398 lines
63 KiB
XML
398 lines
63 KiB
XML
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<mods:titleInfo id="13228125F2294BDFBE8FC276EA1E0419">
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<mods:title id="5AD82D0B448588E1120AC1DEDB8605C8">Integrative taxonomic revision of the African taxa of the Racotis Moore, 1887 generic complex (Lepidoptera, Geometridae, Ennominae, Boarmiini)</mods:title>
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<mods:namePart id="3559797A3F9A031CD57841DD9D546D76">László, Gyula M.</mods:namePart>
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<mods:namePart id="22E28DB19A32F2DE3BFC7510C1ABF359">Hausmann, Axel</mods:namePart>
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<mods:namePart id="DFE9AB5D9206FBB89A617AC83FFBD7E1">Karisch, Timm</mods:namePart>
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<mods:title id="AAAEAFEF024620B88D606553C31E1C60">Zootaxa</mods:title>
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<mods:part id="4617231893109707647CA720F13DF3BE">
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<mods:date id="85E75CB899B2AC4A01676EB8BD8914B9">2023</mods:date>
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<mods:number id="B12C0C87D1AD8BBF806B11259D1414F9">2023-06-26</mods:number>
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<mods:number id="80C44A416A666E6378239E31359100B4">5308</mods:number>
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<mods:url id="3411B2505DAC82F5F11179683370018E">http://dx.doi.org/10.11646/zootaxa.5308.1.1</mods:url>
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<mods:classification id="DC5FF3BA17339364CD19C6A4411DED08">journal article</mods:classification>
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<subSubSection id="C358A6B65D0CFF8D57F555B6FE3BFD49" box="[151,412,690,717]" pageId="49" pageNumber="50" type="nomenclature">
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<paragraph id="8BFDF53D5D0CFF8D57F555B6FE3BFD49" blockId="49.[151,412,690,717]" box="[151,412,690,717]" pageId="49" pageNumber="50">
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<heading id="D0B542515D0CFF8D57F555B6FE3BFD49" bold="true" box="[151,412,690,717]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
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<emphasis id="B936292F5D0CFF8D57F555B6FE3BFD49" bold="true" box="[151,412,690,717]" pageId="49" pageNumber="50">
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<taxonomicName id="4C428EBE5D0CFF8D57F555B6FEB1FD49" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,278,690,717]" family="Geometridae" genus="Zebracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" rank="genus" status="subgen. nov.">
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<emphasis id="B936292F5D0CFF8D57F555B6FEB1FD49" bold="true" box="[151,278,690,717]" italics="true" pageId="49" pageNumber="50">Zebracotis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20594545D0CFF8D567C55B7FE3BFD49" box="[286,412,691,717]" pageId="49" pageNumber="50" rank="subGenus">subgen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358A6B65D0CFF8D57F555FCFDF3FC8B" box="[151,596,759,784]" pageId="49" pageNumber="50" type="type_taxon">
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<paragraph id="8BFDF53D5D0CFF8D57F555FCFDF3FC8B" blockId="49.[151,596,759,784]" box="[151,596,759,784]" pageId="49" pageNumber="50">
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<typeStatus id="54F94B9F5D0CFF8D57F555FCFF6EFC94" box="[151,201,760,784]" pageId="49" pageNumber="50">Type</typeStatus>
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species:
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<taxonomicName id="4C428EBE5D0CFF8D564455FCFDF3FC8B" ID-CoL="9483P" authority="Warren, 1899" authorityName="Warren" authorityYear="1899" box="[294,596,759,783]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="zebrina">
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<emphasis id="B936292F5D0CFF8D564455FCFE62FC8B" box="[294,453,759,783]" italics="true" pageId="49" pageNumber="50">Racotis zebrina</emphasis>
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<bibRefCitation id="EFD388CC5D0CFF8D56A955FCFDF3FC8B" author="Warren, W." box="[459,596,759,783]" pageId="49" pageNumber="50" pagination="1 - 66" refId="ref85194" refString="Warren, W. (1899) New species and genera of the families Drepanulidae, Thyrididae, Uraniidae, Epiplemidae, and Geometridae from the Old-World regions. Novitates Zoologicae, 6 (1), 1 - 66." type="journal article" year="1899">Warren, 1899</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358A6B65D0CFF8D57F55439FD8CF8A7" pageId="49" pageNumber="50" type="discussion">
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<paragraph id="8BFDF53D5D0CFF8D57F55439FD8CF8A7" blockId="49.[151,1437,829,1827]" pageId="49" pageNumber="50">
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<emphasis id="B936292F5D0CFF8D57F55439FEC5FCD3" bold="true" box="[151,354,829,855]" pageId="49" pageNumber="50">Genetic analyses</emphasis>
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(Text fig. 3). The subgenus was recovered as a well-supported sister clade of the subgenus
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<taxonomicName id="4C428EBE5D0CFF8D57F55465FEB1FCFF" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,278,865,891]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="B936292F5D0CFF8D57F55465FEB1FCFF" box="[151,278,865,891]" italics="true" pageId="49" pageNumber="50">Afroracotis</emphasis>
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</taxonomicName>
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with high support in both analyses (BS: 55; PP: 97). The DNA barcode-based infrageneric classification of the subgenus, however, is rather inconsistent often conflicting with the clearly distinguished morphotaxa. Based on certain constant morphological characters discussed below, the subgenus contains ten taxa (nine species and one subspecies), while only seven BINs in the complex were assigned in BOLD. The topologies of the BI and ML trees are nearly identical and both are rather incongruent with the morphology-based taxon delineations, where a single morphological taxon is often split into two separate genetic clusters without any diagnostic characters. Such an example is
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<taxonomicName id="4C428EBE5D0CFF8D56E0533EFE41FBD7" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[386,486,1081,1108]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="milesi" status="sp. nov.">
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<emphasis id="B936292F5D0CFF8D56E0533EFE41FBD7" box="[386,486,1081,1108]" italics="true" pageId="49" pageNumber="50">A. milesi</emphasis>
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</taxonomicName>
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<emphasis id="B936292F5D0CFF8D5692533DFD95FBD7" bold="true" box="[496,562,1081,1107]" pageId="49" pageNumber="50">
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<taxonomicNameLabel id="A20594545D0CFF8D5692533DFD95FBD7" box="[496,562,1081,1107]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, which was recovered as paraphyletic: in a cluster of exclusively Gabonese specimens BIN-sharing with
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<taxonomicName id="4C428EBE5D0CFF8D5683535AFDF2FBF3" authorityName="Warren" authorityYear="1899" box="[481,597,1117,1144]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="zebrina">
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<emphasis id="B936292F5D0CFF8D5683535AFDF2FBF3" box="[481,597,1117,1144]" italics="true" pageId="49" pageNumber="50">A. zebrina</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C428EBE5D0CFF8D5506535AFD44FBF3" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[612,739,1117,1144]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="chaineyi" status="sp. nov.">
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<emphasis id="B936292F5D0CFF8D5506535AFD44FBF3" box="[612,739,1117,1144]" italics="true" pageId="49" pageNumber="50">A. chaineyi</emphasis>
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</taxonomicName>
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<emphasis id="B936292F5D0CFF8D55895359FC8CFBF3" bold="true" box="[747,811,1117,1143]" pageId="49" pageNumber="50">
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<taxonomicNameLabel id="A20594545D0CFF8D55895359FC8CFBF3" box="[747,811,1117,1143]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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and
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<taxonomicName id="4C428EBE5D0CFF8D5401535AFB9DFBF3" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[867,1082,1117,1144]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subSpecies" species="lydiae" subSpecies="orientalis">
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<emphasis id="B936292F5D0CFF8D5401535AFB9DFBF3" box="[867,1082,1117,1144]" italics="true" pageId="49" pageNumber="50">A. lydiae orientalis</emphasis>
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</taxonomicName>
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<emphasis id="B936292F5D0CFF8D53205359FB29FBF3" bold="true" box="[1090,1166,1117,1143]" pageId="49" pageNumber="50">ssp. n.</emphasis>
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and in a distinct cluster with its own BIN URI comprising
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<taxonomicName id="4C428EBE5D0CFF8D554D5386FD33FB1F" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[559,660,1153,1180]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="milesi">
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<emphasis id="B936292F5D0CFF8D554D5386FD33FB1F" box="[559,660,1153,1180]" italics="true" pageId="49" pageNumber="50">A. milesi</emphasis>
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</taxonomicName>
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samples from
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<collectingCountry id="F355B5AD5D0CFF8D54215385FC3CFB1F" box="[835,923,1153,1179]" name="Zambia" pageId="49" pageNumber="50">Zambia</collectingCountry>
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,
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<collectingCountry id="F355B5AD5D0CFF8D54C85385FBB8FB1F" box="[938,1055,1153,1179]" name="Cameroon" pageId="49" pageNumber="50">Cameroon</collectingCountry>
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and
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<collectingCountry id="F355B5AD5D0CFF8D533E5385FB0DFB1F" box="[1116,1194,1153,1179]" name="Gabon" pageId="49" pageNumber="50">Gabon</collectingCountry>
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. Another example of such incongruency between genetics and morphology is
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<taxonomicName id="4C428EBE5D0CFF8D547753A2FCDFFB3B" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[789,888,1189,1216]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="lydiae" status="sp. nov.">
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<emphasis id="B936292F5D0CFF8D547753A2FCDFFB3B" box="[789,888,1189,1216]" italics="true" pageId="49" pageNumber="50">A. lydiae</emphasis>
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</taxonomicName>
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<emphasis id="B936292F5D0CFF8D54E253A1FC67FB3B" bold="true" box="[896,960,1189,1215]" pageId="49" pageNumber="50">
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<taxonomicNameLabel id="A20594545D0CFF8D54E253A1FC67FB3B" box="[896,960,1189,1215]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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which is morphologically subdivided into a West (nominotypical taxon) and East African subspecies (
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<taxonomicName id="4C428EBE5D0CFF8D545653CEFBADFB67" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[820,1034,1225,1252]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subSpecies" species="lydiae" subSpecies="orientalis">
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<emphasis id="B936292F5D0CFF8D545653CEFBADFB67" box="[820,1034,1225,1252]" italics="true" pageId="49" pageNumber="50">A. lydiae orientalis</emphasis>
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</taxonomicName>
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<emphasis id="B936292F5D0CFF8D537053CDFBFBFB67" bold="true" box="[1042,1116,1225,1251]" pageId="49" pageNumber="50">ssp. n.</emphasis>
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), however, the West African subspecies is recovered as a genetically largely separate cluster with its own BIN URI, whereas the East African subspecies is BIN-sharing with the predominantly West African
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<taxonomicName id="4C428EBE5D0CFF8D543A5216FC6EFAAF" authorityName="Warren" authorityYear="1899" box="[856,969,1297,1324]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="zebrina">
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<emphasis id="B936292F5D0CFF8D543A5216FC6EFAAF" box="[856,969,1297,1324]" italics="true" pageId="49" pageNumber="50">A. zebrina</emphasis>
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</taxonomicName>
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, the mainly central African
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<taxonomicName id="4C428EBE5D0CFF8D53985216FAFDFAAF" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[1274,1370,1297,1324]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="milesi" status="sp. nov.">
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<emphasis id="B936292F5D0CFF8D53985216FAFDFAAF" box="[1274,1370,1297,1324]" italics="true" pageId="49" pageNumber="50">A. milesi</emphasis>
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</taxonomicName>
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<emphasis id="B936292F5D0CFF8D523D5215FA3AFAAF" bold="true" box="[1375,1437,1297,1323]" pageId="49" pageNumber="50">
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<taxonomicNameLabel id="A20594545D0CFF8D523D5215FA3AFAAF" box="[1375,1437,1297,1323]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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and the more widespread
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<taxonomicName id="4C428EBE5D0CFF8D56D65232FDB1FACB" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[436,534,1333,1360]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
|
||
<emphasis id="B936292F5D0CFF8D56D65232FDB1FACB" box="[436,534,1333,1360]" italics="true" pageId="49" pageNumber="50">A. smithi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B936292F5D0CFF8D557F5231FDFCFACB" bold="true" box="[541,603,1333,1359]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="A20594545D0CFF8D557F5231FDFCFACB" box="[541,603,1333,1359]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. The only taxa showing genetic features being congruent with morphology are
|
||
<taxonomicName id="4C428EBE5D0CFF8D57A2525EFD79FAF7" authority="(Chainey & Karisch, 2017)" authorityName="László & Hausmann & Karisch" authorityYear="2023" baseAuthorityName="Chainey & Karisch" baseAuthorityYear="2017" box="[192,734,1369,1396]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="madagascariensis">
|
||
<emphasis id="B936292F5D0CFF8D57A2525EFE0FFAF7" box="[192,424,1369,1396]" italics="true" pageId="49" pageNumber="50">A. madagascariensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD388CC5D0CFF8D56D4525DFD70FAF7" author="Chainey, J. & Karisch, T." box="[438,727,1369,1395]" pageId="49" pageNumber="50" pagination="49 - 53" refId="ref83017" refString="Chainey, J. & Karisch, T. (2017) A new species of Racotis Moore, [1887] (Lepidoptera: Geometridae: Ennominae) from Madagascar. Metamorphosis, 28, 49 - 53." type="journal article" year="2017">Chainey & Karisch, 2017</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
which is recovered as a sister clade to all other
|
||
<taxonomicName id="4C428EBE5D0CFF8D538D525DFAC0FAF7" box="[1263,1383,1369,1395]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D0CFF8D538D525DFAC0FAF7" box="[1263,1383,1369,1395]" italics="true" pageId="49" pageNumber="50">Zebracotis</emphasis>
|
||
</taxonomicName>
|
||
taxa in both analyses (BS: 67%, PP: 100%),
|
||
<taxonomicName id="4C428EBE5D0CFF8D5539527AFD64FA13" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[603,707,1405,1432]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ochsei" status="sp. nov.">
|
||
<emphasis id="B936292F5D0CFF8D5539527AFD64FA13" box="[603,707,1405,1432]" italics="true" pageId="49" pageNumber="50">A. ochsei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B936292F5D0CFF8D55A85279FCADFA13" bold="true" box="[714,778,1405,1431]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="A20594545D0CFF8D55A85279FCADFA13" box="[714,778,1405,1431]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C428EBE5D0CFF8D547B527AFC2BFA13" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[793,908,1405,1432]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="takanoi" status="sp. nov.">
|
||
<emphasis id="B936292F5D0CFF8D547B527AFC2BFA13" box="[793,908,1405,1432]" italics="true" pageId="49" pageNumber="50">A. takanoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B936292F5D0CFF8D54F65279FC72FA13" bold="true" box="[916,981,1405,1431]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="A20594545D0CFF8D54F65279FC72FA13" box="[916,981,1405,1431]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C428EBE5D0CFF8D536E527AFB2CFA13" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[1036,1163,1405,1432]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="helicalis" status="sp. nov.">
|
||
<emphasis id="B936292F5D0CFF8D536E527AFB2CFA13" box="[1036,1163,1405,1432]" italics="true" pageId="49" pageNumber="50">A. helicalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B936292F5D0CFF8D53F15279FB74FA13" bold="true" box="[1171,1235,1405,1431]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="A20594545D0CFF8D53F15279FB74FA13" box="[1171,1235,1405,1431]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, although the last one clusters as a sister species to
|
||
<taxonomicName id="4C428EBE5D0CFF8D557352A6FD67FA3F" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[529,704,1441,1468]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subSpecies" species="lydiae" subSpecies="lydiae">
|
||
<emphasis id="B936292F5D0CFF8D557352A6FD67FA3F" box="[529,704,1441,1468]" italics="true" pageId="49" pageNumber="50">A. lydiae lydiae</emphasis>
|
||
</taxonomicName>
|
||
despite their largely dissimilar genital morphology. It is difficult to reliably establish the reason of such an extent of conflicts between genetics and morphology in the
|
||
<taxonomicName id="4C428EBE5D0CFF8D524652C1FA3BFA5B" box="[1316,1436,1477,1503]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D0CFF8D524652C1FA3BFA5B" box="[1316,1436,1477,1503]" italics="true" pageId="49" pageNumber="50">Zebracotis</emphasis>
|
||
</taxonomicName>
|
||
subgenus. Seemingly, DNA barcodes have limited taxonomic relevance in this group, possibly due to repeated genetic introgression. Alternatively, the members of the subgenus may be evolutionary very young taxa “in statu nascendi”, where the speciation has been manifested in an early divergence in genital morphology before a clear differentiation in the external morphology and COI DNA sequences could have taken place. It is also worth noting that the single COI-5P gene region used in the genetic analyses is relatively short and an in-depth genetic analysis based on multiple gene regions (including nDNA) would presumably result in a better resolution of the phylogeny. Overall, as the tree topology of the
|
||
<taxonomicName id="4C428EBE5D0CFF8D554F51C5FD02F95F" box="[557,677,1729,1755]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D0CFF8D554F51C5FD02F95F" box="[557,677,1729,1755]" italics="true" pageId="49" pageNumber="50">Zebracotis</emphasis>
|
||
</taxonomicName>
|
||
subgenus is not entirely congruent with the morphological species concept, the taxa in this subgenus have been delimited based primarily on the remarkable and constant differences observed in the genital morphology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3DA5B55D0FFF8E57FA509CFF45F87C" ID-DOI="http://doi.org/10.5281/zenodo.8110743" ID-Zenodo-Dep="8110743" httpUri="https://zenodo.org/record/8110743/files/figure.png" pageId="50" pageNumber="51" startId="50.[152,224,1944,1968]" subCaptionStartIDs="50.[232,335,1943,1967]" subCaptionStarts="FIGURE 3" targetBox="[297,1092,192,1899]" targetPageId="50">
|
||
<paragraph id="8BFDF53D5D0FFF8E57FA509CFF45F87C" blockId="50.[151,1436,1943,2040]" pageId="50" pageNumber="51">
|
||
<emphasis id="B936292F5D0FFF8E57FA509CFECDF82B" bold="true" box="[152,362,1943,1968]" pageId="50" pageNumber="51">
|
||
TEXT
|
||
<figureCitation id="1379E9B85D0FFF8E578A5093FEC1F82B" box="[232,358,1943,1967]" captionStart="FIGURES 1–20" captionStartId="17.[151,264,1656,1680]" captionTargetBox="[152,1431,183,1623]" captionTargetId="figure-17@17.[151,1436,179,1630]" captionTargetPageId="17" captionText="FIGURES 1–20. Adults, dorsal view. 1. Racotis boarmiaria, LT, ♁, India (NHMUK); 2. Afroracotis squalida squalida, HT of Ophthalmodes squalida, ♀, Nigeria (NHMUK); 3. Ibidem, HT of Boarmia uhligi, ♀, Tanzania (MfN); 4. Ibidem, ♁, Cameroon (ANHRT); 5. Ibidem, ♁, Zambia (ANHRT); 6. Ibidem, ♁, Togo (ANHRT); 7. Ibidem, ♁, Gabon (ANHRT); 8. Ibidem, ♁, Ethiopia (RCDS); 9. Ibidem, ♀, Ethiopia (RCDS); 10. Ibidem, ♀, Cameroon (ANHRT); 11. A. s. voeltzkowii, LT of Boarmia voeltzkowii, ♁, Madagascar (MfN); 12. Ibidem, ♁, Comoros (ZSM); 13. Ibidem, ♁, Madagascar (NHMUK); 14. Ibidem, ♀, Madagascar (NHMUK); 15. A. s. thomensis ssp. n., HT, ♁, São Tomé (ANHRT); 16. Ibidem, PT, ♀, São Tomé (ANHRT); 17. Ibidem, ♀, PT, São Tomé (NHMUK); 18. A. incompletaria incompletaria, ♀, Réunion (ZSM); 19. A. i. atrilunaria, HT of Boarmia atrilunaria, ♀, Mauritius (NHMUK); 20. Ibidem, ♀, Mauritius (ANHRT)." figureDoi="http://doi.org/10.5281/zenodo.8080931" httpUri="https://zenodo.org/record/8080931/files/figure.png" pageId="50" pageNumber="51">FIGURE 3</figureCitation>
|
||
.
|
||
</emphasis>
|
||
ML tree of the subgenus
|
||
<taxonomicName id="4C428EBE5D0FFF8E551B509CFD42F834" box="[633,741,1944,1968]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D0FFF8E551B509CFD42F834" box="[633,741,1944,1968]" italics="true" pageId="50" pageNumber="51">Zebracotis</emphasis>
|
||
</taxonomicName>
|
||
based on analysis of COI barcodes. Bootstrap values are indicated above, posterior probability values below the branches. Samples which have not been confirmed by dissection are labelled as ‘indet’.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C358A6B65D0EFF8F57A55792FA9AFD75" pageId="51" pageNumber="52" type="diagnosis">
|
||
<paragraph id="8BFDF53D5D0EFF8F57A55792FA9AFD75" blockId="51.[151,1437,150,753]" pageId="51" pageNumber="52">
|
||
<emphasis id="B936292F5D0EFF8F57A55792FEE5FF34" bold="true" box="[199,322,150,176]" pageId="51" pageNumber="52">Diagnosis.</emphasis>
|
||
The species of the subgenus are characterized by the broad, triangular forewing, the dark greyish-green colouration of densely speckled scales on the upperside, the well-defined, ovoidal discal spots, and the variably contrasting row of round black patches representing the subterminal line. As the wing pattern varies largely both infra- and interspecifically, the facies of most of the species do not provide reliable distinctive features for species-level identification. In the males of all members of the subgenus, the third abdominal sternite bears a small tuft of fine setae, a character which is also found in
|
||
<taxonomicName id="4C428EBE5D0EFF8F55F0564FFCB5FEE1" baseAuthorityName="Butler" baseAuthorityYear="1878" box="[658,786,331,357]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="squalida">
|
||
<emphasis id="B936292F5D0EFF8F55F0564FFCB5FEE1" box="[658,786,331,357]" italics="true" pageId="51" pageNumber="52">A. squalida</emphasis>
|
||
</taxonomicName>
|
||
and in the monotypic subgenus
|
||
<taxonomicName id="4C428EBE5D0EFF8F531D564FFABBFEE1" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[1151,1308,331,357]" pageId="51" pageNumber="52" rank="subGenus" status="subgen. nov." subGenus="Sokokeracotis">
|
||
<emphasis id="B936292F5D0EFF8F531D564FFABBFEE1" box="[1151,1308,331,357]" italics="true" pageId="51" pageNumber="52">Sokokeracotis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B936292F5D0EFF8F5241564FFA3AFEE1" bold="true" box="[1315,1437,331,357]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A20594545D0EFF8F5241564FFA3AFEE1" box="[1315,1437,331,357]" pageId="51" pageNumber="52" rank="subGenus">subgen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
within
|
||
<taxonomicName id="4C428EBE5D0EFF8F5784566BFEC2FE0D" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[230,357,367,393]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B936292F5D0EFF8F5784566BFEC2FE0D" box="[230,357,367,393]" italics="true" pageId="51" pageNumber="52">Afroracotis</emphasis>
|
||
</taxonomicName>
|
||
, however, it is also present in the sister genus
|
||
<taxonomicName id="4C428EBE5D0EFF8F5413566BFC4BFE0D" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[881,1004,367,393]" class="Insecta" family="Geometridae" genus="Chorocotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B936292F5D0EFF8F5413566BFC4BFE0D" box="[881,1004,367,393]" italics="true" pageId="51" pageNumber="52">Chorocotis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B936292F5D0EFF8F5496566BFBE4FE0D" bold="true" box="[1012,1091,367,393]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A20594545D0EFF8F5496566BFBE4FE0D" box="[1012,1091,367,393]" pageId="51" pageNumber="52" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. The diagnostic male genitalia characters of the subgenus are as follows:
|
||
<taxonomicName id="4C428EBE5D0EFF8F55E25697FD5FFE29" box="[640,760,403,429]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D0EFF8F55E25697FD5FFE29" box="[640,760,403,429]" italics="true" pageId="51" pageNumber="52">Zebracotis</emphasis>
|
||
</taxonomicName>
|
||
has a clasping apparatus similar to that of
|
||
<taxonomicName id="4C428EBE5D0EFF8F53885697FA32FE29" box="[1258,1429,403,429]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="105" phylum="Arthropoda" rank="subGenus" subGenus="Herbuloracotis">
|
||
<emphasis id="B936292F5D0EFF8F53885697FA32FE29" box="[1258,1429,403,429]" italics="true" pageId="51" pageNumber="52">Herbuloracotis</emphasis>
|
||
</taxonomicName>
|
||
, but it has a longer uncus, a medially inverse V-shaped gnathos without a shield-like medial plate, a ventro-medially more concave valva, and a considerably more elongate juxta, medially tapered into a narrow distal projection. The vesica of the
|
||
<taxonomicName id="4C428EBE5D0EFF8F565156FBFE0CFD9D" box="[307,427,511,537]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D0EFF8F565156FBFE0CFD9D" box="[307,427,511,537]" italics="true" pageId="51" pageNumber="52">Zebracotis</emphasis>
|
||
</taxonomicName>
|
||
species bears highly distinctive characters allowing reliable species-level identification. It is characterized by the medium long and moderately inflated main vesica chamber, in most species with a well-developed diverticulum with an apical cornutus of specific shape and a group of cornuti erected distally at the base of the vesica ejaculatorius. The shared characters of the
|
||
<taxonomicName id="4C428EBE5D0EFF8F559D556FFCD0FD01" box="[767,887,619,645]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D0EFF8F559D556FFCD0FD01" box="[767,887,619,645]" italics="true" pageId="51" pageNumber="52">Zebracotis</emphasis>
|
||
</taxonomicName>
|
||
species in the female genitalia are the sclerotised, broad cup-shaped antrum, the absence of the membranous ductus bursae replaced by the variably elongate heavily sclerotised cervix bursae which is fully merged into the corpus bursae forming a sclerotised posterior section. The lack of true ductus bursae clearly separates the
|
||
<taxonomicName id="4C428EBE5D0EFF8F55CA55D3FC87FD75" box="[680,800,727,753]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D0EFF8F55CA55D3FC87FD75" box="[680,800,727,753]" italics="true" pageId="51" pageNumber="52">Zebracotis</emphasis>
|
||
</taxonomicName>
|
||
subgenus from all other lineages of
|
||
<taxonomicName id="4C428EBE5D0EFF8F53DA55D3FA90FD75" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[1208,1335,727,753]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B936292F5D0EFF8F53DA55D3FA90FD75" box="[1208,1335,727,753]" italics="true" pageId="51" pageNumber="52">Afroracotis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358A6B65D0EFF8857F55446FEE7FE55" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
|
||
<paragraph id="8BFDF53D5D0EFF8F57F55446FE8FFCD8" blockId="51.[151,296,834,860]" box="[151,296,834,860]" pageId="51" pageNumber="52">
|
||
<emphasis id="B936292F5D0EFF8F57F55446FE8FFCD8" bold="true" box="[151,296,834,860]" pageId="51" pageNumber="52">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D0EFF8F57F5548FFE20F9F1" blockId="51.[151,1437,906,2013]" pageId="51" pageNumber="52">
|
||
<emphasis id="B936292F5D0EFF8F57F5548FFDC2FC20" bold="true" box="[151,613,906,933]" pageId="51" pageNumber="52">External features of body and wings</emphasis>
|
||
(
|
||
<figureCitation id="1379E9B85D0EFF8F5518548FFCA8FC21" box="[634,783,907,933]" captionStart-0="FIGURES 81–100" captionStart-1="FIGURES 101–120" captionStartId-0="53.[151,264,1614,1638]" captionStartId-1="65.[151,264,1643,1667]" captionTargetBox-0="[176,1414,188,1584]" captionTargetBox-1="[185,1402,188,1614]" captionTargetId-0="figure-17@53.[167,1421,181,1590]" captionTargetId-1="figure-17@65.[176,1412,181,1620]" captionTargetPageId-0="53" captionTargetPageId-1="65" captionText-0="FIGURES 81–100. Adults, dorsal view. 81. Afroracotis zebrina, LT of Racotis zebrina, ♁, Nigeria (NHMUK); 82. Ibidem, ♁, Guinea (ANHRT); 83. Ibidem, ♀, Ivory Coast (ANHRT); 84. Ibidem, ♀, Liberia (ANHRT); 85. A. chaineyi sp. n., HT, ♁, Gabon (ANHRT); 86. Ibidem, PT, ♁, Cameroon (ZSM); 87. Ibidem, PT, ♁, Zambia (ANHRT); 88. Ibidem, PT, ♀, Guinea (ANHRT); 89. A. lydiae lydiae sp. n., HT, ♁, Liberia (ANHRT); 90. Ibidem, PT, ♁, Cameroon (ZSM); 91. Ibidem, PT, ♀, Guinea (ANHRT); 92. Ibidem, PT, ♀, Bioko (ZSM); 93. A. lydiae orientalis ssp. n., HT, ♁, Tanzania (ANHRT); 94. Ibidem, PT, ♁, Mozambique (ANHRT); 95. Ibidem, PT, ♀, Tanzania (ANHRT); 96. Ibidem, PT, ♀, Mozambique (ANHRT); 97. A. madagascariensis, HT of Racotis madagascariensis, ♁, Madagascar (NHMUK); 98. Ibidem, PT, ♁, Madagascar (NHMUK); 99. Ibidem, PT, ♀, Madagascar (NHMUK); 100. Ibidem, ♀, Madagascar (ANHRT)." captionText-1="FIGURES 101–120. Adults, dorsal view (all in coll. ANHRT, unless otherwise stated). 101. A. smithi sp. n., HT, ♁, Zambia; 102. Ibidem, PT, ♁, Gabon; 103. Ibidem, PT, ♀, Zambia; 104. Ibidem, PT, ♀, Cameroon (ZSM); 105. A. milesi sp. n., HT, ♁, Gabon; 106. Ibidem, PT, ♁, Cameroon; 107. Ibidem, PT, ♀, Cameroon; 108. Ibidem, PT, ♁, Zambia; 109. A. ochsei sp. n., HT, ♁, Gabon; 110. Ibidem, PT, ♁, Gabon; 111. Ibidem, PT, ♀, Gabon; 112. Ibidem, PT, ♁, Gabon; 113. A. helicalis sp. n., HT, ♁, Gabon; 114. Ibidem, PT, ♁, Gabon; 115. Ibidem, PT, ♀, Gabon; 116. A. takanoi sp. n., HT, ♁, Sierra Leone; 117. Ibidem, PT, ♁, Guinea; 118. Ibidem, PT, ♁, Liberia; 119. Ibidem, PT, ♀, Liberia; 120. A. staudei sp. n., HT, ♁, Kenya (RCHS)." figureDoi-0="http://doi.org/10.5281/zenodo.8080939" figureDoi-1="http://doi.org/10.5281/zenodo.8080943" httpUri-0="https://zenodo.org/record/8080939/files/figure.png" httpUri-1="https://zenodo.org/record/8080943/files/figure.png" pageId="51" pageNumber="52">Figs 81–119</figureCitation>
|
||
,
|
||
<figureCitation id="1379E9B85D0EFF8F5442548EFC20FC21" box="[800,903,906,933]" captionStart-0="FIGURES 171–190" captionStart-1="FIGURES 191–210" captionStartId-0="80.[151,264,1643,1667]" captionStartId-1="81.[151,264,1641,1665]" captionTargetBox-0="[182,1405,190,1613]" captionTargetBox-1="[151,1433,188,1610]" captionTargetId-0="figure-21@80.[175,1413,181,1619]" captionTargetId-1="figure-17@81.[151,1436,181,1617]" captionTargetPageId-0="80" captionTargetPageId-1="81" captionText-0="FIGURES 171–190. Adults, ventral view. 171. Afroracotis atriclava, ST of Cleora atriclava, ♁, Madagascar (ZSM); 172. Ibidem, ST, ♀, Madagascar (ZSM); 173. A. turlini sp. n., PT, ♁, Rwanda (ZSM); 174. Ibidem, PT, ♀, Rwanda (ZSM); 175. A. zebrina, LT of Racotis zebrina, ♁, Nigeria (NHMUK); 176. A. zebrina, ♁, Guinea (ANHRT); 177. Ibidem, ♁, Liberia (ANHRT); 178. Ibidem, ♀, Ivory Coast (ANHRT); 179. A. chaineyi sp. n., HT, ♁, Gabon (ANHRT); 180. Ibidem, PT, ♁, Cameroon (ZSM); 181. Ibidem, PT, ♀, Bioko (ZSM); 182. A. lydiae lydiae sp. n., PT, ♁, Liberia (ANHRT); 183. Ibidem, PT, ♀, Guinea (ANHRT); 184. A. lydiae orientalis ssp. n., HT, ♁, Tanzania (ANHRT); 185. Ibidem, PT, ♀, Tanzania (ANHRT); 186. A. madagascariensis, ♁, Madagascar (NHMUK); 187. Ibidem, ♀, Madagascar (ANHRT); 188. A. smithi sp. n., HT, ♁, Zambia (ANHRT); 189. Ibidem, PT, ♀, Zambia (ANHRT); 190. Ibidem, PT, ♀, Cameroon (ZSM)." captionText-1="FIGURES 191–210. Adults, ventral view (all in coll. ANHRT, unless otherwise stated). 191. Afroracotis milesi sp. n., HT, ♁, Gabon; 192. Ibidem, PT, ♁, Cameroon; 193. Ibidem, PT, ♁, Zambia; 194. Ibidem, PT, ♀, Gabon; 195. A. ochsei sp. n., HT, ♁, Gabon; 196. Ibidem, PT, ♁, Gabon; 197. Ibidem, PT, ♀, Gabon; 198. A. helicalis sp. n., HT, ♁, Gabon; 199. Ibidem, PT, ♁, Gabon; 200. Ibidem, PT, ♀, Gabon; 201. A. takanoi sp. n., HT, ♁, Sierra Leone; 202. Ibidem, PT, ♁, Guinea; 203. Ibidem, PT, ♀, Ivory Coast; 204. Ibidem, PT, ♀, Liberia; 205. A. staudei sp. n., HT, ♁, Kenya (RCHS); 206. Racotis boarmiaria, LT, ♁, India (NHMUK); 207. Chorocotis ugandaria, HT of Boarmia ugandaria, ♁, Uganda (NHMUK); 208. Ibidem, ♀, Cameroon (NHMUK); 209. Colocleora breijeri, ♁, RSA (RCDS); 210. C. incauta, ♁, RSA (NHMUK)." figureDoi-0="http://doi.org/10.5281/zenodo.8080952" figureDoi-1="http://doi.org/10.5281/zenodo.8080954" httpUri-0="https://zenodo.org/record/8080952/files/figure.png" httpUri-1="https://zenodo.org/record/8080954/files/figure.png" pageId="51" pageNumber="52">175–204</figureCitation>
|
||
). Forewing length
|
||
<quantity id="4CBA58D85D0EFF8F530E548FFB49FC21" box="[1132,1262,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" pageId="51" pageNumber="52" unit="mm" value="20.5" valueMax="24.0" valueMin="17.0">17–24 mm</quantity>
|
||
. Male antenna bipectinate-fasciculate, rami moderately long, distal section filiform, varying from 1/2 to 2/5 of length of antenna. Female antenna filiform. Head moderately large, proboscis well-developed, labial palp short, length 1.2–1.5 times the diameter of eye, porrect, first and second segment dilated, third segment rather short and thin. Compound eyes moderately large. Frons, vertex, collar (patagium), tegula, thorax and abdomen colouration as of wing upperside. Legs rather long, colouration as of wing underside, index of spurs 0-2-4. Forewing broad, triangular, costa straight, apical section slightly arched, termen straight to slightly arched, ventral margin almost straight. Ground colour varying inter- and intraspecifically from dark rusty-brown to greenish-grey usually with mossy tinge, densely speckled with groups of brownish-grey scales, basal and medial area often lighter than terminal one, postmedial area with series of large dark blotches. Transverse lines diffuse, interrupted. Basal, subbasal and antemedial line poorly detectable, interrupted, often partly deleted, more or less straight, or gently arched. Medial line often interrupted, shadow-like, slightly S-curved, postmedial line diffuse, slightly wavy, subterminal line represented by a row of patches of variable size, terminal line narrow, often interrupted, consisting of black dashes or arches between veins. Fringe (cilia) short, concolourous with forewing. Discal spot well-developed, round or ovoid with lighter inner area. Hindwing apex rounded, termen evenly arcuate, slightly scalloped, anal margin straight. Ground colour as of forewing, densely speckled, transverse lines diffuse, shadow-like, basal line evenly arched, wavy or sharply angled medially, medial line undulate, postmedial line interrupted, terminal line sometimes interrupted, consisting of black arches between veins. Discal spot as of forewing. Fringe (cilia) short, concolourous with hindwing. Wing underside with pale brownish-grey basal and medial area speckled densely with greyish-brown scales, terminal area with broad, dark transverse band. Traces of transverse lines present, diffuse, shadow-like, discal spot and postmedial patches well-defined.
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D0EFF8F57A5517BFAFBF811" blockId="51.[151,1437,906,2013]" pageId="51" pageNumber="52">
|
||
<emphasis id="B936292F5D0EFF8F57A5517BFED0F91C" bold="true" box="[199,375,1662,1689]" pageId="51" pageNumber="52">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="1379E9B85D0EFF8F56E8517BFD89F91D" box="[394,558,1662,1689]" captionStart-0="FIGURES 251–255" captionStart-1="FIGURES 256–261" captionStart-2="FIGURES 262–267" captionStart-3="FIGURES 268–273" captionStart-4="FIGURES 274–279" captionStart-5="FIGURES 280–284" captionStartId-0="88.[151,264,1832,1856]" captionStartId-1="89.[151,264,1832,1856]" captionStartId-2="90.[151,264,1842,1866]" captionStartId-3="91.[151,264,1844,1868]" captionStartId-4="92.[151,264,1818,1842]" captionStartId-5="93.[151,264,1824,1848]" captionTargetBox-0="[158,1431,190,1802]" captionTargetBox-1="[151,1433,187,1802]" captionTargetBox-2="[151,1436,187,1812]" captionTargetBox-3="[151,1436,182,1815]" captionTargetBox-4="[151,1433,187,1788]" captionTargetBox-5="[155,1431,189,1794]" captionTargetId-0="figure-21@88.[152,1435,181,1808]" captionTargetId-1="figure-17@89.[151,1436,181,1808]" captionTargetId-2="figure-21@90.[151,1436,181,1818]" captionTargetId-3="figure-17@91.[151,1436,181,1821]" captionTargetId-4="figure-21@92.[151,1436,181,1794]" captionTargetId-5="figure-17@93.[151,1436,181,1800]" captionTargetPageId-0="88" captionTargetPageId-1="89" captionTargetPageId-2="90" captionTargetPageId-3="91" captionTargetPageId-4="92" captionTargetPageId-5="93" captionText-0="FIGURES 251–255. Male genitalia, a: clasping apparatus, b: aedeagus. 251. Afroracotis muscivirens sp. n., PT, Ivory Coast, slide No. LG 4617 (ZSM); 252. A. atriclava, ST of Cleora atriclava, Madagascar, slide No. LG 5039 (ZSM); 253. A. turlini sp. n., HT, Rwanda, slide No. LG 4619 (ZSM). 254. Afroracotis zebrina, Ivory Coast, slide No. LG 4176 (ANHRT); 255. Ibidem, Liberia, slide No. LG 4965 (ANHRT)." captionText-1="FIGURES 256–261. Male genitalia, a: clasping apparatus, b: aedeagus. 256. Afroracotis zebrina, Ivory Coast, slide No. LG 4785 (ZSM); 257. A. chaineyi sp. n., HT, Gabon, slide No. LG 5405 (ANHRT); 258. Ibidem, PT, Nigeria, slide No. LG 4092 (NHMUK); 259. Ibidem, PT, Zambia, slide No. LG 4971 (ANHRT); 260. A. lydiae lydiae sp. n., HT, Liberia, slide No. LG 5788 (ANHRT); 261. Ibidem, PT, Gabon, slide No. LG 4824 (ANHRT)." captionText-2="FIGURES 262–267. Male genitalia, a: clasping apparatus, b: aedeagus. 262. Afroracotis lydiae lydiae sp. n., PT, Princípe, slide No. LG 5789 (ANHRT); 263. A. l. orientalis ssp. n., HT, Tanzania, slide No.LG 4172 (ANHRT); 264. Ibidem, Mozambique, slide No. LG 4932 (ANHRT); 265. Ibidem, Rwanda, slide No. LG 5536 (RCHS); 266. A. madagascariensis, Madagascar, slide No. LG 4102 (NHMUK); 267. Ibidem, Madagascar, slide No. LG 4787 (ZSM)." captionText-3="FIGURES 268–273. Male genitalia, a: clasping apparatus, b: aedeagus. 268. Afroracotis madagascariensis, Madagascar, slide No. LG 4078 (ANHRT); 269. A. smithi sp. n., HT, Zambia, slide No. LG 4972 (ANHRT); 270. Ibidem, PT, Gabon, slide No. LG 5398 (ANHRT); 271. Ibidem, PT, Kenya, slide No. LG 5549 (RCHS); 272. Afroracotis milesi sp. n., HT, Gabon, slide No. LG 5619 (ANHRT); 273. Ibidem, PT, Uganda, slide No. DS 228-2019 (RCDS)." captionText-4="FIGURES 274–279. Male genitalia, a: clasping apparatus, b: aedeagus. 274. Afroracotis milesi sp. n., PT, Zambia, slide No. LG 4061 (ANHRT); 275. A. ochsei sp. n., HT, Gabon, slide No. LG 5406 (ANHRT); 276. Ibidem, PT, Cameroon, slide No. LG 4413 (ZSM); 277. Ibidem, PT, Gabon, slide No. LG 5414 (ANHRT); 278. A. helicalis sp. n., HT, Gabon, slide No. LG 5621 (ANHRT); 279. Ibidem, PT, Gabon, slide No. LG 5412 (ANHRT)." captionText-5="FIGURES 280–284. Male genitalia, a: clasping apparatus, b: aedeagus. 280. Afroracotis helicalis sp. n., PT, Gabon, slide No. LG 5399 (ANHRT); 281. A. takanoi sp. n., HT, Sierra Leone, slide No. LG 4810 (ANHRT); 282. Ibidem, PT, Guinea, slide No. LG 5445 (ANHRT); 283. Ibidem, PT, Liberia, slide No. LG 4982 (ANHRT); 284. A. staudei sp. n., HT, Kenya, slide No. LG 5540 (RCHS)." figureDoi-0="http://doi.org/10.5281/zenodo.8080974" figureDoi-1="http://doi.org/10.5281/zenodo.8080976" figureDoi-2="http://doi.org/10.5281/zenodo.8080978" figureDoi-3="http://doi.org/10.5281/zenodo.8080983" figureDoi-4="http://doi.org/10.5281/zenodo.8080985" figureDoi-5="http://doi.org/10.5281/zenodo.8080987" httpUri-0="https://zenodo.org/record/8080974/files/figure.png" httpUri-1="https://zenodo.org/record/8080976/files/figure.png" httpUri-2="https://zenodo.org/record/8080978/files/figure.png" httpUri-3="https://zenodo.org/record/8080983/files/figure.png" httpUri-4="https://zenodo.org/record/8080985/files/figure.png" httpUri-5="https://zenodo.org/record/8080987/files/figure.png" pageId="51" pageNumber="52">Figs 254–283</figureCitation>
|
||
). Uncus simple, moderately long, distally tapered, apex narrowly rounded. Tegumen short and broad, gnathos well-developed, apically rounded. Valva short, broad at base, distally tapered, costal margin slightly concave, sclerotised, sparsely setose, without apical projection.Ventral margin of valva convex basally and subapically, smoothly concave medially; sacculus short and broad, weakly sclerotised, without process. Juxta weakly sclerotised, distally tapered, elongate shield-like without arms. Vinculum moderately long and broad, V-shaped, apically rounded.Aedeagus rather thick, short to medium long, straight or slightly curved, without carina. Vesica thick, relatively long, in some species with protractile diverticulum armed with a robust apical cornutus, and, in all but one species (
|
||
<taxonomicName id="4C428EBE5D0EFF8F56F0507FFDA2F811" authorityName="Warren" authorityYear="1899" box="[402,517,1915,1941]" class="Insecta" family="Geometridae" genus="Racotis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="zebrina">
|
||
<emphasis id="B936292F5D0EFF8F56F0507FFDA2F811" box="[402,517,1915,1941]" italics="true" pageId="51" pageNumber="52">A. zebrina</emphasis>
|
||
</taxonomicName>
|
||
) with a small group of cornuti situated near the base of vesica ejaculatorius.
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D0EFF8857A5509BFEE7FE55" blockId="51.[151,1437,906,2013]" lastBlockId="52.[151,1437,151,465]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
|
||
<emphasis id="B936292F5D0EFF8F57A5509BFE2BF83C" bold="true" box="[199,396,1950,1977]" pageId="51" pageNumber="52">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="1379E9B85D0EFF8F56F9509BFD9DF83D" box="[411,570,1951,1977]" captionStart-0="FIGURES 320–327" captionStart-1="FIGURES 328–335" captionStart-2="FIGURES 336–339" captionStartId-0="99.[151,264,1778,1802]" captionStartId-1="100.[151,264,1757,1781]" captionStartId-2="101.[151,264,1013,1037]" captionTargetBox-0="[208,1389,194,1748]" captionTargetBox-1="[197,1392,189,1727]" captionTargetBox-2="[191,1397,187,983]" captionTargetId-0="figure-17@99.[186,1401,181,1754]" captionTargetId-1="figure-21@100.[190,1398,181,1733]" captionTargetId-2="figure-17@101.[184,1404,181,989]" captionTargetPageId-0="99" captionTargetPageId-1="100" captionTargetPageId-2="101" captionText-0="FIGURES 320–327. Female genitalia. 320. Afroracotis zebrina, Liberia, slide No.: LG 5436 (ANHRT); 321. Ibidem, Ivory Coast, slide No.: LG 4743 (ZSM); 322. A. chaineyi sp. n., PT, Guinea, slide No.: LG 5446 (ANHRT); 323. Ibidem, PT, Bioko, slide No.: LG 5073 (ZSM); 324. Ibidem, PT, Zambia, slide No.: LG 5432 (ANHRT); 325. Ibidem, PT, DRC, slide No.: LG 5054 (ZSM); 326. A. lydiae lydiae sp. n., PT, Guinea, slide No.: LG 5439 (ANHRT); 327 Ibidem, PT, Gabon, gen. slide No.: LG 5404 (ANHRT)." captionText-1="FIGURES 328–335. Female genitalia. 328. Afroracotis lydiae orientalis ssp. n., PT, Tanzania, slide No.: LG 4173 (ANHRT); 329. Ibidem, PT, Mozambique, slide No.: LG 4955 (ANHRT); 330. A. madagascariensis, Madagascar, slide No.: LG 4750 (ZSM); 331. A. smithi sp. n., PT, Zambia, slide No.: LG 5400 (ANHRT); 332. Ibidem, PT, Cameroon, slide No.: LG 5074 (ZSM); 333. A. milesi sp. n., PT, Gabon, slide No.: LG 5411 (ANHRT); 334. Ibidem, PT, Cameroon, slide No.: LG 5437 (ANHRT); 335. A. ochsei sp. n., PT, Gabon, gen. slide No.: LG 5623 (ANHRT)." captionText-2="FIGURES 336–339. Female genitalia. 336. Afroracotis helicalis sp. n., PT, Gabon, slide No.: LG 5624 (ANHRT); 337. A. takanoi sp. n., PT, Liberia, slide No.: LG 5438 (ANHRT); 338. Ibidem, PT, Ivory Coast, slide No.: LG 4809 (ANHRT); 339. Chorocotis ugandaria, Cameroon, slide No.: NHMUK 010317818 (NHMUK)." figureDoi-0="http://doi.org/10.5281/zenodo.8081003" figureDoi-1="http://doi.org/10.5281/zenodo.8081005" figureDoi-2="http://doi.org/10.5281/zenodo.8081007" httpUri-0="https://zenodo.org/record/8081003/files/figure.png" httpUri-1="https://zenodo.org/record/8081005/files/figure.png" httpUri-2="https://zenodo.org/record/8081007/files/figure.png" pageId="51" pageNumber="52">Figs 320–338</figureCitation>
|
||
). Ovipositor medium long, papilla analis narrow triangular, apically rounded, sparsely setose, apophysis posterioris moderately long. Eighth tergite medium long, trapezoidal, distal margin slightly arcuate, proximal margin undulate with a shallow medial depression, anterior apophysis shorter than posterior apophysis. Ostium bursae broad, membranous, antrum moderately sclerotised, short, funnel-, cup-, goblet-shaped or inverse trapezoidal. Distal section of ductus bursae membranous, very short and narrow, may be fully reduced; proximal part tubular or funnel-shaped, varying in length interspecifically, directly continued in sclerotised distal part of corpus bursae. Cervix bursae well-developed, heavily sclerotised, fully integrated into corpus bursae forming a sclerotised, longitudinally striate distal section of bursa copulatrix. Membranous anterior part of corpus bursae varied in size interspecifically, may be almost fully reduced; corpus bursae with a strongly sclerotised, smooth-surfaced disc-like or quadrangular plate supposedly serving as receptive area of cornutus vesicae during copulation; signum absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358A6B65D09FF8857F55526FD85FC17" pageId="52" pageNumber="53" type="multiple">
|
||
<paragraph id="8BFDF53D5D09FF8857F55526FEF0FDB9" blockId="52.[151,343,546,573]" box="[151,343,546,573]" pageId="52" pageNumber="53">
|
||
<emphasis id="B936292F5D09FF8857F55526FEF0FDB9" bold="true" box="[151,343,546,573]" pageId="52" pageNumber="53">Species content</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F55563FD5DFDFB" blockId="52.[151,991,615,915]" box="[151,762,615,639]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F55563FD27FDFB" authority="(Warren, 1899)" authorityName="László & Hausmann & Karisch" authorityYear="2023" baseAuthorityName="Warren" baseAuthorityYear="1899" box="[151,640,615,639]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="zebrina" status="comb. nov." subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D09FF8857F55563FE47FDFB" box="[151,480,615,639]" italics="true" pageId="52" pageNumber="53">Afroracotis (Zebracotis) zebrina</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD388CC5D09FF88568F5563FDDEFDFB" author="Warren, W." box="[493,633,615,639]" pageId="52" pageNumber="53" pagination="1 - 66" refId="ref85194" refString="Warren, W. (1899) New species and genera of the families Drepanulidae, Thyrididae, Uraniidae, Epiplemidae, and Geometridae from the Old-World regions. Novitates Zoologicae, 6 (1), 1 - 66." type="journal article" year="1899">Warren, 1899</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B936292F5D09FF8855E45563FD5DFDFB" bold="true" box="[646,762,615,639]" pageId="52" pageNumber="53">
|
||
<taxonomicNameLabel id="A20594545D09FF8855E45563FD5DFDFB" box="[646,762,615,639]" pageId="52" pageNumber="53" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F55582FD8BFD1A" blockId="52.[151,991,615,915]" box="[151,556,645,670]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F55582FE4AFD19" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,493,645,670]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="chaineyi" status="sp. nov." subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D09FF8857F55582FE4AFD19" box="[151,493,645,670]" italics="true" pageId="52" pageNumber="53">
|
||
Afroracotis (Zebracotis)
|
||
<emphasis id="B936292F5D09FF8856F65581FE4AFD19" bold="true" box="[404,493,645,669]" pageId="52" pageNumber="53">chaineyi</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20594545D09FF8856915582FD8BFD1A" box="[499,556,646,670]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F555A1FDB6FD39" blockId="52.[151,991,615,915]" box="[151,529,676,701]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F555A1FE75FD38" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,466,676,701]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="lydiae" status="sp. nov." subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D09FF8857F555A1FE75FD38" box="[151,466,676,701]" italics="true" pageId="52" pageNumber="53">
|
||
Afroracotis (Zebracotis)
|
||
<emphasis id="B936292F5D09FF8856F655A0FE75FD38" bold="true" box="[404,466,676,700]" pageId="52" pageNumber="53">lydiae</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20594545D09FF8856BA55A1FDB6FD39" box="[472,529,677,701]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F555C7FD23FD5F" blockId="52.[151,991,615,915]" box="[151,644,707,731]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F555C7FD9CFD5F" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,571,707,731]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="62" phylum="Arthropoda" rank="subSpecies" species="lydiae" status="subsp. nov." subGenus="Zebracotis" subSpecies="orientalis">
|
||
<emphasis id="B936292F5D09FF8857F555C7FD9CFD5F" box="[151,571,707,731]" italics="true" pageId="52" pageNumber="53">
|
||
Afroracotis (Zebracotis)
|
||
<emphasis id="B936292F5D09FF8856F655C7FD9CFD5F" bold="true" box="[404,571,707,731]" pageId="52" pageNumber="53">lydiae orientalis</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
ssp. n.
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F555E6FC78FD7E" blockId="52.[151,991,615,915]" box="[151,991,738,762]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F555E6FCC2FD7E" authority="(Chainey & Karisch, 2017)" authorityName="László & Hausmann & Karisch" authorityYear="2023" baseAuthorityName="Chainey & Karisch" baseAuthorityYear="2017" box="[151,869,738,762]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="madagascariensis" status="comb. nov." subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D09FF8857F555E6FDECFD7E" box="[151,587,738,762]" italics="true" pageId="52" pageNumber="53">Afroracotis (Zebracotis) madagascariensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD388CC5D09FF88553A55E6FCF9FD7E" author="Chainey, J. & Karisch, T." box="[600,862,738,762]" pageId="52" pageNumber="53" pagination="49 - 53" refId="ref83017" refString="Chainey, J. & Karisch, T. (2017) A new species of Racotis Moore, [1887] (Lepidoptera: Geometridae: Ennominae) from Madagascar. Metamorphosis, 28, 49 - 53." type="journal article" year="2017">Chainey & Karisch, 2017</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B936292F5D09FF88540955E6FC78FD7E" bold="true" box="[875,991,738,762]" pageId="52" pageNumber="53">
|
||
<taxonomicNameLabel id="A20594545D09FF88540955E6FC78FD7E" box="[875,991,738,762]" pageId="52" pageNumber="53" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F55405FDB3FC9D" blockId="52.[151,991,615,915]" box="[151,532,768,793]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F55405FE72FC9C" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,469,768,793]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="63" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov." subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D09FF8857F55405FE72FC9C" box="[151,469,768,793]" italics="true" pageId="52" pageNumber="53">
|
||
Afroracotis (Zebracotis)
|
||
<emphasis id="B936292F5D09FF8856F65404FE72FC9C" bold="true" box="[404,469,768,792]" pageId="52" pageNumber="53">smithi</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20594545D09FF8856B95405FDB3FC9D" box="[475,532,769,793]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F5541BFDB6FCB3" blockId="52.[151,991,615,915]" box="[151,529,799,823]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F5541BFE75FCB3" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,466,799,823]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="milesi" status="sp. nov." subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D09FF8857F5541BFE75FCB3" box="[151,466,799,823]" italics="true" pageId="52" pageNumber="53">
|
||
Afroracotis (Zebracotis)
|
||
<emphasis id="B936292F5D09FF8856F6541BFE75FCB3" bold="true" box="[404,466,799,823]" pageId="52" pageNumber="53">milesi</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20594545D09FF8856BA541BFDB6FCB3" box="[472,529,799,823]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F5543AFDB2FCD2" blockId="52.[151,991,615,915]" box="[151,533,829,854]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F5543AFE71FCD1" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,470,829,854]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="ochsei" status="sp. nov." subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D09FF8857F5543AFE71FCD1" box="[151,470,829,854]" italics="true" pageId="52" pageNumber="53">
|
||
Afroracotis (Zebracotis)
|
||
<emphasis id="B936292F5D09FF8856F65439FE71FCD1" bold="true" box="[404,470,829,853]" pageId="52" pageNumber="53">ochsei</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20594545D09FF8856BE543AFDB2FCD2" box="[476,533,830,854]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F55459FD8DFCF1" blockId="52.[151,991,615,915]" box="[151,554,860,885]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F55459FE4CFCF0" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,491,860,885]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="helicalis" status="sp. nov." subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D09FF8857F55459FE4CFCF0" box="[151,491,860,885]" italics="true" pageId="52" pageNumber="53">
|
||
Afroracotis (Zebracotis)
|
||
<emphasis id="B936292F5D09FF8856F65458FE4CFCF0" bold="true" box="[404,491,860,884]" pageId="52" pageNumber="53">helicalis</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20594545D09FF8856935459FD8DFCF1" box="[497,554,861,885]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BFDF53D5D09FF8857F5547FFD85FC17" blockId="52.[151,991,615,915]" box="[151,546,891,915]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C428EBE5D09FF8857F5547FFE44FC17" authority="László & Hausmann & Karisch, 2023" authorityName="László & Hausmann & Karisch" authorityYear="2023" box="[151,483,891,915]" class="Insecta" family="Geometridae" genus="Afroracotis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="69" phylum="Arthropoda" rank="species" species="takanoi" status="sp. nov." subGenus="Zebracotis">
|
||
<emphasis id="B936292F5D09FF8857F5547FFE44FC17" box="[151,483,891,915]" italics="true" pageId="52" pageNumber="53">
|
||
Afroracotis (Zebracotis)
|
||
<emphasis id="B936292F5D09FF8856F6547FFE44FC17" bold="true" box="[404,483,891,915]" pageId="52" pageNumber="53">takanoi</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20594545D09FF88568B547FFD85FC17" box="[489,546,891,915]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |