treatments-xml/data/03/CF/87/03CF87A93136B519FF17FF00F6D934B5.xml

<document ID-DOI="10.11646/zootaxa.5231.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7575308" ID-ZooBank="EBD53535-397E-4894-B0A4-9AC3CC38DF3C" checkinTime="1674811458118" checkinUser="plazi" docAuthor="Lelej, Arkady S. &amp; Williams, Kevin A." docDate="2023" docId="03CF87A93136B519FF17FF00F6D934B5" docLanguage="en" docName="zootaxa.5231.2.1.pdf" docOrigin="Zootaxa 5231 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Smicromyrmini Bischoff 1920" docType="key" docVersion="1" lastPageNumber="108" masterDocId="FFF6FFD13133B51EFF80FF97F345374A" masterDocTitle="Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)" masterLastPageNumber="140" masterPageNumber="101" pageNumber="106" updateTime="1674840100621" updateUser="valdenar" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lelej, Arkady S.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7501-0981</mods:nameIdentifier>
<mods:affiliation>Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia. lelej @ biosoil. ru; https: // orcid. org / 0000 - 0001 - 7501 - 0981</mods:affiliation>
<mods:nameIdentifier type="email">lelej@biosoil.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Williams, Kevin A.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6073-1070</mods:nameIdentifier>
<mods:affiliation>Plant Pest Diagnostic Center, California Department of Food &amp; Agriculture, 3294 Meadowview Road, Sacramento, CA 95832, USA. Kevin. Williams @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 6073 - 1070</mods:affiliation>
<mods:nameIdentifier type="email">illiams@cdfa.ca.gov</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-27</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5231</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>101</mods:start>
<mods:end>140</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5231.2.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5231.2.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7575308</mods:identifier>
<mods:identifier type="ZooBank">EBD53535-397E-4894-B0A4-9AC3CC38DF3C</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03CF87A93136B519FF17FF00F6D934B5" httpUri="http://treatment.plazi.org/id/03CF87A93136B519FF17FF00F6D934B5" lastPageId="7" lastPageNumber="108" pageId="5" pageNumber="106">
<subSubSection box="[151,914,151,177]" pageId="5" pageNumber="106" type="nomenclature">
<paragraph blockId="5.[151,914,151,177]" box="[151,914,151,177]" pageId="5" pageNumber="106">
<heading bold="true" box="[151,914,151,177]" fontSize="11" level="1" pageId="5" pageNumber="106" reason="1">
<emphasis bold="true" box="[151,914,151,177]" pageId="5" pageNumber="106">
Key to the genera and subgenera of the tribe 
<taxonomicName authorityName="Bischoff" authorityYear="1920" box="[716,914,151,177]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="tribe" tribe="Smicromyrmini">Smicromyrmini</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,1080,223,250]" pageId="5" pageNumber="106" type="multiple">
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1080,223,250]" pageId="5" pageNumber="106">
<emphasis bold="true" box="[151,223,223,249]" pageId="5" pageNumber="106">Males</emphasis>
(unknown in 
<taxonomicName box="[379,541,223,249]" class="Insecta" family="Mutillidae" genus="Guineomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis box="[379,541,223,249]" italics="true" pageId="5" pageNumber="106">Guineomutilla</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Lelej" authorityYear="1980" box="[555,690,223,249]" class="Insecta" genus="Nuristanilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis box="[555,690,223,249]" italics="true" pageId="5" pageNumber="106">Nuristanilla</emphasis>
</taxonomicName>
, and 
<taxonomicName authorityName="Lelej &amp; Williams" authorityYear="2023" box="[752,906,223,249]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis box="[752,906,223,249]" italics="true" pageId="5" pageNumber="106">Smicromyrme</emphasis>
</taxonomicName>
(
<taxonomicName box="[922,1060,223,249]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="131" phylum="Arthropoda" rank="subGenus" subGenus="Rhombotilla">
<emphasis box="[922,1060,223,249]" italics="true" pageId="5" pageNumber="106">Rhombotilla</emphasis>
</taxonomicName>
))
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="107" pageId="5" pageNumber="106" type="key">
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,256,278]" pageId="5" pageNumber="106">1. Apterous........................................................................................... 2</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,285,307]" pageId="5" pageNumber="106">- Alates.............................................................................................. 3</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
2. T1 long with distinct dorsal face, with lateral white spots. T2 black with medial subbasal white spot and two postero-lateral white spots. Tegula long, projecting over mesoscuto-scutellar suture.—Volsella short, half gonostylus length, ventrally with dense setae longer than basal gonostylus width (Ori)..................................... 10. 
<taxonomicName authority="Lelej, 1993" authorityName="Lelej" authorityYear="1993" box="[1226,1436,373,395]" class="Insecta" family="Mutillidae" genus="Indratilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1226,1321,373,395]" italics="true" pageId="5" pageNumber="106">Indratilla</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,373,395]" pageId="5" pageNumber="106" pagination="233 - 236" refId="ref26666" refString="Lelej, A. S. (1993) A new genus of velvet ants from Sri Lanka (Insecta, Hymenoptera, Mutillidae). Spixiana, 16 (3), 233 - 236." type="journal article" year="1993">Lelej, 1993</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
- T1 short, without distinct dorsal face and lateral white spots. T2 ferruginous with black posterior third and posterior white fringe. Tegula short, scarcely projecting over mesoscuto-scutellar suture (Afr)............. 9. 
<taxonomicName authority="Arnold, 1946" authorityName="Arnold" authorityYear="1946" box="[1165,1436,432,454]" class="Insecta" genus="Gynandrotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1165,1304,432,454]" italics="true" pageId="5" pageNumber="106">Gynandrotilla</emphasis>
<bibRefCitation author="Arnold, G." box="[1308,1436,432,454]" pageId="5" pageNumber="106" pagination="49 - 97" refId="ref24967" refString="Arnold, G. (1946) New species of African Hymenoptera. No. 6. Occasional Papers of the National Museum of Southern Rhodesia, 12, 49 - 97." type="journal article" year="1946">Arnold, 1946</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,461,483]" pageId="5" pageNumber="106">3. S2 with short lateral felt lines, sometimes reduced to a few small punctures...................................... 4</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,491,513]" pageId="5" pageNumber="106">- S2 without any trace of lateral felt lines.................................................................. 16</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,520,542]" pageId="5" pageNumber="106">4. Mandible strongly widened apically, preapical inner teeth equal to or larger than apical tooth......................... 5</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
- Mandible not widened apically, preapical inner teeth distinctly smaller than apical tooth, second preapical tooth sometimes (
<taxonomicName authorityName="Bischoff" authorityYear="1920" box="[234,419,579,600]" class="Insecta" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis box="[234,419,579,600]" italics="true" pageId="5" pageNumber="106">Pseudocephalotilla</emphasis>
</taxonomicName>
) larger than apical tooth............................................................... 6
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
5. Inner eye margin with deep notch. Stigmatic cell 0.7 × distance between base of stigmatic cell and origin of 
<emphasis box="[1265,1292,608,629]" italics="true" pageId="5" pageNumber="106">RS</emphasis>
on vein 
<emphasis box="[1375,1398,608,629]" italics="true" pageId="5" pageNumber="106">Sc</emphasis>
. T2 width 1.2 × propodeal width (Ori)............................................... 26. 
<taxonomicName authority="Lelej, 1995" authorityName="Lelej" authorityYear="1995" box="[1171,1436,637,659]" class="Insecta" family="Mutillidae" genus="Tsunekimyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1171,1321,637,659]" italics="true" pageId="5" pageNumber="106">Tsunekimyrme</emphasis>
Lelej, 1995
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
- Inner eye margin with weak notch. Stigmatic cell 1.4 × distance between base of stigmatic cell and origin of 
<emphasis box="[1266,1293,667,688]" italics="true" pageId="5" pageNumber="106">RS</emphasis>
on vein 
<emphasis box="[1376,1399,667,688]" italics="true" pageId="5" pageNumber="106">Sc</emphasis>
. T2 width 1.5 × propodeal width (Afr, Ori, Pal).............................. 5. 
<taxonomicName authority="Lelej" authorityName="Lelej" authorityYear="1980" box="[1045,1180,696,718]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1045,1125,696,718]" italics="true" pageId="5" pageNumber="106">Dentilla</emphasis>
Lelej
</taxonomicName>
in 
<bibRefCitation author="Lelej, A. S. &amp; Kabakov, O. N." box="[1211,1436,696,718]" pageId="5" pageNumber="106" pagination="141 - 156" refId="ref27122" refString="Lelej, A. S. &amp; Kabakov, O. N. (1980) On Bradynobaenidae and Mutillidae (Hymenoptera) of Afghanistan. Entomologicheskoe Obozrenie, 59 (1), 181 - 196. [in Russian, English translation: (1981) Entomological Review, 59 (1), 141 - 156]" type="journal article" year="1980">Lelej &amp; Kabakov, 1980</bibRefCitation>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,725,747]" pageId="5" pageNumber="106">6. Flagellomeres overlapped, F1–5 with row of tyloids or inner part of F1–7(10) with apophyses (Afr)................... 7</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,755,777]" pageId="5" pageNumber="106">- Flagellomeres not overlapped, F1–5 without row of tyloids and inner part without apophyses......................... 9</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,784,806]" pageId="5" pageNumber="106">
7. Flagellomeres overlapped, F1–5 with row of tyloids. T7 with cuticular yellowish spot (Afr).. 2. 
<taxonomicName authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" box="[1164,1436,784,806]" class="Insecta" genus="Antennotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1164,1288,784,806]" italics="true" pageId="5" pageNumber="106">Antennotilla</emphasis>
<bibRefCitation author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[1294,1436,784,806]" pageId="5" pageNumber="106" refId="ref25275" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv f ¸ r Naturgeschichte, 86 A (1 - 3) &amp; 86 A (4 - 5), 1 - 480 (1920) &amp; 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,813,835]" pageId="5" pageNumber="106">- F1–7(10) with apophyses. T7 black, without cuticular yellowish spot............................................ 8</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
8. Double series of apophyses different: longer one ~6 × flagellomere diameter, shorter one equal to flagellomere length. Mandible bidentate, with strong subbasal tooth ventrally (Afr)............................... 4. 
<taxonomicName authority="Nonveiller, 1993" authorityName="Nonveiller" authorityYear="1993" box="[1145,1436,872,894]" class="Insecta" family="Mutillidae" genus="Ctenoceraea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1145,1269,872,893]" italics="true" pageId="5" pageNumber="106">Ctenoceraea</emphasis>
<bibRefCitation author="Nonveiller, G." box="[1275,1436,872,894]" pageId="5" pageNumber="106" pagination="15 - 18" refId="ref28072" refString="Nonveiller, G. (1993) Studies on African Mutillidae. XIV. Ctenoceraea, a new genus with pectinate antennae and description of the type-species C. pectinella sp. n. (Hymenoptera). Revue Francaise d'Entomologie, New Series, 15 (1), 15 - 18." type="journal article" year="1993">Nonveiller, 1993</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
- Double series of apophyses more or less equal in length. Mandible tridentate, with ventral basal carina, without subbasal tooth (Afr)................................................................... 19. 
<taxonomicName authority="Latreille, 1825" authorityName="Latreille" authorityYear="1825" box="[1133,1436,930,953]" class="Insecta" family="Mutillidae" genus="Psammotherma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1133,1289,930,952]" italics="true" pageId="5" pageNumber="106">Psammotherma</emphasis>
<bibRefCitation author="Latreille, P. A." box="[1295,1436,931,953]" pageId="5" pageNumber="106" refId="ref26366" refString="Latreille, P. A. (1825) Famillies Naturelles du Regne Animal, exposees succinctement et dans un Ordre Analytique, avec l'Indication de leurs Genres. Bailliere, Paris, 570 pp. https: // doi. org / 10.5962 / bhl. title. 16094" type="book" year="1825">Latreille, 1825</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,960,982]" pageId="5" pageNumber="106">9. Genital volsella with basal external lobe and long narrow cuspis; basivolsella with long setae....................... 10</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,989,1011]" pageId="5" pageNumber="106">- Genital volsella without basal external lobe or long narrow cuspis; basivolsella with setae short or absent.............. 14</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1018,1041]" pageId="5" pageNumber="106">
10. Mandible bidentate at apex (Ori, Pal)............................................. 12. 
<taxonomicName authority="Lelej, 1995" authorityName="Lelej" authorityYear="1995" box="[1166,1436,1018,1041]" class="Insecta" family="Mutillidae" genus="Mickelomyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1166,1321,1018,1040]" italics="true" pageId="5" pageNumber="106">Mickelomyrme</emphasis>
Lelej, 1995
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1048,1070]" pageId="5" pageNumber="106">- Mandible tridentate at apex............................................................................ 11</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1077,1099]" pageId="5" pageNumber="106">11. Mesopleuron beneath with one or two pairs of precoxal denticles or tubercles.................................... 12</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1107,1129]" pageId="5" pageNumber="106">- Mesopleuron beneath without precoxal denticles or tubercles................................................. 13</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
12. Hypostomal bridge without tooth; eye broadly oval; prementum evenly convex or weakly medio-longitudinally carinate; oblique metapleural suture running anterodorsally from endophragmal pit; mesopleron beneath with precoxal paired transverse/oblique carinae (may be toothed mesally) (Ori, Pal).............................................. 13. 
<taxonomicName authority="Lelej, 1985" authorityName="Lelej" authorityYear="1985" box="[1246,1436,1194,1217]" class="Insecta" family="Mutillidae" genus="Nemka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1246,1318,1194,1216]" italics="true" pageId="5" pageNumber="106">Nemka</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1325,1436,1195,1217]" pageId="5" pageNumber="106" refId="ref26618" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) Fauny SSSR i Sopredelnykh Stran [The Velvet Ants (Hymenoptera, Mutillidae) of the USSR and Neighbouring Countries]. Nauka, Leningrad, 268 pp. [in Russian]" type="book" year="1985">Lelej, 1985</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
- Hypostomal bridge with tooth; eye subcircular; prementum with posteromedian domelike elevation; oblique metapleural suture running horizontally from endophragmal pit; mesopleuron beneath with precoxal paired transverse tubercles and teeth (Afr)............................................................................. 25. 
<taxonomicName authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" box="[1193,1436,1282,1305]" class="Insecta" genus="Sulcotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1193,1288,1282,1304]" italics="true" pageId="5" pageNumber="106">Sulcotilla</emphasis>
<bibRefCitation author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[1294,1436,1283,1305]" pageId="5" pageNumber="106" refId="ref25275" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv f ¸ r Naturgeschichte, 86 A (1 - 3) &amp; 86 A (4 - 5), 1 - 480 (1920) &amp; 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
13. Mandible beneath with large subventral tooth, distance between subventral tooth and dorsal mandibular carina equal or more than basal mandibular width. Apex of volsellar cuspis curved inside (Pal)............ 23. 
<taxonomicName authority="Ovtchinnikov, 2002" authorityName="Ovtchinnikov" authorityYear="2002" box="[1131,1436,1341,1363]" class="Insecta" family="Mutillidae" genus="Skorikovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1131,1239,1341,1363]" italics="true" pageId="5" pageNumber="106">Skorikovia</emphasis>
<bibRefCitation author="Ovtchinnikov, S. V." box="[1246,1436,1341,1363]" pageId="5" pageNumber="106" refId="ref28527" refString="Ovtchinnikov, S. V. (2002) Materials to the fauna of velvet ants (Hymenoptera, Mutillidae) of Kyrghizia. In: Zonstein, S. L. (Ed.), Entomological Investigations in Kyrgystan. Vol. 22. Kyrghyz Society of Entomology, Bishkek, pp. 79 - 104. [in Russian]" type="book" year="2002">Ovtchinnikov, 2002</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
- Mandible beneath with small subventral tooth, distance between subventral tooth and dorsal mandibular carina much less than basal mandibular width. Apex of volsellar cuspis narrowly straight (Afr, Pal)............................................................................................... 21. 
<taxonomicName authority="Lelej" authorityName="Lelej" box="[962,1159,1429,1451]" class="Insecta" family="Mutillidae" genus="Rasnitsynitilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[962,1105,1429,1451]" italics="true" pageId="5" pageNumber="106">Rasnitsynitilla</emphasis>
Lelej
</taxonomicName>
in 
<bibRefCitation author="Lelej, A. S. &amp; van Harten, A." box="[1190,1436,1429,1451]" pageId="5" pageNumber="106" pagination="1 - 50" refId="ref27327" refString="Lelej, A. S. &amp; van Harten, A. (2006) A review of the Mutillidae (Hymenoptera) of Yemen. Zootaxa, 1226 (1), 1 - 50. https: // doi. org / 10.11646 / zootaxa. 1226.1.1" type="journal article" year="2006">Lelej &amp; van Harten, 2006</bibRefCitation>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1458,1481]" pageId="5" pageNumber="106">
14. Penis valve shortened, swollen apex curved down and oriented outside (Afr)........ 20. 
<taxonomicName authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" box="[1100,1436,1458,1481]" class="Insecta" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1100,1288,1458,1480]" italics="true" pageId="5" pageNumber="106">Pseudocephalotilla</emphasis>
<bibRefCitation author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[1294,1436,1459,1481]" pageId="5" pageNumber="106" refId="ref25275" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv f ¸ r Naturgeschichte, 86 A (1 - 3) &amp; 86 A (4 - 5), 1 - 480 (1920) &amp; 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1488,1510]" pageId="5" pageNumber="106">- Penis valve not shortened, apex not swollen and not oriented outside........................................... 15</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
15. F1 length equal its maximal width, and 0.6 × F2. Tegula not projecting or at most scarcely projecting over mesoscuto-scutellar suture (Afr, Ori, Pal)...................................................... 24. 
<taxonomicName authority="Thomson, 1870" authorityName="Thomson" authorityYear="1870" box="[1135,1436,1547,1569]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1135,1280,1547,1568]" italics="true" pageId="5" pageNumber="106">Smicromyrme</emphasis>
<bibRefCitation author="Thomson, C. G." box="[1286,1436,1547,1569]" pageId="5" pageNumber="106" pagination="202 - 251" refId="ref30270" refString="Thomson, C. G. (1870) Ofversigt af Sveriges rofsteklar. Opuscula Entomologica, 1 (2), 202 - 251." type="journal article" year="1870">Thomson, 1870</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
15a. Mandible beneath without subventral tooth, if with inconspicuous one then without excision (Pal)....................................................................................... 24.1. Subgenus 
<taxonomicName authority="Schwartz, 1984" authorityName="Schwartz" authorityYear="1984" box="[1169,1436,1605,1627]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="subGenus" subGenus="Astomyrme">
<emphasis bold="true" box="[1169,1281,1605,1626]" italics="true" pageId="5" pageNumber="106">Astomyrme</emphasis>
<bibRefCitation author="Schwartz, F." box="[1287,1436,1605,1627]" pageId="5" pageNumber="106" pagination="21 - 25" refId="ref29541" refString="Schwartz, F. (1984) Description d'un genre nouveau, disjoint de Smicromyrme, et d'une espece nouvelle du sud de la France (Hymenoptera, Mutillidae). L'Entomologiste, 40, 21 - 25." type="journal article" year="1984">Schwartz, 1984</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1635,1657]" pageId="5" pageNumber="106">- Mandible beneath with well developed subventral tooth separated by excision................................... 15b</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1664,1686]" pageId="5" pageNumber="106">15b. Mandible bidentate. Clypeus more or less flattened........................................................ 15c</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1693,1715]" pageId="5" pageNumber="106">
- Mandible tridentate. Clypeus concave. (Afr, Ori, Pal).......................... 24.3. Subgenus 
<taxonomicName authority="Lelej, 1985" authorityName="Lelej" authorityYear="1985" box="[1221,1436,1693,1715]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="subGenus" subGenus="Erimyrme">
<emphasis bold="true" box="[1221,1321,1693,1714]" italics="true" pageId="5" pageNumber="106">Erimyrme</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,1693,1715]" pageId="5" pageNumber="106" refId="ref26618" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) Fauny SSSR i Sopredelnykh Stran [The Velvet Ants (Hymenoptera, Mutillidae) of the USSR and Neighbouring Countries]. Nauka, Leningrad, 268 pp. [in Russian]" type="book" year="1985">Lelej, 1985</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1722,1745]" pageId="5" pageNumber="106">
15c. Clypeus without preapical teeth (Ori, Pal).................................. 24.2. Subgenus 
<taxonomicName authority="Lelej, 1985" authorityName="Lelej" authorityYear="1985" box="[1216,1436,1722,1745]" class="Insecta" family="Mutillidae" genus="Eremotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1216,1321,1722,1744]" italics="true" pageId="5" pageNumber="106">Eremotilla</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,1723,1745]" pageId="5" pageNumber="106" refId="ref26618" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) Fauny SSSR i Sopredelnykh Stran [The Velvet Ants (Hymenoptera, Mutillidae) of the USSR and Neighbouring Countries]. Nauka, Leningrad, 268 pp. [in Russian]" type="book" year="1985">Lelej, 1985</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1752,1774]" pageId="5" pageNumber="106">
- Clypeus with two preapical teeth or tubercles (Pal)..................... 24.5. Subgenus 
<taxonomicName authority="Thomson, 1870" authorityName="Thomson" authorityYear="1870" box="[1141,1436,1752,1774]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1141,1280,1752,1773]" italics="true" pageId="5" pageNumber="106">Smicromyrme</emphasis>
<bibRefCitation author="Thomson, C. G." box="[1286,1436,1752,1774]" pageId="5" pageNumber="106" pagination="202 - 251" refId="ref30270" refString="Thomson, C. G. (1870) Ofversigt af Sveriges rofsteklar. Opuscula Entomologica, 1 (2), 202 - 251." type="journal article" year="1870">Thomson, 1870</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
15 (-). F1 length 0.5–0.6 × its maximal width, and 0.3–0.4 × F2. Tegula projecting over mesoscuto-scutellar suture for 1/5–1/4 of its length (Ori, Pal).................................................................. 22. 
<taxonomicName authority="Lelej, 1995" authorityName="Lelej" authorityYear="1995" box="[1226,1436,1810,1833]" class="Insecta" family="Mutillidae" genus="Sinotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1226,1320,1810,1832]" italics="true" pageId="5" pageNumber="106">Sinotilla</emphasis>
Lelej, 1995
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1840,1862]" pageId="5" pageNumber="106">16. Mandible strongly widened apically, preapical inner teeth equal to or larger than apical tooth........................ 17</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1436,1869,1891]" pageId="5" pageNumber="106">- Mandible narrowed apically, preapical inner teeth considerably smaller than apical tooth........................... 18</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
17. Mandible not excised beneath, without tooth or lobe near base. F1 length 0.7 × maximal width (
<figureCitation box="[1146,1203,1899,1921]" captionStart="FIGURES 1–6" captionStartId="9.[152,270,1647,1671]" captionTargetBox="[151,1436,342,1619]" captionTargetId="figure-105@9.[151,1436,342,1621]" captionTargetPageId="9" captionText="FIGURES 1–6. 1. Andreimyrme long Lelej, 1995, ♁, holotype, lateral habitus; 2–3. A. rong Williams in Okayasu et al., 2021, ♀, holotype, habitus (2. Dorsal view, 3. Lateral view); 4a, 4b. Antennotilla phoebe (Péringuey, 1899), ♁: (4a. Lateral view, 4b. Dorsal view); 5–6. Corytilla pallidipes Arnold, 1956, ♁ (5. Head, lateral view, 6. Lateral habitus). (Photographs by V. Loktionov (1), D. Brothers (4b), and K. Williams (2, 3, 4a, 5, 6))." figureDoi="http://doi.org/10.5281/zenodo.7575310" httpUri="https://zenodo.org/record/7575310/files/figure.png" pageId="5" pageNumber="106">Fig. 5</figureCitation>
) (Ori, Pal)...................................................................................... 1. 
<taxonomicName authority="Lelej, 1995" authorityName="Lelej" authorityYear="1995" box="[1176,1436,1928,1950]" class="Insecta" family="Mutillidae" genus="Andreimyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1176,1320,1928,1950]" italics="true" pageId="5" pageNumber="106">Andreimyrme</emphasis>
Lelej, 1995
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" pageId="5" pageNumber="106">
- Mandible excised beneath, with large subbasal lobe. F1 length 1.5–1.6 × maximal width (Afr, Ori, Pal).............................................................................................. 7. 
<taxonomicName authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[1149,1436,1986,2009]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1149,1280,1986,2008]" italics="true" pageId="5" pageNumber="106">Ephutomma</emphasis>
<bibRefCitation author="Ashmead, W. H." box="[1286,1436,1987,2009]" pageId="5" pageNumber="106" pagination="45 - 60" refId="ref25074" refString="Ashmead, W. H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45 - 60." type="journal article" year="1899">Ashmead, 1899</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="5.[151,1437,223,2038]" box="[151,1437,2016,2038]" pageId="5" pageNumber="106">18. Mandible beneath without large subbasal tooth or lobe, at most with small projection in basal half................... 19.</paragraph>
<paragraph blockId="6.[151,1437,151,701]" box="[151,1436,151,173]" pageId="6" pageNumber="107">- Mandible beneath with large subbasal lobe or tooth......................................................... 20</paragraph>
<paragraph blockId="6.[151,1437,151,701]" pageId="6" pageNumber="107">
19. Flagellomeres overlapped. Penis valve shortened, apex curved down and oriented outside. T2 with central spot of pale setae (Afr)...................................................... 16. 
<taxonomicName authority="Lelej" authorityName="Lelej" box="[976,1159,210,232]" class="Insecta" family="Mutillidae" genus="Paglianotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[976,1104,210,232]" italics="true" pageId="6" pageNumber="107">Paglianotilla</emphasis>
Lelej
</taxonomicName>
in 
<bibRefCitation author="Lelej, A. S. &amp; van Harten, A." box="[1190,1436,210,232]" pageId="6" pageNumber="107" pagination="1 - 50" refId="ref27327" refString="Lelej, A. S. &amp; van Harten, A. (2006) A review of the Mutillidae (Hymenoptera) of Yemen. Zootaxa, 1226 (1), 1 - 50. https: // doi. org / 10.11646 / zootaxa. 1226.1.1" type="journal article" year="2006">Lelej &amp; van Harten, 2006</bibRefCitation>
</paragraph>
<paragraph blockId="6.[151,1437,151,701]" pageId="6" pageNumber="107">
- Flagellomeres not overlapped. Penis valve not shortened, apex not oriented outside. T2 without central spot of pale setae (Ori)......................................................................... 14. 
<taxonomicName authority="Lelej, 2005" authorityName="Lelej" authorityYear="2005" box="[1204,1436,268,291]" class="Insecta" family="Mutillidae" genus="Nordeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1204,1321,268,290]" italics="true" pageId="6" pageNumber="107">Nordeniella</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,269,291]" pageId="6" pageNumber="107" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej, 2005</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="6.[151,1437,151,701]" pageId="6" pageNumber="107">
20. Brachypterous: forewing slightly projecting over T2. Distance between base of stigmatic cell and origin of 
<emphasis box="[1247,1274,298,319]" italics="true" pageId="6" pageNumber="107">RS</emphasis>
on vein 
<emphasis box="[1356,1379,298,319]" italics="true" pageId="6" pageNumber="107">Sc</emphasis>
equal or even less than stigmatic cell length. Propodeum dorsally with two medial sharp teeth (Ori).................................................................................................... 11. 
<taxonomicName authority="Lelej, 2005" authorityName="Lelej" authorityYear="2005" box="[1181,1436,357,379]" class="Insecta" family="Mutillidae" genus="Karunaratnea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1181,1321,357,378]" italics="true" pageId="6" pageNumber="107">Karunaratnea</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,357,379]" pageId="6" pageNumber="107" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej, 2005</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="6.[151,1437,151,701]" pageId="6" pageNumber="107">
- Wings well developed: forewing touch metasomal apex. Distance between base of stigmatic cell and origin of 
<emphasis box="[1301,1328,386,407]" italics="true" pageId="6" pageNumber="107">RS</emphasis>
on vein 
<emphasis box="[1414,1437,386,407]" italics="true" pageId="6" pageNumber="107">Sc</emphasis>
more than stigmatic cell length. Propodeum dorsally without medial sharp teeth.................................. 21
</paragraph>
<paragraph blockId="6.[151,1437,151,701]" pageId="6" pageNumber="107">21. Genital volsella without wide external basal lobe, inner and lower margins of volsella with setae not longer than volsellar cuspis............................................................................................. 22</paragraph>
<paragraph blockId="6.[151,1437,151,701]" box="[151,1436,503,525]" pageId="6" pageNumber="107">- Genital volsella with more or less wide external basal lobe (basivolsella) with setae longer than volsellar cuspis......... 23</paragraph>
<paragraph blockId="6.[151,1437,151,701]" box="[151,1436,532,555]" pageId="6" pageNumber="107">
22. Vertex with raised subtriangular carinated platform (Afr)................................. 3. 
<taxonomicName authority="Arnold, 1956" authorityName="Arnold" authorityYear="1956" box="[1217,1436,532,555]" class="Insecta" genus="Corytilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1217,1304,532,554]" italics="true" pageId="6" pageNumber="107">Corytilla</emphasis>
<bibRefCitation author="Arnold, G." box="[1308,1436,533,555]" pageId="6" pageNumber="107" pagination="52 - 77" refId="ref25000" refString="Arnold, G. (1956) New species of African Hymenoptera. No. 12. Occasional Papers of the National Museum of Southern Rhodesia, 3 (21 B), 52 - 77." type="journal article" year="1956">Arnold, 1956</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="6.[151,1437,151,701]" box="[151,1436,562,584]" pageId="6" pageNumber="107">
- Vertex without raised subtriangular carinated platform (Ori, Pal)............................ 6. 
<taxonomicName authority="Lelej, 1995" authorityName="Lelej" authorityYear="1995" box="[1224,1436,562,584]" class="Insecta" family="Mutillidae" genus="Ephucilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1224,1321,562,584]" italics="true" pageId="6" pageNumber="107">Ephucilla</emphasis>
Lelej, 1995
</taxonomicName>
</paragraph>
<paragraph blockId="6.[151,1437,151,701]" pageId="6" pageNumber="107">
23. Mandible beneath usually not excised, with rounded subventral lobe. Thickened basivolsellar setae not reaching the apex of gonostylus (Afr, Ori, Pal)........................................................ 18. 
<taxonomicName authority="Andre, 1902" authorityName="Andre" authorityYear="1902" box="[1204,1436,620,643]" class="Insecta" family="Mutillidae" genus="Promecilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1204,1312,620,642]" italics="true" pageId="6" pageNumber="107">Promecilla</emphasis>
<bibRefCitation author="Andre, E." box="[1316,1436,621,643]" pageId="6" pageNumber="107" pagination="1 - 77" refId="ref24877" refString="Andre, E. (1902 b [&quot; 1903 &quot;]) Fam. Mutillidae. In: Wytsman, P., Genera Insectorum. Fasc. 11. s. n., Bruxelles, pp. 1 - 77 + 3 pls." type="journal article" year="1902">André, 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="6.[151,1437,151,701]" pageId="6" pageNumber="107">
- Mandible beneath deeply excised, with large subventral tooth. Thickened basivolsellar setae usually protruding beyond apex of gonostylus (in 
<emphasis box="[369,469,679,700]" italics="true" pageId="6" pageNumber="107">scutellaris</emphasis>
species-group not longer than volsellar cuspis) (Afr, Ori, Pal)... 17. 
<taxonomicName authority="Schuster, 1949" authorityName="Schuster" authorityYear="1949" box="[1157,1436,679,701]" class="Insecta" family="Mutillidae" genus="Physetopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1157,1288,679,701]" italics="true" pageId="6" pageNumber="107">Physetopoda</emphasis>
<bibRefCitation author="Schuster, R. M." box="[1295,1436,679,701]" pageId="6" pageNumber="107" pagination="59 - 140" refId="ref29418" refString="Schuster, R. M. (1949) Contribution towards a monograph of the Mutillidae of the Neotropical region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, New Series, 29 (3 - 4), 59 - 140." type="journal article" year="1949">Schuster, 1949</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[151,1432,748,774]" pageId="6" pageNumber="107" type="multiple">
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1432,748,774]" pageId="6" pageNumber="107">
<emphasis bold="true" box="[151,249,748,774]" pageId="6" pageNumber="107">Females</emphasis>
(unknown in 
<taxonomicName authorityName="Bischoff" authorityYear="1920" box="[405,548,748,774]" class="Insecta" genus="Antennotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis box="[405,548,748,774]" italics="true" pageId="6" pageNumber="107">Antennotilla</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Nonveiller" authorityYear="1993" box="[564,707,748,774]" class="Insecta" family="Mutillidae" genus="Ctenoceraea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis box="[564,707,748,774]" italics="true" pageId="6" pageNumber="107">Ctenoceraea</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Arnold" authorityYear="1946" box="[721,879,748,774]" class="Insecta" genus="Gynandrotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis box="[721,879,748,774]" italics="true" pageId="6" pageNumber="107">Gynandrotilla</emphasis>
</taxonomicName>
, 
<taxonomicName box="[893,1046,748,774]" class="Insecta" family="Mutillidae" genus="Paglianotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis box="[893,1046,748,774]" italics="true" pageId="6" pageNumber="107">Paglianotilla</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Latreille" authorityYear="1825" box="[1061,1242,748,774]" class="Insecta" family="Mutillidae" genus="Psammotherma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis box="[1061,1242,748,774]" italics="true" pageId="6" pageNumber="107">Psammotherma</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Lelej" authorityYear="1995" box="[1257,1423,748,774]" class="Insecta" family="Mutillidae" genus="Tsunekimyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis box="[1257,1423,748,774]" italics="true" pageId="6" pageNumber="107">Tsunekimyrme</emphasis>
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="108" pageId="6" pageNumber="107" type="key">
<key pageId="6" pageNumber="107">
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,781,803]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,781,803]" pageId="6" pageNumber="107">1. T6 convex, without distinct pygidial plate, with glabrous shiny area widened posterad.............................. 2</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">- T6 with distinct flattened pygidial plate, carinated laterally, longitudinally striate, granulous or with glabrous shiny apical part, or entirely glabrous, shiny.............................................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,869,891]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,869,891]" pageId="6" pageNumber="107">2. Scutellar scale more or less developed.................................................................... 3</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,898,920]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,898,920]" pageId="6" pageNumber="107">- Scutellar scale totally lacking........................................................................... 6</keyLead>
</paragraph>
</keyStep>
</key>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
3. Scutellar scale strongly developed. Mesosomal dorsum with parallel sides or slightly widened posterad.—F1 distinctly longer than F2. T6 not carinated even apically (Ori)....................................... 11. 
<taxonomicName authority="Lelej, 2005" authorityName="Lelej" authorityYear="2005" box="[1181,1436,957,979]" class="Insecta" family="Mutillidae" genus="Karunaratnea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1181,1321,957,978]" italics="true" pageId="6" pageNumber="107">Karunaratnea</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,957,979]" pageId="6" pageNumber="107" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,986,1008]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,986,1008]" pageId="6" pageNumber="107">- Scutellar scale weak, but distinct. Mesosomal dorsum often narrowed posterad .................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
4. T6 with glabrous shiny medial part, not carinated laterad, considerably convergent basally with basolateral tufts of dense pale setae (Ori, Pal).................................................................... 22. 
<taxonomicName authority="Lelej, 1995" authorityName="Lelej" authorityYear="1995" box="[1237,1436,1044,1067]" class="Insecta" family="Mutillidae" genus="Sinotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1237,1321,1044,1066]" italics="true" pageId="6" pageNumber="107">Sinotilla</emphasis>
Lelej, 1995
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,1074,1096]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,1074,1096]" pageId="6" pageNumber="107">- T6 with elongated shiny glabrous pygidial plate, weakly carinate laterad......................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,1103,1125]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,1103,1125]" pageId="6" pageNumber="107">
5. F1 equals its apical width and length of F2. T1 short, without lateral pale spots (Ori, Pal)......... 6. 
<taxonomicName authority="Lelej, 1995" authorityName="Lelej" authorityYear="1995" box="[1224,1436,1103,1125]" class="Insecta" family="Mutillidae" genus="Ephucilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1224,1321,1103,1125]" italics="true" pageId="6" pageNumber="107">Ephucilla</emphasis>
Lelej, 1995
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
- F1 longer than its apical width and 1.3 × longer than F2. T1 long with distinct dorsal face, with lateral white spots (Ori)................................................................................... 10. 
<taxonomicName authority="Lelej, 1993" authorityName="Lelej" authorityYear="1993" box="[1226,1436,1162,1184]" class="Insecta" family="Mutillidae" genus="Indratilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1226,1321,1162,1184]" italics="true" pageId="6" pageNumber="107">Indratilla</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,1162,1184]" pageId="6" pageNumber="107" pagination="233 - 236" refId="ref26666" refString="Lelej, A. S. (1993) A new genus of velvet ants from Sri Lanka (Insecta, Hymenoptera, Mutillidae). Spixiana, 16 (3), 233 - 236." type="journal article" year="1993">Lelej, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,1191,1213]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,1191,1213]" pageId="6" pageNumber="107">6. F1 1.6 × its maximum width. Mesosoma elongated, distinctly narrowed posterad.................................. 7</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
- F1 1.0–1.1 × its maximum width. Mesosoma not elongated, with more or less parallel sides or widened posterad (Ori).................................................................................. 14. 
<taxonomicName authority="Lelej, 2005" authorityName="Lelej" authorityYear="2005" box="[1204,1436,1250,1272]" class="Insecta" family="Mutillidae" genus="Nordeniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1204,1321,1250,1272]" italics="true" pageId="6" pageNumber="107">Nordeniella</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,1250,1272]" pageId="6" pageNumber="107" refId="ref26959" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
7. T1 with distinct anterior and dorsal surfaces, upper edge of the truncation projecting in the center above. T6 with pygidial plate convex, non sculptured, apical half weakly carinated laterally (Afr)..................... 8. 
<taxonomicName authority="Suarez, 1977" authorityName="Suarez" authorityYear="1977" box="[1161,1436,1308,1330]" class="Insecta" family="Mutillidae" genus="Guineomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1161,1305,1308,1330]" italics="true" pageId="6" pageNumber="107">Guineomutilla</emphasis>
<bibRefCitation author="Suarez, F. J." box="[1311,1436,1309,1330]" pageId="6" pageNumber="107" pagination="213 - 223" refId="ref30025" refString="Suarez, F. J. (1977) Dos nuevos Mutilidos Etiopicos y propuesta de un genero nuevo para uno de ellos (Hymenoptera, Mutillidae). Nouvelle Revue d'Entomologie, 7 (2), 213 - 223." type="journal article" year="1977">Suárez, 1977</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
- T1 gently sloping without dorsal surface and truncation. T6 with convex pygidial plate non sculptured and not carinated even in apical half (Afr, Ori, Pal)....................................................... 18. 
<taxonomicName authority="Andre, 1902" authorityName="Andre" authorityYear="1902" box="[1200,1436,1367,1389]" class="Insecta" family="Mutillidae" genus="Promecilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1200,1311,1367,1389]" italics="true" pageId="6" pageNumber="107">Promecilla</emphasis>
<bibRefCitation author="Andre, E." box="[1316,1436,1367,1389]" pageId="6" pageNumber="107" pagination="1 - 77" refId="ref24877" refString="Andre, E. (1902 b [&quot; 1903 &quot;]) Fam. Mutillidae. In: Wytsman, P., Genera Insectorum. Fasc. 11. s. n., Bruxelles, pp. 1 - 77 + 3 pls." type="journal article" year="1902">André, 1902</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
8. T2 with central yellowish cuticular spot, without spots of pale setae (Pal)..................................................................................................... 15. 
<taxonomicName authority="Lelej" authorityName="Lelej" authorityYear="1980" box="[996,1180,1426,1448]" class="Insecta" genus="Nuristanilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[996,1125,1426,1448]" italics="true" pageId="6" pageNumber="107">Nuristanilla</emphasis>
Lelej
</taxonomicName>
in 
<bibRefCitation author="Lelej, A. S. &amp; Kabakov, O. N." box="[1211,1436,1426,1448]" pageId="6" pageNumber="107" pagination="141 - 156" refId="ref27122" refString="Lelej, A. S. &amp; Kabakov, O. N. (1980) On Bradynobaenidae and Mutillidae (Hymenoptera) of Afghanistan. Entomologicheskoe Obozrenie, 59 (1), 181 - 196. [in Russian, English translation: (1981) Entomological Review, 59 (1), 141 - 156]" type="journal article" year="1980">Lelej &amp; Kabakov, 1980</bibRefCitation>
</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
- T2 without pale cuticular spot, with one or three basal spots of pale setae, rarely (some 
<taxonomicName authorityName="Lelej" authorityYear="1980" box="[1064,1143,1455,1476]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis box="[1064,1143,1455,1476]" italics="true" pageId="6" pageNumber="107">Dentilla</emphasis>
</taxonomicName>
) with basal band of pale setae ................................................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,1514,1536]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,1514,1536]" pageId="6" pageNumber="107">9. Scutellar scale lacking................................................................................ 10</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,1543,1565]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,1543,1565]" pageId="6" pageNumber="107">- Scutellar scale more or less developed........................................................................ 11</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,1572,1595]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,1572,1595]" pageId="6" pageNumber="107">
10. Mandible wide, tridentate apically (
<figureCitation box="[568,626,1573,1595]" captionStart="FIGURES 7–16" captionStartId="10.[152,270,1712,1736]" captionTargetBox="[151,1436,484,1688]" captionTargetId="figure-103@10.[151,1436,478,1688]" captionTargetPageId="10" captionText="FIGURES 7–16. 7–10. Ctenoceraea pectinella Nonveiller, 1993, ♁, holotype (7. Antenna, 8. Forewing, 9. Genitalia, dorsal view, 10. Genitalia, lateral view); 11–12. Dentilla rasnitsyni Lelej in Lelej &amp; van Harten, 2011, ♁, paratype (11. Lateral habitus, 12. Head, face view); 13–16. D. curtiventris (André, 1901): 13. Dorsal habitus, ♀, 14. Lateral habitus, ♀, 15. Lateral habitus, ♁, 16. Head, face view. (Photographs by K. Williams, 7–10 from Nonveiller 1993)." figureDoi="http://doi.org/10.5281/zenodo.7575312" httpUri="https://zenodo.org/record/7575312/files/figure.png" pageId="6" pageNumber="107">Fig. 7</figureCitation>
) (Ori, Pal)................................. 1. 
<taxonomicName authority="Lelej, 1995" authorityName="Lelej" authorityYear="1995" box="[1167,1436,1572,1595]" class="Insecta" family="Mutillidae" genus="Andreimyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1167,1319,1572,1594]" italics="true" pageId="6" pageNumber="107">Andreimyrme</emphasis>
Lelej, 1995
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,1602,1624]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,1602,1624]" pageId="6" pageNumber="107">
- Mandible falcate and edentate with acute apex (Afr)..................................... 3. 
<taxonomicName authority="Arnold, 1956" authorityName="Arnold" authorityYear="1956" box="[1216,1436,1602,1624]" class="Insecta" genus="Corytilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1216,1303,1602,1624]" italics="true" pageId="6" pageNumber="107">Corytilla</emphasis>
<bibRefCitation author="Arnold, G." box="[1308,1436,1602,1624]" pageId="6" pageNumber="107" pagination="52 - 77" refId="ref25000" refString="Arnold, G. (1956) New species of African Hymenoptera. No. 12. Occasional Papers of the National Museum of Southern Rhodesia, 3 (21 B), 52 - 77." type="journal article" year="1956">Arnold, 1956</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
11. Head widened posterad, posteriorly angulate. Pronotum dorsally with medial longitudinal depression (sulcus) laterally bordered by tubercles (Afr)............................................................. 25. 
<taxonomicName authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" box="[1193,1436,1660,1683]" class="Insecta" genus="Sulcotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1193,1288,1660,1682]" italics="true" pageId="6" pageNumber="107">Sulcotilla</emphasis>
<bibRefCitation author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[1294,1436,1661,1683]" pageId="6" pageNumber="107" refId="ref25275" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv f ¸ r Naturgeschichte, 86 A (1 - 3) &amp; 86 A (4 - 5), 1 - 480 (1920) &amp; 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" box="[151,1436,1690,1712]" pageId="6" pageNumber="107">
<keyLead box="[151,1436,1690,1712]" pageId="6" pageNumber="107">- Head not widened posterad, posteriorly rouded. Pronotum dorsally without medial longitudinal depression (sulcus)...... 12</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
12. Pygidial plate narrow, usually less than 0.3 × T6 width, narrowed and generally closed dorsally, at least 2 × higher than wide; usually with long setae laterally (
<figureCitation box="[544,625,1749,1771]" captionStart="FIGURES 110–115" captionStartId="33.[152,270,1115,1139]" captionTargetBox="[151,1433,181,1091]" captionTargetId="figure-17@33.[151,1436,181,1091]" captionTargetPageId="33" captionText="FIGURES 110–115. Smicromyrmini, ♀, pygidial plate (T6). 110.Andreimyrme rong Williams in Okayasu et al., 2021, holotype; 111. Dentilla curtiventris (André, 1901); 112. Ephucilla thalia (Mickel, 1933); 113. Ephutomma fletcheri (Turner, 1911); 114. Karunaratnea palatupanae Lelej, 2005, paratype; 115. Nemka viduata (Pallas, 1773). (Photographs by K. Williams andA. Lelej (114))." figureDoi="http://doi.org/10.5281/zenodo.7575342" httpUri="https://zenodo.org/record/7575342/files/figure.png" pageId="6" pageNumber="107">Fig. 113</figureCitation>
)................................................................ 13
</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
- Pygidial plate generally at least 0.5 × T6 width, usually open dorsally, its shape variable: elongate with more or less parallel lateral sides (
<figureCitation box="[353,437,1807,1829]" captionStart="FIGURES 104–109" captionStartId="32.[152,270,1978,2002]" captionTargetBox="[151,1436,609,1955]" captionTargetId="figure-211@32.[151,1436,597,1955]" captionTargetPageId="32" captionText="FIGURES 104–109. Smicromyrme (s. str.) rufipes (Fabricius, 1787): 104. ♀, dorsal habitus, 105. ♀, lateral habitus, 106. ♁, lateral habitus, 107. ♁, f. nigra, lateral habitus, 108. ♁, head, face view, 109. ♀, pygidial plate. (Photographs by K. Williams)." figureDoi="http://doi.org/10.5281/zenodo.7575340" httpUri="https://zenodo.org/record/7575340/files/figure.png" pageId="6" pageNumber="107">Fig. 109</figureCitation>
), broadly ovate (at most 1.7 × higher than wide, 
<figureCitation box="[905,987,1807,1829]" captionStart="FIGURES 110–115" captionStartId="33.[152,270,1115,1139]" captionTargetBox="[151,1433,181,1091]" captionTargetId="figure-17@33.[151,1436,181,1091]" captionTargetPageId="33" captionText="FIGURES 110–115. Smicromyrmini, ♀, pygidial plate (T6). 110.Andreimyrme rong Williams in Okayasu et al., 2021, holotype; 111. Dentilla curtiventris (André, 1901); 112. Ephucilla thalia (Mickel, 1933); 113. Ephutomma fletcheri (Turner, 1911); 114. Karunaratnea palatupanae Lelej, 2005, paratype; 115. Nemka viduata (Pallas, 1773). (Photographs by K. Williams andA. Lelej (114))." figureDoi="http://doi.org/10.5281/zenodo.7575342" httpUri="https://zenodo.org/record/7575342/files/figure.png" pageId="6" pageNumber="107">Fig. 115</figureCitation>
), or triangular in shape (widened dorsally, 
<figureCitation captionStart="FIGURES 110–115" captionStartId="33.[152,270,1115,1139]" captionTargetBox="[151,1433,181,1091]" captionTargetId="figure-17@33.[151,1436,181,1091]" captionTargetPageId="33" captionText="FIGURES 110–115. Smicromyrmini, ♀, pygidial plate (T6). 110.Andreimyrme rong Williams in Okayasu et al., 2021, holotype; 111. Dentilla curtiventris (André, 1901); 112. Ephucilla thalia (Mickel, 1933); 113. Ephutomma fletcheri (Turner, 1911); 114. Karunaratnea palatupanae Lelej, 2005, paratype; 115. Nemka viduata (Pallas, 1773). (Photographs by K. Williams andA. Lelej (114))." figureDoi="http://doi.org/10.5281/zenodo.7575342" httpUri="https://zenodo.org/record/7575342/files/figure.png" pageId="6" pageNumber="107">Fig. 111</figureCitation>
).............................................................................................. 14
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="6" pageNumber="107">
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
13. Eye enlarged, minimal distance between eyes 1.1–1.3 × longitudinal eye diameter. Scutellar scale generally lowly produced (Afr, Ori, Pal)............................................................... 7. 
<taxonomicName authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[1156,1436,1895,1917]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1156,1280,1895,1917]" italics="true" pageId="6" pageNumber="107">Ephutomma</emphasis>
<bibRefCitation author="Ashmead, W. H." box="[1286,1436,1895,1917]" pageId="6" pageNumber="107" pagination="45 - 60" refId="ref25074" refString="Ashmead, W. H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45 - 60." type="journal article" year="1899">Ashmead, 1899</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
- Eye moderate in size, minimal distance between eyes 1.6–1.8 × longitudinal eye diameter. Scutellar scale usually erect, welldeveloped, nail-like (Afr, Ori, Pal).............................................. 17. 
<taxonomicName authority="Schuster, 1949" authorityName="Schuster" authorityYear="1949" box="[1163,1436,1954,1976]" class="Insecta" family="Mutillidae" genus="Physetopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1163,1288,1954,1976]" italics="true" pageId="6" pageNumber="107">Physetopoda</emphasis>
<bibRefCitation author="Schuster, R. M." box="[1295,1436,1954,1976]" pageId="6" pageNumber="107" pagination="59 - 140" refId="ref29418" refString="Schuster, R. M. (1949) Contribution towards a monograph of the Mutillidae of the Neotropical region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, New Series, 29 (3 - 4), 59 - 140." type="journal article" year="1949">Schuster, 1949</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph blockId="6.[151,1437,748,2035]" pageId="6" pageNumber="107">
<keyLead pageId="6" pageNumber="107">
<keyStep pageId="6" pageNumber="107">
14. T2 with one or three spots of pale setae disposed in central horizontal line. T1 posterodorsdally with small medial spot of pale setae (Afr)............................................................ 20. 
<taxonomicName authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" box="[1100,1436,2012,2035]" class="Insecta" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1100,1288,2012,2034]" italics="true" pageId="6" pageNumber="107">Pseudocephalotilla</emphasis>
<bibRefCitation author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[1294,1436,2013,2035]" pageId="6" pageNumber="107" refId="ref25275" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv f ¸ r Naturgeschichte, 86 A (1 - 3) &amp; 86 A (4 - 5), 1 - 480 (1920) &amp; 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" box="[151,1436,151,173]" pageId="7" pageNumber="108">- T2 with one or three spots of pale setae disposed basally. T1 posterodorsdally at most with fringe of pale setae.......... 15</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" box="[151,1436,180,203]" pageId="7" pageNumber="108">
15. Pygidial plate short-oval (Ori, Pal)..................................................... 13. 
<taxonomicName authority="Lelej, 1985" authorityName="Lelej" authorityYear="1985" box="[1246,1436,180,203]" class="Insecta" family="Mutillidae" genus="Nemka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1246,1320,180,202]" italics="true" pageId="7" pageNumber="108">Nemka</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,181,203]" pageId="7" pageNumber="108" refId="ref26618" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) Fauny SSSR i Sopredelnykh Stran [The Velvet Ants (Hymenoptera, Mutillidae) of the USSR and Neighbouring Countries]. Nauka, Leningrad, 268 pp. [in Russian]" type="book" year="1985">Lelej, 1985</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" box="[151,1436,210,232]" pageId="7" pageNumber="108">- Pygidial plate elongated, with parallel sides, or widened basally............................................... 16</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">
16. T2 with three basal spots of pale setae disposed in horizontal line and with at most median apical patch of pale setae. Scutellar scale more than 2 × as wide as long (Ori, Pal)..................................... 12. 
<taxonomicName authority="Lelej, 1995" authorityName="Lelej" authorityYear="1995" box="[1163,1436,268,291]" class="Insecta" family="Mutillidae" genus="Mickelomyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1163,1320,268,290]" italics="true" pageId="7" pageNumber="108">Mickelomyrme</emphasis>
Lelej, 1995
</taxonomicName>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">- T2 with one basal medial spot of pale setae, if lateral pale spots present, then they are disposed posterior to median patch, and with apical band or fringe of pale setae. Scutellar scale usually only slightly wider than its length.................... 17</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">
17. Pronotum with protruding humeral part, distinctly wider than propodeum (Afr, Ori, Pal).............................................................................................. 5. 
<taxonomicName authority="Lelej" authorityName="Lelej" authorityYear="1980" box="[1045,1180,386,408]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1045,1125,386,408]" italics="true" pageId="7" pageNumber="108">Dentilla</emphasis>
Lelej
</taxonomicName>
in 
<bibRefCitation author="Lelej, A. S. &amp; Kabakov, O. N." box="[1211,1436,386,408]" pageId="7" pageNumber="108" pagination="141 - 156" refId="ref27122" refString="Lelej, A. S. &amp; Kabakov, O. N. (1980) On Bradynobaenidae and Mutillidae (Hymenoptera) of Afghanistan. Entomologicheskoe Obozrenie, 59 (1), 181 - 196. [in Russian, English translation: (1981) Entomological Review, 59 (1), 141 - 156]" type="journal article" year="1980">Lelej &amp; Kabakov, 1980</bibRefCitation>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" box="[151,1436,415,437]" pageId="7" pageNumber="108">- Pronotum without protruding humeral part, usually narrower or at most scarcely wider than propodeum............... 18</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">
18 (-). Basal medial spot of pale setae on T2 not enlarged: distance between spot and band of pale setae of tergum more than spot diameter (Afr, Ori, Pal).................................................... 24. 
<taxonomicName authority="Thomson, 1870" authorityName="Thomson" authorityYear="1870" box="[1135,1436,474,496]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1135,1280,474,495]" italics="true" pageId="7" pageNumber="108">Smicromyrme</emphasis>
<bibRefCitation author="Thomson, C. G." box="[1286,1436,474,496]" pageId="7" pageNumber="108" pagination="202 - 251" refId="ref30270" refString="Thomson, C. G. (1870) Ofversigt af Sveriges rofsteklar. Opuscula Entomologica, 1 (2), 202 - 251." type="journal article" year="1870">Thomson, 1870</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">
18a. Metasomal segment 1 constricted posterad. Pygidial plate rhomboid with granulated sculpture (Pal)....................................................................................... 24.4. Subgenus 
<taxonomicName authority="Nagy, 1966" authorityName="Nagy" authorityYear="1966" box="[1197,1436,532,555]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="subGenus" subGenus="Rhombotilla">
<emphasis bold="true" box="[1197,1320,532,554]" italics="true" pageId="7" pageNumber="108">Rhombotilla</emphasis>
<bibRefCitation author="Nagy, C. G." box="[1326,1436,533,555]" pageId="7" pageNumber="108" pagination="113 - 117" refId="ref27878" refString="Nagy, C. G. (1966) Ein neues Mutilliden-Subgenus aus Rumanien (Hymenoptera). Reichenbachia, 6 (13 - 34), 113 - 117." type="journal article" year="1966">Nagy, 1966</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">- Metasomal segment 1 not constricted posterad. Pygidial plate another shape, longitudinally striate with striae reaching posterior margin or apically smooth............................................................................ 18b</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" box="[151,1436,620,643]" pageId="7" pageNumber="108">
18b. Pygidial plate triangular, conspicuously widened basally (Ori, Pal)............... 24.2. Subgenus 
<taxonomicName authority="Lelej, 1985" authorityName="Lelej" authorityYear="1985" box="[1216,1436,620,643]" class="Insecta" family="Mutillidae" genus="Eremotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1216,1321,620,642]" italics="true" pageId="7" pageNumber="108">Eremotilla</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,621,643]" pageId="7" pageNumber="108" refId="ref26618" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) Fauny SSSR i Sopredelnykh Stran [The Velvet Ants (Hymenoptera, Mutillidae) of the USSR and Neighbouring Countries]. Nauka, Leningrad, 268 pp. [in Russian]" type="book" year="1985">Lelej, 1985</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" box="[151,1436,650,672]" pageId="7" pageNumber="108">- Pygidial plate elongated, often with more or less parallel sides............................................... 18c</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" box="[151,1436,679,701]" pageId="7" pageNumber="108">
18c. Pygidial plate with striae reaching posterior margin (Pal)................ 24.5. Subgenus 
<taxonomicName authority="Thomson, 1870" authorityName="Thomson" authorityYear="1870" box="[1141,1436,679,701]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1141,1280,679,700]" italics="true" pageId="7" pageNumber="108">Smicromyrme</emphasis>
<bibRefCitation author="Thomson, C. G." box="[1286,1436,679,701]" pageId="7" pageNumber="108" pagination="202 - 251" refId="ref30270" refString="Thomson, C. G. (1870) Ofversigt af Sveriges rofsteklar. Opuscula Entomologica, 1 (2), 202 - 251." type="journal article" year="1870">Thomson, 1870</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" box="[151,1436,709,731]" pageId="7" pageNumber="108">- Pygidial plate with striae not reaching posterior margin, apical portion smooth.................................. 18d</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">
18d. Mesosoma with well developed anterodorsal and posterodorsal angles, dorsally more or less flattened, in lateral view (Pal)................................................................... 24.1. Subgenus 
<taxonomicName authority="Schwartz, 1984" authorityName="Schwartz" authorityYear="1984" box="[1169,1436,767,789]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="subGenus" subGenus="Astomyrme">
<emphasis bold="true" box="[1169,1281,767,788]" italics="true" pageId="7" pageNumber="108">Astomyrme</emphasis>
<bibRefCitation author="Schwartz, F." box="[1287,1436,767,789]" pageId="7" pageNumber="108" pagination="21 - 25" refId="ref29541" refString="Schwartz, F. (1984) Description d'un genre nouveau, disjoint de Smicromyrme, et d'une espece nouvelle du sud de la France (Hymenoptera, Mutillidae). L'Entomologiste, 40, 21 - 25." type="journal article" year="1984">Schwartz, 1984</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">
- Mesosoma with rounded anterodorsal and posterodorsal angles, dorsally weakly arcuate, in lateral view (Afr, Ori, Pal).......................................................................... 24.3. Subgenus 
<taxonomicName authority="Lelej, 1985" authorityName="Lelej" authorityYear="1985" box="[1221,1436,826,848]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="subGenus" subGenus="Erimyrme">
<emphasis bold="true" box="[1221,1321,826,847]" italics="true" pageId="7" pageNumber="108">Erimyrme</emphasis>
<bibRefCitation author="Lelej, A. S." box="[1327,1436,826,848]" pageId="7" pageNumber="108" refId="ref26618" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) Fauny SSSR i Sopredelnykh Stran [The Velvet Ants (Hymenoptera, Mutillidae) of the USSR and Neighbouring Countries]. Nauka, Leningrad, 268 pp. [in Russian]" type="book" year="1985">Lelej, 1985</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">18 (-). Basal medial spot of pale setae on T2 enlarged, distance between spot and band of tergum equal to spot diameter or less than it ................................................................................................. 19</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">
19. Pygidial plate apically distinctly angulate, lateral carina distinctly widened apically. Clypeus without median basal tubercle (Afr, Pal)................................................. 21. 
<taxonomicName authority="Lelej" authorityName="Lelej" box="[962,1159,943,965]" class="Insecta" family="Mutillidae" genus="Rasnitsynitilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[962,1105,943,965]" italics="true" pageId="7" pageNumber="108">Rasnitsynitilla</emphasis>
Lelej
</taxonomicName>
in 
<bibRefCitation author="Lelej, A. S. &amp; van Harten, A." box="[1190,1436,943,965]" pageId="7" pageNumber="108" pagination="1 - 50" refId="ref27327" refString="Lelej, A. S. &amp; van Harten, A. (2006) A review of the Mutillidae (Hymenoptera) of Yemen. Zootaxa, 1226 (1), 1 - 50. https: // doi. org / 10.11646 / zootaxa. 1226.1.1" type="journal article" year="2006">Lelej &amp; van Harten, 2006</bibRefCitation>
</paragraph>
<paragraph blockId="7.[151,1436,151,1024]" pageId="7" pageNumber="108">
- Pygidial plate apically rounded, lateral carina not widened apically. Clypeus with median basal tubercle (Pal).................................................................................... 23. 
<taxonomicName authority="Ovtchinnikov, 2002" authorityName="Ovtchinnikov" authorityYear="2002" box="[1125,1436,1002,1024]" class="Insecta" family="Mutillidae" genus="Skorikovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1125,1239,1002,1024]" italics="true" pageId="7" pageNumber="108">Skorikovia</emphasis>
<bibRefCitation author="Ovtchinnikov, S. V." box="[1246,1436,1002,1024]" pageId="7" pageNumber="108" refId="ref28527" refString="Ovtchinnikov, S. V. (2002) Materials to the fauna of velvet ants (Hymenoptera, Mutillidae) of Kyrghizia. In: Zonstein, S. L. (Ed.), Entomological Investigations in Kyrgystan. Vol. 22. Kyrghyz Society of Entomology, Bishkek, pp. 79 - 104. [in Russian]" type="book" year="2002">Ovtchinnikov, 2002</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>