75FD26" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFD67FA75FD26" pageId="10" pageNumber="211">
- Male antennae with 6570 apical cones. Tergites glabrous. Legs light brown. Prefemur mesal margin with indentations like the femur (
<figureCitation id="1379F895E3008C3EA35DFD0BFE97FD25" box="[295,382,708,729]" captionStart="FIGURE 13" captionStartId="19.[151,250,1951,1973]" captionTargetBox="[191,1401,195,1934]" captionTargetId="figure@19.[191,1401,193,1934]" captionTargetPageId="19" captionText="FIGURE 13. Sphaerobelum denticulatum n. sp., holotype (MHNG); A: 9th left leg; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left posterior telopod, anterior view; E: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601100/files/figure.png" pageId="10" pageNumber="211">Fig. 13A</figureCitation>
).....................................................................
<taxonomicName id="4C429F93E3008C3EA6B0FD0BFAB6FD26" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1226,1375,708,730]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
<emphasis id="B9363802E3008C3EA6B0FD0BFAB6FD26" box="[1226,1375,708,730]" italics="true" pageId="10" pageNumber="211">S. denticulatum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3008C3EA71CFD0BFA75FD26" bold="true" box="[1382,1436,708,730]" pageId="10" pageNumber="211">
<taxonomicNameLabel id="A2058579E3008C3EA71CFD0BFA75FD26" box="[1382,1436,708,730]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFD2DFA75FCB2" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFD2DFA72FCEF" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFD2DFA72FCEF" pageId="10" pageNumber="211">6. Male antennae red or brown, with&gt;70 apical cones. Tergites matte. First laterotergite weakly projecting posterior-laterally. ...................................................................................................... 7</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFCD3FA75FCB2" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFCD3FA75FCB2" pageId="10" pageNumber="211">
- Male antennae yellow, with 5055 apical cones. Tergites shiny, with setae. First laterotergite strongly projecting posterior-laterally.................................................................................
<taxonomicName id="4C429F93E3008C3EA695FCF7FAB6FCB2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1263,1375,824,846]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
<emphasis id="B9363802E3008C3EA695FCF7FAB6FCB2" box="[1263,1375,824,846]" italics="true" pageId="10" pageNumber="211">S. spinatum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3008C3EA71CFCF7FA75FCB2" bold="true" box="[1382,1436,824,846]" pageId="10" pageNumber="211">
<taxonomicNameLabel id="A2058579E3008C3EA71CFCF7FA75FCB2" box="[1382,1436,824,846]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFC9AFA72FC3D" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFC9AFA75FC74" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFC9AFA75FC74" pageId="10" pageNumber="211">
7. Tergites glabrous, matte. Endotergum bristles only protruding slightly midway towards margin. Anterior telopod: telopoditomeres 3 and 4 not well-separated in posterior view (
<figureCitation id="1379F895E3008C3EA0C5FCBDFCE1FC7B" box="[703,776,882,903]" captionStart="FIGURE 21" captionStartId="30.[151,250,1950,1972]" captionTargetBox="[174,1418,193,1936]" captionTargetId="figure@30.[174,1418,193,1936]" captionTargetPageId="30" captionText="FIGURE 21. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="10" pageNumber="211">Figs 21</figureCitation>
BD).....................................
<taxonomicName id="4C429F93E3008C3EA778FCBDFAB6FC74" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1282,1375,882,904]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov.">
<emphasis id="B9363802E3008C3EA778FCBDFAB6FC74" box="[1282,1375,882,904]" italics="true" pageId="10" pageNumber="211">S. nigrum</emphasis>
</taxonomicName>
<emphasis id="B9363802E3008C3EA71CFCBDFA75FC74" bold="true" box="[1382,1436,882,904]" pageId="10" pageNumber="211">
<taxonomicNameLabel id="A2058579E3008C3EA71CFCBDFA75FC74" box="[1382,1436,882,904]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFC40FA72FC3D" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFC40FA72FC3D" pageId="10" pageNumber="211">- Tergites hairy, with pits. Endotergum bristles long, protruding above tergite margin. Anterior telopod: telopoditomeres 3 and 4 well separated........................................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFC06FA75FB93" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFC06FA75FBE4" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFC06FA75FBE4" pageId="10" pageNumber="211">
8. Colour dark olive, posterior tergal margins with a broad, light-brown band, antennae and legs conspicuously orange. Prefemur without lateral projection. Apical margins of vulval operculum equally converging towards a mesal tip. Tip of laterotergite 1 slightly extended, well-rounded. Male antennomere 6 thickened......................
<taxonomicName id="4C429F93E3008C3EA62BFBCDFA75FBE4" authority="Semenyuk et al., 2018" authorityName="Semenyuk" authorityYear="2018" box="[1105,1436,1026,1048]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="cattiense">
<emphasis id="B9363802E3008C3EA62BFBCDFB57FBE4" box="[1105,1214,1026,1048]" italics="true" pageId="10" pageNumber="211">S. cattiense</emphasis>
<bibRefCitation id="EFD399E1E3008C3EA6BEFBCCFA75FBE4" author="Semenyuk, I. &amp; Golovatch, S. I. &amp; Wesener, T." box="[1220,1436,1026,1048]" pageId="10" pageNumber="211" pagination="535 - 550" refId="ref25362" refString="Semenyuk, I., Golovatch, S. I. &amp; Wesener, T. (2018) Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida: Zephroniidae). Zootaxa, 4459 (3), 535 - 550. https: // doi. org / 10.11646 / zootaxa. 4459.3.7" type="journal article" year="2018">
Semenyuk
<emphasis id="B9363802E3008C3EA74AFBCCFAB3FBE4" box="[1328,1370,1026,1048]" italics="true" pageId="10" pageNumber="211">et al</emphasis>
., 2018
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFBEFFA75FB93" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFBEFFA75FB93" pageId="10" pageNumber="211">
- Colour dark chestnut brown, posterior tergal margins black, legs dark brown, tips light brown. Lateral margin of prefemur with a sharp projection juxtaposed to coxal process. Apical margin of vulval operculum recessed centrally. Tip of laterotergite 1 slightly extended, sharp. Male antennomere 6 axe-shaped.......................
<taxonomicName id="4C429F93E3008C3EA66CFB96FA75FB93" authority="Semenyuk et al., 2018" authorityName="Semenyuk" authorityYear="2018" box="[1046,1436,1113,1135]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="konkakinhense">
<emphasis id="B9363802E3008C3EA66CFB96FB57FB93" box="[1046,1214,1113,1135]" italics="true" pageId="10" pageNumber="211">S. konkakinhense</emphasis>
<bibRefCitation id="EFD399E1E3008C3EA6BEFB95FA75FB93" author="Semenyuk, I. &amp; Golovatch, S. I. &amp; Wesener, T." box="[1220,1436,1113,1135]" pageId="10" pageNumber="211" pagination="535 - 550" refId="ref25362" refString="Semenyuk, I., Golovatch, S. I. &amp; Wesener, T. (2018) Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida: Zephroniidae). Zootaxa, 4459 (3), 535 - 550. https: // doi. org / 10.11646 / zootaxa. 4459.3.7" type="journal article" year="2018">
Semenyuk
<emphasis id="B9363802E3008C3EA74AFB95FAB3FB93" box="[1328,1370,1113,1135]" italics="true" pageId="10" pageNumber="211">et al</emphasis>
., 2018
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFBB8FA72FB55" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFBB8FA72FB70" blockId="10.[151,1436,217,2032]" box="[151,1435,1143,1164]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFBB8FA72FB70" box="[151,1435,1143,1164]" pageId="10" pageNumber="211">9. Posterior telopod without 'hook'......................................................................... 10</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFB5BFA72FB55" blockId="10.[151,1436,217,2032]" box="[151,1435,1172,1193]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFB5BFA72FB55" box="[151,1435,1172,1193]" pageId="10" pageNumber="211">
- Posterior telopod: telopoditomere 4 apically always with a short recurved 'hook' (
<figureCitation id="1379F895E3008C3EA196FB5BFBABFB55" box="[1004,1090,1172,1193]" captionStart="FIGURE 21" captionStartId="30.[151,250,1950,1972]" captionTargetBox="[174,1418,193,1936]" captionTargetId="figure@30.[174,1418,193,1936]" captionTargetPageId="30" captionText="FIGURE 21. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="10" pageNumber="211">Fig. 21E</figureCitation>
)........................... 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFB7FFA72FAC5" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFB7FFA75FAFC" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFB7FFA75FAFC" pageId="10" pageNumber="211">
10. Tergites covered with very long setae. Setae longer than width of tergites (
<figureCitation id="1379F895E3008C3EA1D9FB7FFC04FB39" box="[931,1005,1200,1221]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="10" pageNumber="211">Fig. 6E</figureCitation>
). Immovable finger of posterior telopod with very long mesal extension (
<figureCitation id="1379F895E3008C3EA39DFB02FDD7FB1E" box="[487,574,1229,1250]" captionStart="FIGURE 18" captionStartId="25.[151,250,1426,1448]" captionTargetBox="[151,1436,193,1405]" captionTargetId="figure@25.[151,1436,193,1405]" captionTargetPageId="25" captionText="FIGURE 18. Sphaerobelum lachneeis n. sp., holotype (MHNG); A: left anterior telopod, anterior view; B: left anterior telopod, posterior view; C: left posterior telopod, anterior view; D: left posterior telopod, posterior view. Abbreviations: Syn- Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601111/files/figure.png" pageId="10" pageNumber="211">Fig. 18C</figureCitation>
). Antennae green. Anterior telopod suture between telopoditomeres 3 and 4 weak, not visible in anterior view (
<figureCitation id="1379F895E3008C3EA3D5FB25FDCEFB03" box="[431,551,1258,1279]" captionStart="FIGURE 18" captionStartId="25.[151,250,1426,1448]" captionTargetBox="[151,1436,193,1405]" captionTargetId="figure@25.[151,1436,193,1405]" captionTargetPageId="25" captionText="FIGURE 18. Sphaerobelum lachneeis n. sp., holotype (MHNG); A: left anterior telopod, anterior view; B: left anterior telopod, posterior view; C: left posterior telopod, anterior view; D: left posterior telopod, posterior view. Abbreviations: Syn- Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601111/files/figure.png" pageId="10" pageNumber="211">Figs 18A, B</figureCitation>
)..........................................................
<taxonomicName id="4C429F93E3008C3EA696FB25FAB6FAFC" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1260,1375,1258,1280]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="lachneeis" status="sp. nov.">
<emphasis id="B9363802E3008C3EA696FB25FAB6FAFC" box="[1260,1375,1258,1280]" italics="true" pageId="10" pageNumber="211">S. lachneeis</emphasis>
</taxonomicName>
<emphasis id="B9363802E3008C3EA71CFB25FA75FAFC" bold="true" box="[1382,1436,1258,1280]" pageId="10" pageNumber="211">
<taxonomicNameLabel id="A2058579E3008C3EA71CFB25FA75FAFC" box="[1382,1436,1258,1280]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFAC8FA72FAC5" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFAC8FA72FAC5" pageId="10" pageNumber="211">- Tergites covered with short setae. Immovable finger of posterior telopod swollen, but not with such a long lateral extension. Anterior telopod: telopoditomeres 3 and 4 well separated...................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFA8EFA75FA1B" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFA8EFA75FA6C" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFA8EFA75FA6C" pageId="10" pageNumber="211">
11. Podomere 3 of anterior telopod distally with a lateral extension. Posterior telopod: telopoditomere 4 distally with an incurved process overlapping with process of podomere 2. Marginal setae of terga not reaching posterior margin. Tergite posterior margin well-rounded, not tesselated.....................................................
<taxonomicName id="4C429F93E3008C3EA6ECFAB5FA75FA6C" authority="Attems, 1953" authorityName="Attems" authorityYear="1953" box="[1174,1436,1402,1424]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="separatum">
<emphasis id="B9363802E3008C3EA6ECFAB5FAFAFA6C" box="[1174,1299,1402,1424]" italics="true" pageId="10" pageNumber="211">S. separatum</emphasis>
<bibRefCitation id="EFD399E1E3008C3EA761FAB4FA75FA6C" author="Attems, C." box="[1307,1436,1403,1424]" pageId="10" pageNumber="211" pagination="133 - 230" refId="ref23903" refString="Attems, C. (1953) Myriopoden von Indochina, Expedition von Dr. C. Dawydoff (1938 - 1939). Memoires du Museum national d'histoire naturelle, N. S., serie A, 5 (3), 133 - 230." type="journal article" year="1953">Attems, 1953</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDFA57FA75FA1B" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFA57FA75FA1B" pageId="10" pageNumber="211">- Podomere 3 of anterior telopod cylindrical, without lateral extension. Posterior telopod: telopoditomere 4 without incurved process, but with 3 separate spines. Marginal setae of terga reaching posterior margin. Tergite posterior margin tesselated...................................................................................................... 12.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDFA20FA75F9A7" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDFA20FA75F9DD" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDFA20FA75F9DD" pageId="10" pageNumber="211">
12. Antennae with&gt;80 apical cones. Legs: femur
<date id="FFFCC2D0E3008C3EA0E6FA20FD5DF9F8" box="[668,692,1519,1540]" pageId="10" pageNumber="211">2x</date>
longer than wide. Tarsi with 7 or 8 ventral spines. Antennae and legs yellow. Vulval operculum well-rounded, not extending.............................
<taxonomicName id="4C429F93E3008C3EA181F9C4FA75F9DD" authority="Wongthamwanich et al., 2012" authorityName="Wongthamwanich" authorityYear="2012" box="[1019,1436,1547,1569]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="truncatum">
<emphasis id="B9363802E3008C3EA181F9C4FB9FF9DD" box="[1019,1142,1547,1569]" italics="true" pageId="10" pageNumber="211">S. truncatum</emphasis>
<bibRefCitation id="EFD399E1E3008C3EA606F9C3FA75F9DD" author="Wongthamwanich, N. &amp; Panha, S. &amp; Sierwald, P. &amp; Wesener, T. &amp; Thirakhupt, K." box="[1148,1436,1547,1569]" pageId="10" pageNumber="211" pagination="29 - 43" refId="ref26080" refString="Wongthamwanich, N., Panha, S., Sierwald, P., Wesener, T. &amp; Thirakhupt, K. (2012) A new species of the giant pill-millipede genus Sphaerobelum Verhoeff, 1924 from northern Thailand, with an extensive description and molecular characters (Diplopoda: Sphaerotheriida: Zephroniidae). Zootaxa, 3220, 29 - 43" type="journal article" year="2012">
Wongthamwanich
<emphasis id="B9363802E3008C3EA74AF9C3FAB3F9DD" box="[1328,1370,1547,1569]" italics="true" pageId="10" pageNumber="211">et al</emphasis>
., 2012
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDF9E6FA75F9A7" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDF9E6FA75F9A7" pageId="10" pageNumber="211">
- Antennae with &lt;70 apical cones. Legs: femur 1.7 times longer than wide. Tarsi with 710 ventral spines. Antennae and legs orange-red............................................................................
<taxonomicName id="4C429F93E3008C3EA6A8F98AFAB6F9A7" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1234,1375,1605,1627]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="peterjaegeri" status="sp. nov.">
<emphasis id="B9363802E3008C3EA6A8F98AFAB6F9A7" box="[1234,1375,1605,1627]" italics="true" pageId="10" pageNumber="211">S. peterjaegeri</emphasis>
</taxonomicName>
<emphasis id="B9363802E3008C3EA71CF98AFA75F9A7" bold="true" box="[1382,1436,1605,1627]" pageId="10" pageNumber="211">
<taxonomicNameLabel id="A2058579E3008C3EA71CF98AFA75F9A7" box="[1382,1436,1605,1627]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDF9ACFA72F968" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDF9ACFA75F984" blockId="10.[151,1436,217,2032]" box="[151,1436,1635,1656]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDF9ACFA75F984" box="[151,1436,1635,1656]" pageId="10" pageNumber="211">13. Tergites glabrous, at least posterior 2/3 of each tergite....................................................... 14.</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDF9B0FA72F968" blockId="10.[151,1436,217,2032]" box="[151,1435,1663,1685]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDF9B0FA72F968" box="[151,1435,1663,1685]" pageId="10" pageNumber="211">
- Tergites covered with short setae. Dirty olive-brown. Head punctured and densely hirsute........
<taxonomicName id="4C429F93E3008C3EA6E3F9B0FA72F968" authority="Verhoeff, 1924" authorityName="Verhoeff" authorityYear="1924" box="[1177,1435,1663,1685]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="hirsutum">
<emphasis id="B9363802E3008C3EA6E3F9B0FAEFF969" box="[1177,1286,1663,1685]" italics="true" pageId="10" pageNumber="211">S. hirsutum</emphasis>
<bibRefCitation id="EFD399E1E3008C3EA776F9B0FA72F968" author="Verhoeff, K. W." box="[1292,1435,1663,1684]" pageId="10" pageNumber="211" pagination="1 - 142" refId="ref25447" refString="Verhoeff, K. W. (1924) Results of Dr. E. Mjoberg's Swedish scientific expedition to Australia 1910 - 1913. 34. Myriapoda: Diplopoda. Arkif for Zoologi, 16 (5), 1 - 142, pls. 1 - 5." type="journal article" year="1924">Verhoeff, 1924</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B6FCB5E3008C3EA2EDF953FA72F8F4" pageId="10" pageNumber="211">
<paragraph id="8BFDE410E3008C3EA2EDF953FA75F932" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDF953FA75F932" pageId="10" pageNumber="211">
14. Anal shield with a short, or completely reduced locking carina. Posterior telopod: immovable finger, swollen at inner and mesal margin. Telopoditomere 4 at mid-point strongly curved and overlapping immovable finger (
<figureCitation id="1379F895E3008C3EA6F4F976FB0FF932" box="[1166,1254,1721,1742]" captionStart="FIGURE 22" captionStartId="33.[151,250,1922,1944]" captionTargetBox="[168,1421,193,1901]" captionTargetId="figure@33.[166,1421,193,1901]" captionTargetPageId="33" captionText="FIGURE 22. Sphaerobelum splendidum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601119/files/figure.png" pageId="10" pageNumber="211">Fig. 22G</figureCitation>
)............ 15
</keyLead>
</paragraph>
<paragraph id="8BFDE410E3008C3EA2EDF919FA72F8F4" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
<keyLead id="30B34725E3008C3EA2EDF919FA72F8F4" pageId="10" pageNumber="211">
- Anal shield with long locking carina,&gt;
<date id="FFFCC2D0E3008C3EA021F919FD90F917" box="[603,633,1750,1771]" pageId="10" pageNumber="211">3X</date>
length of tergite carina. Posterior telopod: immovable finger, swollen only at inner margin; mesal margin straight.....................................................
<taxonomicName id="4C429F93E3008C3EA6F8F93CFA72F8F4" authority="Verhoeff, 1924" authorityName="Verhoeff" authorityYear="1924" box="[1154,1435,1779,1801]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="clavigerum">
<emphasis id="B9363802E3008C3EA6F8F93CFAEFF8F5" box="[1154,1286,1779,1801]" italics="true" pageId="10" pageNumber="211">S. clavigerum</emphasis>
<bibRefCitation id="EFD399E1E3008C3EA776F93CFA72F8F4" author="Verhoeff, K. W." box="[1292,1435,1779,1800]" pageId="10" pageNumber="211" pagination="1 - 142" refId="ref25447" refString="Verhoeff, K. W. (1924) Results of Dr. E. Mjoberg's Swedish scientific expedition to Australia 1910 - 1913. 34. Myriapoda: Diplopoda. Arkif for Zoologi, 16 (5), 1 - 142, pls. 1 - 5." type="journal article" year="1924">Verhoeff, 1924</bibRefCitation>
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15. Anterior telopod: telopoditomeres 3 and 4 clearly separated (
<figureCitation id="1379F895E3008C3EA152F8DFFC99F8D9" box="[808,880,1808,1829]" captionStart="FIGURE 21" captionStartId="30.[151,250,1950,1972]" captionTargetBox="[174,1418,193,1936]" captionTargetId="figure@30.[174,1418,193,1936]" captionTargetPageId="30" captionText="FIGURE 21. Sphaerobelum nigrum n. sp., holotype (SMF); A: coxa of second left leg with gonopore; B: right anterior telopod, anterior view; C: right anterior telopod, posterior view; D: right anterior telopod, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601117/files/figure.png" pageId="10" pageNumber="211">Figs 21</figureCitation>
BD). Antennae and legs brown.........................................................................................................
<taxonomicName id="4C429F93E3008C3EA6AFF8E3FAB6F8BE" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1237,1375,1836,1858]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="splendidum" status="sp. nov.">
<emphasis id="B9363802E3008C3EA6AFF8E3FAB6F8BE" box="[1237,1375,1836,1858]" italics="true" pageId="10" pageNumber="211">S. splendidum</emphasis>
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<taxonomicNameLabel id="A2058579E3008C3EA71CF8E2FA75F8BF" box="[1382,1436,1837,1859]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
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- Anterior telopod: telopoditomeres 3 and 4 separation only partly visible in anterior view (
<figureCitation id="1379F895E3008C3EA641F885FB7CF8A3" box="[1083,1173,1866,1887]" captionStart="FIGURE 22" captionStartId="33.[151,250,1922,1944]" captionTargetBox="[168,1421,193,1901]" captionTargetId="figure@33.[166,1421,193,1901]" captionTargetPageId="33" captionText="FIGURE 22. Sphaerobelum splendidum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601119/files/figure.png" pageId="10" pageNumber="211">Fig. 22D</figureCitation>
). Antennae and legs red..................................................................................................... 16
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<paragraph id="8BFDE410E3008C3EA2EDF84BFA75F84A" blockId="10.[151,1436,217,2032]" pageId="10" pageNumber="211">
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16. Process of telopoditomere 2 projecting as high as basal 1/3 of telopoditomere 3+4 (
<figureCitation id="1379F895E3008C3EA671F84BFB8BF865" box="[1035,1122,1924,1945]" captionStart="FIGURE 22" captionStartId="33.[151,250,1922,1944]" captionTargetBox="[168,1421,193,1901]" captionTargetId="figure@33.[166,1421,193,1901]" captionTargetPageId="33" captionText="FIGURE 22. Sphaerobelum splendidum n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601119/files/figure.png" pageId="10" pageNumber="211">Fig. 22F</figureCitation>
). Anal shield with a single short locking carinae. Palpi of gnathochilarium, sensory cones located in single field........................
<taxonomicName id="4C429F93E3008C3EA689F86FFAB6F84A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1267,1375,1952,1974]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="laoticum" status="sp. nov.">
<emphasis id="B9363802E3008C3EA689F86FFAB6F84A" box="[1267,1375,1952,1974]" italics="true" pageId="10" pageNumber="211">S. laoticum</emphasis>
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<emphasis id="B9363802E3008C3EA71CF86FFA75F84A" bold="true" box="[1382,1436,1952,1974]" pageId="10" pageNumber="211">
<taxonomicNameLabel id="A2058579E3008C3EA71CF86FFA75F84A" box="[1382,1436,1952,1974]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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- Process of telopoditomere 2 projecting as high as basal 3/4 of telopoditomere 3+4 (
<figureCitation id="1379F895E3008C3EA184F871FBBFF82F" box="[1022,1110,1982,2003]" captionStart="FIGURE 25" captionStartId="41.[151,250,1885,1907]" captionTargetBox="[207,1378,195,1868]" captionTargetId="figure@41.[207,1378,195,1868]" captionTargetPageId="41" captionText="FIGURE 25. Sphaerobelum schwendingeri n. sp. AC female paratype (ZMUC), DG male holotype (MHNG). A: coxa of first right leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: right posterior telopod, anterior view; G: right posterior telopod, posterior view. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601125/files/figure.png" pageId="10" pageNumber="211">Fig. 25E</figureCitation>
). Anal shield without locking carinae. Palpi of gnathochilarium, sensory cones located in several clusters........................
<taxonomicName id="4C429F93E3008C3EA6C1F815FAB6F80C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1211,1375,2010,2032]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
<emphasis id="B9363802E3008C3EA6C1F815FAB6F80C" box="[1211,1375,2010,2032]" italics="true" pageId="10" pageNumber="211">S. schwendingeri</emphasis>
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