158 lines
16 KiB
XML
158 lines
16 KiB
XML
<document ID-DOI="10.11646/zootaxa.5231.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7575308" ID-ZooBank="EBD53535-397E-4894-B0A4-9AC3CC38DF3C" approvalRequired="228" approvalRequired_for_taxonomicNames="8" approvalRequired_for_textStreams="89" approvalRequired_for_treatments="131" checkinTime="1674811458118" checkinUser="plazi" docAuthor="Lelej, Arkady S. & Williams, Kevin A." docDate="2023" docId="03CF87A9313DB511FF17F92BF07A345F" docLanguage="en" docName="zootaxa.5231.2.1.pdf" docOrigin="Zootaxa 5231 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Karunaratnea Lelej 2005" docType="treatment" docVersion="2" lastPageNumber="116" masterDocId="FFF6FFD13133B51EFF80FF97F345374A" masterDocTitle="Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)" masterLastPageNumber="140" masterPageNumber="101" pageNumber="115" updateTime="1674812169880" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lelej, Arkady S.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-7501-0981</mods:nameIdentifier>
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<mods:affiliation>Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia. lelej @ biosoil. ru; https: // orcid. org / 0000 - 0001 - 7501 - 0981</mods:affiliation>
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<mods:nameIdentifier type="email">lelej@biosoil.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Williams, Kevin A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-6073-1070</mods:nameIdentifier>
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<mods:affiliation>Plant Pest Diagnostic Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832, USA. Kevin. Williams @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 6073 - 1070</mods:affiliation>
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<mods:nameIdentifier type="email">illiams@cdfa.ca.gov</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-27</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5231</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>101</mods:start>
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<mods:end>140</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5231.2.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5231.2.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7575308</mods:identifier>
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<mods:identifier type="ZooBank">EBD53535-397E-4894-B0A4-9AC3CC38DF3C</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7575372" ID-Zenodo-Dep="7575372" LSID="urn:lsid:plazi:treatment:03CF87A9313DB511FF17F92BF07A345F" httpUri="http://treatment.plazi.org/id/03CF87A9313DB511FF17F92BF07A345F" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115">
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<subSubSection box="[151,509,1724,1750]" pageId="14" pageNumber="115" type="nomenclature">
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<paragraph blockId="14.[151,509,1724,1786]" box="[151,509,1724,1750]" pageId="14" pageNumber="115">
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<heading bold="true" box="[151,509,1724,1750]" fontSize="11" level="1" pageId="14" pageNumber="115" reason="1">
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<emphasis bold="true" box="[151,509,1724,1750]" pageId="14" pageNumber="115">
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11.
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<taxonomicName authority="Lelej, 2005" authorityName="Lelej" authorityYear="2005" box="[194,509,1724,1750]" class="Insecta" family="Mutillidae" genus="Karunaratnea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[194,365,1724,1750]" italics="true" pageId="14" pageNumber="115">Karunaratnea</emphasis>
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<bibRefCitation author="Lelej, A. S." box="[372,509,1724,1750]" pageId="14" pageNumber="115" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej, 2005</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,350,1760,1786]" pageId="14" pageNumber="115" type="description">
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<paragraph blockId="14.[151,509,1724,1786]" box="[151,350,1760,1786]" pageId="14" pageNumber="115">
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(
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<figureCitation box="[159,289,1760,1786]" captionStart="FIGURES 26–34" captionStartId="16.[152,270,1753,1777]" captionTargetBox="[151,1436,183,1728]" captionTargetId="figure-16@16.[151,1436,181,1729]" captionTargetPageId="16" captionText="FIGURES 26–34. 26. Guineomutilla mateui Suárez, 1977, ♀, dorsal habitus; 27. Gynandrotilla ferruginea Arnold, 1946, ♁, dorsal habitus; 28–29. Indratilla gynandromorpha Lelej, 1993: 28. ♀, dorsal habitus, 29. ♁, holotype, dorsal habitus; 30–34. Karunaratnea palatupanae Lelej, 2005: 30–31. Paratype, ♀ (30. Dorsal habitus, 31. Lateral habitus); 32–34. Holotype, ♁: 32. Dorsal habitus, 33. Head, face view, 34. Genitalia, ventral view. (Photographs by D. Brothers (26, 27), V. Loktionov (28, 29), and A. Lelej (30–34))." figureDoi="http://doi.org/10.5281/zenodo.7575316" httpUri="https://zenodo.org/record/7575316/files/figure.png" pageId="14" pageNumber="115">Figs 30–34</figureCitation>
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,
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<figureCitation box="[300,342,1760,1786]" captionStart="FIGURES 110–115" captionStartId="33.[152,270,1115,1139]" captionTargetBox="[151,1433,181,1091]" captionTargetId="figure-17@33.[151,1436,181,1091]" captionTargetPageId="33" captionText="FIGURES 110–115. Smicromyrmini, ♀, pygidial plate (T6). 110.Andreimyrme rong Williams in Okayasu et al., 2021, holotype; 111. Dentilla curtiventris (André, 1901); 112. Ephucilla thalia (Mickel, 1933); 113. Ephutomma fletcheri (Turner, 1911); 114. Karunaratnea palatupanae Lelej, 2005, paratype; 115. Nemka viduata (Pallas, 1773). (Photographs by K. Williams andA. Lelej (114))." figureDoi="http://doi.org/10.5281/zenodo.7575342" httpUri="https://zenodo.org/record/7575342/files/figure.png" pageId="14" pageNumber="115">114</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="115" type="reference_group">
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<paragraph blockId="14.[151,1436,1828,1883]" pageId="14" pageNumber="115">
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<treatmentCitationGroup pageId="14" pageNumber="115">
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<treatmentCitation author="Lelej, A. S." box="[151,471,1828,1852]" page="145" pageId="14" pageNumber="115" year="2005">
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<taxonomicName authority="Lelej, 2005: 145" authorityName="Lelej" authorityPageNumber="145" authorityYear="2005" box="[151,471,1828,1852]" class="Insecta" family="Mutillidae" genus="Karunaratnea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,295,1829,1852]" italics="true" pageId="14" pageNumber="115">Karunaratnea</emphasis>
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<bibRefCitation author="Lelej, A. S." box="[302,471,1828,1852]" pageId="14" pageNumber="115" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej, 2005: 145</bibRefCitation>
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</taxonomicName>
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, 148 (key), 181 (description), ♁,
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<specimenCount box="[813,833,1829,1852]" pageId="14" pageNumber="115" type="female">♀</specimenCount>
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;
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<treatmentCitation author="Lelej, A. S. & Brothers, D. J." box="[846,1117,1828,1853]" page="30" pageId="14" pageNumber="115" year="2008">
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<bibRefCitation author="Lelej, A. S. & Brothers, D. J." box="[846,1117,1828,1853]" pageId="14" pageNumber="115" pagination="1 - 79" refId="ref27054" refString="Lelej, A. S. & Brothers, D. J. (2008) The genus-group names of Mutillidae (Hymenoptera) and their type species, with a new genus, new name, new synonymies, new combinations and lectotypifications. Zootaxa, 1889 (1), 1 - 79. https: // doi. org / 10.11646 / zootaxa. 1889.1.1" type="journal article" year="2008">Lelej & Brothers 2008: 30</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Brothers, D. J. & Lelej, A. S." box="[1129,1399,1828,1853]" page="95" pageId="14" pageNumber="115" year="2017">
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<bibRefCitation author="Brothers, D. J. & Lelej, A. S." box="[1129,1399,1828,1853]" pageId="14" pageNumber="115" pagination="1 - 97" refId="ref25501" refString="Brothers, D. J. & Lelej, A. S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1 - 97. https: // doi. org / 10.3897 / jhr. 60.20091" type="journal article" year="2017">Brothers & Lelej 2017: 95</bibRefCitation>
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</treatmentCitation>
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, ♁,
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<specimenCount box="[199,219,1860,1883]" pageId="14" pageNumber="115" type="female">♀</specimenCount>
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;
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<treatmentCitation author="Pagliano, G. & Brothers, D. J. & Cambra, R. & Lelej, A. S. & Lo Cascio, P. & Matteini Palmerini, M. & Scaramozzino, P. L. & Williams, K. A. & Romano, M." box="[232,489,1859,1883]" page="174" pageId="14" pageNumber="115" year="2020">
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<bibRefCitation author="Pagliano, G. & Brothers, D. J. & Cambra, R. & Lelej, A. S. & Lo Cascio, P. & Matteini Palmerini, M. & Scaramozzino, P. L. & Williams, K. A. & Romano, M." box="[232,489,1859,1883]" pageId="14" pageNumber="115" pagination="5 - 425" refId="ref28648" refString="Pagliano, G., Brothers, D. J., Cambra, R., Lelej, A. S., Lo Cascio, P., Matteini Palmerini, M., Scaramozzino, P. L., Williams, K. A. & Romano, M. (2020 [" 2018 "]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torin o, 36 (1 - 2), 5 - 425." type="journal article" year="2020">
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Pagliano
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<emphasis box="[327,378,1859,1883]" italics="true" pageId="14" pageNumber="115">et al.</emphasis>
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2020: 174
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</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,1316,1929,1958]" pageId="14" pageNumber="115" type="type_taxon">
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<paragraph blockId="14.[151,1436,1929,2027]" box="[151,1316,1929,1958]" pageId="14" pageNumber="115">
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<emphasis bold="true" box="[151,309,1929,1955]" pageId="14" pageNumber="115">Type species:</emphasis>
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<taxonomicName authority="Lelej, 2005" authorityName="Lelej" authorityYear="2005" box="[316,759,1929,1956]" class="Insecta" family="Mutillidae" genus="Karunaratnea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="palatupanae">
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<emphasis box="[316,621,1929,1955]" italics="true" pageId="14" pageNumber="115">Karunaratnea palatupanae</emphasis>
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<bibRefCitation author="Lelej, A. S." box="[628,759,1929,1956]" pageId="14" pageNumber="115" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej, 2005</bibRefCitation>
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</taxonomicName>
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, ♁,
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<specimenCount box="[805,826,1930,1956]" pageId="14" pageNumber="115" type="female">♀</specimenCount>
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(
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<typeStatus box="[840,940,1929,1955]" pageId="14" pageNumber="115">holotype</typeStatus>
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is male), by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115" type="diagnosis">
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<paragraph blockId="14.[151,1436,1929,2027]" lastBlockId="15.[151,1437,150,789]" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115">
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<emphasis bold="true" box="[199,314,1965,1991]" pageId="14" pageNumber="115">Diagnosis</emphasis>
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. MALE. Head angulate behind the eyes. Mandible tridentate, deeply excised beneath with large subbasal tooth. Clypeus rather flattened with projecting midpart of anterior margin. Ocelli small, POD much less than OOD. Scape distinctly bicarinate beneath, flagellomeres not shortened, F1 1.5 × longer its width and slightly shorter than F2. Pronotum angulate. Notauli and parapsids developed in posterior half. Tegula slightly projected over mesoscuto-scutellar suture. Parascutal carina weak. Mesoscutellum simple. Mesopleuron beneath without precoxal ridge. Propodeum shortened, dorsum with two medial sharp tubercles bordered medial elongate cell. Metacoxa carinate inside. Wing shortened, slightly infuscated, stigmatic cell approximately equal to distance between base of stigmatic cell and origin of
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<emphasis box="[462,493,331,357]" italics="true" pageId="15" pageNumber="116">RS</emphasis>
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on vein
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<emphasis box="[592,618,331,357]" italics="true" pageId="15" pageNumber="116">Sc</emphasis>
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. T2 with long lateral felt lines, S2 without any trace of felt lines. T7 and S8 punctured throughout. Volsella slightly longer than penial valve, cuspis with tuft of short setae inside and ventral basal setae no longer than cuspis length. FEMALE. Fore tarsus without external comb of spines. Mandible slender bidentate. Clypeus with transverse lamellate carina and small medial basal tubercle. Genal carina weak and forming tubercle on hypostomal carina. F1 1.25 × longer than its maximum width. Mesosoma slightly divergent posterad, shortened, length equal its maximum width. Posterior propodeal slope abrupt. Scutellar scale well developed. Mid and hind tibia with two row of spines. T6 with shiny smooth medial part not carinated laterally but bordered by pale setae. T2 with posterior narrow fringe of pale setae widened medially, basally with or without medial patch of pale setae. T3 with band of pale setae.
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</paragraph>
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<paragraph blockId="15.[151,1437,150,789]" pageId="15" pageNumber="116">
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<emphasis bold="true" box="[199,508,654,681]" pageId="15" pageNumber="116">Diversity and distribution</emphasis>
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. Four species (one based on females only, two based on males only, and one known from both sexes) are recognized from
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<collectingRegion box="[579,753,691,717]" country="Oman" name="Shamal ash Sharqiyah" pageId="15" pageNumber="116">Oriental region</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="116" type="discussion">
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<paragraph blockId="15.[151,1437,150,789]" pageId="15" pageNumber="116">
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<emphasis bold="true" box="[199,308,727,753]" pageId="15" pageNumber="116">Remarks</emphasis>
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.
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<bibRefCitation author="Lelej, A. S." box="[321,460,726,753]" pageId="15" pageNumber="116" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej (2005)</bibRefCitation>
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suggested that many undescribed species occur in
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<collectingCountry box="[1026,1084,727,753]" name="India" pageId="15" pageNumber="116">India</collectingCountry>
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and
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<collectingCountry box="[1137,1249,727,753]" name="Sri Lanka" pageId="15" pageNumber="116">Sri Lanka</collectingCountry>
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, so no keys have been produced for separating the members of
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<taxonomicName authorityName="Lelej" authorityYear="2005" box="[665,824,763,788]" class="Insecta" family="Mutillidae" genus="Karunaratnea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="genus">
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<emphasis box="[665,824,763,788]" italics="true" pageId="15" pageNumber="116">Karunaratnea</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |