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<document id="F2D3722FD461E61BD01CFF2FAF8C7808" ID-CLB-Dataset="24794" ID-DOI="10.11646/zootaxa.4701.1.4" ID-GBIF-Dataset="ca29e500-63fd-4f87-b24b-c41cdfa2cf1f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3557825" ID-ZooBank="338B950C-1590-4E98-8254-83DD1AD39259" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575043589075" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03CF87DF697BFFE515C2AEB9FE86F9D0" docLanguage="en" docName="zootaxa.4701.1.4.pdf" docOrigin="Zootaxa 4701 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Veraphis sakaii Jałoszyński 2019, sp. n." docType="treatment" docVersion="7" lastPageNumber="64" masterDocId="FFF6FFA7697FFFE31555A80CFFA6FFA7" masterDocTitle="Four new species and new records of Veraphis Casey in Japan (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="64" masterPageNumber="54" pageNumber="54" updateTime="1698769603799" updateUser="ExternalLinkService">
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<mods:title id="ED09524E088616F12F1AB9C7DEDC7A3F">Four new species and new records of Veraphis Casey in Japan (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="E9BCA70831763738768F7EB61AF085C2">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="2852372DE6B7F05E0F2F92DFB3214D4E">text</mods:typeOfResource>
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<mods:date id="648A5DB46785A190FBACA9C678E9DDD4">2019</mods:date>
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<mods:number id="521C7EF43945EA1373D985A0AB957AB2">2019-11-26</mods:number>
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<mods:identifier id="368EB2B7DE0910962F245EB77E61725B" type="DOI">10.11646/zootaxa.4701.1.4</mods:identifier>
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<treatment id="03CF87DF697BFFE515C2AEB9FE86F9D0" ID-GBIF-Taxon="160605361" LSID="urn:lsid:plazi:treatment:03CF87DF697BFFE515C2AEB9FE86F9D0" httpUri="http://treatment.plazi.org/id/03CF87DF697BFFE515C2AEB9FE86F9D0" lastPageId="6" pageId="4">
<subSubSection id="C37C6542697BFFE715C2AEB9FE3CF977" box="[151,410,1717,1744]" pageId="4" type="nomenclature">
<paragraph id="8BD936C9697BFFE715C2AEB9FE3CF977" blockId="4.[151,490,1717,1780]" box="[151,410,1717,1744]" pageId="4">
<heading id="D09181A5697BFFE715C2AEB9FE3CF977" bold="true" box="[151,410,1717,1744]" fontSize="11" level="1" pageId="4" reason="1">
<emphasis id="B912EADB697BFFE715C2AEB9FE3CF977" bold="true" box="[151,410,1717,1744]" pageId="4">
<taxonomicName id="4C664D4A697BFFE715C2AEB9FEF6F977" ID-CoL="8YC74" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,336,1717,1744]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="sakaii" status="sp. nov.">
<emphasis id="B912EADB697BFFE715C2AEB9FEF6F977" bold="true" box="[151,336,1717,1744]" italics="true" pageId="4">Veraphis sakaii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157A0697BFFE7140DAEBAFE3CF977" box="[344,410,1718,1744]" pageId="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C6542697BFFE715C2AED6FE4CF953" box="[151,490,1753,1780]" pageId="4" type="description">
<paragraph id="8BD936C9697BFFE715C2AED6FE4CF953" blockId="4.[151,490,1717,1780]" box="[151,490,1753,1780]" pageId="4">
(
<figureCitation id="135D2A4C697BFFE715CAAED6FF4EF953" box="[159,232,1754,1780]" captionStart="FIGURES 14" captionStartId="4.[151,264,1575,1600]" captionTargetBox="[155,1426,854,1517]" captionTargetId="figure@4.[151,1436,841,1526]" captionTargetPageId="4" captionText="FIGURES 14. Dorsal habitus of males. Veraphis loebli sp. n. (1); V. mutsuensis sp. n. (2); V. aomoriensis sp. n. (3); V. sakaii sp. n. (4)." figureDoi="http://doi.org/10.5281/zenodo.3557827" httpUri="https://zenodo.org/record/3557827/files/figure.png" pageId="4">Figs 4</figureCitation>
,
<figureCitation id="135D2A4C697BFFE715A1AED6FE9DF953" box="[244,315,1754,1780]" captionStart="FIGURES 512" captionStartId="5.[151,264,1966,1991]" captionTargetBox="[184,1424,314,1942]" captionTargetId="figure@5.[151,1436,301,1946]" captionTargetPageId="5" captionText="FIGURES 512. Aedeagus in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views. Veraphis loebli sp. n. (56); V. mutsuensis sp. n. (78); V. aomoriensis sp. n. (910); V. sakaii sp. n. (1112)." figureDoi="http://doi.org/10.5281/zenodo.3557829" httpUri="https://zenodo.org/record/3557829/files/figure.png" pageId="4">1112</figureCitation>
,
<figureCitation id="135D2A4C697BFFE71412AED6FE29F953" box="[327,399,1754,1780]" captionStart="FIGURES 1320" captionStartId="7.[151,264,1920,1945]" captionTargetBox="[176,1432,248,1881]" captionTargetId="figure@7.[151,1436,229,1901]" captionTargetPageId="7" captionText="FIGURES 1320. Aedeagus in ventral (13, 15, 17, 19) and lateral (14, 16, 18, 20) views. Veraphis loebli sp. n. (1314); V. mutsuensis sp. n. (1516); V. aomoriensis sp. n. (1718); V. sakaii sp. n. (1920)." figureDoi="http://doi.org/10.5281/zenodo.3557831" httpUri="https://zenodo.org/record/3557831/files/figure.png" pageId="4">1920</figureCitation>
,
<figureCitation id="135D2A4C697BFFE714CEAED6FE1FF953" box="[411,441,1754,1780]" captionStart="FIGURES 2124" captionStartId="8.[151,264,1456,1481]" captionTargetBox="[169,1427,432,1413]" captionTargetId="figure@8.[151,1436,404,1436]" captionTargetPageId="8" captionText="FIGURES 2124. Fore leg of male. Veraphis loebli sp. n. (21); V. mutsuensis sp. n. (22); V. aomoriensis sp. n. (23); V. sakaii sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3557833" httpUri="https://zenodo.org/record/3557833/files/figure.png" pageId="4">24</figureCitation>
,
<figureCitation id="135D2A4C697BFFE71490AED5FE44F953" box="[453,482,1753,1780]" captionStart="FIGURE 25" captionStartId="9.[151,250,1702,1727]" captionTargetBox="[151,1436,696,1668]" captionTargetId="figure@9.[151,1436,692,1683]" captionTargetPageId="9" captionText="FIGURE 25. Distribution of Veraphis in Japan." figureDoi="http://doi.org/10.5281/zenodo.3557835" httpUri="https://zenodo.org/record/3557835/files/figure.png" pageId="4">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C6542697BFFE715C2AF2EFDE9F823" pageId="4" type="materials_examined">
<paragraph id="8BD936C9697BFFE715C2AF2EFDE9F823" blockId="4.[151,1436,1825,2032]" pageId="4">
<emphasis id="B912EADB697BFFE715C2AF2EFEE3F89B" bold="true" box="[151,325,1825,1852]" pageId="4">Type material.</emphasis>
<materialsCitation id="3B0E3C94697BFFE71419AF2EFDE9F823" ID-GBIF-Occurrence="2460169389" collectingDate="1997-05-24" collectionCode="EUMJ" collectorName="M. Sakai &amp; P. Jaloszynski" country="Japan" location="Shikoku" pageId="4" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B912EADB697BFFE71419AF2EFC99F89B" bold="true" box="[332,831,1825,1852]" pageId="4">
<typeStatus id="54DD886B697BFFE71419AF2EFE1BF89B" box="[332,445,1826,1852]" pageId="4" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3717659697BFFE7149CAF2DFD85F89B" box="[457,547,1825,1852]" name="Japan" pageId="4">JAPAN</collectingCountry>
(
<location id="8EB96012697BFFE71764AF2DFD3EF89B" LSID="urn:lsid:plazi:treatment:03CF87DF697BFFE515C2AEB9FE86F9D0:8EB96012697BFFE71764AF2DFD3EF89B" box="[561,664,1825,1852]" country="Japan" name="Shikoku" pageId="4">Shikoku</location>
, Kôchi Pref.):
</emphasis>
<specimenCount id="9D60FD40697BFFE71613AF2EFCFBF89B" box="[838,861,1826,1852]" pageId="4" type="male"></specimenCount>
, two labels: “[Kôchi: JAPAN] / Kamioriwatashi / Yusuhara-cho /
<date id="FFD81009697BFFE71412AF4AFE61F8C7" box="[327,455,1862,1888]" pageId="4" value="1997-05-24">
<collectingDate id="EF9CE9E1697BFFE71412AF4AFE61F8C7" box="[327,455,1862,1888]" pageId="4" value="1997-05-24">24. V. 1997</collectingDate>
</date>
/
<collectorName id="2693531F697BFFE71489AF4AFD9DF8C7" box="[476,571,1862,1888]" pageId="4">M.Sakai</collectorName>
/ (
<emphasis id="B912EADB697BFFE7170FAF4BFD0AF8C7" box="[602,684,1862,1888]" italics="true" pageId="4">in litter</emphasis>
)” [white, printed], “
<taxonomicName id="4C664D4A697BFFE716CCAF4AFBBCF8C7" authorityName="Casey" authorityYear="1897" box="[921,1050,1862,1888]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="genus">
<emphasis id="B912EADB697BFFE716CCAF4AFBBCF8C7" box="[921,1050,1862,1888]" italics="true" pageId="4">VERAPHIS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C664D4A697BFFE71165AF4AFBD5F8C7" authorityName="Jałoszyński" authorityYear="2019" box="[1072,1139,1862,1888]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="sakaii">
<emphasis id="B912EADB697BFFE71165AF4AFBD5F8C7" box="[1072,1139,1862,1888]" italics="true" pageId="4">sakaii</emphasis>
</taxonomicName>
m. /
<collectorName id="2693531F697BFFE711F9AF4AFAECF8C7" box="[1196,1354,1862,1888]" pageId="4">P. Jałoszyński</collectorName>
, 2019 /
<typeStatus id="54DD886B697BFFE715C2AF66FEE7F823" box="[151,321,1898,1924]" pageId="4" type="holotype">HOLOTYPUS</typeStatus>
” [red, printed] (
<collectionCode id="ED77AE0C697BFFE714A1AF66FDE5F823" box="[500,579,1898,1924]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4">EUMJ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37C6542697BFFE61592AF81FD39FF70" lastPageId="5" pageId="4" type="diagnosis">
<paragraph id="8BD936C9697BFFE61592AF81FD39FF70" blockId="4.[151,1436,1825,2032]" lastBlockId="5.[151,1436,153,287]" lastPageId="5" pageId="4">
<emphasis id="B912EADB697BFFE71592AF81FEE4F80F" bold="true" box="[199,322,1933,1960]" pageId="4">Diagnosis.</emphasis>
Male protrochanter unmodified; male protibia with subapical fin-like projection about as broad as half of tibial width; median lobe of aedeagus in lateral view weakly curved; in ventral view gradually and strongly broadening from base to subapical region, with convex sides; apical region subtriangular with rounded apex, broad- er than long; ventral diaphragm slightly shorter than half of median lobe, distant from basal margin of aedeagus; parameres in lateral view straight in distal 2/3.
</paragraph>
</subSubSection>
<subSubSection id="C37C6542697AFFE51592A8ECFE38FC48" lastPageId="6" pageId="5" type="description">
<paragraph id="8BD936C9697AFFE61592A8ECFD9FFEB8" blockId="5.[151,1436,153,287]" pageId="5">
<emphasis id="B912EADB697AFFE61592A8ECFEFFFF5C" bold="true" box="[199,345,224,251]" pageId="5">Description.</emphasis>
BL
<quantity id="4C9E9B2C697AFFE614DCA8ECFE56FF5C" box="[393,496,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="5" unit="mm" value="1.35">1.35 mm</quantity>
. Body of male (
<figureCitation id="135D2A4C697AFFE617F6A8EDFD40FF5C" box="[675,742,225,251]" captionStart="FIGURES 14" captionStartId="4.[151,264,1575,1600]" captionTargetBox="[155,1426,854,1517]" captionTargetId="figure@4.[151,1436,841,1526]" captionTargetPageId="4" captionText="FIGURES 14. Dorsal habitus of males. Veraphis loebli sp. n. (1); V. mutsuensis sp. n. (2); V. aomoriensis sp. n. (3); V. sakaii sp. n. (4)." figureDoi="http://doi.org/10.5281/zenodo.3557827" httpUri="https://zenodo.org/record/3557827/files/figure.png" pageId="5">Fig. 4</figureCitation>
) flattened, slender, moderately light brown, covered with yel- lowish vestiture, tarsi slightly lighter.
</paragraph>
<caption id="DF196641697AFFE615C2AFA2FCB5F84C" ID-DOI="http://doi.org/10.5281/zenodo.3557829" ID-Zenodo-Dep="3557829" httpUri="https://zenodo.org/record/3557829/files/figure.png" pageId="5" startId="5.[151,264,1966,1991]" targetBox="[184,1424,314,1942]" targetPageId="5">
<paragraph id="8BD936C9697AFFE615C2AFA2FCB5F84C" blockId="5.[151,1436,1966,2027]" pageId="5">
<emphasis id="B912EADB697AFFE615C2AFA2FEE1F860" bold="true" box="[151,327,1966,1991]" pageId="5">FIGURES 512.</emphasis>
Aedeagus in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views.
<taxonomicName id="4C664D4A697AFFE61684AFA3FBCFF860" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[977,1129,1967,1991]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B912EADB697AFFE61684AFA3FBCFF860" box="[977,1129,1967,1991]" italics="true" pageId="5">Veraphis loebli</emphasis>
</taxonomicName>
<emphasis id="B912EADB697AFFE6113AAFA3FB01F860" bold="true" box="[1135,1191,1967,1991]" pageId="5">
<taxonomicNameLabel id="A22157A0697AFFE6113AAFA3FB01F860" box="[1135,1191,1967,1991]" pageId="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(56);
<taxonomicName id="4C664D4A697AFFE611A4AFA3FADEF861" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[1265,1400,1967,1990]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="mutsuensis" status="sp. nov.">
<emphasis id="B912EADB697AFFE611A4AFA3FADEF861" box="[1265,1400,1967,1990]" italics="true" pageId="5">V. mutsuensis</emphasis>
</taxonomicName>
<emphasis id="B912EADB697AFFE6102BAFA3FF0AF84C" bold="true" pageId="5">
<taxonomicNameLabel id="A22157A0697AFFE6102BAFA3FF0AF84C" pageId="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(78);
<taxonomicName id="4C664D4A697AFFE615A3AFDFFE2DF84D" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[246,395,2003,2026]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="aomoriensis" status="sp. nov.">
<emphasis id="B912EADB697AFFE615A3AFDFFE2DF84D" box="[246,395,2003,2026]" italics="true" pageId="5">V. aomoriensis</emphasis>
</taxonomicName>
<emphasis id="B912EADB697AFFE614C4AFDFFE6CF84C" bold="true" box="[401,458,2003,2027]" pageId="5">
<taxonomicNameLabel id="A22157A0697AFFE614C4AFDFFE6CF84C" box="[401,458,2003,2027]" pageId="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(910);
<taxonomicName id="4C664D4A697AFFE61777AFDFFDD1F84C" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[546,631,2003,2027]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="sakaii" status="sp. nov.">
<emphasis id="B912EADB697AFFE61777AFDFFDD1F84C" box="[546,631,2003,2027]" italics="true" pageId="5">V. sakaii</emphasis>
</taxonomicName>
<emphasis id="B912EADB697AFFE61728AFDFFD10F84C" bold="true" box="[637,694,2003,2027]" pageId="5">
<taxonomicNameLabel id="A22157A0697AFFE61728AFDFFD10F84C" box="[637,694,2003,2027]" pageId="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(1112).
</paragraph>
</caption>
<paragraph id="8BD936C96979FFE51592A895FAE8FE08" blockId="6.[151,1437,152,1655]" pageId="6">
Head broadest at large, moderately convex eyes, HL
<quantity id="4C9E9B2C6979FFE51646A894FCDEFF14" box="[787,888,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="6" unit="mm" value="0.15">0.15 mm</quantity>
, HW
<quantity id="4C9E9B2C6979FFE516EDA895FBB9FF14" box="[952,1055,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="6" unit="mm" value="0.24">0.24 mm</quantity>
; vertex with pair of small postero- median pits, each prolonged by shallow but distinct longitudinal groove extending anterad to posteromesal margin of weakly elevated supraantennal tubercle; area between grooves distinctly flattened and depressed in relation to convex sides of vertex and frons. Punctures on sides of vertex fine and shallow, inconspicuous, area between grooves with uneven surface; setae short, sparse and suberect, except for a pair of conspicuously long and erect thin setae, each inserted just behind eye. Antennae slender, with distinctly demarcated trimerous club, AnL
<quantity id="4C9E9B2C6979FFE511C5A940FB51FEC0" box="[1168,1271,332,359]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="6" unit="mm" value="0.53">0.53 mm</quantity>
; antennomeres III distinctly elongate; III distinctly transverse; IV about as long as broad; V indistinctly elongate, VIX each dis- tinctly transverse (VIII not narrower, but shorter than VII), XI as broad as long and slightly narrower than X.
</paragraph>
<paragraph id="8BD936C96979FFE51592A9B5FEF6FD20" blockId="6.[151,1437,152,1655]" pageId="6">
Pronotum nearly semielliptical, broadest at base; PL
<quantity id="4C9E9B2C6979FFE51664A9B5FC3DFE74" box="[817,923,441,467]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" unit="mm" value="0.3">0.30 mm</quantity>
, PW
<quantity id="4C9E9B2C6979FFE5168BA9B5FBEFFE74" box="[990,1097,441,467]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" unit="mm" value="0.4">0.40 mm</quantity>
; anterior margin broadly and evenly rounded; lateral margins strongly rounded in anterior third, in posterior half nearly straight and parallel, hind angles obtuse and blunt; posterior margin indistinctly bisinuate. Pronotal base with distinct transverse groove with barely marked median pit, groove laterally reaching lateral pair of elongate and deep impressions; lateral pronotal margins narrowly carinate in posterior half. Punctures on pronotal disc fine and inconspicuous; setae short and sparse, suberect.
</paragraph>
<paragraph id="8BD936C96979FFE51592AA9DFAADFD54" blockId="6.[151,1437,152,1655]" pageId="6">
Elytra slightly more convex than pronotum, together oval, broadest slightly in front of middle; EL
<quantity id="4C9E9B2C6979FFE51057AA9DFAC1FD0C" box="[1282,1383,657,683]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="6" unit="mm" value="0.73">0.73 mm</quantity>
, EW
<quantity id="4C9E9B2C6979FFE515C2AAB8FF58FD68" box="[151,254,692,719]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="6" unit="mm" value="0.53">0.53 mm</quantity>
, EI 1.38; humeral calli well-marked, elongate. Surface of elytra less glossy than pronotum, covered with fine, superficial, inconspicuous punctures; setae distinctly thicker and denser than those on pronotum.
</paragraph>
<paragraph id="8BD936C96979FFE51592AAF1FEA6FC9C" blockId="6.[151,1437,152,1655]" pageId="6">
Legs moderately long and slender; protrochanter unmodified, protibia with small subapical fin-like projection (
<figureCitation id="135D2A4C6979FFE515F5AB2DFF52FC9C" box="[160,244,801,827]" captionStart="FIGURES 2124" captionStartId="8.[151,264,1456,1481]" captionTargetBox="[169,1427,432,1413]" captionTargetId="figure@8.[151,1436,404,1436]" captionTargetPageId="8" captionText="FIGURES 2124. Fore leg of male. Veraphis loebli sp. n. (21); V. mutsuensis sp. n. (22); V. aomoriensis sp. n. (23); V. sakaii sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3557833" httpUri="https://zenodo.org/record/3557833/files/figure.png" pageId="6">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="8BD936C96979FFE51592AB49FB04FC6C" blockId="6.[151,1437,152,1655]" pageId="6">
Aedeagus (
<figureCitation id="135D2A4C6979FFE51411AB49FE63FCF8" box="[324,453,837,863]" captionStart="FIGURES 512" captionStartId="5.[151,264,1966,1991]" captionTargetBox="[184,1424,314,1942]" captionTargetId="figure@5.[151,1436,301,1946]" captionTargetPageId="5" captionText="FIGURES 512. Aedeagus in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views. Veraphis loebli sp. n. (56); V. mutsuensis sp. n. (78); V. aomoriensis sp. n. (910); V. sakaii sp. n. (1112)." figureDoi="http://doi.org/10.5281/zenodo.3557829" httpUri="https://zenodo.org/record/3557829/files/figure.png" pageId="6">Figs 1112</figureCitation>
,
<figureCitation id="135D2A4C6979FFE51485AB49FDB1FCF8" box="[464,535,837,863]" captionStart="FIGURES 1320" captionStartId="7.[151,264,1920,1945]" captionTargetBox="[176,1432,248,1881]" captionTargetId="figure@7.[151,1436,229,1901]" captionTargetPageId="7" captionText="FIGURES 1320. Aedeagus in ventral (13, 15, 17, 19) and lateral (14, 16, 18, 20) views. Veraphis loebli sp. n. (1314); V. mutsuensis sp. n. (1516); V. aomoriensis sp. n. (1718); V. sakaii sp. n. (1920)." figureDoi="http://doi.org/10.5281/zenodo.3557831" httpUri="https://zenodo.org/record/3557831/files/figure.png" pageId="6">1920</figureCitation>
) elongate but relatively stout; AeL
<quantity id="4C9E9B2C6979FFE516F7AB49FBAFFCF8" box="[930,1033,837,863]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" unit="mm" value="0.3">0.30 mm</quantity>
; in ventral view median lobe gradu- ally broadening from base to subapical 1/4, with subtriangular, short and rounded apical portion; ventral diaphragm about as long as 1/3 of median lobe and with its proximal margin distant from base of aedeagus; in lateral view median lobe weakly curved; parameres straight in distal 2/3, each with two tiny apical setae.
</paragraph>
<paragraph id="8BD936C96979FFE51592ABD9FE38FC48" blockId="6.[151,1437,152,1655]" box="[199,414,981,1007]" pageId="6">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C37C65426979FFE51592ABF4FD47FBB4" box="[199,737,1016,1043]" pageId="6" type="distribution">
<paragraph id="8BD936C96979FFE51592ABF4FD47FBB4" blockId="6.[151,1437,152,1655]" box="[199,737,1016,1043]" pageId="6">
<emphasis id="B912EADB6979FFE51592ABF4FEC7FBB4" bold="true" box="[199,353,1016,1043]" pageId="6">Distribution.</emphasis>
<collectingCountry id="F37176596979FFE51432ABF5FE0CFBB4" box="[359,426,1017,1043]" name="Japan" pageId="6">Japan</collectingCountry>
, western Shikoku (
<figureCitation id="135D2A4C6979FFE517D5ABF5FD72FBB4" box="[640,724,1016,1043]" captionStart="FIGURE 25" captionStartId="9.[151,250,1702,1727]" captionTargetBox="[151,1436,696,1668]" captionTargetId="figure@9.[151,1436,692,1683]" captionTargetPageId="9" captionText="FIGURE 25. Distribution of Veraphis in Japan." figureDoi="http://doi.org/10.5281/zenodo.3557835" httpUri="https://zenodo.org/record/3557835/files/figure.png" pageId="6">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C65426979FFE51592AC11FC81FBFC" pageId="6" type="etymology">
<paragraph id="8BD936C96979FFE51592AC11FC81FBFC" blockId="6.[151,1437,152,1655]" pageId="6">
<emphasis id="B912EADB6979FFE51592AC11FEE8FB90" bold="true" box="[199,334,1053,1079]" pageId="6">Etymology.</emphasis>
This species is dedicated to the famous Japanese coleopterist Masahiro Sakai, who collected many rare and interesting Scydmaeninae, especially on Shikoku.
</paragraph>
</subSubSection>
<subSubSection id="C37C65426979FFE51592AC69FE86F9D0" pageId="6" type="discussion">
<paragraph id="8BD936C96979FFE51592AC69FE86F9D0" blockId="6.[151,1437,152,1655]" pageId="6">
<emphasis id="B912EADB6979FFE51592AC69FE9DFBD8" bold="true" box="[199,315,1125,1151]" pageId="6">Remarks.</emphasis>
<taxonomicName id="4C664D4A6979FFE51410AC69FE55FBD8" authorityName="Jałoszyński" authorityYear="2019" box="[325,499,1125,1151]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="sakaii">
<emphasis id="B912EADB6979FFE51410AC69FE55FBD8" box="[325,499,1125,1151]" italics="true" pageId="6">Veraphis sakaii</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C664D4A6979FFE517FDAC6AFCACFBD8" baseAuthorityName="K. Sawada" baseAuthorityYear="1962" box="[680,778,1126,1151]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B912EADB6979FFE517FDAC6AFCACFBD8" box="[680,778,1126,1151]" italics="true" pageId="6">japonica</emphasis>
</taxonomicName>
group of species, which is characterized by the antennomeres XI and protrochanters unmodified in males, protibiae with a subapical fin-like projection; male metaventrite with median elongate impression; and aedeagus lacking apical setae, with apical portion of median lobe in lateral view demarcated by a deep ventral emargination. This group comprises
<taxonomicName id="4C664D4A6979FFE516E9ACDEFAB2FB4B" authority="(K. Sawada, 1962)" authorityName="K. Sawada" authorityYear="1962" baseAuthorityName="K. Sawada" baseAuthorityYear="1962" box="[956,1300,1233,1260]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B912EADB6979FFE516E9ACDEFB9FFB4C" box="[956,1081,1234,1259]" italics="true" pageId="6">V. japonica</emphasis>
(K.
<bibRefCitation id="EFF74B386979FFE5113EACDDFAAAFB4B" author="Sawada, K." box="[1131,1292,1233,1260]" pageId="6" pagination="19 - 20" refId="ref6032" refString="Sawada, K. (1962) Sur deux formes nouvelles des Cephenniini du Japan (Coleopt. Scydmaenidae). The Entomological Review of Japan, 14, 19 - 20." type="journal article" year="1962">Sawada, 1962</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B912EADB6979FFE5104AACDEFF1EFAA8" italics="true" pageId="6">V. hisamat- sui</emphasis>
<bibRefCitation id="EFF74B386979FFE51594ACF9FDB2FAA8" author="Jaloszynski, P. &amp; Hoshina, H." box="[193,532,1268,1295]" pageId="6" pagination="101 - 136" refId="ref5900" refString="Jaloszynski, P. &amp; Hoshina, H. (2005) Revision of Palearctic species of Veraphis Casey (Coleoptera, Scydmaenidae). Entomological Review of Japan, 60, 101 - 136." type="journal article" year="2005">Jałoszyński &amp; Hoshina, 2005</bibRefCitation>
,
<taxonomicName id="4C664D4A6979FFE51777ACFAFBEEFAA8" authority="Jaloszynski &amp; Hoshina, 2005" authorityName="Jaloszynski &amp; Hoshina" authorityYear="2005" box="[546,1096,1268,1295]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="odaigaharensis">
<emphasis id="B912EADB6979FFE51777ACFAFD4AFAA8" box="[546,748,1269,1295]" italics="true" pageId="6">V. odaigaharensis</emphasis>
<bibRefCitation id="EFF74B386979FFE517A0ACF9FBEEFAA8" author="Jaloszynski, P. &amp; Hoshina, H." box="[757,1096,1268,1295]" pageId="6" pagination="101 - 136" refId="ref5900" refString="Jaloszynski, P. &amp; Hoshina, H. (2005) Revision of Palearctic species of Veraphis Casey (Coleoptera, Scydmaenidae). Entomological Review of Japan, 60, 101 - 136." type="journal article" year="2005">Jałoszyński &amp; Hoshina, 2005</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C664D4A6979FFE51103ACFAFE99FA94" authority="Jaloszynski &amp; Hoshina, 2005" authorityName="Jaloszynski &amp; Hoshina" authorityYear="2005" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="tottoriensis">
<emphasis id="B912EADB6979FFE51103ACFAFB54FAA8" box="[1110,1266,1270,1295]" italics="true" pageId="6">V. tottoriensis</emphasis>
<bibRefCitation id="EFF74B386979FFE511A9ACF9FE99FA94" author="Jaloszynski, P. &amp; Hoshina, H." pageId="6" pagination="101 - 136" refId="ref5900" refString="Jaloszynski, P. &amp; Hoshina, H. (2005) Revision of Palearctic species of Veraphis Casey (Coleoptera, Scydmaenidae). Entomological Review of Japan, 60, 101 - 136." type="journal article" year="2005">Jałoszyński &amp; Hoshina, 2005</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C664D4A6979FFE5141EAD16FD1CFA94" authority="Jaloszynski, 2014 a" authorityName="Jaloszynski" authorityYear="2014" box="[331,698,1305,1331]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="yoshitomii">
<emphasis id="B912EADB6979FFE5141EAD16FE7BFA94" box="[331,477,1305,1331]" italics="true" pageId="6">V. yoshitomii</emphasis>
<bibRefCitation id="EFF74B386979FFE514B0AD15FD1CFA94" author="Jaloszynski, P." box="[485,698,1305,1331]" pageId="6" pagination="589 - 595" refId="ref5798" refString="Jaloszynski, P. (2014 a) Veraphis yoshitomii sp. n. (Coleoptera, Staphylinidae, Scydmaeninae) from Japan. Zootaxa, 3860 (6), 589 - 595. https: // doi. org / 10.11646 / zootaxa. 3860.6.7" type="journal article" year="2014">Jałoszyński, 2014a</bibRefCitation>
</taxonomicName>
(all known to occur in
<collectingCountry id="F37176596979FFE5169DAD15FBAAFA94" box="[968,1036,1305,1331]" name="Japan" pageId="6">Japan</collectingCountry>
),
<taxonomicName id="4C664D4A6979FFE51175AD16FACDFA94" authority="Jaloszynski, 2009" authorityName="Jaloszynski" authorityYear="2009" box="[1056,1387,1305,1331]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B912EADB6979FFE51175AD16FB3CFA94" box="[1056,1178,1305,1331]" italics="true" pageId="6">V. modesta</emphasis>
<bibRefCitation id="EFF74B386979FFE511F6AD15FACDFA94" author="Jaloszynski, P." box="[1187,1387,1305,1331]" pageId="6" pagination="43 - 48" refId="ref5675" refString="Jaloszynski, P. (2009) Two new species of Veraphis Casey from China (Coleoptera, Scydmaenidae). Genus, 20 (1), 43 - 48." type="journal article" year="2009">Jałoszyński, 2009</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C664D4A6979FFE515C2AD32FD90FAFF" authority="Jaloszynski, 2013" authorityName="Jaloszynski" authorityYear="2013" box="[151,566,1341,1368]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="assingi">
<emphasis id="B912EADB6979FFE515C2AD32FEA5FAF0" box="[151,259,1342,1367]" italics="true" pageId="6">V. assingi</emphasis>
<bibRefCitation id="EFF74B386979FFE51459AD31FE72FAF0" author="Jaloszynski, P." box="[268,468,1341,1367]" pageId="6" pagination="495 - 503" refId="ref5753" refString="Jaloszynski, P. (2013) Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa, 3609 (5), 495 - 503. https: // doi. org / 10.11646 / zootaxa. 3609.5.5" type="journal article" year="2013">Jałoszyński, 2013</bibRefCitation>
(
<collectingCountry id="F37176596979FFE514B0AD31FD88FAFF" box="[485,558,1341,1368]" name="China" pageId="6">China</collectingCountry>
)
</taxonomicName>
. Also
<taxonomicName id="4C664D4A6979FFE51728AD32FC5AFAF0" authority="Jaloszynski, 2012" authorityName="Jaloszynski" authorityYear="2012" box="[637,1020,1341,1367]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="qinghaiensis">
<emphasis id="B912EADB6979FFE51728AD32FC8EFAF0" box="[637,808,1341,1367]" italics="true" pageId="6">V. qinghaiensis</emphasis>
<bibRefCitation id="EFF74B386979FFE51665AD31FC5AFAF0" author="Jaloszynski, P." box="[816,1020,1341,1367]" pageId="6" pagination="63 - 68" refId="ref5707" refString="Jaloszynski, P. (2012) Two new species of Veraphis Casey (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa, 3322 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 3322.1.4" type="journal article" year="2012">Jałoszyński, 2012</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C664D4A6979FFE5115DAD32FAC1FAF0" authority="Jaloszynski, 2012" authorityName="Jaloszynski" authorityYear="2012" box="[1032,1383,1341,1367]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="calcarifer">
<emphasis id="B912EADB6979FFE5115DAD32FB35FAF0" box="[1032,1171,1341,1367]" italics="true" pageId="6">V. calcarifer</emphasis>
<bibRefCitation id="EFF74B386979FFE511C9AD31FAC1FAF0" author="Jaloszynski, P." box="[1180,1383,1341,1367]" pageId="6" pagination="63 - 68" refId="ref5707" refString="Jaloszynski, P. (2012) Two new species of Veraphis Casey (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa, 3322 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 3322.1.4" type="journal article" year="2012">Jałoszyński, 2012</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C664D4A6979FFE515C2AD6EFDECFADB" authority="Jaloszynski, 2013" authorityName="Jaloszynski" authorityYear="2013" box="[151,586,1377,1404]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="gansuana">
<emphasis id="B912EADB6979FFE515C2AD6EFE86FADC" box="[151,288,1378,1403]" italics="true" pageId="6">V. gansuana</emphasis>
<bibRefCitation id="EFF74B386979FFE51473AD6DFE4BFADC" author="Jaloszynski, P." box="[294,493,1377,1403]" pageId="6" pagination="495 - 503" refId="ref5753" refString="Jaloszynski, P. (2013) Three new species of Eutheiini (Coleoptera, Staphylinidae, Scydmaeninae) from China. Zootaxa, 3609 (5), 495 - 503. https: // doi. org / 10.11646 / zootaxa. 3609.5.5" type="journal article" year="2013">Jałoszyński, 2013</bibRefCitation>
(
<collectingCountry id="F37176596979FFE514A9AD6DFDE4FADB" box="[508,578,1377,1404]" name="China" pageId="6">China</collectingCountry>
)
</taxonomicName>
have similar aedeagi, but they have modified male protrochanters.
<taxonomicName id="4C664D4A6979FFE5106FAD6DFF7CFA38" authorityName="Jałoszyński" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="sakaii">
<emphasis id="B912EADB6979FFE5106FAD6DFF7CFA38" italics="true" pageId="6">Veraphis sakaii</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C664D4A6979FFE51438AD8AFE4CFA38" baseAuthorityName="K. Sawada" baseAuthorityYear="1962" box="[365,490,1414,1439]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B912EADB6979FFE51438AD8AFE4CFA38" box="[365,490,1414,1439]" italics="true" pageId="6">V. japonica</emphasis>
</taxonomicName>
in a much shorter apical portion of the median lobe, which is broader than long (distinctly elongate in
<taxonomicName id="4C664D4A6979FFE5143CADA6FE41FA64" baseAuthorityName="K. Sawada" baseAuthorityYear="1962" box="[361,487,1450,1475]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B912EADB6979FFE5143CADA6FE41FA64" box="[361,487,1450,1475]" italics="true" pageId="6">V. japonica</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C664D4A6979FFE51769ADA6FD77FA64" authorityName="Jaloszynski &amp; Hoshina" authorityYear="2005" box="[572,721,1449,1475]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="hisamatsui">
<emphasis id="B912EADB6979FFE51769ADA6FD77FA64" box="[572,721,1449,1475]" italics="true" pageId="6">V. hisamatsui</emphasis>
</taxonomicName>
,
<taxonomicName id="4C664D4A6979FFE517B5ADA6FC01FA64" authorityName="Jaloszynski &amp; Hoshina" authorityYear="2005" box="[736,935,1449,1475]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="odaigaharensis">
<emphasis id="B912EADB6979FFE517B5ADA6FC01FA64" box="[736,935,1449,1475]" italics="true" pageId="6">V. odaigaharensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C664D4A6979FFE5168BADA6FBD1FA64" authorityName="Jaloszynski &amp; Hoshina" authorityYear="2005" box="[990,1143,1450,1475]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="tottoriensis">
<emphasis id="B912EADB6979FFE5168BADA6FBD1FA64" box="[990,1143,1450,1475]" italics="true" pageId="6">V. tottoriensis</emphasis>
</taxonomicName>
in the aedeagus gradually broadening from base to subapical region (with distinctly concave sides in submedian region in these three species); from
<taxonomicName id="4C664D4A6979FFE51580ADFEFEC3F9AC" authorityName="Jaloszynski" authorityYear="2014" box="[213,357,1521,1547]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="yoshitomii">
<emphasis id="B912EADB6979FFE51580ADFEFEC3F9AC" box="[213,357,1521,1547]" italics="true" pageId="6">V. yoshitomii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C664D4A6979FFE514C9ADFEFDA0F9AC" authorityName="Jaloszynski" authorityYear="2013" box="[412,518,1522,1547]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="assingi">
<emphasis id="B912EADB6979FFE514C9ADFEFDA0F9AC" box="[412,518,1522,1547]" italics="true" pageId="6">V. assingi</emphasis>
</taxonomicName>
in a symmetrical apex of the median lobe (strongly asymmetrical in
<taxonomicName id="4C664D4A6979FFE51059ADFEFA3BF9AC" authorityName="Jaloszynski" authorityYear="2014" box="[1292,1437,1521,1547]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="yoshitomii">
<emphasis id="B912EADB6979FFE51059ADFEFA3BF9AC" box="[1292,1437,1521,1547]" italics="true" pageId="6">V. yoshitomii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C664D4A6979FFE51593AE1AFE96F988" authorityName="Jaloszynski" authorityYear="2013" box="[198,304,1558,1583]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="assingi">
<emphasis id="B912EADB6979FFE51593AE1AFE96F988" box="[198,304,1558,1583]" italics="true" pageId="6">V. assingi</emphasis>
</taxonomicName>
); and from
<taxonomicName id="4C664D4A6979FFE514E0AE1AFD8BF988" authorityName="Jaloszynski" authorityYear="2009" box="[437,557,1557,1583]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B912EADB6979FFE514E0AE1AFD8BF988" box="[437,557,1557,1583]" italics="true" pageId="6">V. modesta</emphasis>
</taxonomicName>
in a rounded apex of the median lobe and parameres longer than half length of the median lobe (apex shallowly and narrowly emarginate and parameres shorter than half length of median lobe in
<taxonomicName id="4C664D4A6979FFE515C2AE52FEB6F9D0" authorityName="Jaloszynski" authorityYear="2009" box="[151,272,1629,1655]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B912EADB6979FFE515C2AE52FEB6F9D0" box="[151,272,1629,1655]" italics="true" pageId="6">V. modesta</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>